156 lines
18 KiB
XML
156 lines
18 KiB
XML
<document id="A16B6D20A3D67466701F92B5E1A75325" ID-DOI="10.5281/zenodo.13203160" ID-ISSN="1407-8619" ID-Zenodo-Dep="13203160" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722657382176" checkinUser="felipe" docAuthor="Fanti, Fabrizio & Müller, Patrick" docDate="2022" docId="03DAD52BFFABFFFAFD32BCF85D6BA46D" docLanguage="en" docName="BalticJColeopterol.22.2.331-380.pdf" docOrigin="Baltic Journal of Coleopterology 22 (2)" docStyle="DocumentStyle:9720869F1FB170B2A509257D1FA394CD.2:BalticJColeop.2022.journal_article" docStyleId="9720869F1FB170B2A509257D1FA394CD" docStyleName="BalticJColeop.2022.journal_article" docStyleVersion="2" docTitle="Sanaungulus dunlopi Fanti & Müller, 2022, sp. nov." docType="treatment" docVersion="2" lastPageNumber="347" masterDocId="FFE3AD53FFA5FFEAFF9ABD325F46A170" masterDocTitle="Fossil Cantharidae from the Cretaceous Burmese (Kachin) amber of the Patrick Müller collection, and taxonomic information" masterLastPageNumber="380" masterPageNumber="331" pageNumber="345" updateTime="1722697113453" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-4.0">
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<mods:mods id="18C31C95BBE6F245719DFEF34D6768C6" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="63FDE99D911B6E818C197507E7B580BA">
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<mods:title id="DAA4337F90F34A86F674AF798594D089">Fossil Cantharidae from the Cretaceous Burmese (Kachin) amber of the Patrick Müller collection, and taxonomic information</mods:title>
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</mods:titleInfo>
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<mods:name id="2A138B515DEC85A8440496F1FD555FE1" type="personal">
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<mods:role id="91E3173D86DCE705E1CFF1F9815AC375">
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<mods:roleTerm id="9B8D71680340970DAF83CD5C9EBB86AF">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="75603E399800454D0B53A96897693282">Fanti, Fabrizio</mods:namePart>
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</mods:name>
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<mods:name id="490BC0D8923E14969C39F79A7D7E9103" type="personal">
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<mods:roleTerm id="9AF8A943A938BD9818EFF0E5F5D15AEE">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="783B0140CC8517B7C395507A2FCA34D1">Müller, Patrick</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="D85BA04D4FFC351F50B32B9E7D58DDE9">text</mods:typeOfResource>
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<mods:titleInfo id="BE65585CDC9602CFBF0ECB35BCAB0DE1">
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<mods:title id="D4C882E88563333781F65962258C87D2">Baltic Journal of Coleopterology</mods:title>
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</mods:titleInfo>
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<mods:part id="61CC37DAC172CB5C062B51CC4476531C">
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<mods:date id="39032E560F3B6B0C2888D0D3EF53B9A2">2022</mods:date>
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<mods:detail id="52E355E365DCCD8AF01036870F3D7DEA" type="volume">
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<mods:number id="A7B4997E21AD82E361202415B88D0E3D">22</mods:number>
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</mods:detail>
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<mods:detail id="3D7A416669EFF0CF63D1F99A40C95131" type="issue">
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<mods:number id="4C5F04C81A87495008478E29E9552395">2</mods:number>
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</mods:detail>
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<mods:extent id="7D7E17BBF6DDF28257D18B02D43600C4" unit="page">
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<mods:start id="3012E52FAD4810680E222A4F9E6E97BC">331</mods:start>
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<mods:end id="6165AFB6FCA37E9C09FD18F429A3409D">380</mods:end>
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<mods:classification id="4C0D07661FEC8D2C0116E22AD5265F5D">journal article</mods:classification>
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<mods:identifier id="1350DE1E7E1D411307129B06A48AC1D8" type="DOI">10.5281/zenodo.13203160</mods:identifier>
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<mods:identifier id="D7AF06F47E8CAE0CC3E57B22985DF08A" type="ISSN">1407-8619</mods:identifier>
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<mods:identifier id="620318A44C813CC1C9E4AF02CBB93A93" type="Zenodo-Dep">13203160</mods:identifier>
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</mods:mods>
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<treatment id="03DAD52BFFABFFFAFD32BCF85D6BA46D" LSID="urn:lsid:plazi:treatment:03DAD52BFFABFFFAFD32BCF85D6BA46D" httpUri="http://treatment.plazi.org/id/03DAD52BFFABFFFAFD32BCF85D6BA46D" lastPageId="16" lastPageNumber="347" pageId="14" pageNumber="345">
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<subSubSection id="C36937B6FFABFFE4FD32BCF85CB4A092" box="[680,1010,458,482]" pageId="14" pageNumber="345" type="nomenclature">
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<paragraph id="8BCC643DFFABFFE4FD32BCF85CB4A092" blockId="14.[680,1010,458,514]" box="[680,1010,458,482]" pageId="14" pageNumber="345">
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<heading id="D084D351FFABFFE4FD32BCF85CB4A092" bold="true" box="[680,1010,458,482]" fontSize="10" level="2" pageId="14" pageNumber="345" reason="4">
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<emphasis id="B907B82FFFABFFE4FD32BCF85CB4A092" bold="true" box="[680,1010,458,482]" pageId="14" pageNumber="345">
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<taxonomicName id="4C731FBEFFABFFE4FD32BCF85CD3A092" authorityName="Fanti & Müller" authorityYear="2022" box="[680,917,458,482]" class="Insecta" family="Cantharidae" genus="Sanaungulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="345" phylum="Arthropoda" rank="species" species="dunlopi" status="sp. nov.">
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<emphasis id="B907B82FFFABFFE4FD32BCF85CD3A092" bold="true" box="[680,917,458,482]" italics="true" pageId="14" pageNumber="345">Sanaungulus dunlopi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A2340554FFABFFE4FC07BCF85CB4A092" box="[925,1010,458,482]" pageId="14" pageNumber="345" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C36937B6FFABFFE4FD32BCD95DBEA372" box="[680,760,491,514]" pageId="14" pageNumber="345" type="description">
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<paragraph id="8BCC643DFFABFFE4FD32BCD95DBEA372" blockId="14.[680,1010,458,514]" box="[680,760,491,514]" pageId="14" pageNumber="345">
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(
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<figureCitation id="134878B8FFABFFE4FD2ABCD95DA9A372" box="[688,751,491,514]" captionStart="Fig" captionStartId="15.[113,152,1639,1662]" captionTargetBox="[113,1134,385,1610]" captionTargetId="figure-9@15.[113,1134,385,1610]" captionTargetPageId="15" captionText="Fig. 8. Sanaungulus dunlopi sp. nov. in Burmese amber. A: Holotype, dorsal view; B: Holotype, ventrolateral view." figureDoi="http://doi.org/10.5281/zenodo.13203189" httpUri="https://zenodo.org/record/13203189/files/figure.png" pageId="14" pageNumber="345">Fig. 8</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C36937B6FFABFFE4FD32BF135C81A218" pageId="14" pageNumber="345" type="materials_examined">
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<paragraph id="8BCC643DFFABFFE4FD32BF135B13A3CA" blockId="14.[680,1172,545,698]" pageId="14" pageNumber="345">
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<materialsCitation id="3B1B6E60FFABFFE4FD32BF135B13A3CA" collectingDate="2021-12" collectionCode="BUB" county="Female" location="Catalog" municipality="Burmese" pageId="14" pageNumber="345" specimenCode="BUB3886" specimenCount="1" stateProvince="Kachin" typeStatus="holotype">
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<emphasis id="B907B82FFFABFFE4FD32BF135C52A349" bold="true" box="[680,788,545,569]" pageId="14" pageNumber="345">
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<typeStatus id="54C8DA9FFFABFFE4FD32BF135C49A349" box="[680,783,545,569]" pageId="14" pageNumber="345" type="holotype">Holotype</typeStatus>
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.
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</emphasis>
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<collectingCounty id="62AD1CB1FFABFFE4FC82BF105C21A349" box="[792,871,546,569]" pageId="14" pageNumber="345">Female</collectingCounty>
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, adult specimen in a
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<collectingMunicipality id="6BA8FE47FFABFFE4FBADBF105BD2A349" box="[1079,1172,546,569]" pageId="14" pageNumber="345">Burmese</collectingMunicipality>
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(
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<collectingRegion id="49B7AADFFFABFFE4FD2ABF705C46A329" box="[688,768,578,601]" country="Myanmar" name="Kachin" pageId="14" pageNumber="345">Kachin</collectingRegion>
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) amber piece:
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<location id="8EAC32E6FFABFFE4FC03BF705CAAA329" LSID="urn:lsid:plazi:treatment:03DAD52BFFABFFFAFD32BCF85D6BA46D:8EAC32E6FFABFFE4FC03BF705CAAA329" box="[921,1004,578,601]" county="Female" municipality="Burmese" name="Catalog" pageId="14" pageNumber="345" stateProvince="Kachin">Catalog</location>
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number SNSBBSPG
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<date id="FFCD42FDFFABFFE4FD6EBF505C1EA309" box="[756,856,610,633]" pageId="14" pageNumber="345" value="2021-12">
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<collectingDate id="EF89BB15FFABFFE4FD6EBF505C1EA309" box="[756,856,610,633]" pageId="14" pageNumber="345" value="2021-12">2021 XII</collectingDate>
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</date>
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<quantity id="4C8BC9D8FFABFFE4FCFBBF505CDFA309" box="[865,921,610,633]" metricMagnitude="0" metricUnit="m" metricValue="1.7271999999999998" pageId="14" pageNumber="345" unit="in" value="68.0">68 in</quantity>
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the
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<location id="8EAC32E6FFABFFE4FC56BF505C51A3E9" LSID="urn:lsid:plazi:treatment:03DAD52BFFABFFFAFD32BCF85D6BA46D:8EAC32E6FFABFFE4FC56BF505C51A3E9" county="Female" municipality="Burmese" name="Bayerische Staatssammlung" pageId="14" pageNumber="345" stateProvince="Kachin">Bayerische Staatssammlung</location>
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für
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<location id="8EAC32E6FFABFFE4FCD9BFB05C90A3E9" LSID="urn:lsid:plazi:treatment:03DAD52BFFABFFFAFD32BCF85D6BA46D:8EAC32E6FFABFFE4FCD9BFB05C90A3E9" box="[835,982,642,665]" county="Female" municipality="Burmese" name="Palaontologie" pageId="14" pageNumber="345" stateProvince="Kachin">Paläontologie</location>
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und
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<location id="8EAC32E6FFABFFE4FB90BFB05B2AA3E9" LSID="urn:lsid:plazi:treatment:03DAD52BFFABFFFAFD32BCF85D6BA46D:8EAC32E6FFABFFE4FB90BFB05B2AA3E9" box="[1034,1132,642,665]" county="Female" municipality="Burmese" name="Geologie" pageId="14" pageNumber="345" stateProvince="Kachin">Geologie</location>
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(ex
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<location id="8EAC32E6FFABFFE4FD32BF915C42A3CA" LSID="urn:lsid:plazi:treatment:03DAD52BFFABFFFAFD32BCF85D6BA46D:8EAC32E6FFABFFE4FD32BF915C42A3CA" box="[680,772,675,698]" county="Female" municipality="Burmese" name="Muller's" pageId="14" pageNumber="345" stateProvince="Kachin">Müller’s</location>
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collection number:
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<specimenCode id="DBD5CC46FFABFFE4FC43BF915B0FA3CA" box="[985,1097,675,698]" collectionCode="BUB" pageId="14" pageNumber="345">BUB3886</specimenCode>
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).
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</materialsCitation>
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</paragraph>
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<paragraph id="8BCC643DFFABFFE4FD32BFEB5CBDA261" blockId="14.[680,1172,729,785]" pageId="14" pageNumber="345">
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<emphasis id="B907B82FFFABFFE4FD32BFEB5C07A381" bold="true" box="[680,833,729,753]" pageId="14" pageNumber="345">Type locality.</emphasis>
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<collectingCountry id="F36424ADFFABFFE4FCD0BFE85CF0A381" box="[842,950,730,753]" name="Myanmar" pageId="14" pageNumber="345">Myanmar</collectingCountry>
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:
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<collectingRegion id="49B7AADFFFABFFE4FC5BBFE85B0DA381" box="[961,1099,730,753]" country="Myanmar" name="Kachin" pageId="14" pageNumber="345">Kachin state</collectingRegion>
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, Myitkyina District, Hukawng Valley.
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</paragraph>
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<paragraph id="8BCC643DFFABFFE4FD32BE025C81A218" blockId="14.[680,1172,816,872]" pageId="14" pageNumber="345">
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<emphasis id="B907B82FFFABFFE4FD32BE025C7AA238" bold="true" box="[680,828,816,840]" pageId="14" pageNumber="345">Type horizon.</emphasis>
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Lowermost Cenomanian (98.79 ± 0.62 Ma), midCretaceous.
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</paragraph>
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</subSubSection>
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<subSubSection id="C36937B6FFABFFE4FD32BEBA5B77A513" pageId="14" pageNumber="345" type="diagnosis">
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<paragraph id="8BCC643DFFABFFE4FD32BEBA5B77A513" blockId="14.[680,1172,904,1123]" pageId="14" pageNumber="345">
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<emphasis id="B907B82FFFABFFE4FD32BEBA5CDBA2D0" bold="true" box="[680,925,904,928]" pageId="14" pageNumber="345">Differential diagnosis.</emphasis>
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The new species is characterized by an elongated pronotum with a strongly rounded anterior margin, and it is equipped with robust antennomeres. The genus
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<taxonomicName id="4C731FBEFFABFFE4FD32B9395C6AA552" authorityName="Fanti, Damgaard & Ellenberger" authorityYear="2018" box="[680,812,1035,1058]" class="Insecta" family="Cantharidae" genus="Sanaungulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="345" phylum="Arthropoda" rank="genus">
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<emphasis id="B907B82FFFABFFE4FD32B9395C6AA552" box="[680,812,1035,1058]" italics="true" pageId="14" pageNumber="345">Sanaungulus</emphasis>
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</taxonomicName>
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is characterized by very short elytra often with deep punctuation arranged in striae, and by a head restricted behind eyes.
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</paragraph>
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</subSubSection>
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<subSubSection id="C36937B6FFABFFFAFD32B9B15D52A289" lastPageId="16" lastPageNumber="347" pageId="14" pageNumber="345" type="description">
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<paragraph id="8BCC643DFFABFFE4FD32B9B15C65A46C" blockId="14.[680,1172,1155,1308]" pageId="14" pageNumber="345">
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<emphasis id="B907B82FFFABFFE4FD32B9B15C72A5EB" bold="true" box="[680,820,1155,1179]" pageId="14" pageNumber="345">Description.</emphasis>
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Adult, winged, slender. Female, defined on the shape of the last abdominal segments, and on the basis of short antennae. Body length about
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<quantity id="4C8BC9D8FFABFFE4FCABB9D75CC7A58C" box="[817,897,1253,1276]" metricMagnitude="-3" metricUnit="m" metricValue="3.2" pageId="14" pageNumber="345" unit="mm" value="3.2">3.2 mm</quantity>
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(the body is curled). Entirely brown.
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</paragraph>
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<paragraph id="8BCC643DFFABFFFAFD32B80E5D52A289" blockId="14.[680,1172,1340,1689]" lastBlockId="16.[151,644,155,1017]" lastPageId="16" lastPageNumber="347" pageId="14" pageNumber="345">Head short, constricted (triangularshaped) behind the eyes, smooth, without punctuation, fitted with few long setae. Eyes very prominent, large, roundish, inserted in the lateral part of the head, interocular dorsal distance about 1.3 times greater than eye diameter. Mandibles elongated, falciform. Maxillary palps 4 segmented, with the last palpomere securiform.Antennae 11 segmented, filiform, robust, short, slightly surpassing the middle of the elytra; scape very elongated and clubshaped; antennomere II short; antennomeres III–VIII subequal in length, very robust, without antennal process, enlarged at the external corner which is rounded; antennomere IX approximately as long as previous ones, slightly narrower; antennomere X narrower than previous one, slightly enlarged at the external corner; antennomere XI eleongated, strongly squared apically; antennomeres pubescent. Pronotum very elongated, very slightly narrower than head, anterior margin strongly rounded and bordered, posterior margin rather straight and slightly bordered, corners rather acute, sides straight and slightly bordered, pronotum disc not flat and very slightly wrinkled with shallow punctuation. Scutellum triangularshaped with pointed apex. Elytra very short which reveals many abdominal segments, wider than pronotum at humery, strongly dehiscent posteriorly, rounded apexes, surface strongly ruguose equipped with several short setae. Metathoracic wings transparent, very short, very slightly surpassing the elytra. Sternum elongated with posterior margin rounded, sternites strongly transverse, robust, equipped with short setae; last tergite short, wide, emarginate at apex; last sternite transverse, robust, short, as long as last tergite, with the posterior margin straight. Legs thin, pro and mesolegs short, metalegs elongated and slender, pubescent; coxae massive; trochanters elongated and rounded apically; femora nearly straight, slightly sturdier than tibiae (expecially the metafemora); tibiae cylindrical, thin, without spurs near the apex, shorter than femora. Tarsal formula 5 5 5; first tarsomere elongated and thin; second tarsomere slightly shorter than first; third tarsomere slightly elongated and triangularshaped; fourth strongly bilobed with the lobes very long; fifth tarsomere elongated, flat and enlarged apically; claws simple, without denticle at the base.</paragraph>
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</subSubSection>
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||
<caption id="DF0C34B5FFAAFFE5FFEBBB555E79A7EE" ID-DOI="http://doi.org/10.5281/zenodo.13203189" ID-Zenodo-Dep="13203189" httpUri="https://zenodo.org/record/13203189/files/figure.png" pageId="15" pageNumber="346" startId="15.[113,152,1639,1662]" targetBox="[113,1134,385,1610]" targetPageId="15" targetType="figure">
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||
<paragraph id="8BCC643DFFAAFFE5FFEBBB555E79A7EE" blockId="15.[113,1134,1639,1694]" pageId="15" pageNumber="346">
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||
Fig. 8.
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<taxonomicName id="4C731FBEFFAAFFE5FF27BB5A5ED9A70F" authorityName="Fanti & Müller" authorityYear="2022" box="[189,415,1640,1663]" class="Insecta" family="Cantharidae" genus="Sanaungulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="346" phylum="Arthropoda" rank="species" species="dunlopi" status="sp. nov.">
|
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<emphasis id="B907B82FFFAAFFE5FF27BB5A5ED9A70F" box="[189,415,1640,1663]" italics="true" pageId="15" pageNumber="346">Sanaungulus dunlopi</emphasis>
|
||
</taxonomicName>
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||
<taxonomicNameLabel id="A2340554FFAAFFE5FE3FBB555EB1A70E" box="[421,503,1639,1662]" pageId="15" pageNumber="346" rank="species">sp. nov.</taxonomicNameLabel>
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in Burmese amber. A: Holotype, dorsal view; B: Holotype, ventrolateral view.
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</paragraph>
|
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</caption>
|
||
<subSubSection id="C36937B6FFB5FFFAFF0DB92A5D59A5D6" pageId="16" pageNumber="347" type="etymology">
|
||
<paragraph id="8BCC643DFFB5FFFAFF0DB92A5DC2A500" blockId="16.[151,644,1048,1136]" pageId="16" pageNumber="347">
|
||
<emphasis id="B907B82FFFB5FFFAFF0DB92A5E52A540" bold="true" box="[151,276,1048,1072]" pageId="16" pageNumber="347">Etymology.</emphasis>
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||
Named in honor of Jason A. Dunlop (Museum für Naturkunde,
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<collectingRegion id="49B7AADFFFB5FFFAFE3FB90B5EAFA520" box="[421,489,1081,1104]" country="Germany" name="Berlin" pageId="16" pageNumber="347">Berlin</collectingRegion>
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), a wellknown specialist of fossil arachnids and their relatives.
|
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</paragraph>
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<paragraph id="8BCC643DFFB5FFFAFF0DB9BD5D59A5D6" blockId="16.[151,543,1167,1191]" box="[151,543,1167,1191]" pageId="16" pageNumber="347">
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<emphasis id="B907B82FFFB5FFFAFF0DB9BD5E7CA5D7" bold="true" box="[151,314,1167,1191]" pageId="16" pageNumber="347">Syninclusions.</emphasis>
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Air bubbles, detritus.
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</paragraph>
|
||
</subSubSection>
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<subSubSection id="C36937B6FFB5FFFAFF0DB9F75D6BA46D" pageId="16" pageNumber="347" type="discussion">
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<paragraph id="8BCC643DFFB5FFFAFF0DB9F75D6BA46D" blockId="16.[151,643,1221,1309]" pageId="16" pageNumber="347">
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<emphasis id="B907B82FFFB5FFFAFF0DB9F75E41A5AD" bold="true" box="[151,263,1221,1245]" pageId="16" pageNumber="347">Remarks.</emphasis>
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Piece of clear, golden amber that measures 10x9x
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<quantity id="4C8BC9D8FFB5FFFAFEDCB9D45ED9A58D" box="[326,415,1254,1277]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="16" pageNumber="347" unit="mm" value="3.5">3.5 mm</quantity>
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. In the inclusion the right antenna and a leg are not visible.
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</paragraph>
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</subSubSection>
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</treatment>
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||
</document> |