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<document id="CA77656603043B0063AC6C03C6D8DD7C" ID-CLB-Dataset="5602" ID-DOI="10.11646/zootaxa.3796.2.4" ID-GBIF-Dataset="e0da9034-33f3-4508-aae5-c5dbeb2ee4c3" ID-ISSN="1175-5326" ID-Zenodo-Dep="4915173" ID-ZooBank="D64A7C50-26ED-4D39-B376-2200CCCB138E" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1623198611048" checkinUser="felipe" docAuthor="Badano, Davide &amp; Pantaleoni, Roberto Antonio" docDate="2014" docId="03DA9D6FFF9AFFFD92EBD385FABFF9BF" docLanguage="en" docName="zt03796p319.pdf" docOrigin="Zootaxa 3796 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Puer Lefebvre 1842" docType="treatment" docVersion="6" lastPageNumber="295" masterDocId="FFE3E517FF92FFF5927CD037FFC1FFC9" masterDocTitle="The Larvae of European Ascalaphidae (Neuroptera)" masterLastPageNumber="319" masterPageNumber="287" pageNumber="295" updateTime="1699006410788" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="B56024F9E2C95B54865015524546B17C">The Larvae of European Ascalaphidae (Neuroptera)</mods:title>
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<mods:namePart id="AE092B87CB203CFE2488C75B8B53875E">Badano, Davide</mods:namePart>
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<mods:namePart id="724E269E97F3044F2333B8CDD03F73CE">Pantaleoni, Roberto Antonio</mods:namePart>
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<treatment id="03DA9D6FFF9AFFFD92EBD385FABFF9BF" ID-DOI="http://doi.org/10.5281/zenodo.4915183" ID-GBIF-Taxon="183403924" ID-Zenodo-Dep="4915183" LSID="urn:lsid:plazi:treatment:03DA9D6FFF9AFFFD92EBD385FABFF9BF" httpUri="http://treatment.plazi.org/id/03DA9D6FFF9AFFFD92EBD385FABFF9BF" lastPageNumber="295" pageId="8" pageNumber="295">
<subSubSection id="C3697FF2FF9AFFFD92EBD385FE53FC04" box="[151,402,946,973]" pageId="8" pageNumber="295" type="nomenclature">
<paragraph id="8BCC2C79FF9AFFFD92EBD385FE53FC04" blockId="8.[151,402,946,973]" box="[151,402,946,973]" pageId="8" pageNumber="295">
<heading id="D0849B15FF9AFFFD92EBD385FE53FC04" bold="true" box="[151,402,946,973]" fontSize="11" level="1" pageId="8" pageNumber="295" reason="1">
<taxonomicName id="4C7357FAFF9AFFFD92EBD385FE53FC04" ID-CoL="75BD" authority="Lefebvre, 1842" authorityName="Lefebvre" authorityYear="1842" box="[151,402,946,973]" class="Insecta" family="Ascalaphidae" genus="Puer" kingdom="Animalia" order="Neuroptera" pageId="8" pageNumber="295" phylum="Arthropoda" rank="genus">
<emphasis id="B907F06BFF9AFFFD92EBD385FE53FC04" bold="true" box="[151,402,946,973]" pageId="8" pageNumber="295">
<emphasis id="B907F06BFF9AFFFD92EBD385FF13FC05" bold="true" box="[151,210,946,972]" italics="true" pageId="8" pageNumber="295">Puer</emphasis>
Lefèbvre, 1842
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3697FF2FF9AFFFD92EBD3CEFD25FA2F" pageId="8" pageNumber="295" type="diagnosis">
<paragraph id="8BCC2C79FF9AFFFD92EBD3CEFDE3FA08" blockId="8.[151,1440,1017,1654]" pageId="8" pageNumber="295">
<emphasis id="B907F06BFF9AFFFD92EBD3CEFED0FBDB" bold="true" box="[151,273,1017,1042]" inLineHeading="true" pageId="8" pageNumber="295" reason="1">Diagnosis.</emphasis>
Head capsule one and a half times wider than long; antennae thin, longer than the ocular tubercles; ocular tubercles small, flattened, wider than long and forward directed; opening gap of the jaws larger than 180°; mandibles closely approximated at the insertion; mandibles narrow, armed with three pairs of teeth and with numerous pseudo-teeth; median tooth larger than the other teeth and closer to the apical tooth; base of the mandible covered with hair-like dolichasters; labial palpi four-articulated, segments 24 as long as the basal width of the mandible, last segment swollen; thorax provided with elongated and flattened scolus-like processes; mesothorax bearing two pairs of scolus-like processes of which the anterior ones are bent at 90°; metathorax with an anterior pair of large scolus-like processes and a posterior pair of very short tubercle-like processes; first pair of abdominal spiracles placed on the dorsal side, the following seven pairs on ventral side; abdomen with eight pairs of dorsal scolus-like setiferous processes, ventral series absent except a pair of tubercle-like setiferous processes on the first abdominal segment; VIII sternite without odontoid processes; IX sternite with two short rastra each bearing three digging setae (
<figureCitation id="134830FCFF9AFFFD9347D5B2FE52FA57" box="[315,403,1413,1438]" captionStart="FIGURE 2" captionStartId="5.[151,249,1866,1888]" captionTargetBox="[214,1374,688,1811]" captionTargetId="figure-236@5.[204,1382,673,1817]" captionTargetPageId="5" captionText="FIGURE 2. VIII and IX abdominal sternites of 3rd instar larva of Ascalaphidae. A, Ascalaphus festivus; B, Bubopsis agrionoides; C, Puer maculatus; D, Deleproctophylla australis; E, Libelloides macaronius. Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4915181" httpUri="https://zenodo.org/record/4915181/files/figure.png" pageId="8" pageNumber="295">Fig. 2C</figureCitation>
); body thickly covered with short, pale dolichasters (
<figureCitation id="134830FCFF9AFFFD9199D5B2FBFCFA57" box="[997,1085,1413,1438]" captionStart="FIGURE 3" captionStartId="6.[151,249,1300,1322]" captionTargetBox="[194,1362,201,1269]" captionTargetId="figure-247@6.[184,1403,193,1276]" captionTargetPageId="6" captionText="FIGURE 3. Chaetotaxy of the VIII abdominal sternite of 3rd instar larva of Ascalaphidae. A, Ascalaphus festivus; B, Bubopsis agrionoides; C, Puer maculatus; D, Deleproctophylla australis; E, Libelloides longicornis. Scale bar: 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4915185" httpUri="https://zenodo.org/record/4915185/files/figure.png" pageId="8" pageNumber="295">Fig. 3C</figureCitation>
); long and thin dolichasters are present on the setiferous processes.
</paragraph>
<paragraph id="8BCC2C79FF9AFFFD92BBD5FAFD25FA2F" blockId="8.[151,1440,1017,1654]" box="[199,740,1485,1510]" pageId="8" pageNumber="295">
<emphasis id="B907F06BFF9AFFFD92BBD5FAFE60FA2F" bold="true" box="[199,417,1485,1510]" inLineHeading="true" pageId="8" pageNumber="295" reason="1">Examined species.</emphasis>
<taxonomicName id="4C7357FAFF9AFFFD93DBD5F9FD25FA2F" authority="(Olivier, 1789)" baseAuthorityName="Olivier" baseAuthorityYear="1790" box="[423,740,1485,1510]" class="Insecta" family="Ascalaphidae" genus="Puer" kingdom="Animalia" order="Neuroptera" pageId="8" pageNumber="295" phylum="Arthropoda" rank="species" species="maculatus">
<emphasis id="B907F06BFF9AFFFD93DBD5F9FDF7FA2F" box="[423,566,1485,1510]" italics="true" pageId="8" pageNumber="295">P. maculatus</emphasis>
(Olivier, 1789)
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="C3697FF2FF9AFFFD92BBD5C6FABFF9BF" pageId="8" pageNumber="295" type="discussion">
<paragraph id="8BCC2C79FF9AFFFD92BBD5C6FABFF9BF" blockId="8.[151,1440,1017,1654]" pageId="8" pageNumber="295">
<emphasis id="B907F06BFF9AFFFD92BBD5C6FE8EF9C3" bold="true" box="[199,335,1521,1546]" inLineHeading="true" pageId="8" pageNumber="295" reason="1">Comments.</emphasis>
The genus
<taxonomicName id="4C7357FAFF9AFFFD939ED5C5FDD9F9C0" authorityName="Lefebvre" authorityYear="1842" box="[482,536,1522,1545]" class="Insecta" family="Ascalaphidae" genus="Puer" kingdom="Animalia" order="Neuroptera" pageId="8" pageNumber="295" phylum="Arthropoda" rank="genus">
<emphasis id="B907F06BFF9AFFFD939ED5C5FDD9F9C0" box="[482,536,1522,1545]" italics="true" pageId="8" pageNumber="295">Puer</emphasis>
</taxonomicName>
is exclusive of the Western Palaearctic Region, comprising two species:
<taxonomicName id="4C7357FAFF9AFFFD97FBD5C5FECBF9E7" baseAuthorityName="Olivier" baseAuthorityYear="1790" class="Insecta" family="Ascalaphidae" genus="Puer" kingdom="Animalia" order="Neuroptera" pageId="8" pageNumber="295" phylum="Arthropoda" rank="species" species="maculatus">
<emphasis id="B907F06BFF9AFFFD97FBD5C5FECBF9E7" italics="true" pageId="8" pageNumber="295">P. maculatus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C7357FAFF9AFFFD933CD621FD71F9E7" authority="van der Weele, 1909" authorityName="van der Weele" authorityYear="1909" box="[320,688,1557,1582]" class="Insecta" family="Ascalaphidae" genus="Puer" kingdom="Animalia" order="Neuroptera" pageId="8" pageNumber="295" phylum="Arthropoda" rank="species" species="algericus">
<emphasis id="B907F06BFF9AFFFD933CD621FE05F9E7" box="[320,452,1557,1582]" italics="true" pageId="8" pageNumber="295">P. algericus</emphasis>
<bibRefCitation id="EFE25188FF9AFFFD93B7D621FD71F9E7" author="van der Weele, H. W." box="[459,688,1557,1582]" pageId="8" pageNumber="295" pagination="31 - 32" refId="ref17602" refString="van der Weele, H. W. (1909) Ascalaphiden monographisch bearbeitet. In: Collections Zoologiques du Baron Edm. de Selys Longchamps, Catalogue Systematique et Descriptif, 1908 8, 1 - 326. [date of distr.: 5. I. 1909 cf. Cowley, J. (1937) The dates of distribution of the fascicles of the &quot; Catalogue des collections de Selys Longchamps. Journal of the Society for the Bibliography of Natural History, 1, 31 - 32.]" type="journal article" year="1909">van der Weele, 1909</bibRefCitation>
</taxonomicName>
reported for South-Western Europe, North Africa and Middle East (U. Aspöck &amp; H.
<bibRefCitation id="EFE25188FF9AFFFD9318D60EFE3CF99B" author="Aspock, U. &amp; Aspock, H." box="[356,509,1593,1618]" pageId="8" pageNumber="295" pagination="5 - 11" refId="ref15533" refString="Aspock, U. &amp; Aspock, H. (1987) Wiederentdeckung von Puer maculatus (Olivier) in Europa (Neuropteroidea: Planipennia: Ascalaphidae). Zeitschrift der Arbeitsgemeinschaft Osterreichischer Entomologen, 39, 5 - 11." type="journal article" year="1987">Aspöck 1987</bibRefCitation>
; H.
<bibRefCitation id="EFE25188FF9AFFFD9052D60EFCC8F99B" author="Aspock, H. &amp; Holzel, H. &amp; Aspock, U." box="[558,777,1593,1618]" pageId="8" pageNumber="295" pagination="1 - 606" refId="ref15410" refString="Aspock, H., Holzel, H. &amp; Aspock, U. (2001) Kommentierter Katalog der Neuropterida (Insecta: Raphidioptera, Megaloptera, Neuroptera) der Westpalaarktis. Denisia, 2, 1 - 606." type="journal article" year="2001">
Aspöck
<emphasis id="B907F06BFF9AFFFD90F0D60DFD04F99B" box="[652,709,1593,1618]" italics="true" pageId="8" pageNumber="295">et al.</emphasis>
2001
</bibRefCitation>
;
<bibRefCitation id="EFE25188FF9AFFFD916AD60EFB8FF99B" author="Badano, D. &amp; Pantaleoni, R. A." box="[790,1102,1593,1618]" pageId="8" pageNumber="295" pagination="51 - 57" refId="ref15724" refString="Badano, D. &amp; Pantaleoni, R. A. (2012) Agadirius trojani gen. et sp. nov.: a new owlfly (Neuroptera: Ascalaphidae) from Morocco. Zootaxa, 3270, 51 - 57." type="journal article" year="2012">Badano &amp; Pantaleoni 2012</bibRefCitation>
). The larva of this genus was exclusively treated by
<bibRefCitation id="EFE25188FF9AFFFD93E8D669FDEFF9BF" author="Hagen, H. A." box="[404,558,1629,1654]" pageId="8" pageNumber="295" pagination="33 - 62" refId="ref16310" refString="Hagen, H. A. (1873) Die Larven von Ascalaphus. Stettiner Entomologische Zeitung, 34, 33 - 62." type="journal article" year="1873">Hagen (1873)</bibRefCitation>
thought the validity of his account was questioned (H.
<bibRefCitation id="EFE25188FF9AFFFD96E6D66AFAB0F9BF" author="Aspock, H. &amp; Aspock, U. &amp; Holzel, H." box="[1178,1393,1629,1654]" pageId="8" pageNumber="295" refId="ref15298" refString="Aspock, H., Aspock, U. &amp; Holzel, H. (1980) Die Neuropteren Europas. Eine zusammenfassende Darstellung der Systematik, Okologie und Chorologie der Neuropteroidea (Megaloptera, Raphidioptera, Planipennia) Europas. Goecke and Evers, Krefeld, Germany, 495 + 355 pp." type="book" year="1980">
Aspöck
<emphasis id="B907F06BFF9AFFFD968AD669FAEEF9BF" box="[1270,1327,1629,1654]" italics="true" pageId="8" pageNumber="295">et al.</emphasis>
1980
</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
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