163 lines
18 KiB
XML
163 lines
18 KiB
XML
<document id="AA168818E8CA1370A8705508141246A9" ID-DOI="10.5281/zenodo.13203160" ID-ISSN="1407-8619" ID-Zenodo-Dep="13203160" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722657382176" checkinUser="felipe" docAuthor="Fanti, Fabrizio & Müller, Patrick" docDate="2022" docId="03DAD52BFFAEFFE6FFEBBDA85CC7A534" docLanguage="en" docName="BalticJColeopterol.22.2.331-380.pdf" docOrigin="Baltic Journal of Coleopterology 22 (2)" docStyle="DocumentStyle:9720869F1FB170B2A509257D1FA394CD.2:BalticJColeop.2022.journal_article" docStyleId="9720869F1FB170B2A509257D1FA394CD" docStyleName="BalticJColeop.2022.journal_article" docStyleVersion="2" docTitle="Hukawngichthyurus ditaddeoi Fanti & Müller, 2022, sp. nov." docType="treatment" docVersion="3" lastPageNumber="343" masterDocId="FFE3AD53FFA5FFEAFF9ABD325F46A170" masterDocTitle="Fossil Cantharidae from the Cretaceous Burmese (Kachin) amber of the Patrick Müller collection, and taxonomic information" masterLastPageNumber="380" masterPageNumber="331" pageNumber="342" updateTime="1722697113453" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-4.0">
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<mods:mods id="29B9C0DE193F6A39E8E48F5FE03B87AC" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="C95FF48CE5DE828D0BDF1C52F79FD9CC">
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<mods:title id="18A620EA5DB56B894EF7F15147D9E49C">Fossil Cantharidae from the Cretaceous Burmese (Kachin) amber of the Patrick Müller collection, and taxonomic information</mods:title>
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<mods:namePart id="97B6BA1C60C5A8D2546D53FF34F7A1FD">Fanti, Fabrizio</mods:namePart>
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<mods:namePart id="CF6761910502006AE5DFFF9A3C1CE4C1">Müller, Patrick</mods:namePart>
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<mods:typeOfResource id="889CB54F6A82541E1DF2BF05943A3B1C">text</mods:typeOfResource>
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<mods:title id="20879E3F2DBCCD29FAA316A7FAA63C9A">Baltic Journal of Coleopterology</mods:title>
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</mods:titleInfo>
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<mods:part id="7E32A85D5AD415A88DE1DF66E4BF3E37">
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<mods:date id="E80C1B2BDD112CD14B7D12E91CDB340F">2022</mods:date>
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<mods:detail id="76693ACB5EE0C5CE1A197E6FCEDF7662" type="volume">
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<mods:number id="E1EE4F6896EEF6C2A7438349B248D95E">22</mods:number>
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<mods:identifier id="B93790E3E4C03408118CB4105028C181" type="DOI">10.5281/zenodo.13203160</mods:identifier>
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<mods:identifier id="B315891579D74569530B6B3A5DC04945" type="ISSN">1407-8619</mods:identifier>
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<treatment id="03DAD52BFFAEFFE6FFEBBDA85CC7A534" ID-DOI="http://doi.org/10.5281/zenodo.13199795" ID-Zenodo-Dep="13199795" LSID="urn:lsid:plazi:treatment:03DAD52BFFAEFFE6FFEBBDA85CC7A534" httpUri="http://treatment.plazi.org/id/03DAD52BFFAEFFE6FFEBBDA85CC7A534" lastPageId="12" lastPageNumber="343" pageId="11" pageNumber="342">
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<subSubSection id="C36937B6FFAEFFE1FFEBBDA85D5DA1C2" box="[113,539,154,178]" pageId="11" pageNumber="342" type="nomenclature">
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<paragraph id="8BCC643DFFAEFFE1FFEBBDA85D5DA1C2" blockId="11.[113,539,154,210]" box="[113,539,154,178]" pageId="11" pageNumber="342">
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<heading id="D084D351FFAEFFE1FFEBBDA85D5DA1C2" bold="true" box="[113,539,154,178]" fontSize="10" level="2" pageId="11" pageNumber="342" reason="4">
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<emphasis id="B907B82FFFAEFFE1FFEBBDA85D5DA1C2" bold="true" box="[113,539,154,178]" pageId="11" pageNumber="342">
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<taxonomicName id="4C731FBEFFAEFFE1FFEBBDA85EF9A1C2" authorityName="Fanti & Müller" authorityYear="2022" box="[113,447,154,178]" class="Insecta" family="Cantharidae" genus="Hukawngichthyurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="342" phylum="Arthropoda" rank="species" species="ditaddeoi" status="sp. nov.">
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||
<emphasis id="B907B82FFFAEFFE1FFEBBDA85EF9A1C2" bold="true" box="[113,447,154,178]" italics="true" pageId="11" pageNumber="342">Hukawngichthyurus ditaddeoi</emphasis>
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||
</taxonomicName>
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<taxonomicNameLabel id="A2340554FFAEFFE1FE5FBDA85D5DA1C2" box="[453,539,154,178]" pageId="11" pageNumber="342" rank="species">sp. nov.</taxonomicNameLabel>
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||
</emphasis>
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||
</heading>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C36937B6FFAEFFE1FFEBBD895F87A1A2" box="[113,193,187,210]" pageId="11" pageNumber="342" type="description">
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||
<paragraph id="8BCC643DFFAEFFE1FFEBBD895F87A1A2" blockId="11.[113,539,154,210]" box="[113,193,187,210]" pageId="11" pageNumber="342">
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(
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<figureCitation id="134878B8FFAEFFE1FFE3BD895FFFA1A2" box="[121,185,187,210]" captionStart="Fig" captionStartId="11.[113,152,1636,1659]" captionTargetBox="[113,1134,382,1607]" captionTargetId="figure-98@11.[113,1134,382,1607]" captionTargetPageId="11" captionText="Fig. 6. Hukawngichthyurus ditaddeoi sp. nov. in Burmese amber. A: Holotype, ventrolateral view; B: Holotype, dorsolateral view." figureDoi="http://doi.org/10.5281/zenodo.13203185" httpUri="https://zenodo.org/record/13203185/files/figure.png" pageId="11" pageNumber="342">Fig. 6</figureCitation>
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)
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C36937B6FFAEFFE6FFEBBDC35EF0A1A3" lastPageId="12" lastPageNumber="343" pageId="11" pageNumber="342" type="materials_examined">
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||
<paragraph id="8BCC643DFFAEFFE1FFEBBDC35D8FA182" blockId="11.[113,605,241,328]" lastBlockId="11.[643,1134,155,242]" pageId="11" pageNumber="342">
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<materialsCitation id="3B1B6E60FFAEFFE1FFEBBDC35D8FA182" collectingDate="2021-12-17" collectionCode="BUB" county="Probable" location="Catalog" municipality="Burmese" pageId="11" pageNumber="342" specimenCode="BUB 4326" specimenCount="1" stateProvince="Kachin" typeStatus="holotype">
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<emphasis id="B907B82FFFAEFFE1FFEBBDC35FA5A079" bold="true" box="[113,227,241,265]" pageId="11" pageNumber="342">
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<typeStatus id="54C8DA9FFFAEFFE1FFEBBDC35F9DA079" box="[113,219,241,265]" pageId="11" pageNumber="342" type="holotype">Holotype</typeStatus>
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.
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</emphasis>
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<collectingCounty id="62AD1CB1FFAEFFE1FF77BDC35E08A078" box="[237,334,241,264]" pageId="11" pageNumber="342">Probable</collectingCounty>
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female, adult specimen in a
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<collectingMunicipality id="6BA8FE47FFAEFFE1FF3FBC235E42A058" box="[165,260,273,296]" pageId="11" pageNumber="342">Burmese</collectingMunicipality>
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||
(
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<collectingRegion id="49B7AADFFFAEFFE1FE8EBC235E22A058" box="[276,356,273,296]" country="Myanmar" name="Kachin" pageId="11" pageNumber="342">Kachin</collectingRegion>
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) amber piece:
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<location id="8EAC32E6FFAEFFE1FD90BC235D1BA058" LSID="urn:lsid:plazi:treatment:03DAD52BFFAEFFE6FFEBBDA85CC7A534:8EAC32E6FFAEFFE1FD90BC235D1BA058" box="[522,605,273,296]" county="Probable" municipality="Burmese" name="Catalog" pageId="11" pageNumber="342" stateProvince="Kachin">Catalog</location>
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number SNSBBSPG
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<date id="FFCD42FDFFAEFFE1FEC4BC035EA6A038" box="[350,480,305,328]" pageId="11" pageNumber="342" value="2021-12-17">
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<collectingDate id="EF89BB15FFAEFFE1FEC4BC035EA6A038" box="[350,480,305,328]" pageId="11" pageNumber="342" value="2021-12-17">
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2021 XII
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<quantity id="4C8BC9D8FFAEFFE1FE5FBC035EA6A038" box="[453,480,305,328]" metricMagnitude="-1" metricUnit="m" metricValue="4.318" pageId="11" pageNumber="342" unit="in" value="17.0">17</quantity>
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</collectingDate>
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</date>
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in the
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<location id="8EAC32E6FFAEFFE1FDB3BC035CC5A1C2" LSID="urn:lsid:plazi:treatment:03DAD52BFFAEFFE6FFEBBDA85CC7A534:8EAC32E6FFAEFFE1FDB3BC035CC5A1C2" box="[553,899,155,328]" county="Probable" municipality="Burmese" name="Bayerische Staatssammlung" pageId="11" pageNumber="342" stateProvince="Kachin">Bayerische Staatssammlung</location>
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für
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<location id="8EAC32E6FFAEFFE1FC34BDA95B07A1C2" LSID="urn:lsid:plazi:treatment:03DAD52BFFAEFFE6FFEBBDA85CC7A534:8EAC32E6FFAEFFE1FC34BDA95B07A1C2" box="[942,1089,155,178]" county="Probable" municipality="Burmese" name="Palaontologie" pageId="11" pageNumber="342" stateProvince="Kachin">Paläontologie</location>
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und
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<location id="8EAC32E6FFAEFFE1FD19BD895DA2A1A2" LSID="urn:lsid:plazi:treatment:03DAD52BFFAEFFE6FFEBBDA85CC7A534:8EAC32E6FFAEFFE1FD19BD895DA2A1A2" box="[643,740,187,210]" county="Probable" municipality="Burmese" name="Geologie" pageId="11" pageNumber="342" stateProvince="Kachin">Geologie</location>
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(ex
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<location id="8EAC32E6FFAEFFE1FC95BD895C2FA1A2" LSID="urn:lsid:plazi:treatment:03DAD52BFFAEFFE6FFEBBDA85CC7A534:8EAC32E6FFAEFFE1FC95BD895C2FA1A2" box="[783,873,187,210]" county="Probable" municipality="Burmese" name="Muller's" pageId="11" pageNumber="342" stateProvince="Kachin">Müller’s</location>
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collection number:
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<specimenCode id="DBD5CC46FFAEFFE1FBADBD895DFBA182" collectionCode="BUB" pageId="11" pageNumber="342">BUB 4326</specimenCode>
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).
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||
</materialsCitation>
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||
</paragraph>
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||
<paragraph id="8BCC643DFFAEFFE1FD19BC235C93A038" blockId="11.[643,1134,273,328]" pageId="11" pageNumber="342">
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||
<emphasis id="B907B82FFFAEFFE1FD19BC235C5AA059" bold="true" box="[643,796,273,297]" pageId="11" pageNumber="342">Type locality.</emphasis>
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<collectingCountry id="F36424ADFFAEFFE1FCBEBC235CD6A058" box="[804,912,273,296]" name="Myanmar" pageId="11" pageNumber="342">Myanmar</collectingCountry>
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:
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<collectingRegion id="49B7AADFFFAEFFE1FC01BC235B63A058" box="[923,1061,273,296]" country="Myanmar" name="Kachin" pageId="11" pageNumber="342">Kachin state</collectingRegion>
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, Myitkyina District, Hukawng Valley.
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</paragraph>
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<caption id="DF0C34B5FFAEFFE1FFEBBB565D4EA7EB" ID-DOI="http://doi.org/10.5281/zenodo.13203185" ID-Zenodo-Dep="13203185" httpUri="https://zenodo.org/record/13203185/files/figure.png" pageId="11" pageNumber="342" startId="11.[113,152,1636,1659]" targetBox="[113,1134,382,1607]" targetPageId="11" targetType="figure">
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||
<paragraph id="8BCC643DFFAEFFE1FFEBBB565D4EA7EB" blockId="11.[113,1134,1636,1691]" pageId="11" pageNumber="342">
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||
Fig. 6.
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<taxonomicName id="4C731FBEFFAEFFE1FF59BB565D4EA70B" authorityName="Fanti & Müller" authorityYear="2022" box="[195,520,1636,1659]" class="Insecta" family="Cantharidae" genus="Hukawngichthyurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="342" phylum="Arthropoda" rank="species" species="ditaddeoi" status="sp. nov.">
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<emphasis id="B907B82FFFAEFFE1FF59BB565D4EA70B" box="[195,520,1636,1659]" italics="true" pageId="11" pageNumber="342">Hukawngichthyurus ditaddeoi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A2340554FFAEFFE1FD88BB565D21A70B" box="[530,615,1636,1659]" pageId="11" pageNumber="342" rank="species">sp. nov.</taxonomicNameLabel>
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in Burmese amber. A: Holotype, ventrolateral view; B: Holotype, dorsolateral view.
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</paragraph>
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</caption>
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<paragraph id="8BCC643DFFA9FFE6FF0DBDA95EF0A1A3" blockId="12.[151,643,155,211]" pageId="12" pageNumber="343">
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<emphasis id="B907B82FFFA9FFE6FF0DBDA95E6CA1C3" bold="true" box="[151,298,155,179]" pageId="12" pageNumber="343">Type horizon.</emphasis>
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Lowermost Cenomanian (98.79 ± 0.62 Ma), midCretaceous.
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</paragraph>
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</subSubSection>
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||
<subSubSection id="C36937B6FFA9FFE6FF0DBDC15E86A0BF" pageId="12" pageNumber="343" type="diagnosis">
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<paragraph id="8BCC643DFFA9FFE6FF0DBDC15E86A0BF" blockId="12.[151,643,243,463]" pageId="12" pageNumber="343">
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<emphasis id="B907B82FFFA9FFE6FF0DBDC15EDFA07B" bold="true" box="[151,409,243,267]" pageId="12" pageNumber="343">Differential diagnosis.</emphasis>
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The relatively small body size, the pronotum irregular with a convex draft, and the last urites in a kind of “fish tail” make this new species belong to the genus
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<emphasis id="B907B82FFFA9FFE6FF0DBC455E32A0FE" box="[151,372,375,398]" italics="true" pageId="12" pageNumber="343">
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<taxonomicName id="4C731FBEFFA9FFE6FF0DBC455E36A0FE" authorityName="Fanti & Ellenberger" authorityYear="2018" box="[151,368,375,398]" class="Insecta" family="Cantharidae" genus="Hukawngichthyurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="343" phylum="Arthropoda" rank="genus">Hukawngichthyurus</taxonomicName>
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.
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</emphasis>
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The new species is distinguishable from other
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<taxonomicName id="4C731FBEFFA9FFE6FEE9BCAA5D01A0DF" authorityName="Fanti & Ellenberger" authorityYear="2018" box="[371,583,408,431]" class="Insecta" family="Cantharidae" genus="Hukawngichthyurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="343" phylum="Arthropoda" rank="genus">
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<emphasis id="B907B82FFFA9FFE6FEE9BCAA5D01A0DF" box="[371,583,408,431]" italics="true" pageId="12" pageNumber="343">Hukawngichthyurus</emphasis>
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</taxonomicName>
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by its longer pronotum and elytra.
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C36937B6FFA9FFE6FF0DBCDD5C2CA3CB" pageId="12" pageNumber="343" type="description">
|
||
<paragraph id="8BCC643DFFA9FFE6FF0DBCDD5D0FA338" blockId="12.[151,643,495,584]" pageId="12" pageNumber="343">
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||
<emphasis id="B907B82FFFA9FFE6FF0DBCDD5E6EA377" bold="true" box="[151,296,495,519]" pageId="12" pageNumber="343">Description.</emphasis>
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||
Adult, alate, robust. Probable female, for the wide abdomen. Entirely brown with darker elytra. Body length:
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<quantity id="4C8BC9D8FFA9FFE6FE6ABF035D03A338" box="[496,581,561,584]" metricMagnitude="-3" metricUnit="m" metricValue="4.9" pageId="12" pageNumber="343" unit="mm" value="4.9">4.9 mm</quantity>
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||
.
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||
</paragraph>
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||
<paragraph id="8BCC643DFFA9FFE6FF0DBF5B5C2CA3CB" blockId="12.[151,644,617,1690]" lastBlockId="12.[680,1172,155,699]" pageId="12" pageNumber="343">Head elongated, convex in the anterior part and almost flat posteriorly, slightly restricted behind the eyes, surface finely punctated. Eyes large and prominent, subelliptical, inserted laterally to the head. Rostrum elongated and robust. Mandibles falciform. Maxillary palps 4 segmented, last palpomere securiform with rounded apex. Labial palps 3 segmented, last palpomere stout and slightly securiform.Antennae 11 segmented, short, filiform with the central antennomeres enlarged, reaching almost the half of elytra, insertion of the antennae close to the eyes and in their lower part; scape robust, elongated, clubshaped; antennomere II very short, moniliformtriangular; antennomere III moniliformglobular, slightly irregular; antennomeres IV– Velongated, robust; antennomeresVI–X filiform, less sturdier than previous ones; antennomere XI filiform, rounded at apex; all antennomeres with not erected and very short setae. Pronotum elongated, slightly narrower than head, anterior margin slightly undulated, posterior margin strongly undulated and slightly bordered, sides straight, surface irregular with a central convex draft. Scutellum triangular. Elytra wider than pronotum, parallelsided, rounded at apex, rather elongated and revealing the last two abdominal segments, surface without punctation, equipped with few little and thin striae. Metathoracic wings transparent, folded, very long, surpassing the elytra and the last tergites. Metasternum large, stout, subquadrate; sternites wide, transverse. Last urite with two asymmetrical apical lobes in a kind of fish tail, one of which is curved and very long with the base slightly enlarged and the center thinner and newly enlarged at apex, while the other one is very short and sturdier. Legs robust, long, with pubescence; coxae elongated and robust; trochanters slightly elongated, globularshaped with apex rounded; femora robust and slightly curved, sturdier and longer than tibiae; tibiae cylindrical, thin, without spurs at apex. Tarsal formula 5 5 5; first tarsomere elongated, about 1.2 times longer than second; second tarsomere thin at base and enlarged apically; third tarsomere triangular and shorter than second; fourth tarsomere bilobed with the lobes very long and wide; fifth tarsomere elongated with wider sides; claws simple without denticles or lobes.</paragraph>
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||
</subSubSection>
|
||
<subSubSection id="C36937B6FFA9FFE6FD32BFE95C32A2FB" pageId="12" pageNumber="343" type="etymology">
|
||
<paragraph id="8BCC643DFFA9FFE6FD32BFE95DB4A243" blockId="12.[680,1172,731,819]" pageId="12" pageNumber="343">
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||
<emphasis id="B907B82FFFA9FFE6FD32BFE95C6FA383" bold="true" box="[680,809,731,755]" pageId="12" pageNumber="343">Etymology.</emphasis>
|
||
Named in honor of Vincenzo Di Taddeo (Avigliana,
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<collectingCountry id="F36424ADFFA9FFE6FCE8BFCE5CE3A263" box="[882,933,764,787]" name="Italy" pageId="12" pageNumber="343">Italy</collectingCountry>
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||
), dear friend of the first author.
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</paragraph>
|
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<paragraph id="8BCC643DFFA9FFE6FD32BE615C32A2FB" blockId="12.[680,1172,851,907]" pageId="12" pageNumber="343">
|
||
<emphasis id="B907B82FFFA9FFE6FD32BE615C05A21B" bold="true" box="[680,835,851,875]" pageId="12" pageNumber="343">Syninclusions.</emphasis>
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Air bubbles (few), detritus (many botanical remains).
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</paragraph>
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</subSubSection>
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<subSubSection id="C36937B6FFA9FFE6FD32BE995CC7A534" pageId="12" pageNumber="343" type="discussion">
|
||
<paragraph id="8BCC643DFFA9FFE6FD32BE995CC7A534" blockId="12.[680,1172,939,1092]" pageId="12" pageNumber="343">
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<emphasis id="B907B82FFFA9FFE6FD32BE995C51A2B3" bold="true" box="[680,791,939,963]" pageId="12" pageNumber="343">Remarks.</emphasis>
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Piece of clear amber that measures 13x12x
|
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<quantity id="4C8BC9D8FFA9FFE6FD63BEF95C71A292" box="[761,823,971,994]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="12" pageNumber="343" unit="mm" value="3.0">3 mm</quantity>
|
||
. The inclusion is complete except for the missing right metathoracic leg. Furthermore, the right antenna is preserved until the eighth antennomere.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |