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<document ID-DOI="10.1515/vzoo-2016-0005" ID-ISSN="2073-2333" ID-Zenodo-Dep="6453486" approvalRequired="5" approvalRequired_for_taxonomicNames="3" approvalRequired_for_textStreams="2" checkinTime="1649779108492" checkinUser="felipe" docAuthor="Korneyev, V. A." docDate="2016" docId="03DA9357B63DFF87CDD1FA3FFE1B1573" docLanguage="en" docName="VestnikZoologii.50.1.39-48.pdf" docOrigin="Vestnik Zoologii 50 (1)" docSource="http://dx.doi.org/10.1515/vzoo-2016-0005" docStyle="DocumentStyle:9FE6554C034D0A4D8EA9CEAE255F4368.2:VestnikZoologii.2014-2019.journal_article" docStyleId="9FE6554C034D0A4D8EA9CEAE255F4368" docStyleName="VestnikZoologii.2014-2019.journal_article" docStyleVersion="2" docTitle="Toxopyrgotini Korneyev 2016, trib. n." docType="treatment" docVersion="2" lastPageId="6" lastPageNumber="45" masterDocId="FFE3EB2FB638FF81CD7FFF8CFFDC1279" masterDocTitle="New Taxa And Synonymy In The Family Pyrgotidae (Diptera, Tephritoidea). I. Tribes Prodalmanniini And Toxopyrgotini" masterLastPageNumber="48" masterPageNumber="39" pageId="5" pageNumber="44" updateTime="1649779555460" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>New Taxa And Synonymy In The Family Pyrgotidae (Diptera, Tephritoidea). I. Tribes Prodalmanniini And Toxopyrgotini</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Korneyev, V. A.</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
<mods:title>Vestnik Zoologii</mods:title>
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<mods:part>
<mods:date>2016</mods:date>
<mods:detail type="pubDate">
<mods:number>2016-02-01</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>50</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>1</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>39</mods:start>
<mods:end>48</mods:end>
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<mods:url>http://dx.doi.org/10.1515/vzoo-2016-0005</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.1515/vzoo-2016-0005</mods:identifier>
<mods:identifier type="ISSN">2073-2333</mods:identifier>
<mods:identifier type="Zenodo-Dep">6453486</mods:identifier>
</mods:mods>
<treatment LSID="urn:lsid:plazi:treatment:03DA9357B63DFF87CDD1FA3FFE1B1573" httpUri="http://treatment.plazi.org/id/03DA9357B63DFF87CDD1FA3FFE1B1573" lastPageId="6" lastPageNumber="45" pageId="5" pageNumber="44">
<subSubSection box="[174,517,1459,1486]" pageId="5" pageNumber="44" type="nomenclature">
<paragraph blockId="5.[174,517,1459,1486]" box="[174,517,1459,1486]" pageId="5" pageNumber="44">
<heading box="[174,517,1459,1486]" fontSize="11" level="2" pageId="5" pageNumber="44" reason="4">
<emphasis bold="true" box="[174,517,1459,1486]" pageId="5" pageNumber="44">
Tribe
<taxonomicName authority="Korneyev, 2016" authorityName="Korneyev" authorityYear="2016" box="[248,427,1459,1485]" class="Insecta" family="Pyrgotidae" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="44" phylum="Arthropoda" rank="tribe" status="trib. n." tribe="Toxopyrgotini">Toxopyrgotini</taxonomicName>
<taxonomicNameLabel box="[433,517,1459,1486]" pageId="5" pageNumber="44" rank="tribe">trib. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection lastPageId="6" lastPageNumber="45" pageId="5" pageNumber="44" type="materials_examined">
<paragraph blockId="5.[174,638,1491,1518]" box="[174,638,1491,1518]" pageId="5" pageNumber="44">
<typeStatus box="[174,233,1492,1518]" pageId="5" pageNumber="44">Type</typeStatus>
genus:
<taxonomicName authority="Hendel, 1914" authorityName="Hendel" authorityYear="1914" box="[322,634,1491,1518]" class="Insecta" family="Pyrgotidae" genus="Toxopyrgota" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="44" phylum="Arthropoda" rank="genus">
<emphasis box="[322,469,1492,1517]" italics="true" pageId="5" pageNumber="44">Toxopyrgota</emphasis>
<bibRefCitation author="Hendel, F." box="[475,634,1491,1518]" pageId="5" pageNumber="44" pagination="77 - 117" refId="ref4315" refString="Hendel, F. 1914. Neue Beitrage zur Kenntnis der Pyrgotinen. Archiv fur Naturgeschichte (1913), 79 A (11), 77 - 117 + 1 Taf." type="journal article" year="1914">Hendel, 1914</bibRefCitation>
</taxonomicName>
.
</paragraph>
<paragraph blockId="5.[174,1233,1539,1854]" pageId="5" pageNumber="44">
D i a g n o s i s. Ocelli absent; frontal setae present; antennae very short, pedicel with deep incision (P); face with short, partly fused antennal grooves; epistome not developed: subcranial membrane between face and clypeus very large, but non-sclerotized (P); wing with subcostal vein entire and reaching costal vein at acute angle (?A); costal vein with humeral and subcostal break (P); cell cup closed by arcuate crossvein without posteroapical lobe (A); abdomen with sternites 1 and 2 fused (
<collectionCode box="[738,775,1700,1726]" country="France" name="Museum national d'Histoire Naturelle, Laboratiore de Paleontologie" pageId="5" pageNumber="44">SA</collectionCode>
with other
<taxonomicName box="[910,1042,1699,1726]" class="Insecta" family="Pyrgotidae" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="40" phylum="Arthropoda" rank="subFamily" subFamily="Pyrgotinae">Pyrgotinae</taxonomicName>
); male genitalia: phallic guide not developed (P); epandrium densely setose dorsally (P); inner surstylus with at least one prensiseta (thickened seta) (P); female: oviscape unmodified; eversible membrane without taeniae (
<collectionCode box="[444,481,1796,1822]" country="France" name="Museum national d'Histoire Naturelle, Laboratiore de Paleontologie" pageId="5" pageNumber="44">SA</collectionCode>
with Pyrgotini and Toxurini); aculeus small, as long as oviscape width at aperture (
<collectionCode box="[388,425,1828,1854]" country="France" name="Museum national d'Histoire Naturelle, Laboratiore de Paleontologie" pageId="5" pageNumber="44">SA</collectionCode>
with Pyrgotini and Toxurini), with sternite 8 inconspicuous (
<collectionCode box="[1137,1174,1828,1854]" country="France" name="Museum national d'Histoire Naturelle, Laboratiore de Paleontologie" pageId="5" pageNumber="44">SA</collectionCode>
with
</paragraph>
<paragraph blockId="6.[166,1225,1159,1235]" pageId="6" pageNumber="45">
Fig. 3036.
<taxonomicName authorityName="Hendel" authorityYear="1914" box="[283,500,1160,1182]" class="Insecta" family="Pyrgotidae" genus="Toxopyrgota" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="45" phylum="Arthropoda" rank="species" species="inclinata">
<emphasis box="[283,500,1160,1182]" italics="true" pageId="6" pageNumber="45">Toxopyrgota inclinata</emphasis>
</taxonomicName>
,
<specimenCount box="[512,527,1159,1182]" pageId="6" pageNumber="45" type="female"></specimenCount>
(30, 35, 36) and Ò (3134): 30, 31 — habitus, left; 32, 33 — head, left and anterior view; 34 — Ò mid coxa, hind coxa and hind trochanter; 35 — head and thorax, dorsal view; 36 — wing.
</paragraph>
<paragraph blockId="6.[166,1225,1296,1802]" pageId="6" pageNumber="45">
Pyrgotini and Toxurini), without anal slit (
<collectionCode box="[665,702,1296,1322]" country="France" name="Museum national d'Histoire Naturelle, Laboratiore de Paleontologie" pageId="6" pageNumber="45">SA</collectionCode>
with Pyrgotini and Toxurini); ventral receptacle scepter-like (
<collectionCode box="[378,415,1328,1354]" country="France" name="Museum national d'Histoire Naturelle, Laboratiore de Paleontologie" pageId="6" pageNumber="45">SA</collectionCode>
with other
<taxonomicName box="[550,683,1328,1355]" class="Insecta" family="Pyrgotidae" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="40" phylum="Arthropoda" rank="subFamily" subFamily="Pyrgotinae">Pyrgotinae</taxonomicName>
).
</paragraph>
<paragraph blockId="6.[166,1225,1296,1802]" pageId="6" pageNumber="45">
<materialsCitation location="Afrotropical Region" pageId="6" pageNumber="45" specimenCount="1">
This tribe is represented by its
<typeStatus box="[586,636,1361,1387]" pageId="6" pageNumber="45">type</typeStatus>
genus,
<taxonomicName authority="Hendel, 1914" authorityName="Hendel" authorityYear="1914" box="[727,1039,1360,1386]" class="Insecta" family="Pyrgotidae" genus="Toxopyrgota" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="45" phylum="Arthropoda" rank="genus">
<emphasis box="[727,874,1361,1386]" italics="true" pageId="6" pageNumber="45">Toxopyrgota</emphasis>
<bibRefCitation author="Hendel, F." box="[882,1039,1360,1386]" pageId="6" pageNumber="45" pagination="77 - 117" refId="ref4315" refString="Hendel, F. 1914. Neue Beitrage zur Kenntnis der Pyrgotinen. Archiv fur Naturgeschichte (1913), 79 A (11), 77 - 117 + 1 Taf." type="journal article" year="1914">Hendel, 1914</bibRefCitation>
</taxonomicName>
from the
<location LSID="urn:lsid:plazi:treatment:03DA9357B63DFF87CDD1FA3FFE1B1573:8EAC749AB63EFF87C9F8FADCFE8317F3" name="Afrotropical Region" pageId="6" pageNumber="45">Afrotropical Region</location>
</materialsCitation>
.
</paragraph>
<paragraph blockId="6.[166,1225,1296,1802]" pageId="6" pageNumber="45">
T a x o n o m i c p o s i t i o n.
<taxonomicName authority="Korneyev, 2016" authorityName="Korneyev" authorityYear="2016" box="[551,726,1424,1450]" class="Insecta" family="Pyrgotidae" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="45" phylum="Arthropoda" rank="tribe" status="trib. n." tribe="Toxopyrgotini">Toxopyrgotini</taxonomicName>
share the modified structure of eversible membrane and aculeus (see above
<emphasis bold="true" box="[580,787,1456,1482]" pageId="6" pageNumber="45">synapomorphies</emphasis>
) with Pyrgotini and Toxurini, clearly differing from them by several characters keeping their plesiomorphic mode: antenna with a deep pedicellar incision (notch), membranous area between face and clypeus instead of sclerotized epistome, male genitalia with inner surstylus bearing prensiseta. It is therefore believed to be a sister-group to the Pyrgotini + Toxurini lineage. This is a sufficient cause to establish the new tribe.
</paragraph>
<paragraph blockId="6.[166,1225,1296,1802]" pageId="6" pageNumber="45">
The setose pedicel, large subcranial membrane, vestigial proboscis and cell cup closed by arcuate vein are the reasons to hypothesize possible relationships of the
<taxonomicName authority="Korneyev, 2016" authorityName="Korneyev" authorityYear="2016" box="[1050,1224,1680,1706]" class="Insecta" family="Pyrgotidae" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="45" phylum="Arthropoda" rank="tribe" status="trib. n." tribe="Toxopyrgotini">Toxopyrgotini</taxonomicName>
<taxonomicNameLabel box="[166,249,1712,1739]" pageId="6" pageNumber="45" rank="tribe">trib. n.</taxonomicNameLabel>
with the enigmatic family
<taxonomicName authorityName="Bigot" authorityYear="1882" box="[582,749,1712,1738]" class="Insecta" family="Ctenostylidae" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="45" phylum="Arthropoda" rank="family">Ctenostylidae</taxonomicName>
; however, most of these characters are reductions strongly subject to homoplasy and thus having very small weight for phylogenetic reconstructions.
</paragraph>
</subSubSection>
</treatment>
</document>