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<document id="D6BC454CA4B36E5375E8CCD0CA1D5979" ID-CLB-Dataset="20127" ID-DOI="10.5852/ejt.2022.806.1705" ID-GBIF-Dataset="0c5965bd-8762-46b1-ad73-61d1347db34b" ID-ISSN="2118-9773" ID-Zenodo-Dep="6384569" ID-ZooBank="4991BA30-038F-4B86-983B-E4E738C56759" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1648212006250" checkinUser="felipe" docAuthor="Cassar, Thomas, Mifsud, David &amp; Decae, Arthur E." docDate="2022" docId="03DA934D0A6ED151941A3B24FE63F921" docLanguage="en" docName="ejt-806_90-112.pdf" docOrigin="European Journal of Taxonomy 806 (1)" docSource="http://zoobank.org/4991ba30-038f-4b86-983b-e4e738c56759" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Nemesia maltensis Cassar &amp; Mifsud &amp; Decae 2022, sp. nov." docType="treatment" docUuid="BB791B03-F14E-4D15-9338-EE2E02244C4A" docUuidSource="ZooBank" docVersion="8" lastPageNumber="106" masterDocId="FFE3EB350A65D14196623D1BFFBCFFF3" masterDocTitle="The Nemesia trapdoor spider fauna of the Maltese archipelago, with the description of two new species (Araneae, Mygalomorphae, Nemesiidae)" masterLastPageNumber="112" masterPageNumber="90" pageNumber="101" updateTime="1699318544272" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="63E248D09F56DB2AA12C05345A1EBE87">The Nemesia trapdoor spider fauna of the Maltese archipelago, with the description of two new species (Araneae, Mygalomorphae, Nemesiidae)</mods:title>
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<mods:namePart id="B4B5BC78223E0A09497C5EF09C95BAF7">Cassar, Thomas</mods:namePart>
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<mods:affiliation id="08BFBADA33FFC6C6E085B58DA661A918">Department of Biology, Faculty of Science, University of Malta, Msida MSD 2080, Malta. Institute of Earth Systems, Division of Rural Sciences and Food Systems, University of Malta, Msida MSD 2080, Malta. Natural History Museum Rotterdam, Westzeedijk 345 (Museumpark), 3015 AA Rotterdam, The Netherlands.</mods:affiliation>
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<mods:namePart id="94829E0284DF4F880AE0582C7E8727EA">Mifsud, David</mods:namePart>
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<mods:namePart id="B4C1BA9EAB9DE3F57D81BE4F75EBDC26">Decae, Arthur E.</mods:namePart>
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<taxonomicName id="4C7359D80A6ED14A941A3B24FCE6F9AA" ID-CoL="98ZJF" authority="Cassar &amp; Mifsud &amp; Decae, 2022" authorityName="Cassar &amp; Mifsud &amp; Decae" authorityYear="2022" box="[632,858,1598,1625]" class="Arachnida" family="Nemesiidae" genus="Nemesia" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="101" phylum="Arthropoda" rank="species" species="maltensis" status="sp. nov.">
<emphasis id="B907FE490A6ED14A941A3B24FCE6F9AA" bold="true" box="[632,858,1598,1625]" italics="true" pageId="11" pageNumber="101">Nemesia maltensis</emphasis>
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<taxonomicNameLabel id="A23443320A6ED14A95033B5BFC00F9A9" box="[865,956,1600,1626]" pageId="11" pageNumber="101" rank="species">sp. nov.</taxonomicNameLabel>
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<paragraph id="8BCC225B0A6ED14A94BA3B9EFCE7F953" blockId="11.[370,1219,1598,1696]" box="[728,859,1669,1696]" pageId="11" pageNumber="101">
<figureCitation id="13483EDE0A6ED14A94BA3B9EFCE7F953" box="[728,859,1669,1696]" captionStart-0="Figs 5463" captionStart-1="Figs 6469" captionStart-2="Figs 7073" captionStartId-0="13.[189,242,1739,1765]" captionStartId-1="14.[189,242,1816,1842]" captionStartId-2="15.[189,242,1886,1912]" captionTargetBox-0="[189,1398,965,1691]" captionTargetBox-1="[189,1398,1125,1769]" captionTargetBox-2="[189,1398,1037,1843]" captionTargetId-0="figure-258@13.[189,1398,965,1691]" captionTargetId-1="figure-378@14.[189,1398,1125,1769]" captionTargetId-2="figure-227@15.[189,1398,1037,1843]" captionTargetPageId-0="13" captionTargetPageId-1="14" captionTargetPageId-2="15" captionText-0="Figs 5463. Nemesia maltensis sp. nov. holotype, ♂ (TC.004, NHMR). 54. Dorsal habitus (note distinct dark flanks of cephalic part). 55. Prosoma in lateral view (note dark margin of the carapace and cephalic part not elevated). 56. Dorso-distal right palp (note strong development of PTR). 57. Prolateral left metatarsus and distal tibia leg I (note CF with dense brush of short bristles, short TA and smoothly curved TS). 58. Spinnerets, lateral view showing macula (MAC) on external proximal article of PLS. 5963. Right palpal organ in four different positions rotated clock-wise by 90°. 59. Ventral view. 60. Prolateral view. 61. Dorsal view. 62. Retrolateral view. 63. Embolus tip at high magnification, note short row of tiny denticles (den) and bevelled tip." captionText-1="Figs 6469. Nemesia maltensis sp. nov. paratype, ♀ (TC.012, NHMR). 64. Dorsal habitus, note maculae on external femora I &amp; II (m). 65. Left lateral habitus, note numerous maculae on legs arrow indications (m) and the low carapace profile (lp). 66. Ventral anterior prosoma, note indistinct central division (cd) in labial furrow. 67. Prolateral patella III, note three spines (sp) in a row. 68. Ventral spinnerets, note absence of distinct light coloured spigot fields. 69. Spermathecae, ventral view, note median part of the receptacles being folded and twisted (ft)." captionText-2="Figs 7073. Nemesia maltensis sp. nov. observations. 70. Natural habitat. 71. Borrow top cut open, note virtual absence of a silk lining of the burrow shaft and the compacted soil from which the burrow walls derive most of their structural integrity. 72. Natural burrow entrance flush with the soil surface, note absence of trapdoor. 73. Two surface openings of the bifurcated burrow seen in an observation container." figureDoi-0="http://doi.org/10.5281/zenodo.6384585" figureDoi-1="http://doi.org/10.5281/zenodo.6384587" figureDoi-2="http://doi.org/10.5281/zenodo.6384589" httpUri-0="https://zenodo.org/record/6384585/files/figure.png" httpUri-1="https://zenodo.org/record/6384587/files/figure.png" httpUri-2="https://zenodo.org/record/6384589/files/figure.png" pageId="11" pageNumber="101">Figs 5473</figureCitation>
</paragraph>
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<emphasis id="B907FE490A6ED14A96DF3BDCFE8AF912" bold="true" box="[189,310,1735,1761]" pageId="11" pageNumber="101">Diagnosis</emphasis>
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<paragraph id="8BCC225B0A6ED14D96DF3BEDFEC9FD70" blockId="11.[188,1399,1781,2019]" lastBlockId="12.[188,1399,265,643]" lastPageId="12" lastPageNumber="102" pageId="11" pageNumber="101">
<taxonomicName id="4C7359D80A6ED14A96DF3BEDFE23F8FC" authorityName="Cassar &amp; Mifsud &amp; Decae" authorityYear="2022" box="[189,415,1781,1807]" class="Magnoliopsida" family="Scrophulariaceae" genus="Nemesia" kingdom="Plantae" order="Lamiales" pageId="11" pageNumber="101" phylum="Tracheophyta" rank="species" species="maltensis">
<emphasis id="B907FE490A6ED14A96DF3BEDFE23F8FC" box="[189,415,1781,1807]" italics="true" pageId="11" pageNumber="101">Nemesia maltensis</emphasis>
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groups out with a species-complex that is common in the central and eastern Mediterranean Basin (
<bibRefCitation id="EFE25FAA0A6ED14A97A73A02FDE5F8C0" author="Decae A. E." box="[453,601,1817,1843]" pageId="11" pageNumber="101" pagination="24 - 28" refId="ref12742" refString="Decae A. E. 2012. Geography-related sub-generic diversity within the Mediterranean trapdoor spider genus Nemesia (Araneae, Mygalomorphae, Nemesiidae). Arachnologische Mitteilungen 43: 24 - 28. https: // doi. org / 10.5431 / aramit 4304" type="journal article" year="2012">Decae 2012</bibRefCitation>
). In this species-complex the palpal organ of males has striae (fine ribs) on the proximal embolus (
<figureCitation id="13483EDE0A6ED14A94573A27FD7CF8A4" box="[565,704,1852,1879]" captionStart="Figs 5463" captionStartId="13.[189,242,1739,1765]" captionTargetBox="[189,1398,965,1691]" captionTargetId="figure-258@13.[189,1398,965,1691]" captionTargetPageId="13" captionText="Figs 5463. Nemesia maltensis sp. nov. holotype, ♂ (TC.004, NHMR). 54. Dorsal habitus (note distinct dark flanks of cephalic part). 55. Prosoma in lateral view (note dark margin of the carapace and cephalic part not elevated). 56. Dorso-distal right palp (note strong development of PTR). 57. Prolateral left metatarsus and distal tibia leg I (note CF with dense brush of short bristles, short TA and smoothly curved TS). 58. Spinnerets, lateral view showing macula (MAC) on external proximal article of PLS. 5963. Right palpal organ in four different positions rotated clock-wise by 90°. 59. Ventral view. 60. Prolateral view. 61. Dorsal view. 62. Retrolateral view. 63. Embolus tip at high magnification, note short row of tiny denticles (den) and bevelled tip." figureDoi="http://doi.org/10.5281/zenodo.6384585" httpUri="https://zenodo.org/record/6384585/files/figure.png" pageId="11" pageNumber="101">Figs 5962</figureCitation>
), and females have tube shaped, tripartite spermathecae in which the median part is folded and twisted (
<figureCitation id="13483EDE0A6ED14A94813A44FC80F889" box="[739,828,1887,1914]" captionStart="Figs 6469" captionStartId="14.[189,242,1816,1842]" captionTargetBox="[189,1398,1125,1769]" captionTargetId="figure-378@14.[189,1398,1125,1769]" captionTargetPageId="14" captionText="Figs 6469. Nemesia maltensis sp. nov. paratype, ♀ (TC.012, NHMR). 64. Dorsal habitus, note maculae on external femora I &amp; II (m). 65. Left lateral habitus, note numerous maculae on legs arrow indications (m) and the low carapace profile (lp). 66. Ventral anterior prosoma, note indistinct central division (cd) in labial furrow. 67. Prolateral patella III, note three spines (sp) in a row. 68. Ventral spinnerets, note absence of distinct light coloured spigot fields. 69. Spermathecae, ventral view, note median part of the receptacles being folded and twisted (ft)." figureDoi="http://doi.org/10.5281/zenodo.6384587" httpUri="https://zenodo.org/record/6384587/files/figure.png" pageId="11" pageNumber="101">Fig. 69</figureCitation>
). Within this species-complex,
<taxonomicName id="4C7359D80A6ED14A92D73A7BFAF5F88A" authorityName="Cassar &amp; Mifsud &amp; Decae" authorityYear="2022" box="[1205,1353,1887,1913]" class="Magnoliopsida" family="Scrophulariaceae" genus="Nemesia" kingdom="Plantae" order="Lamiales" pageId="11" pageNumber="101" phylum="Tracheophyta" rank="species" species="maltensis">
<emphasis id="B907FE490A6ED14A92D73A7BFAF5F88A" box="[1205,1353,1887,1913]" italics="true" pageId="11" pageNumber="101">N. maltensis</emphasis>
</taxonomicName>
fits in a central Mediterranean subgroup that has informally been named the
<taxonomicName id="4C7359D80A6ED14A92593A99FB75F86F" authorityName="Ausserer" authorityYear="1871" box="[1083,1225,1922,1948]" class="Magnoliopsida" family="Scrophulariaceae" genus="Nemesia" kingdom="Plantae" order="Lamiales" pageId="11" pageNumber="101" phylum="Tracheophyta" rank="species" species="maculatipes">
<emphasis id="B907FE490A6ED14A92593A99FB75F86F" box="[1083,1225,1922,1948]" italics="true" pageId="11" pageNumber="101">maculatipes</emphasis>
</taxonomicName>
group (
<bibRefCitation id="EFE25FAA0A6ED14A93493A99FE81F833" author="Decae A. &amp; Pantini P. &amp; Isaia M." pageId="11" pageNumber="101" pagination="525 - 540" refId="ref12786" refString="Decae A., Pantini P. &amp; Isaia M. 2015. A new species-complex within the trapdoor spider genus Nemesia Audouin 1826 distributed in northern and central Italy, with descriptions of three new species (Araneae, Mygalomorphae, Nemesiidae). Zootaxa 4059 (3): 525 - 540. https: // doi. org / 10.11646 / zootaxa. 4059.3.5" type="journal article" year="2015">
Decae
<emphasis id="B907FE490A6ED14A96DF3ABDFF4CF84C" box="[189,240,1957,1983]" italics="true" pageId="11" pageNumber="101">et al</emphasis>
. 2015
</bibRefCitation>
). Species in this subgroup, such as
<taxonomicName id="4C7359D80A6ED14A94B13ABDFB0AF833" authority="Ausserer, 1871" authorityName="Ausserer" authorityYear="1871" box="[723,1206,1957,1984]" class="Magnoliopsida" family="Scrophulariaceae" genus="Nemesia" kingdom="Plantae" order="Lamiales" pageId="11" pageNumber="101" phylum="Tracheophyta" rank="species" species="maculatipes">
<emphasis id="B907FE490A6ED14A94B13ABDFC3EF84C" box="[723,898,1957,1983]" italics="true" pageId="11" pageNumber="101">N. maculatipes</emphasis>
Ausserer, 1871 (
<collectingRegion id="49B7ECB90A6ED14A92243ABEFB12F833" box="[1094,1198,1957,1984]" country="Italy" name="Sardegna" pageId="11" pageNumber="101">Sardinia</collectingRegion>
)
</taxonomicName>
,
<taxonomicName id="4C7359D80A6ED14A92A23ABDFEE0F810" authority="(Costa, 1835)" baseAuthorityName="Costa" baseAuthorityYear="1835" class="Magnoliopsida" family="Scrophulariaceae" genus="Nemesia" kingdom="Plantae" order="Lamiales" pageId="11" pageNumber="101" phylum="Tracheophyta" rank="species" species="meridionalis">
<emphasis id="B907FE490A6ED14A92A23ABDFAC9F84C" box="[1216,1397,1957,1983]" italics="true" pageId="11" pageNumber="101">N. meridionalis</emphasis>
(Costa, 1835)
</taxonomicName>
(southern
<collectingCountry id="F36462CB0A6ED14A97BE3AD3FDAAF810" box="[476,534,1992,2019]" name="Italy" pageId="11" pageNumber="101">Italy</collectingCountry>
),
<taxonomicName id="4C7359D80A6ED14A944F3AD2FC39F810" authority="Fage, 1917" authorityName="Fage" authorityYear="1917" box="[557,901,1992,2019]" class="Magnoliopsida" family="Scrophulariaceae" genus="Nemesia" kingdom="Plantae" order="Lamiales" pageId="11" pageNumber="101" phylum="Tracheophyta" rank="species" species="sanzoi">
<emphasis id="B907FE490A6ED14A944F3AD2FD27F811" box="[557,667,1993,2018]" italics="true" pageId="11" pageNumber="101">N. sanzoi</emphasis>
Fage, 1917 (
<collectingRegion id="49B7ECB90A6ED14A95543AD3FCC2F810" box="[822,894,1992,2019]" country="Italy" name="Sicilia" pageId="11" pageNumber="101">Sicily</collectingRegion>
)
</taxonomicName>
and
<taxonomicName id="4C7359D80A6ED14A95A33AD2FACAF810" authority="Herman, 1879" authorityName="Herman" authorityYear="1879" box="[961,1398,1992,2019]" class="Magnoliopsida" family="Scrophulariaceae" genus="Nemesia" kingdom="Plantae" order="Lamiales" pageId="11" pageNumber="101" phylum="Tracheophyta" rank="species" species="pannonica">
<emphasis id="B907FE490A6ED14A95A33AD2FBDCF811" box="[961,1120,1993,2018]" italics="true" pageId="11" pageNumber="101">N. pannonica</emphasis>
Herman, 1879 (
<collectingCountry id="F36462CB0A6ED14A93423AD3FAD2F810" box="[1312,1390,1992,2019]" name="Serbia" pageId="11" pageNumber="101">Serbia</collectingCountry>
)
</taxonomicName>
have conspicuous dark coloured patches (maculae) on legs and/or the external proximal article of the PLS (
<figureCitation id="13483EDE0A69D14D97663C37FEDEFEB4" box="[260,354,300,327]" captionStart="Figs 5463" captionStartId="13.[189,242,1739,1765]" captionTargetBox="[189,1398,965,1691]" captionTargetId="figure-258@13.[189,1398,965,1691]" captionTargetPageId="13" captionText="Figs 5463. Nemesia maltensis sp. nov. holotype, ♂ (TC.004, NHMR). 54. Dorsal habitus (note distinct dark flanks of cephalic part). 55. Prosoma in lateral view (note dark margin of the carapace and cephalic part not elevated). 56. Dorso-distal right palp (note strong development of PTR). 57. Prolateral left metatarsus and distal tibia leg I (note CF with dense brush of short bristles, short TA and smoothly curved TS). 58. Spinnerets, lateral view showing macula (MAC) on external proximal article of PLS. 5963. Right palpal organ in four different positions rotated clock-wise by 90°. 59. Ventral view. 60. Prolateral view. 61. Dorsal view. 62. Retrolateral view. 63. Embolus tip at high magnification, note short row of tiny denticles (den) and bevelled tip." figureDoi="http://doi.org/10.5281/zenodo.6384585" httpUri="https://zenodo.org/record/6384585/files/figure.png" pageId="12" pageNumber="102">Figs 58</figureCitation>
,
<figureCitation id="13483EDE0A69D14D97133C37FE51FEB4" box="[369,493,300,327]" captionStart="Figs 6469" captionStartId="14.[189,242,1816,1842]" captionTargetBox="[189,1398,1125,1769]" captionTargetId="figure-378@14.[189,1398,1125,1769]" captionTargetPageId="14" captionText="Figs 6469. Nemesia maltensis sp. nov. paratype, ♀ (TC.012, NHMR). 64. Dorsal habitus, note maculae on external femora I &amp; II (m). 65. Left lateral habitus, note numerous maculae on legs arrow indications (m) and the low carapace profile (lp). 66. Ventral anterior prosoma, note indistinct central division (cd) in labial furrow. 67. Prolateral patella III, note three spines (sp) in a row. 68. Ventral spinnerets, note absence of distinct light coloured spigot fields. 69. Spermathecae, ventral view, note median part of the receptacles being folded and twisted (ft)." figureDoi="http://doi.org/10.5281/zenodo.6384587" httpUri="https://zenodo.org/record/6384587/files/figure.png" pageId="12" pageNumber="102">6465, 67</figureCitation>
). Within the
<emphasis id="B907FE490A69D14D94E83C37FBB9FEB5" box="[650,1029,300,326]" italics="true" pageId="12" pageNumber="102">
<taxonomicName id="4C7359D80A69D14D94E83C37FCA4FEB5" authorityName="Ausserer" authorityYear="1871" box="[650,792,300,326]" class="Magnoliopsida" family="Scrophulariaceae" genus="Nemesia" kingdom="Plantae" order="Lamiales" pageId="12" pageNumber="102" phylum="Tracheophyta" rank="species" species="maculatipes">maculatipes</taxonomicName>
group
<taxonomicName id="4C7359D80A69D14D95133C36FBB9FEB5" authorityName="Cassar &amp; Mifsud &amp; Decae" authorityYear="2022" box="[881,1029,300,326]" class="Magnoliopsida" family="Scrophulariaceae" genus="Nemesia" kingdom="Plantae" order="Lamiales" pageId="12" pageNumber="102" phylum="Tracheophyta" rank="species" species="maltensis">N. maltensis</taxonomicName>
</emphasis>
keys out with
<taxonomicName id="4C7359D80A69D14D92DF3C36FACAFEB5" baseAuthorityName="Costa" baseAuthorityYear="1835" box="[1213,1398,300,326]" class="Magnoliopsida" family="Scrophulariaceae" genus="Nemesia" kingdom="Plantae" order="Lamiales" pageId="12" pageNumber="102" phylum="Tracheophyta" rank="species" species="meridionalis">
<emphasis id="B907FE490A69D14D92DF3C36FACAFEB5" box="[1213,1398,300,326]" italics="true" pageId="12" pageNumber="102">N. meridionalis</emphasis>
</taxonomicName>
on grounds of a similar morphology of the distal embolus in males being ornamented with a short row of tiny denticles (
<figureCitation id="13483EDE0A69D14D97AB3C69FD9CFE7F" box="[457,544,370,396]" captionStart="Figs 5463" captionStartId="13.[189,242,1739,1765]" captionTargetBox="[189,1398,965,1691]" captionTargetId="figure-258@13.[189,1398,965,1691]" captionTargetPageId="13" captionText="Figs 5463. Nemesia maltensis sp. nov. holotype, ♂ (TC.004, NHMR). 54. Dorsal habitus (note distinct dark flanks of cephalic part). 55. Prosoma in lateral view (note dark margin of the carapace and cephalic part not elevated). 56. Dorso-distal right palp (note strong development of PTR). 57. Prolateral left metatarsus and distal tibia leg I (note CF with dense brush of short bristles, short TA and smoothly curved TS). 58. Spinnerets, lateral view showing macula (MAC) on external proximal article of PLS. 5963. Right palpal organ in four different positions rotated clock-wise by 90°. 59. Ventral view. 60. Prolateral view. 61. Dorsal view. 62. Retrolateral view. 63. Embolus tip at high magnification, note short row of tiny denticles (den) and bevelled tip." figureDoi="http://doi.org/10.5281/zenodo.6384585" httpUri="https://zenodo.org/record/6384585/files/figure.png" pageId="12" pageNumber="102">Fig. 63</figureCitation>
cf.
<bibRefCitation id="EFE25FAA0A69D14D94313C69FCF4FE7E" author="Isaia M. &amp; Decae A." box="[595,840,370,397]" pageId="12" pageNumber="102" pagination="280 - 284" refId="ref12959" refString="Isaia M. &amp; Decae A. 2012. Revalidation of Nemesia meridionalis Costa, 1835 (Araneae, Mygalomorphae, Nemesiidae), and first description of the male. Bulletin of the British Arachnological Society 15 (8): 280 - 284." type="journal article" year="2012">Isaia &amp; Decae 2012</bibRefCitation>
: fig. 4). This character is not known from any other
<taxonomicName id="4C7359D80A69D14D97613C8CFE73FE43" box="[259,463,406,432]" class="Magnoliopsida" family="Scrophulariaceae" genus="Nemesia" kingdom="Plantae" order="Lamiales" pageId="12" pageNumber="102" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="B907FE490A69D14D97613C8CFED4FE43" box="[259,360,407,432]" italics="true" pageId="12" pageNumber="102">Nemesia</emphasis>
species.
</taxonomicName>
Males of
<taxonomicName id="4C7359D80A69D14D942E3C8CFD5CFE43" authorityName="Cassar &amp; Mifsud &amp; Decae" authorityYear="2022" box="[588,736,406,432]" class="Magnoliopsida" family="Scrophulariaceae" genus="Nemesia" kingdom="Plantae" order="Lamiales" pageId="12" pageNumber="102" phylum="Tracheophyta" rank="species" species="maltensis">
<emphasis id="B907FE490A69D14D942E3C8CFD5CFE43" box="[588,736,406,432]" italics="true" pageId="12" pageNumber="102">N. maltensis</emphasis>
</taxonomicName>
can be distinguished from those of
<taxonomicName id="4C7359D80A69D14D92F43C8CFAF3FE43" baseAuthorityName="Costa" baseAuthorityYear="1835" box="[1174,1359,406,432]" class="Magnoliopsida" family="Scrophulariaceae" genus="Nemesia" kingdom="Plantae" order="Lamiales" pageId="12" pageNumber="102" phylum="Tracheophyta" rank="species" species="meridionalis">
<emphasis id="B907FE490A69D14D92F43C8CFAF3FE43" box="[1174,1359,406,432]" italics="true" pageId="12" pageNumber="102">N. meridionalis</emphasis>
</taxonomicName>
by the absence of an abrupt narrowing of the distal embolus (
<figureCitation id="13483EDE0A69D14D95163CA2FC46FE27" box="[884,1018,441,468]" captionStart="Figs 5463" captionStartId="13.[189,242,1739,1765]" captionTargetBox="[189,1398,965,1691]" captionTargetId="figure-258@13.[189,1398,965,1691]" captionTargetPageId="13" captionText="Figs 5463. Nemesia maltensis sp. nov. holotype, ♂ (TC.004, NHMR). 54. Dorsal habitus (note distinct dark flanks of cephalic part). 55. Prosoma in lateral view (note dark margin of the carapace and cephalic part not elevated). 56. Dorso-distal right palp (note strong development of PTR). 57. Prolateral left metatarsus and distal tibia leg I (note CF with dense brush of short bristles, short TA and smoothly curved TS). 58. Spinnerets, lateral view showing macula (MAC) on external proximal article of PLS. 5963. Right palpal organ in four different positions rotated clock-wise by 90°. 59. Ventral view. 60. Prolateral view. 61. Dorsal view. 62. Retrolateral view. 63. Embolus tip at high magnification, note short row of tiny denticles (den) and bevelled tip." figureDoi="http://doi.org/10.5281/zenodo.6384585" httpUri="https://zenodo.org/record/6384585/files/figure.png" pageId="12" pageNumber="102">Figs 5962</figureCitation>
cf.
<bibRefCitation id="EFE25FAA0A69D14D92483CA2FAA7FE20" author="Isaia M. &amp; Decae A." box="[1066,1307,441,467]" pageId="12" pageNumber="102" pagination="280 - 284" refId="ref12959" refString="Isaia M. &amp; Decae A. 2012. Revalidation of Nemesia meridionalis Costa, 1835 (Araneae, Mygalomorphae, Nemesiidae), and first description of the male. Bulletin of the British Arachnological Society 15 (8): 280 - 284." type="journal article" year="2012">Isaia &amp; Decae 2012</bibRefCitation>
: fig. 3), females of
<taxonomicName id="4C7359D80A69D14D97223CC6FE6EFE05" authorityName="Cassar &amp; Mifsud &amp; Decae" authorityYear="2022" box="[320,466,476,502]" class="Magnoliopsida" family="Scrophulariaceae" genus="Nemesia" kingdom="Plantae" order="Lamiales" pageId="12" pageNumber="102" phylum="Tracheophyta" rank="species" species="maltensis">
<emphasis id="B907FE490A69D14D97223CC6FE6EFE05" box="[320,466,476,502]" italics="true" pageId="12" pageNumber="102">N. maltensis</emphasis>
</taxonomicName>
can be distinguished from those of
<taxonomicName id="4C7359D80A69D14D951A3CC6FB92FE05" baseAuthorityName="Costa" baseAuthorityYear="1835" box="[888,1070,476,502]" class="Magnoliopsida" family="Scrophulariaceae" genus="Nemesia" kingdom="Plantae" order="Lamiales" pageId="12" pageNumber="102" phylum="Tracheophyta" rank="species" species="meridionalis">
<emphasis id="B907FE490A69D14D951A3CC6FB92FE05" box="[888,1070,476,502]" italics="true" pageId="12" pageNumber="102">N. meridionalis</emphasis>
</taxonomicName>
by the relatively flat profile of the carapace (lateral view
<figureCitation id="13483EDE0A69D14D947B3CE4FDCCFDE9" box="[537,624,511,538]" captionStart="Figs 6469" captionStartId="14.[189,242,1816,1842]" captionTargetBox="[189,1398,1125,1769]" captionTargetId="figure-378@14.[189,1398,1125,1769]" captionTargetPageId="14" captionText="Figs 6469. Nemesia maltensis sp. nov. paratype, ♀ (TC.012, NHMR). 64. Dorsal habitus, note maculae on external femora I &amp; II (m). 65. Left lateral habitus, note numerous maculae on legs arrow indications (m) and the low carapace profile (lp). 66. Ventral anterior prosoma, note indistinct central division (cd) in labial furrow. 67. Prolateral patella III, note three spines (sp) in a row. 68. Ventral spinnerets, note absence of distinct light coloured spigot fields. 69. Spermathecae, ventral view, note median part of the receptacles being folded and twisted (ft)." figureDoi="http://doi.org/10.5281/zenodo.6384587" httpUri="https://zenodo.org/record/6384587/files/figure.png" pageId="12" pageNumber="102">Fig. 65</figureCitation>
), lack of sharply defined light coloured spigot fields on the PMS and PLS (
<figureCitation id="13483EDE0A69D14D97563F39FE35FDCF" box="[308,393,546,572]" captionStart="Figs 6469" captionStartId="14.[189,242,1816,1842]" captionTargetBox="[189,1398,1125,1769]" captionTargetId="figure-378@14.[189,1398,1125,1769]" captionTargetPageId="14" captionText="Figs 6469. Nemesia maltensis sp. nov. paratype, ♀ (TC.012, NHMR). 64. Dorsal habitus, note maculae on external femora I &amp; II (m). 65. Left lateral habitus, note numerous maculae on legs arrow indications (m) and the low carapace profile (lp). 66. Ventral anterior prosoma, note indistinct central division (cd) in labial furrow. 67. Prolateral patella III, note three spines (sp) in a row. 68. Ventral spinnerets, note absence of distinct light coloured spigot fields. 69. Spermathecae, ventral view, note median part of the receptacles being folded and twisted (ft)." figureDoi="http://doi.org/10.5281/zenodo.6384587" httpUri="https://zenodo.org/record/6384587/files/figure.png" pageId="12" pageNumber="102">Fig. 68</figureCitation>
cf.
<bibRefCitation id="EFE25FAA0A69D14D97DB3F39FD16FDCE" author="Isaia M. &amp; Decae A." box="[441,682,546,573]" pageId="12" pageNumber="102" pagination="280 - 284" refId="ref12959" refString="Isaia M. &amp; Decae A. 2012. Revalidation of Nemesia meridionalis Costa, 1835 (Araneae, Mygalomorphae, Nemesiidae), and first description of the male. Bulletin of the British Arachnological Society 15 (8): 280 - 284." type="journal article" year="2012">Isaia &amp; Decae 2012</bibRefCitation>
: fig. 5), and absence of a strong colour contrast between the labium and the sternum and the indistinct central division of the labial furrow (
<figureCitation id="13483EDE0A69D14D92303F5DFB1AFD93" box="[1106,1190,582,608]" captionStart="Figs 6469" captionStartId="14.[189,242,1816,1842]" captionTargetBox="[189,1398,1125,1769]" captionTargetId="figure-378@14.[189,1398,1125,1769]" captionTargetPageId="14" captionText="Figs 6469. Nemesia maltensis sp. nov. paratype, ♀ (TC.012, NHMR). 64. Dorsal habitus, note maculae on external femora I &amp; II (m). 65. Left lateral habitus, note numerous maculae on legs arrow indications (m) and the low carapace profile (lp). 66. Ventral anterior prosoma, note indistinct central division (cd) in labial furrow. 67. Prolateral patella III, note three spines (sp) in a row. 68. Ventral spinnerets, note absence of distinct light coloured spigot fields. 69. Spermathecae, ventral view, note median part of the receptacles being folded and twisted (ft)." figureDoi="http://doi.org/10.5281/zenodo.6384587" httpUri="https://zenodo.org/record/6384587/files/figure.png" pageId="12" pageNumber="102">Fig. 66</figureCitation>
cf.
<bibRefCitation id="EFE25FAA0A69D14D92B03F5DFF47FD70" author="Isaia M. &amp; Decae A." pageId="12" pageNumber="102" pagination="280 - 284" refId="ref12959" refString="Isaia M. &amp; Decae A. 2012. Revalidation of Nemesia meridionalis Costa, 1835 (Araneae, Mygalomorphae, Nemesiidae), and first description of the male. Bulletin of the British Arachnological Society 15 (8): 280 - 284." type="journal article" year="2012">Isaia &amp; Decae 2012</bibRefCitation>
: figs 78).
</paragraph>
</subSubSection>
<subSubSection id="C36971D00A69D14D96DF3FB0FC95FCE4" pageId="12" pageNumber="102" type="etymology">
<paragraph id="8BCC225B0A69D14D96DF3FB0FEF8FD36" blockId="12.[189,324,683,709]" box="[189,324,683,709]" pageId="12" pageNumber="102">
<heading id="D08495370A69D14D96DF3FB0FEF8FD36" bold="true" box="[189,324,683,709]" fontSize="11" level="3" pageId="12" pageNumber="102" reason="3">
<emphasis id="B907FE490A69D14D96DF3FB0FEF8FD36" bold="true" box="[189,324,683,709]" pageId="12" pageNumber="102">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8BCC225B0A69D14D96DF3FC2FC95FCE4" blockId="12.[189,1398,729,791]" pageId="12" pageNumber="102">
The name refers to
<collectingCountry id="F36462CB0A69D14D97CF3FC2FE48FD00" box="[429,500,729,755]" name="Malta" pageId="12" pageNumber="102">Malta</collectingCountry>
, the largest island in the Maltese Archipelago and the only Mediterranean island where the species is currently known to occur.
</paragraph>
</subSubSection>
<subSubSection id="C36971D00A69D14D96DF3E25FCF2FBA6" pageId="12" pageNumber="102" type="materials_examined">
<paragraph id="8BCC225B0A69D14D96DF3E25FED0FCA4" blockId="12.[189,364,829,856]" box="[189,364,829,856]" pageId="12" pageNumber="102">
<heading id="D08495370A69D14D96DF3E25FED0FCA4" bold="true" box="[189,364,829,856]" fontSize="11" level="3" pageId="12" pageNumber="102" reason="3">
<emphasis id="B907FE490A69D14D96DF3E25FED0FCA4" bold="true" box="[189,364,829,856]" pageId="12" pageNumber="102">Type material</emphasis>
</heading>
</paragraph>
<paragraph id="8BCC225B0A69D14D96883E77FAF3FC59" blockId="12.[189,1359,876,938]" pageId="12" pageNumber="102">
<materialsCitation id="3B1B28060A69D14D96883E77FAF3FC59" ID-GBIF-Occurrence="3734085306" collectingDate="2020-09-08" collectingDateMax="2020-09-13" collectingDateMin="2020-09-08" collectionCode="NHMR" collectorName="T. Cassar" country="Malta" latitude="35.856" location="Buskett" longLatPrecision="71" longitude="14.396" pageId="12" pageNumber="102" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<emphasis id="B907FE490A69D14D96883E77FEE0FC75" bold="true" box="[234,348,876,902]" pageId="12" pageNumber="102">
<typeStatus id="54C89CF90A69D14D96883E77FEE0FC75" box="[234,348,876,902]" pageId="12" pageNumber="102" type="holotype">Holotype</typeStatus>
</emphasis>
<collectingCountry id="F36462CB0A69D14D96DF3E8BFEA3FC59" box="[189,287,912,938]" name="Malta" pageId="12" pageNumber="102">MALTA</collectingCountry>
<specimenCount id="9D75E9D20A69D14D975A3E8BFEEDFC59" box="[312,337,912,938]" pageId="12" pageNumber="102" type="male"></specimenCount>
;
<location id="8EAC74800A69D14D973F3E94FE06FC5A" LSID="urn:lsid:plazi:treatment:03DA934D0A6ED151941A3B24FE63F921:8EAC74800A69D14D973F3E94FE06FC5A" box="[349,442,911,937]" country="Malta" latitude="35.856" longLatPrecision="71" longitude="14.396" name="Buskett" pageId="12" pageNumber="102">Buskett</location>
;
<date id="FFCD049B0A69D14D97A43E94FD3DFC5A" box="[454,641,911,937]" pageId="12" pageNumber="102" value="2020-09-08" valueMax="2020-09-13" valueMin="2020-09-08">
<collectingDate id="EF89FD730A69D14D97A43E94FD3DFC5A" box="[454,641,911,937]" pageId="12" pageNumber="102" value="2020-09-08" valueMax="2020-09-13" valueMin="2020-09-08">813 Sep. 2020</collectingDate>
</date>
;
<geoCoordinate id="EE47449C0A69D14D94EF3E94FCB5FC59" box="[653,777,911,938]" degrees="35.856" direction="north" orientation="latitude" pageId="12" pageNumber="102" precision="55" value="35.856">35.856° N</geoCoordinate>
,
<geoCoordinate id="EE47449C0A69D14D95773E94FC31FC59" box="[789,909,911,938]" degrees="14.396" direction="east" orientation="longitude" pageId="12" pageNumber="102" precision="55" value="14.396">14.396° E</geoCoordinate>
;
<collectorName id="2686478D0A69D14D95FB3E8BFBBAFC5A" box="[921,1030,911,938]" pageId="12" pageNumber="102">T. Cassar</collectorName>
leg.; (no. TC.004);
<collectionCode id="ED62BA9E0A69D14D928D3E8BFAF7FC59" box="[1263,1355,912,938]" country="Iceland" httpUri="http://grbio.org/cool/1rf3-4kh4" name="Natural History Museum, Reykjavik" pageId="12" pageNumber="102">NHMR</collectionCode>
.
</materialsCitation>
</paragraph>
<paragraph id="8BCC225B0A69D14D96883ECAFCF2FBA6" blockId="12.[189,1398,977,1109]" pageId="12" pageNumber="102">
<materialsCitation id="3B1B28060A69D14D96883ECAFB77FBFD" ID-GBIF-Occurrence="3734085312" collectingDate="2020-09-08" collectingDateMax="2020-09-13" collectingDateMin="2020-09-08" collectionCode="NHMR" collectorName="Cassar" country="Malta" latitude="35.856" location="MALTA" longLatPrecision="71" longitude="14.396" pageId="12" pageNumber="102" specimenCount="4" specimenCount-male="4" typeStatus="paratype">
<emphasis id="B907FE490A69D14D96883ECAFEDBFC18" bold="true" box="[234,359,977,1003]" pageId="12" pageNumber="102">
<typeStatus id="54C89CF90A69D14D96883ECAFEDBFC18" box="[234,359,977,1003]" pageId="12" pageNumber="102" type="paratype">Paratypes</typeStatus>
</emphasis>
<collectingCountry id="F36462CB0A69D14D96DF3EEFFEA3FBFD" box="[189,287,1012,1038]" name="Malta" pageId="12" pageNumber="102">MALTA</collectingCountry>
<specimenCount id="9D75E9D20A69D14D97583EEFFE3CFBFD" box="[314,384,1012,1038]" pageId="12" pageNumber="102" type="male">4 ♂♂</specimenCount>
; same collection data as for holotype; (no. TC.005 to TC.008);
<collectionCode id="ED62BA9E0A69D14D92113EEFFB77FBFD" box="[1139,1227,1012,1038]" collectionName="Netherlands, Rotterdam, Natuurhistorisch Museum" country="Iceland" httpUri="http://grbio.org/cool/1rf3-4kh4" name="Natural History Museum, Reykjavik" pageId="12" pageNumber="102">NHMR</collectionCode>
</materialsCitation>
<materialsCitation id="3B1B28060A69D14D92853EEFFC05FBC1" ID-GBIF-Occurrence="3734085310" collectingDate="2020-09-08" collectingDateMax="2020-09-13" collectingDateMin="2020-09-08" collectionCode="NHMW" collectorName="Cassar" country="Malta" latitude="35.856" location="Buskett" longLatPrecision="71" longitude="14.396" pageId="12" pageNumber="102" specimenCount="2" specimenCount-male="2" typeStatus="paratype">
<specimenCount id="9D75E9D20A69D14D92853EEFFA91FBFD" box="[1255,1325,1012,1038]" pageId="12" pageNumber="102" type="male">2 ♂♂</specimenCount>
; same collection data as for holotype; (no. TC.009, TC.010);
<collectionCode id="ED62BA9E0A69D14D953B3903FC05FBC1" box="[857,953,1048,1074]" collectionName="Austria, Wien, Naturhistorisches Museum Wien" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="12" pageNumber="102" type="Museum">NHMW</collectionCode>
</materialsCitation>
<materialsCitation id="3B1B28060A69D14D95BA390CFCF6FBA6" ID-GBIF-Occurrence="3734085304" collectingDate="2021-03-14" collectingDateMax="2020-09-13" collectingDateMin="2020-09-08" collectionCode="NHMR" collectorName="Cassar" country="Malta" latitude="35.856" location="Buskett" longLatPrecision="71" longitude="14.396" pageId="12" pageNumber="102" specimenCount="2" specimenCount-female="2" typeStatus="paratype">
<specimenCount id="9D75E9D20A69D14D95BA390CFBA2FBC1" box="[984,1054,1047,1074]" pageId="12" pageNumber="102" type="female">2 ♀♀</specimenCount>
; same collection data as for holotype;
<date id="FFCD049B0A69D14D97513921FE68FBA7" box="[307,468,1082,1109]" pageId="12" pageNumber="102" value="2021-03-14">
<collectingDate id="EF89FD730A69D14D97513921FE68FBA7" box="[307,468,1082,1109]" pageId="12" pageNumber="102" value="2021-03-14">14 Mar. 2021</collectingDate>
</date>
; (no. TC.012, TC.013);
<collectionCode id="ED62BA9E0A69D14D948C3920FCF6FBA6" box="[750,842,1083,1109]" collectionName="Netherlands, Rotterdam, Natuurhistorisch Museum" country="Iceland" httpUri="http://grbio.org/cool/1rf3-4kh4" name="Natural History Museum, Reykjavik" pageId="12" pageNumber="102">NHMR</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C36971D00A69D15196DF3960FE21FB03" lastPageId="16" lastPageNumber="106" pageId="12" pageNumber="102" type="description">
<paragraph id="8BCC225B0A69D14D96DF3960FEF2FB66" blockId="12.[189,334,1147,1173]" box="[189,334,1147,1173]" pageId="12" pageNumber="102">
<heading id="D08495370A69D14D96DF3960FEF2FB66" bold="true" box="[189,334,1147,1173]" fontSize="11" level="3" pageId="12" pageNumber="102" reason="3">
<emphasis id="B907FE490A69D14D96DF3960FEF2FB66" bold="true" box="[189,334,1147,1173]" pageId="12" pageNumber="102">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8BCC225B0A69D14D968839B1FD19FB36" blockId="12.[188,1399,1194,1292]" box="[234,677,1194,1221]" pageId="12" pageNumber="102">
<emphasis id="B907FE490A69D14D968839B1FE21FB37" bold="true" box="[234,413,1194,1220]" pageId="12" pageNumber="102">
Male
<typeStatus id="54C89CF90A69D14D975339B1FE21FB37" box="[305,413,1194,1220]" pageId="12" pageNumber="102" type="holotype">holotype</typeStatus>
</emphasis>
(no. TC-004, NHMR)
</paragraph>
<paragraph id="8BCC225B0A69D14D96DF39D5FD09FAFF" blockId="12.[188,1399,1194,1292]" pageId="12" pageNumber="102">
PRESERVATION AND CONDITION. Specimen 6 months preserved in 70% ethanol, in good condition (
<figureCitation id="13483EDE0A69D14D937039D6FAD5FB1B" box="[1298,1385,1229,1256]" captionStart="Figs 5463" captionStartId="13.[189,242,1739,1765]" captionTargetBox="[189,1398,965,1691]" captionTargetId="figure-258@13.[189,1398,965,1691]" captionTargetPageId="13" captionText="Figs 5463. Nemesia maltensis sp. nov. holotype, ♂ (TC.004, NHMR). 54. Dorsal habitus (note distinct dark flanks of cephalic part). 55. Prosoma in lateral view (note dark margin of the carapace and cephalic part not elevated). 56. Dorso-distal right palp (note strong development of PTR). 57. Prolateral left metatarsus and distal tibia leg I (note CF with dense brush of short bristles, short TA and smoothly curved TS). 58. Spinnerets, lateral view showing macula (MAC) on external proximal article of PLS. 5963. Right palpal organ in four different positions rotated clock-wise by 90°. 59. Ventral view. 60. Prolateral view. 61. Dorsal view. 62. Retrolateral view. 63. Embolus tip at high magnification, note short row of tiny denticles (den) and bevelled tip." figureDoi="http://doi.org/10.5281/zenodo.6384585" httpUri="https://zenodo.org/record/6384585/files/figure.png" pageId="12" pageNumber="102">Fig. 54</figureCitation>
), right bulb detached for study (
<figureCitation id="13483EDE0A69D14D947D39EAFD14FAFF" box="[543,680,1265,1292]" captionStart="Figs 5463" captionStartId="13.[189,242,1739,1765]" captionTargetBox="[189,1398,965,1691]" captionTargetId="figure-258@13.[189,1398,965,1691]" captionTargetPageId="13" captionText="Figs 5463. Nemesia maltensis sp. nov. holotype, ♂ (TC.004, NHMR). 54. Dorsal habitus (note distinct dark flanks of cephalic part). 55. Prosoma in lateral view (note dark margin of the carapace and cephalic part not elevated). 56. Dorso-distal right palp (note strong development of PTR). 57. Prolateral left metatarsus and distal tibia leg I (note CF with dense brush of short bristles, short TA and smoothly curved TS). 58. Spinnerets, lateral view showing macula (MAC) on external proximal article of PLS. 5963. Right palpal organ in four different positions rotated clock-wise by 90°. 59. Ventral view. 60. Prolateral view. 61. Dorsal view. 62. Retrolateral view. 63. Embolus tip at high magnification, note short row of tiny denticles (den) and bevelled tip." figureDoi="http://doi.org/10.5281/zenodo.6384585" httpUri="https://zenodo.org/record/6384585/files/figure.png" pageId="12" pageNumber="102">Figs 5963</figureCitation>
).
</paragraph>
<paragraph id="8BCC225B0A69D14D96DF3829FD9AFA0E" blockId="12.[187,1399,1330,1533]" pageId="12" pageNumber="102">
GENERAL COLORATION. Carapace yellowish with dark flanks of cephalic part and dark margin (
<figureCitation id="13483EDE0A69D14D93723829FAD4FABE" box="[1296,1384,1330,1357]" captionStart="Figs 5463" captionStartId="13.[189,242,1739,1765]" captionTargetBox="[189,1398,965,1691]" captionTargetId="figure-258@13.[189,1398,965,1691]" captionTargetPageId="13" captionText="Figs 5463. Nemesia maltensis sp. nov. holotype, ♂ (TC.004, NHMR). 54. Dorsal habitus (note distinct dark flanks of cephalic part). 55. Prosoma in lateral view (note dark margin of the carapace and cephalic part not elevated). 56. Dorso-distal right palp (note strong development of PTR). 57. Prolateral left metatarsus and distal tibia leg I (note CF with dense brush of short bristles, short TA and smoothly curved TS). 58. Spinnerets, lateral view showing macula (MAC) on external proximal article of PLS. 5963. Right palpal organ in four different positions rotated clock-wise by 90°. 59. Ventral view. 60. Prolateral view. 61. Dorsal view. 62. Retrolateral view. 63. Embolus tip at high magnification, note short row of tiny denticles (den) and bevelled tip." figureDoi="http://doi.org/10.5281/zenodo.6384585" httpUri="https://zenodo.org/record/6384585/files/figure.png" pageId="12" pageNumber="102">Fig. 55</figureCitation>
), chelicerae brown, darker than carapace with dorsal lighter coloured patch, opisthosoma dorsal anterior dark greyish-brown, further light coloured with dark, irregular chevrons and a dark cardiac line (
<figureCitation id="13483EDE0A69D14D93713863FAD6FA60" box="[1299,1386,1400,1427]" captionStart="Figs 5463" captionStartId="13.[189,242,1739,1765]" captionTargetBox="[189,1398,965,1691]" captionTargetId="figure-258@13.[189,1398,965,1691]" captionTargetPageId="13" captionText="Figs 5463. Nemesia maltensis sp. nov. holotype, ♂ (TC.004, NHMR). 54. Dorsal habitus (note distinct dark flanks of cephalic part). 55. Prosoma in lateral view (note dark margin of the carapace and cephalic part not elevated). 56. Dorso-distal right palp (note strong development of PTR). 57. Prolateral left metatarsus and distal tibia leg I (note CF with dense brush of short bristles, short TA and smoothly curved TS). 58. Spinnerets, lateral view showing macula (MAC) on external proximal article of PLS. 5963. Right palpal organ in four different positions rotated clock-wise by 90°. 59. Ventral view. 60. Prolateral view. 61. Dorsal view. 62. Retrolateral view. 63. Embolus tip at high magnification, note short row of tiny denticles (den) and bevelled tip." figureDoi="http://doi.org/10.5281/zenodo.6384585" httpUri="https://zenodo.org/record/6384585/files/figure.png" pageId="12" pageNumber="102">Fig. 54</figureCitation>
), opisthosoma ventral uniform creamy white, palps and legs lighter coloured than body in dorsal view and ventrally lighter coloured than dorsally, sternum and ventral coxae creamy white, labium conspicuously darker than other ventral parts.
</paragraph>
<paragraph id="8BCC225B0A69D14D96DF3B3FFB73F95B" blockId="12.[189,1400,1571,1704]" pageId="12" pageNumber="102">
CARAPACE. Longer than wide (CW/CL 0.8), covered with very fine silvery pubescence, curved bristles concentrated along the margins and posterior on the thoracic part, longitudinal row on the crest of the cephalic part and few forwardly projecting bristles on clypeus. Cephalic not elevated, thoracic part bulging slightly up from the fovea before sloping down to the posterior margin (
<figureCitation id="13483EDE0A69D14D92083B96FB7EF95B" box="[1130,1218,1677,1704]" captionStart="Figs 5463" captionStartId="13.[189,242,1739,1765]" captionTargetBox="[189,1398,965,1691]" captionTargetId="figure-258@13.[189,1398,965,1691]" captionTargetPageId="13" captionText="Figs 5463. Nemesia maltensis sp. nov. holotype, ♂ (TC.004, NHMR). 54. Dorsal habitus (note distinct dark flanks of cephalic part). 55. Prosoma in lateral view (note dark margin of the carapace and cephalic part not elevated). 56. Dorso-distal right palp (note strong development of PTR). 57. Prolateral left metatarsus and distal tibia leg I (note CF with dense brush of short bristles, short TA and smoothly curved TS). 58. Spinnerets, lateral view showing macula (MAC) on external proximal article of PLS. 5963. Right palpal organ in four different positions rotated clock-wise by 90°. 59. Ventral view. 60. Prolateral view. 61. Dorsal view. 62. Retrolateral view. 63. Embolus tip at high magnification, note short row of tiny denticles (den) and bevelled tip." figureDoi="http://doi.org/10.5281/zenodo.6384585" httpUri="https://zenodo.org/record/6384585/files/figure.png" pageId="12" pageNumber="102">Fig. 55</figureCitation>
).
</paragraph>
<paragraph id="8BCC225B0A69D14D96DF3BD4FCF5F8DC" blockId="12.[189,1399,1743,1840]" pageId="12" pageNumber="102">EYES. Ocular-tubercle dome-shaped, clypeus narrow sloping down from eyes, eye-group rectangular, twice as wide as long (EL/PR 0.51), AME less than their diameter apart (dis.AME/dia.AME 0.87), dis. ALEPLE less than ½ dia.ALE (ALEPLE/ALE 0.44).</paragraph>
<paragraph id="8BCC225B0A69D14D96DF3A4CFCC5F867" blockId="12.[188,1399,1878,1940]" pageId="12" pageNumber="102">CHELICERAE. Rastellum with few strong teeth on the apical prolateral corner of chelicerae, prolateral row of furrow teeth, fangs sharp, bent, ventral un-serrated ridge.</paragraph>
<paragraph id="8BCC225B0A68D14C96DF3C12FB4EFE9A" blockId="13.[188,1399,265,361]" pageId="13" pageNumber="103">VENTRAL PROSOMA. Maxillae rounded distal lobe, few spiky cuspules on proximo-anterior margin. Labium: wider than long (LW/LL 0.7), cuspules absent, labial furrow narrow. Sternum: longer than wide (SW/SL 0.8) three pairs round sigilla, posteriors sub-marginal, evenly covered with bristles.</paragraph>
<paragraph id="8BCC225B0A68D14C96DF3C8AFA88FDCB" blockId="13.[189,1399,401,568]" pageId="13" pageNumber="103">
PALPS. Cymbium with distal group of spines, tibia with PTR strongly develop (
<figureCitation id="13483EDE0A68D14C92E03C8AFB61FE5F" box="[1154,1245,401,428]" captionStart="Figs 5463" captionStartId="13.[189,242,1739,1765]" captionTargetBox="[189,1398,965,1691]" captionTargetId="figure-258@13.[189,1398,965,1691]" captionTargetPageId="13" captionText="Figs 5463. Nemesia maltensis sp. nov. holotype, ♂ (TC.004, NHMR). 54. Dorsal habitus (note distinct dark flanks of cephalic part). 55. Prosoma in lateral view (note dark margin of the carapace and cephalic part not elevated). 56. Dorso-distal right palp (note strong development of PTR). 57. Prolateral left metatarsus and distal tibia leg I (note CF with dense brush of short bristles, short TA and smoothly curved TS). 58. Spinnerets, lateral view showing macula (MAC) on external proximal article of PLS. 5963. Right palpal organ in four different positions rotated clock-wise by 90°. 59. Ventral view. 60. Prolateral view. 61. Dorsal view. 62. Retrolateral view. 63. Embolus tip at high magnification, note short row of tiny denticles (den) and bevelled tip." figureDoi="http://doi.org/10.5281/zenodo.6384585" httpUri="https://zenodo.org/record/6384585/files/figure.png" pageId="13" pageNumber="103">Fig. 56</figureCitation>
), proximally inflated (TibW/PTib 0.6), patella spineless, femur longer than tibia (PFem/PTib 1.6) with group of slender spines dorsally on distal half of the article. Palpal organ, proximal bulbous part pyriform extending distally into a slender curved embolus. Embolus curved, distally gradually tapering to a sharp, bevelled tip. Striae on proximal embolus, few tiny denticles just proximal of the embolus tip (
<figureCitation id="13483EDE0A68D14C92FF3F05FA9BFDCB" box="[1181,1319,541,568]" captionStart="Figs 5463" captionStartId="13.[189,242,1739,1765]" captionTargetBox="[189,1398,965,1691]" captionTargetId="figure-258@13.[189,1398,965,1691]" captionTargetPageId="13" captionText="Figs 5463. Nemesia maltensis sp. nov. holotype, ♂ (TC.004, NHMR). 54. Dorsal habitus (note distinct dark flanks of cephalic part). 55. Prosoma in lateral view (note dark margin of the carapace and cephalic part not elevated). 56. Dorso-distal right palp (note strong development of PTR). 57. Prolateral left metatarsus and distal tibia leg I (note CF with dense brush of short bristles, short TA and smoothly curved TS). 58. Spinnerets, lateral view showing macula (MAC) on external proximal article of PLS. 5963. Right palpal organ in four different positions rotated clock-wise by 90°. 59. Ventral view. 60. Prolateral view. 61. Dorsal view. 62. Retrolateral view. 63. Embolus tip at high magnification, note short row of tiny denticles (den) and bevelled tip." figureDoi="http://doi.org/10.5281/zenodo.6384585" httpUri="https://zenodo.org/record/6384585/files/figure.png" pageId="13" pageNumber="103">Figs 5963</figureCitation>
).
</paragraph>
<paragraph id="8BCC225B0A68D14C96DF3F7BFEA7FC9C" blockId="13.[188,1401,607,879]" pageId="13" pageNumber="103">
LEGS. All tarsi with fine ventral scopulae. Leg: I: metatarsus and tibia modified, metatarsus curved, lateral longitudinal rows of spines over the length of the article, CF with dense brush of short bristles, tibia distally widened, TA reduced (
<figureCitation id="13483EDE0A68D14C94083FBEFD7CFD33" box="[618,704,677,704]" captionStart="Figs 5463" captionStartId="13.[189,242,1739,1765]" captionTargetBox="[189,1398,965,1691]" captionTargetId="figure-258@13.[189,1398,965,1691]" captionTargetPageId="13" captionText="Figs 5463. Nemesia maltensis sp. nov. holotype, ♂ (TC.004, NHMR). 54. Dorsal habitus (note distinct dark flanks of cephalic part). 55. Prosoma in lateral view (note dark margin of the carapace and cephalic part not elevated). 56. Dorso-distal right palp (note strong development of PTR). 57. Prolateral left metatarsus and distal tibia leg I (note CF with dense brush of short bristles, short TA and smoothly curved TS). 58. Spinnerets, lateral view showing macula (MAC) on external proximal article of PLS. 5963. Right palpal organ in four different positions rotated clock-wise by 90°. 59. Ventral view. 60. Prolateral view. 61. Dorsal view. 62. Retrolateral view. 63. Embolus tip at high magnification, note short row of tiny denticles (den) and bevelled tip." figureDoi="http://doi.org/10.5281/zenodo.6384585" httpUri="https://zenodo.org/record/6384585/files/figure.png" pageId="13" pageNumber="103">Fig. 57</figureCitation>
cf
<figureCitation id="13483EDE0A68D14C948B3FBEFCFEFD33" box="[745,834,677,704]" captionStart="Figs 2735" captionStartId="8.[187,240,1814,1840]" captionTargetBox="[192,1397,963,1769]" captionTargetId="figure-318@8.[189,1398,963,1769]" captionTargetPageId="8" captionText="Figs 2735. Nemesia arboricola Pocock, 1903, ♂ reference specimen (TD.4, NHMR). 27. Dorsal habitus. 28. Prosoma lateral, note slight elevation of the cephalic part, ocular process and gradual posterior slope of thoracic part. 29. Metatarsus and tibia leg I, note tibial apophysis (TA), tibia spur (TS), clasper field (CF). 30. Prolateral patella III with single spine (SS). 3134. Right palpal organ in four different positions rotated clock-wise by 90°. 31. Ventral view, 32. Prolateral view. 33. Dorsal view. 34. Retrolateral view. 35. Embolus tip at high magnification, note slightly scooped tip." figureDoi="http://doi.org/10.5281/zenodo.6384579" httpUri="https://zenodo.org/record/6384579/files/figure.png" pageId="13" pageNumber="103">Fig. 29</figureCitation>
), TS smoothly upward curved (
<figureCitation id="13483EDE0A68D14C92DC3FBEFAAAFD33" box="[1214,1302,677,704]" captionStart="Figs 5463" captionStartId="13.[189,242,1739,1765]" captionTargetBox="[189,1398,965,1691]" captionTargetId="figure-258@13.[189,1398,965,1691]" captionTargetPageId="13" captionText="Figs 5463. Nemesia maltensis sp. nov. holotype, ♂ (TC.004, NHMR). 54. Dorsal habitus (note distinct dark flanks of cephalic part). 55. Prosoma in lateral view (note dark margin of the carapace and cephalic part not elevated). 56. Dorso-distal right palp (note strong development of PTR). 57. Prolateral left metatarsus and distal tibia leg I (note CF with dense brush of short bristles, short TA and smoothly curved TS). 58. Spinnerets, lateral view showing macula (MAC) on external proximal article of PLS. 5963. Right palpal organ in four different positions rotated clock-wise by 90°. 59. Ventral view. 60. Prolateral view. 61. Dorsal view. 62. Retrolateral view. 63. Embolus tip at high magnification, note short row of tiny denticles (den) and bevelled tip." figureDoi="http://doi.org/10.5281/zenodo.6384585" httpUri="https://zenodo.org/record/6384585/files/figure.png" pageId="13" pageNumber="103">Fig. 57</figureCitation>
), spines present on all articles proximal of tarsus, patella with two prolateral spines. Legs IIII; tarsi without spines, other articles with numerous spines, maculae vague, most prominent on prolateral femur (apical) and patella (central). Leg IV, tibia&gt; femur&gt; metatarsus (Fem4/Met4 = 1.02, Tib4/Met4 = 1.07). Leg formula: 4123. PTC on all tarsi with ventrally two parallel combs of small teeth (genus character), ATC smooth.
</paragraph>
<caption id="DF0C72D30A68D14C96DF3BD0FCBCF828" ID-DOI="http://doi.org/10.5281/zenodo.6384585" ID-Zenodo-Dep="6384585" httpUri="https://zenodo.org/record/6384585/files/figure.png" pageId="13" pageNumber="103" startId="13.[189,242,1739,1765]" targetBox="[189,1398,965,1691]" targetPageId="13">
<paragraph id="8BCC225B0A68D14C96DF3BD0FCBCF828" blockId="13.[188,1399,1739,2012]" pageId="13" pageNumber="103">
<emphasis id="B907FE490A68D14C96DF3BD0FEF5F916" bold="true" box="[189,329,1739,1765]" pageId="13" pageNumber="103">Figs 5463.</emphasis>
<taxonomicName id="4C7359D80A68D14C972D3BD7FD90F916" authority="Cassar &amp; Mifsud &amp; Decae, 2022" authorityName="Cassar &amp; Mifsud &amp; Decae" authorityYear="2022" box="[335,556,1739,1765]" class="Magnoliopsida" family="Scrophulariaceae" genus="Nemesia" kingdom="Plantae" order="Lamiales" pageId="13" pageNumber="103" phylum="Tracheophyta" rank="species" species="maltensis" status="sp. nov.">
<emphasis id="B907FE490A68D14C972D3BD7FD90F916" box="[335,556,1739,1765]" italics="true" pageId="13" pageNumber="103">Nemesia maltensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23443320A68D14C94513BD7FD31F915" box="[563,653,1740,1766]" pageId="13" pageNumber="103" rank="species">sp. nov.</taxonomicNameLabel>
holotype, ♂ (TC.004, NHMR).
<emphasis id="B907FE490A68D14C92613BD0FB9CF916" bold="true" box="[1027,1056,1739,1765]" pageId="13" pageNumber="103">54</emphasis>
. Dorsal habitus (note distinct dark flanks of cephalic part).
<emphasis id="B907FE490A68D14C94733BF5FD92F8FB" bold="true" box="[529,558,1774,1800]" pageId="13" pageNumber="103">55</emphasis>
. Prosoma in lateral view (note dark margin of the carapace and cephalic part not elevated).
<emphasis id="B907FE490A68D14C97C63A0AFE7DF8D8" bold="true" box="[420,449,1809,1835]" pageId="13" pageNumber="103">56</emphasis>
. Dorso-distal right palp (note strong development of PTR).
<emphasis id="B907FE490A68D14C92C13A09FB7CF8DF" bold="true" box="[1187,1216,1810,1836]" pageId="13" pageNumber="103">57</emphasis>
. Prolateral left metatarsus and distal tibia leg I (note CF with dense brush of short bristles, short TA and smoothly curved TS).
<emphasis id="B907FE490A68D14C97303A43FED2F881" bold="true" box="[338,366,1880,1906]" pageId="13" pageNumber="103">58</emphasis>
. Spinnerets, lateral view showing macula (MAC) on external proximal article of PLS.
<emphasis id="B907FE490A68D14C96DD3A60FEBAF866" bold="true" box="[191,262,1915,1941]" pageId="13" pageNumber="103">5963</emphasis>
. Right palpal organ in four different positions rotated clock-wise by 90°.
<emphasis id="B907FE490A68D14C92C43A60FB7FF866" bold="true" box="[1190,1219,1915,1941]" pageId="13" pageNumber="103">59</emphasis>
. Ventral view.
<emphasis id="B907FE490A68D14C96DF3A85FF66F84B" bold="true" box="[189,218,1950,1976]" pageId="13" pageNumber="103">60</emphasis>
. Prolateral view.
<emphasis id="B907FE490A68D14C97C83A85FE7BF84B" bold="true" box="[426,455,1950,1976]" pageId="13" pageNumber="103">61</emphasis>
. Dorsal view.
<emphasis id="B907FE490A68D14C94113A85FD2CF84B" bold="true" box="[627,656,1950,1976]" pageId="13" pageNumber="103">62</emphasis>
. Retrolateral view.
<emphasis id="B907FE490A68D14C951A3A85FC29F84B" bold="true" box="[888,917,1950,1976]" pageId="13" pageNumber="103">63</emphasis>
. Embolus tip at high magnification, note short row of tiny denticles (den) and bevelled tip.
</paragraph>
</caption>
<paragraph id="8BCC225B0A6BD14F96DF3C12FD62FE7F" blockId="14.[189,1399,265,397]" pageId="14" pageNumber="104">
OPISTHOSOMA. Ovoid, anterior narrowing with numerous forward directed bristles, dorsal bristles are backward directed, Spinnerets: PMS knob-shaped, spigots apically concentrated, PLS proximal article with external macula (
<figureCitation id="13483EDE0A6BD14F97A73C54FDA0FE99" box="[453,540,335,362]" captionStart="Figs 5463" captionStartId="13.[189,242,1739,1765]" captionTargetBox="[189,1398,965,1691]" captionTargetId="figure-258@13.[189,1398,965,1691]" captionTargetPageId="13" captionText="Figs 5463. Nemesia maltensis sp. nov. holotype, ♂ (TC.004, NHMR). 54. Dorsal habitus (note distinct dark flanks of cephalic part). 55. Prosoma in lateral view (note dark margin of the carapace and cephalic part not elevated). 56. Dorso-distal right palp (note strong development of PTR). 57. Prolateral left metatarsus and distal tibia leg I (note CF with dense brush of short bristles, short TA and smoothly curved TS). 58. Spinnerets, lateral view showing macula (MAC) on external proximal article of PLS. 5963. Right palpal organ in four different positions rotated clock-wise by 90°. 59. Ventral view. 60. Prolateral view. 61. Dorsal view. 62. Retrolateral view. 63. Embolus tip at high magnification, note short row of tiny denticles (den) and bevelled tip." figureDoi="http://doi.org/10.5281/zenodo.6384585" httpUri="https://zenodo.org/record/6384585/files/figure.png" pageId="14" pageNumber="104">Fig. 58</figureCitation>
), as long as medial + distal articles, spigots spread over ventral proximal and medial articles and on apical distal article.
</paragraph>
<paragraph id="8BCC225B0A6BD14F96DF3CAEFB36FDAF" blockId="14.[188,1399,436,604]" pageId="14" pageNumber="104">MEASUREMENTS. TBL = 12.3; CL = 5.0; CW = 3.8; CP = 3.0; AR = 0.83; PR = 0.83; EL = 0.42; dia.ALE = 0.25; dia.PLE 0.15; dia.AME = 0.15; dia.PME = 0.12; dis.AMEAME = 0.13; dis.ALEPLE = 0.11; SL = 2.6; SW = 1.9; LL = 0.6; LW = 0.8; Palp = 4.9 (0.7 + 1.2 + 1.1 + 1.9); Leg I = 13.2 (2.0 + 2.6 + 2.6 + 2.4 + 3.6); Leg II = 12.5 (2.0 + 2.5 + 2.4 + 2.1 + 3.5); Leg III = 11.9 (2.0 + 3.0 + 2.1 + 1.7 + 3.1); Leg IV = 16.8 (2.3 + 3.9 + 4.2 + 2.4 + 4.0); BuL = 1.12; BuW = 0.39; EmL = 0.51.</paragraph>
<paragraph id="8BCC225B0A6BD14F96DF3F98FDA1FC2A" blockId="14.[188,1399,642,986]" pageId="14" pageNumber="104">VARIATION MALES (n = 7). TBL = 8.712.3 (av. 10.1, sd. 1.2); CL = 3.85.0 (av. 4.4, sd. 0.4); CW = 2.83.8 (av. 3.3, sd. 0.3); CP = 2.13.0 (av. 2.5, sd. 0.3); AR = 0.670.83 (av. 0.74, sd. 0.05); PR = 0.69 0.83 (av. 0.76, sd. 0.05); EL = 0.360.44 (av. 0.40, sd. 0.03); dia.ALE = 0.200.25 (av. 0.22, sd. 0.02); dia.PLE = 0.140.18 (av. 0.17, sd. 0.02); dia.AME = 0.130.15 (av. 0.14, sd. 0.01); dia.PME = 0.11 0.14 (av. 0.13, sd. 0.01); dis.AMEAME = 0.080.14 (av. 0.11, sd. 0.02); dis.ALEPLE = 0.040.11 (av. 0.07, sd. 0.02); SL = 1.92.6 (av. 2.2, sd. 0.2); SW = 1.51.9 (av. 1.7, sd. 0.1); LL = 0.30.6 (av. 0.4, sd. 0.1); LW = 0.60.8 (av. 0.7, sd. 0.1); Palp = 4.04.9 (av. 4.6, sd. 0.3); Leg I = 10.413.0 (av. 11.7, sd. 0.9); Leg II = 9.512.5 (av. 11.0, sd. 1.0); Leg III = 9.011.8 (av. 10.3, sd. 0.9); Leg IV = 13.516.8 (av. 15.1, sd. 1.2); Bul = 0.961.12 (av. 1.06, sd. 0.06); BuW = 0.310.39 (av. 0.36, sd. 0.03); EmL = 0.470.52 (av. 0.52, sd. 0.02).</paragraph>
<caption id="DF0C72D30A6BD14F96DF3A03FD20F810" ID-DOI="http://doi.org/10.5281/zenodo.6384587" ID-Zenodo-Dep="6384587" httpUri="https://zenodo.org/record/6384587/files/figure.png" pageId="14" pageNumber="104" startId="14.[189,242,1816,1842]" targetBox="[189,1398,1125,1769]" targetPageId="14">
<paragraph id="8BCC225B0A6BD14F96DF3A03FD20F810" blockId="14.[187,1399,1816,2019]" pageId="14" pageNumber="104">
<emphasis id="B907FE490A6BD14F96DF3A03FEF5F8C1" bold="true" box="[189,329,1816,1842]" pageId="14" pageNumber="104">Figs 6469.</emphasis>
<taxonomicName id="4C7359D80A6BD14F972C3A02FD96F8C1" authority="Cassar &amp; Mifsud &amp; Decae, 2022" authorityName="Cassar &amp; Mifsud &amp; Decae" authorityYear="2022" box="[334,554,1816,1842]" class="Magnoliopsida" family="Scrophulariaceae" genus="Nemesia" kingdom="Plantae" order="Lamiales" pageId="14" pageNumber="104" phylum="Tracheophyta" rank="species" species="maltensis" status="sp. nov.">
<emphasis id="B907FE490A6BD14F972C3A02FD96F8C1" box="[334,554,1816,1842]" italics="true" pageId="14" pageNumber="104">Nemesia maltensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23443320A6BD14F94533A02FD37F8C0" box="[561,651,1817,1843]" pageId="14" pageNumber="104" rank="species">sp. nov.</taxonomicNameLabel>
paratype, ♀ (TC.012, NHMR).
<emphasis id="B907FE490A6BD14F959E3A03FBA5F8C1" bold="true" box="[1020,1049,1816,1842]" pageId="14" pageNumber="104">64</emphasis>
. Dorsal habitus, note maculae on external femora I &amp; II (m).
<emphasis id="B907FE490A6BD14F947D3A20FD80F8A6" bold="true" box="[543,572,1851,1877]" pageId="14" pageNumber="104">65</emphasis>
. Left lateral habitus, note numerous maculae on legs arrow indications (m) and the low carapace profile (lp).
<emphasis id="B907FE490A6BD14F941E3A44FD25F88A" bold="true" box="[636,665,1887,1913]" pageId="14" pageNumber="104">66</emphasis>
. Ventral anterior prosoma, note indistinct central division (cd) in labial furrow.
<emphasis id="B907FE490A6BD14F97EA3A99FE1FF86F" bold="true" box="[392,419,1922,1948]" pageId="14" pageNumber="104">67</emphasis>
. Prolateral patella III, note three spines (sp) in a row.
<emphasis id="B907FE490A6BD14F92513A99FBF2F86F" bold="true" box="[1075,1102,1922,1948]" pageId="14" pageNumber="104">68</emphasis>
. Ventral spinnerets, note absence of distinct light coloured spigot fields.
<emphasis id="B907FE490A6BD14F948A3ABEFCB9F84C" bold="true" box="[744,773,1957,1983]" pageId="14" pageNumber="104">69</emphasis>
. Spermathecae, ventral view, note median part of the receptacles being folded and twisted (ft).
</paragraph>
</caption>
<paragraph id="8BCC225B0A6AD14E96883C12FD7AFED0" blockId="15.[189,1399,265,394]" box="[234,710,265,291]" pageId="15" pageNumber="105">
<emphasis id="B907FE490A6AD14E96883C12FE01FED0" bold="true" box="[234,445,265,291]" pageId="15" pageNumber="105">
Female
<typeStatus id="54C89CF90A6AD14E972F3C12FE01FED0" box="[333,445,265,291]" pageId="15" pageNumber="105" type="paratype">paratype</typeStatus>
</emphasis>
(no. TC-012, NHMR)
</paragraph>
<paragraph id="8BCC225B0A6AD14E96DF3C37FCE4FE79" blockId="15.[189,1399,265,394]" pageId="15" pageNumber="105">
<emphasis id="B907FE490A6AD14E96DF3C37FF40FEB6" box="[189,252,300,325]" italics="true" pageId="15" pageNumber="105">Note.</emphasis>
The females available for description are small and might not be reproductive adults, the spermathecae of the here described
<typeStatus id="54C89CF90A6AD14E94073C55FD76FE9B" box="[613,714,334,360]" pageId="15" pageNumber="105" type="paratype">paratype</typeStatus>
are however sufficiently developed for study. Specimens, 6 months preserved in 70% ethanol, in perfect condition.
</paragraph>
<paragraph id="8BCC225B0A6AD14E96DF3CAAFE22FDA6" blockId="15.[188,1399,433,597]" pageId="15" pageNumber="105">
GENERAL COLORATION. Carapace cephalic part has wide light brown crest-zone and is generally darker in colour than thoracic part, black margin prominent (
<figureCitation id="13483EDE0A6AD14E95693CC8FCDDFE1D" box="[779,865,467,494]" captionStart="Figs 6469" captionStartId="14.[189,242,1816,1842]" captionTargetBox="[189,1398,1125,1769]" captionTargetId="figure-378@14.[189,1398,1125,1769]" captionTargetPageId="14" captionText="Figs 6469. Nemesia maltensis sp. nov. paratype, ♀ (TC.012, NHMR). 64. Dorsal habitus, note maculae on external femora I &amp; II (m). 65. Left lateral habitus, note numerous maculae on legs arrow indications (m) and the low carapace profile (lp). 66. Ventral anterior prosoma, note indistinct central division (cd) in labial furrow. 67. Prolateral patella III, note three spines (sp) in a row. 68. Ventral spinnerets, note absence of distinct light coloured spigot fields. 69. Spermathecae, ventral view, note median part of the receptacles being folded and twisted (ft)." figureDoi="http://doi.org/10.5281/zenodo.6384587" httpUri="https://zenodo.org/record/6384587/files/figure.png" pageId="15" pageNumber="105">Fig. 64</figureCitation>
), chelicerae bicoloured brown, slightly darker than carapace, abdomen dorsal as in male, palps and legs generally as in male, but dorsal femora lighter coloured and with prominent maculae (
<figureCitation id="13483EDE0A6AD14E94F13F03FCF7FDC0" box="[659,843,536,563]" captionStart="Figs 6469" captionStartId="14.[189,242,1816,1842]" captionTargetBox="[189,1398,1125,1769]" captionTargetId="figure-378@14.[189,1398,1125,1769]" captionTargetPageId="14" captionText="Figs 6469. Nemesia maltensis sp. nov. paratype, ♀ (TC.012, NHMR). 64. Dorsal habitus, note maculae on external femora I &amp; II (m). 65. Left lateral habitus, note numerous maculae on legs arrow indications (m) and the low carapace profile (lp). 66. Ventral anterior prosoma, note indistinct central division (cd) in labial furrow. 67. Prolateral patella III, note three spines (sp) in a row. 68. Ventral spinnerets, note absence of distinct light coloured spigot fields. 69. Spermathecae, ventral view, note median part of the receptacles being folded and twisted (ft)." figureDoi="http://doi.org/10.5281/zenodo.6384587" httpUri="https://zenodo.org/record/6384587/files/figure.png" pageId="15" pageNumber="105">Figs 6465, 67</figureCitation>
), sternum and labium slightly darker coloured than ventral coxae.
</paragraph>
<paragraph id="8BCC225B0A6AD14E96DF3F60FC35FD28" blockId="15.[189,1399,635,731]" pageId="15" pageNumber="105">
CARAPACE. Longer than wide (CW/CL 0.7), sparsely covered with very fine black pubescence, bristles centrally in longitudinal row on crest-zone. Cephalic part only slightly elevated and gradually sloping down over the thoracic part to the posterior margin (
<figureCitation id="13483EDE0A6AD14E95463FDBFCC0FD28" box="[804,892,704,731]" captionStart="Figs 6469" captionStartId="14.[189,242,1816,1842]" captionTargetBox="[189,1398,1125,1769]" captionTargetId="figure-378@14.[189,1398,1125,1769]" captionTargetPageId="14" captionText="Figs 6469. Nemesia maltensis sp. nov. paratype, ♀ (TC.012, NHMR). 64. Dorsal habitus, note maculae on external femora I &amp; II (m). 65. Left lateral habitus, note numerous maculae on legs arrow indications (m) and the low carapace profile (lp). 66. Ventral anterior prosoma, note indistinct central division (cd) in labial furrow. 67. Prolateral patella III, note three spines (sp) in a row. 68. Ventral spinnerets, note absence of distinct light coloured spigot fields. 69. Spermathecae, ventral view, note median part of the receptacles being folded and twisted (ft)." figureDoi="http://doi.org/10.5281/zenodo.6384587" httpUri="https://zenodo.org/record/6384587/files/figure.png" pageId="15" pageNumber="105">Fig. 65</figureCitation>
).
</paragraph>
<paragraph id="8BCC225B0A6AD14E96DF3E1AFEFEFCAC" blockId="15.[189,1399,768,863]" pageId="15" pageNumber="105">EYES. Ocular-tubercle as in male, eye-group rectangular, twice as wide as long (EL/PR 0.53), AME less than their diameter apart (dis.AME/dia.AME 0.92), dis.ALEPLE less than ½ dia.ALE (ALEPLE/ ALE 0.42).</paragraph>
<paragraph id="8BCC225B0A6AD14E96DF3E9DFC34FC31" blockId="15.[188,1398,902,962]" pageId="15" pageNumber="105">CHELICERAE. Stronger developed than in male with distinct rastellum, prolateral row of furrow teeth with six conical teeth, fangs sharp, bent, ventral un-serrated ridge.</paragraph>
<caption id="DF0C72D30A6AD14E96DF3A45FACBF811" ID-DOI="http://doi.org/10.5281/zenodo.6384589" ID-Zenodo-Dep="6384589" httpUri="https://zenodo.org/record/6384589/files/figure.png" pageId="15" pageNumber="105" startId="15.[189,242,1886,1912]" targetBox="[189,1398,1037,1843]" targetPageId="15">
<paragraph id="8BCC225B0A6AD14E96DF3A45FACBF811" blockId="15.[188,1399,1886,2019]" pageId="15" pageNumber="105">
<emphasis id="B907FE490A6AD14E96DF3A45FEF5F88B" bold="true" box="[189,329,1886,1912]" pageId="15" pageNumber="105">Figs 7073.</emphasis>
<taxonomicName id="4C7359D80A6AD14E972D3A44FD91F88B" authority="Cassar &amp; Mifsud &amp; Decae, 2022" authorityName="Cassar &amp; Mifsud &amp; Decae" authorityYear="2022" box="[335,557,1886,1912]" class="Magnoliopsida" family="Scrophulariaceae" genus="Nemesia" kingdom="Plantae" order="Lamiales" pageId="15" pageNumber="105" phylum="Tracheophyta" rank="species" species="maltensis" status="sp. nov.">
<emphasis id="B907FE490A6AD14E972D3A44FD91F88B" box="[335,557,1886,1912]" italics="true" pageId="15" pageNumber="105">Nemesia maltensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23443320A6AD14E94513A44FD32F88A" box="[563,654,1887,1913]" pageId="15" pageNumber="105" rank="species">sp. nov.</taxonomicNameLabel>
observations.
<emphasis id="B907FE490A6AD14E95563A45FCEDF88B" bold="true" box="[820,849,1886,1912]" pageId="15" pageNumber="105">70</emphasis>
. Natural habitat.
<emphasis id="B907FE490A6AD14E92793A45FB84F88B" bold="true" box="[1051,1080,1886,1912]" pageId="15" pageNumber="105">71</emphasis>
. Borrow top cut open, note virtual absence of a silk lining of the burrow shaft and the compacted soil from which the burrow walls derive most of their structural integrity.
<emphasis id="B907FE490A6AD14E94C03ABEFD01F84C" bold="true" box="[674,701,1957,1983]" pageId="15" pageNumber="105">72</emphasis>
. Natural burrow entrance flush with the soil surface, note absence of trapdoor.
<emphasis id="B907FE490A6AD14E97CE3AD3FE75F811" bold="true" box="[428,457,1992,2018]" pageId="15" pageNumber="105">73</emphasis>
. Two surface openings of the bifurcated burrow seen in an observation container.
</paragraph>
</caption>
<paragraph id="8BCC225B0A75D15196DF3C12FB8BFE9B" blockId="16.[189,1398,265,361]" pageId="16" pageNumber="106">
VENTRAL PROSOMA. Maxillae small rounded distal lobe, few knobby cuspules on proximo-anterior margin. Labium: twice as wide as long (LW/LL 2.3), cuspules absent, labial furrow centrally divided (
<figureCitation id="13483EDE0A75D15196AA3C55FEA3FE9A" box="[200,287,334,361]" captionStart="Figs 6469" captionStartId="14.[189,242,1816,1842]" captionTargetBox="[189,1398,1125,1769]" captionTargetId="figure-378@14.[189,1398,1125,1769]" captionTargetPageId="14" captionText="Figs 6469. Nemesia maltensis sp. nov. paratype, ♀ (TC.012, NHMR). 64. Dorsal habitus, note maculae on external femora I &amp; II (m). 65. Left lateral habitus, note numerous maculae on legs arrow indications (m) and the low carapace profile (lp). 66. Ventral anterior prosoma, note indistinct central division (cd) in labial furrow. 67. Prolateral patella III, note three spines (sp) in a row. 68. Ventral spinnerets, note absence of distinct light coloured spigot fields. 69. Spermathecae, ventral view, note median part of the receptacles being folded and twisted (ft)." figureDoi="http://doi.org/10.5281/zenodo.6384587" httpUri="https://zenodo.org/record/6384587/files/figure.png" pageId="16" pageNumber="106">Fig. 66</figureCitation>
). Sternum: longer than wide (SW/SL 0.7), sigilla not distinguished.
</paragraph>
<paragraph id="8BCC225B0A75D15196DF3C9CFE79FDFB" blockId="16.[189,1401,391,521]" pageId="16" pageNumber="106">PALPS. Femur and patella spineless, tibia ventral and prolateral sharp distally pointing spines and dorsal long parallel rows of trichobothria, tarsus, distally pointed, ventral half fully scopulate with group of sharp distally pointing spines, dorsal row trichobothria in central part of article, palpal claw with short proximal row of teeth.</paragraph>
<paragraph id="8BCC225B0A75D15196DF3F33FBE3FD38" blockId="16.[188,1400,551,715]" pageId="16" pageNumber="106">
LEGS. All femora with external maculae (
<figureCitation id="13483EDE0A75D15194D63F3CFCFDFDB1" box="[692,833,551,578]" captionStart="Figs 6469" captionStartId="14.[189,242,1816,1842]" captionTargetBox="[189,1398,1125,1769]" captionTargetId="figure-378@14.[189,1398,1125,1769]" captionTargetPageId="14" captionText="Figs 6469. Nemesia maltensis sp. nov. paratype, ♀ (TC.012, NHMR). 64. Dorsal habitus, note maculae on external femora I &amp; II (m). 65. Left lateral habitus, note numerous maculae on legs arrow indications (m) and the low carapace profile (lp). 66. Ventral anterior prosoma, note indistinct central division (cd) in labial furrow. 67. Prolateral patella III, note three spines (sp) in a row. 68. Ventral spinnerets, note absence of distinct light coloured spigot fields. 69. Spermathecae, ventral view, note median part of the receptacles being folded and twisted (ft)." figureDoi="http://doi.org/10.5281/zenodo.6384587" httpUri="https://zenodo.org/record/6384587/files/figure.png" pageId="16" pageNumber="106">Figs 6465</figureCitation>
). Anterior legs with ventro-prolateral scopulae extending from tip of tarsus to distal patella, ventral spines on tibia and metatarsus. Posterior legs with dense groups of spiny bristles prolateral on distal femur and dorsal patella. Leg III, patella with three prolateral spines (
<figureCitation id="13483EDE0A75D15197EF3F95FE5EFD5A" box="[397,482,654,681]" captionStart="Figs 6469" captionStartId="14.[189,242,1816,1842]" captionTargetBox="[189,1398,1125,1769]" captionTargetId="figure-378@14.[189,1398,1125,1769]" captionTargetPageId="14" captionText="Figs 6469. Nemesia maltensis sp. nov. paratype, ♀ (TC.012, NHMR). 64. Dorsal habitus, note maculae on external femora I &amp; II (m). 65. Left lateral habitus, note numerous maculae on legs arrow indications (m) and the low carapace profile (lp). 66. Ventral anterior prosoma, note indistinct central division (cd) in labial furrow. 67. Prolateral patella III, note three spines (sp) in a row. 68. Ventral spinnerets, note absence of distinct light coloured spigot fields. 69. Spermathecae, ventral view, note median part of the receptacles being folded and twisted (ft)." figureDoi="http://doi.org/10.5281/zenodo.6384587" httpUri="https://zenodo.org/record/6384587/files/figure.png" pageId="16" pageNumber="106">Fig. 67</figureCitation>
) and one retrolateral spine. Leg IV patella spineless, tibia&gt; femur&gt; metatarsus (Fem4/Met4 1,1, Tib4/Met4 1,3). Leg formula: 4123. PTC and ATC as in male.
</paragraph>
<paragraph id="8BCC225B0A75D15196DF3FF1FE0CFC98" blockId="16.[188,1398,746,875]" pageId="16" pageNumber="106">
OPISTHOSOMA. Generally as in male, anterior bristle group less distinct, spinnerets without sharply defined spigot-fields (
<figureCitation id="13483EDE0A75D151973C3E17FE0AFCD4" box="[350,438,780,807]" captionStart="Figs 6469" captionStartId="14.[189,242,1816,1842]" captionTargetBox="[189,1398,1125,1769]" captionTargetId="figure-378@14.[189,1398,1125,1769]" captionTargetPageId="14" captionText="Figs 6469. Nemesia maltensis sp. nov. paratype, ♀ (TC.012, NHMR). 64. Dorsal habitus, note maculae on external femora I &amp; II (m). 65. Left lateral habitus, note numerous maculae on legs arrow indications (m) and the low carapace profile (lp). 66. Ventral anterior prosoma, note indistinct central division (cd) in labial furrow. 67. Prolateral patella III, note three spines (sp) in a row. 68. Ventral spinnerets, note absence of distinct light coloured spigot fields. 69. Spermathecae, ventral view, note median part of the receptacles being folded and twisted (ft)." figureDoi="http://doi.org/10.5281/zenodo.6384587" httpUri="https://zenodo.org/record/6384587/files/figure.png" pageId="16" pageNumber="106">Fig. 68</figureCitation>
). Spermathecae, tripartite, tube shaped, proximal part with dense concentration of pigmented cells, proximally widest, median part folded and twisted (
<figureCitation id="13483EDE0A75D15195BC3E35FB8AFCBA" box="[990,1078,814,841]" captionStart="Figs 6469" captionStartId="14.[189,242,1816,1842]" captionTargetBox="[189,1398,1125,1769]" captionTargetId="figure-378@14.[189,1398,1125,1769]" captionTargetPageId="14" captionText="Figs 6469. Nemesia maltensis sp. nov. paratype, ♀ (TC.012, NHMR). 64. Dorsal habitus, note maculae on external femora I &amp; II (m). 65. Left lateral habitus, note numerous maculae on legs arrow indications (m) and the low carapace profile (lp). 66. Ventral anterior prosoma, note indistinct central division (cd) in labial furrow. 67. Prolateral patella III, note three spines (sp) in a row. 68. Ventral spinnerets, note absence of distinct light coloured spigot fields. 69. Spermathecae, ventral view, note median part of the receptacles being folded and twisted (ft)." figureDoi="http://doi.org/10.5281/zenodo.6384587" httpUri="https://zenodo.org/record/6384587/files/figure.png" pageId="16" pageNumber="106">Fig. 69</figureCitation>
), distal part digitiform with few pigmented cells.
</paragraph>
<paragraph id="8BCC225B0A75D15196DF3E90FDF9FBDD" blockId="16.[188,1399,906,1070]" pageId="16" pageNumber="106">MEASUREMENTS. TBL = 9.7; CL = 3.9; CW = 2.9; CP = 2.3; AR = 0.71; PR = 0.74; EL = 0.39; dia.ALE = 0.18; dia.PLE 0.13; dia.AME = 0.10; dia.PME = 0.10; dis.AMEAME = 0.10; dis.ALEPLE = 0.05; SL = 1.7; SW = 1.3; LL = 0.3; LW = 0.6; Palp = 5.1 (1.1 + 1.1 + 1.1 + 1.8); Leg I = 8.3 (1.0 + 1.3 + 1.6 + 1.8 + 2.6); Leg II = 7.4 (1.0 + 1.3 + 1.4 + 1.5 + 2.2); Leg III = 7.1 (1.1 + 1.6 + 1.1 + 1.3 + 2.0); Leg IV = 11.3 (1.1 + 2.4 + 3.1 + 2.0 + 2.7).</paragraph>
<paragraph id="8BCC225B0A75D15196DF3956FE21FB03" blockId="16.[189,1399,1101,1265]" pageId="16" pageNumber="106">VARIATION FEMALE (n = 2). TBL = 9.2, 9.7; CL = 3.4, 3.9; CW = 2.3, 2.9; CP = 2.0, 2.3; AR = 0.62, 0.71; PR = 0.61, 0.74; EL = 0.33, 0.39; dia.ALE = 0.18, 0.19; dia.PLE = 0.13, 0.16; dia.AME = 0.10, 0.12; dia.PME = 0.10; dis.AMEAME = 0.10, 0.11; dis.ALEPLE = 0.05, 0.08; SL = 1.7, 2.1; SW = 1.3, 1.5; LL = 0.3, 0.4; LW = 0.6, 0.8; Palp = 4.2, 5.1; Leg I = 6.7, 8.3; Leg II = 6.3, 7.4; Leg III = 5.9, 7.1; Leg IV = 9.6, 11.3.</paragraph>
</subSubSection>
<subSubSection id="C36971D00A75D15196DF3814FE63F921" pageId="16" pageNumber="106" type="discussion">
<paragraph id="8BCC225B0A75D15196DF3814FEDFFADA" blockId="16.[189,355,1295,1321]" box="[189,355,1295,1321]" pageId="16" pageNumber="106">
<heading id="D08495370A75D15196DF3814FEDFFADA" bold="true" box="[189,355,1295,1321]" fontSize="11" level="3" pageId="16" pageNumber="106" reason="3">
<emphasis id="B907FE490A75D15196DF3814FEDFFADA" bold="true" box="[189,355,1295,1321]" pageId="16" pageNumber="106">Observations</emphasis>
</heading>
</paragraph>
<paragraph id="8BCC225B0A75D15196DF3825FE63F921" blockId="16.[188,1401,1341,1746]" pageId="16" pageNumber="106">
In the field (
<figureCitation id="13483EDE0A75D15197363826FE12FAA4" box="[340,430,1341,1367]" captionStart="Figs 7073" captionStartId="15.[189,242,1886,1912]" captionTargetBox="[189,1398,1037,1843]" captionTargetId="figure-227@15.[189,1398,1037,1843]" captionTargetPageId="15" captionText="Figs 7073. Nemesia maltensis sp. nov. observations. 70. Natural habitat. 71. Borrow top cut open, note virtual absence of a silk lining of the burrow shaft and the compacted soil from which the burrow walls derive most of their structural integrity. 72. Natural burrow entrance flush with the soil surface, note absence of trapdoor. 73. Two surface openings of the bifurcated burrow seen in an observation container." figureDoi="http://doi.org/10.5281/zenodo.6384589" httpUri="https://zenodo.org/record/6384589/files/figure.png" pageId="16" pageNumber="106">Fig. 70</figureCitation>
), the burrows of this species appeared to be completely lidless, with the burrow entrance opening at the ground surface directly (
<figureCitation id="13483EDE0A75D151949D3844FCE6FA89" box="[767,858,1375,1402]" captionStart="Figs 7073" captionStartId="15.[189,242,1886,1912]" captionTargetBox="[189,1398,1037,1843]" captionTargetId="figure-227@15.[189,1398,1037,1843]" captionTargetPageId="15" captionText="Figs 7073. Nemesia maltensis sp. nov. observations. 70. Natural habitat. 71. Borrow top cut open, note virtual absence of a silk lining of the burrow shaft and the compacted soil from which the burrow walls derive most of their structural integrity. 72. Natural burrow entrance flush with the soil surface, note absence of trapdoor. 73. Two surface openings of the bifurcated burrow seen in an observation container." figureDoi="http://doi.org/10.5281/zenodo.6384589" httpUri="https://zenodo.org/record/6384589/files/figure.png" pageId="16" pageNumber="106">Fig. 72</figureCitation>
). The wall of the burrow is lined with almost imperceptibly thin strands of silk from the inside; but the burrow walls derive most of their structural integrity from the compaction of the soil around them (
<figureCitation id="13483EDE0A75D151952038BFFC25FA4D" box="[834,921,1444,1470]" captionStart="Figs 7073" captionStartId="15.[189,242,1886,1912]" captionTargetBox="[189,1398,1037,1843]" captionTargetId="figure-227@15.[189,1398,1037,1843]" captionTargetPageId="15" captionText="Figs 7073. Nemesia maltensis sp. nov. observations. 70. Natural habitat. 71. Borrow top cut open, note virtual absence of a silk lining of the burrow shaft and the compacted soil from which the burrow walls derive most of their structural integrity. 72. Natural burrow entrance flush with the soil surface, note absence of trapdoor. 73. Two surface openings of the bifurcated burrow seen in an observation container." figureDoi="http://doi.org/10.5281/zenodo.6384589" httpUri="https://zenodo.org/record/6384589/files/figure.png" pageId="16" pageNumber="106">Fig. 71</figureCitation>
). In captivity, once soil was moistened, a female constructed a bifurcated burrow with two shafts opening at the soil surface with no lids (
<figureCitation id="13483EDE0A75D151937038DCFAD4FA12" box="[1298,1384,1479,1505]" captionStart="Figs 7073" captionStartId="15.[189,242,1886,1912]" captionTargetBox="[189,1398,1037,1843]" captionTargetId="figure-227@15.[189,1398,1037,1843]" captionTargetPageId="15" captionText="Figs 7073. Nemesia maltensis sp. nov. observations. 70. Natural habitat. 71. Borrow top cut open, note virtual absence of a silk lining of the burrow shaft and the compacted soil from which the burrow walls derive most of their structural integrity. 72. Natural burrow entrance flush with the soil surface, note absence of trapdoor. 73. Two surface openings of the bifurcated burrow seen in an observation container." figureDoi="http://doi.org/10.5281/zenodo.6384589" httpUri="https://zenodo.org/record/6384589/files/figure.png" pageId="16" pageNumber="106">Fig. 73</figureCitation>
), joining together into one shaft deeper into the soil. Females were collected from burrows some
<quantity id="4C8B8FBE0A75D151934E38F2FACAF9F7" box="[1324,1398,1513,1540]" metricMagnitude="-1" metricUnit="m" metricValue="1.5" pageId="16" pageNumber="106" unit="cm" value="15.0">15 cm</quantity>
deep in the soil, congregated at the edge of a small boulder embedded in deep soil in a wooded area populated by trees of
<taxonomicName id="4C7359D80A75D15197A53B34FDC8F9BB" box="[455,628,1582,1608]" class="Magnoliopsida" family="Lauraceae" genus="Laurus" kingdom="Plantae" order="Laurales" pageId="16" pageNumber="106" phylum="Tracheophyta" rank="species" species="nobilis">
<emphasis id="B907FE490A75D15197A53B34FDC8F9BB" box="[455,628,1582,1608]" italics="true" pageId="16" pageNumber="106">Laurus nobilis</emphasis>
</taxonomicName>
L.,
<taxonomicName id="4C7359D80A75D15194C43B35FCEBF9BB" box="[678,855,1582,1608]" class="Magnoliopsida" family="Oleaceae" genus="Olea" kingdom="Plantae" order="Lamiales" pageId="16" pageNumber="106" phylum="Tracheophyta" rank="species" species="europaea">
<emphasis id="B907FE490A75D15194C43B35FCEBF9BB" box="[678,855,1582,1608]" italics="true" pageId="16" pageNumber="106">Olea europaea</emphasis>
</taxonomicName>
,
<taxonomicName id="4C7359D80A75D15195053B35FBF1F9BB" box="[871,1101,1582,1608]" class="Magnoliopsida" family="Rhamnaceae" genus="Rhamnus" kingdom="Plantae" order="Rosales" pageId="16" pageNumber="106" phylum="Tracheophyta" rank="species" species="alaternus">
<emphasis id="B907FE490A75D15195053B35FBF1F9BB" box="[871,1101,1582,1608]" italics="true" pageId="16" pageNumber="106">Rhamnus alaternus</emphasis>
</taxonomicName>
L. and
<taxonomicName id="4C7359D80A75D15192D23B34FEE6F998" authority="Mill. Burrow" authorityName="Mill. Burrow" class="Pinopsida" family="Pinaceae" genus="Pinus" kingdom="Plantae" order="Pinales" pageId="16" pageNumber="106" phylum="Tracheophyta" rank="species" species="halepensis">
<emphasis id="B907FE490A75D15192D23B34FAC5F9BB" box="[1200,1401,1582,1608]" italics="true" pageId="16" pageNumber="106">Pinus halepensis</emphasis>
Mill. Burrow
</taxonomicName>
entrances always appeared at the soil surface at the edge of the boulder or, in one instance, at the base of the thick stems of an
<taxonomicName id="4C7359D80A75D15194363B68FCACF97E" box="[596,784,1651,1677]" class="Magnoliopsida" family="Acanthaceae" genus="Acanthus" kingdom="Plantae" order="Lamiales" pageId="16" pageNumber="106" phylum="Tracheophyta" rank="species" species="mollis">
<emphasis id="B907FE490A75D15194363B68FCACF97E" box="[596,784,1651,1677]" italics="true" pageId="16" pageNumber="106">Acanthus mollis</emphasis>
</taxonomicName>
L. plant nearby. Males were collected in pitfall traps after several heavy rainfall events which occurred in September the so called “first rains” which bring an end to the dry season.
</paragraph>
</subSubSection>
</treatment>
</document>