341 lines
47 KiB
XML
341 lines
47 KiB
XML
<document id="89AB879E8468896B119980C4178CE594" ID-DOI="10.11646/zootaxa.4012.1.9" ID-GBIF-Dataset="4c0d8376-71b9-42d1-8dbc-08e0ff16338a" ID-ISSN="1175-5326" ID-Zenodo-Dep="244632" ID-ZooBank="55822610-6FE6-4218-A412-A080F3218F51" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461239579310" checkinUser="plazi" docAuthor="Keith, Denis, Sabatinelli, Guido & Uliana, Marco" docDate="2015" docId="03DA87884C34FFECFF0FB476FA1E6BB1" docLanguage="en" docName="zt04012p180.pdf" docOrigin="Zootaxa 4012 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pharaonus Blanchard 1851" docType="key" docVersion="8" lastPageNumber="179" masterDocId="FFE3FFF04C3FFFE0FF98B26BFF826853" masterDocTitle="Synopsis of the genus Pharaonus (Coleoptera: Scarabaeidae: Rutelinae), with descriptions of new taxa" masterLastPageNumber="180" masterPageNumber="167" pageNumber="178" updateTime="1698605019073" updateUser="plazi">
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<mods:title id="33462B89D3683A4F4802DCAB021BB8B4">Synopsis of the genus Pharaonus (Coleoptera: Scarabaeidae: Rutelinae), with descriptions of new taxa</mods:title>
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<mods:namePart id="FABA3E8F5270E92E8B35EA57909F2DFC">Keith, Denis</mods:namePart>
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<mods:namePart id="B12D08F84920EA5E3693A26E790F7116">Sabatinelli, Guido</mods:namePart>
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<mods:namePart id="F0B240DB91CCA275207331E4FA3C60DC">Uliana, Marco</mods:namePart>
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<mods:title id="AF16B22B810071ECFDFA5C8426F1AED5">Zootaxa</mods:title>
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<mods:date id="2AFECA6B2766EBC50ACA58438940A377">2015</mods:date>
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<mods:number id="F40F8A9740BDA8D011E164F37ACD1C00">4012</mods:number>
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<treatment id="03DA87884C34FFECFF0FB476FA1E6BB1" ID-DOI="http://doi.org/10.5281/zenodo.5694718" ID-GBIF-Taxon="119632614" ID-Zenodo-Dep="5694718" LSID="urn:lsid:plazi:treatment:03DA87884C34FFECFF0FB476FA1E6BB1" httpUri="http://treatment.plazi.org/id/03DA87884C34FFECFF0FB476FA1E6BB1" lastPageId="12" lastPageNumber="179" pageId="11" pageNumber="178">
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<subSubSection id="C36965154C34FFEBFF0FB476FC2D6E96" pageId="11" pageNumber="178" type="nomenclature">
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<paragraph id="8BCC369E4C34FFEBFF0FB476FCF66E64" blockId="11.[151,884,1565,1591]" box="[151,884,1565,1591]" pageId="11" pageNumber="178">
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<heading id="D08481F24C34FFEBFF0FB476FCF66E64" bold="true" box="[151,884,1565,1591]" fontSize="11" level="1" pageId="11" pageNumber="178" reason="1">
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<emphasis id="B907EA8C4C34FFEBFF0FB476FCF66E64" bold="true" box="[151,884,1565,1591]" pageId="11" pageNumber="178">
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Key to the species of the genus
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<taxonomicName id="4C734D1D4C34FFEBFD81B476FCF66E64" authority="Blanchard, 1851" authorityName="Blanchard" authorityYear="1851" box="[537,884,1565,1591]" class="Insecta" family="Rutelidae" genus="Pharaonus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="178" phylum="Arthropoda" rank="genus">
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<emphasis id="B907EA8C4C34FFEBFD81B476FD1D6E64" bold="true" box="[537,671,1565,1591]" italics="true" pageId="11" pageNumber="178">Pharaonus</emphasis>
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<bibRefCitation id="EFE24B6F4C34FFEBFD3EB476FCF66E64" author="Blanchard" box="[678,884,1565,1591]" pageId="11" pageNumber="178" refString="Blanchard, C. E. (1851) Ordre des Coleopteres. In: Milne-Edwards, H., Blanchard, C. E. & Lucas, H. (Eds.), Catalogue de la collection Entomologique du Museum d'Histoire Naturelle de Paris. Classe des Insectes. Tome 2. Gide et Baudry, Paris, pp. 129 - 240." type="journal article" year="1851">Blanchard, 1851</bibRefCitation>
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</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BCC369E4C34FFEBFF0FB40EFC2D6E96" blockId="11.[151,1436,1636,1733]" pageId="11" pageNumber="178">
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We only examined female
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<taxonomicName id="4C734D1D4C34FFEBFE55B40EFD6B6E2E" box="[461,745,1636,1661]" class="Insecta" family="Rutelidae" genus="Pharaonus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="178" phylum="Arthropoda" rank="species" species="adelphus" subGenus="Pharaonus">
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<emphasis id="B907EA8C4C34FFEBFE55B40EFE596E2F" box="[461,475,1637,1660]" italics="true" pageId="11" pageNumber="178">P</emphasis>
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. (
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<emphasis id="B907EA8C4C34FFEBFE6DB40FFDF26E2E" box="[501,624,1636,1661]" italics="true" pageId="11" pageNumber="178">Pharaonus</emphasis>
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)
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<emphasis id="B907EA8C4C34FFEBFD1CB40FFD6B6E2E" box="[644,745,1636,1661]" italics="true" pageId="11" pageNumber="178">adelphus</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="4C734D1D4C34FFEBFD62B40EFC546E2E" box="[762,982,1636,1661]" class="Insecta" family="Rutelidae" genus="Pharaonus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="178" phylum="Arthropoda" rank="species" species="varicoloreus" subGenus="Pharaonus">
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<emphasis id="B907EA8C4C34FFEBFD62B40EFC8A6E2F" box="[762,776,1637,1660]" italics="true" pageId="11" pageNumber="178">P</emphasis>
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. (
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<emphasis id="B907EA8C4C34FFEBFCBAB40EFCB26E2F" box="[802,816,1637,1660]" italics="true" pageId="11" pageNumber="178">P</emphasis>
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.)
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<emphasis id="B907EA8C4C34FFEBFCD2B40FFC546E2E" box="[842,982,1636,1661]" italics="true" pageId="11" pageNumber="178">varicoloreus</emphasis>
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</taxonomicName>
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, and
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<taxonomicName id="4C734D1D4C34FFEBFB81B40EFAA66E2E" box="[1049,1316,1636,1661]" class="Insecta" family="Rutelidae" genus="Pharaonus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="178" phylum="Arthropoda" rank="species" species="lederi" subGenus="Tamerlanius">
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<emphasis id="B907EA8C4C34FFEBFB81B40EFBA56E2F" box="[1049,1063,1637,1660]" italics="true" pageId="11" pageNumber="178">P</emphasis>
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. (
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<emphasis id="B907EA8C4C34FFEBFBD9B40FFB4E6E2E" box="[1089,1228,1636,1661]" italics="true" pageId="11" pageNumber="178">Tamerlanius</emphasis>
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)
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<emphasis id="B907EA8C4C34FFEBFB47B40FFAA66E2E" box="[1247,1316,1636,1661]" italics="true" pageId="11" pageNumber="178">lederi</emphasis>
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</taxonomicName>
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. Based on these species, the following secondary sexual characters can be used for determining the sex: in males the antennal clubs are longer, the protarsi stouter, the inner claw of prolegs broader.
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</paragraph>
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</subSubSection>
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<subSubSection id="C36965154C34FFECFF0FB49FFA1E6BB1" lastPageId="12" lastPageNumber="179" pageId="11" pageNumber="178" type="key">
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<keyStep id="30872E3B4C34FFEBFF0FB49FFA1E6FC8" pageId="11" pageNumber="178">
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<paragraph id="8BCC369E4C34FFEBFF0FB49FFA1E6F10" blockId="11.[151,1437,1780,2035]" pageId="11" pageNumber="178">
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<keyLead id="308295AB4C34FFEBFF0FB49FFA1E6F10" pageId="11" pageNumber="178">
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1. Clypeus with anterior margin raised, sides parallel, anterior angles rounded but right. Labrum not visible from above. In males: inner claw of the prolegs narrow and sharp (
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<figureCitation id="13482A1B4C34FFEBFD16B57AFD456F75" box="[654,711,1809,1830]" captionStart="FIGURES 1 – 2" captionStartId="1.[151,264,1730,1752]" captionTargetBox="[250,1348,1407,1701]" captionTargetId="figure@1.[229,1358,1399,1705]" captionTargetPageId="1" captionText="FIGURES 1 – 2. Proclaws in Pharaonus (Pharaonus) (1) and in Pharaonus (Tamerlanius) (2)." httpUri="https://zenodo.org/record/244633/files/figure.png" pageId="11" pageNumber="178">Fig. 1</figureCitation>
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); antennal club longer, at least 1.7 times as long as the rest of the antenna. Species from North Africa to the Caucasus............................................... subgenus
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<taxonomicName id="4C734D1D4C34FFEBFA82B545FA016F10" box="[1306,1411,1838,1859]" class="Insecta" family="Rutelidae" genus="Pharaonus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="178" phylum="Arthropoda" rank="genus">
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<emphasis id="B907EA8C4C34FFEBFA82B545FA016F10" box="[1306,1411,1838,1859]" italics="true" pageId="11" pageNumber="178">Pharaonus</emphasis>
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</taxonomicName>
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||
: 2
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</keyLead>
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||
</paragraph>
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||
<paragraph id="8BCC369E4C34FFEBFF0FB527FA1E6FC8" blockId="11.[151,1437,1780,2035]" pageId="11" pageNumber="178">
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||
<keyLead id="308295AB4C34FFEBFF0FB527FA1E6FC8" pageId="11" pageNumber="178">
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- Clypeus with anterior margin bordered but not raised, sides convergent forward, anterior angles obtuse. Labrum visible from above. In males: inner claw of the prolegs broad, subrectangular, with truncate apex (
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<figureCitation id="13482A1B4C34FFEBFB93B502FBC46F2D" box="[1035,1094,1897,1918]" captionStart="FIGURES 1 – 2" captionStartId="1.[151,264,1730,1752]" captionTargetBox="[250,1348,1407,1701]" captionTargetId="figure@1.[229,1358,1399,1705]" captionTargetPageId="1" captionText="FIGURES 1 – 2. Proclaws in Pharaonus (Pharaonus) (1) and in Pharaonus (Tamerlanius) (2)." httpUri="https://zenodo.org/record/244633/files/figure.png" pageId="11" pageNumber="178">Fig. 2</figureCitation>
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); antennal club shorter, 1.3 times as long or shorter compared to the rest of the antenna. Species from central Asia.................. subgenus
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<emphasis id="B907EA8C4C34FFEBFA94B5EDFA016FC8" box="[1292,1411,1926,1947]" italics="true" pageId="11" pageNumber="178">Tamerlanius</emphasis>
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: 7
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</keyLead>
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</paragraph>
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||
</keyStep>
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<keyStep id="30872E3B4C34FFEBFF0FB5CFFA1E6FA0" pageId="11" pageNumber="178">
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<paragraph id="8BCC369E4C34FFEBFF0FB5CFFA146F85" blockId="11.[151,1437,1780,2035]" pageId="11" pageNumber="178">
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<keyLead id="308295AB4C34FFEBFF0FB5CFFA146F85" pageId="11" pageNumber="178">
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2. (1) Pronotum (except margins) glabrous. Protibia of male tridentate. Black to brownish black, elytra seldom light brown.
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<collectingCountry id="F364760E4C34FFEBFAF7B5CFFA1A6FEA" box="[1391,1432,1956,1977]" name="Iran" pageId="11" pageNumber="178">Iran</collectingCountry>
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,
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<collectingCountry id="F364760E4C34FFEBFF79B5AAFE886F85" box="[225,266,1985,2006]" name="Iraq" pageId="11" pageNumber="178">Iraq</collectingCountry>
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,
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<collectingCountry id="F364760E4C34FFEBFE8BB5AAFED76F85" box="[275,341,1985,2006]" name="Jordan" pageId="11" pageNumber="178">Jordan</collectingCountry>
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,
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<collectingCountry id="F364760E4C34FFEBFEC7B5AAFE2A6F85" box="[351,424,1985,2006]" name="Kuwait" pageId="11" pageNumber="178">Kuwait</collectingCountry>
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,
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<collectingCountry id="F364760E4C34FFEBFE2AB5AAFE656F85" box="[434,487,1985,2006]" name="Syria" pageId="11" pageNumber="178">Syria</collectingCountry>
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,
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<collectingCountry id="F364760E4C34FFEBFE68B5AAFDB66F85" box="[496,564,1985,2006]" name="Turkey" pageId="11" pageNumber="178">Turkey</collectingCountry>
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||
(
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<figureCitation id="13482A1B4C34FFEBFDDAB5AAFD286F85" box="[578,682,1985,2006]" captionStart="FIGURES 7 – 10" captionStartId="5.[151,264,1955,1977]" captionTargetBox="[211,1377,193,1914]" captionTargetId="figure@5.[196,1390,193,1933]" captionTargetPageId="5" captionText="FIGURES 7 – 10. Habitus of Pharaonus (Pharaonus) species. 7: P. fasciculatus, male (Egypt). 8: P. saharicus, male paratype (Tamrit, Tassili n’Ajjer, Algeria). 9: P. varicoloreus, male (environs of Palymra, Syria). 10: P. varicoloreus, female (Birecik, Turkey)." httpUri="https://zenodo.org/record/244635/files/figure.png" pageId="11" pageNumber="178">Figs. 9, 10</figureCitation>
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,
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<figureCitation id="13482A1B4C34FFEBFD2CB5AAFD4F6F85" box="[692,717,1985,2006]" captionStart="FIGURES 15 – 23" captionStartId="8.[151,264,1847,1869]" captionTargetBox="[174,1402,241,1792]" captionTargetId="figure@8.[174,1405,193,1813]" captionTargetPageId="8" captionText="FIGURES 15 – 23. Genitalia of Pharaonus. 15 – 19: aedeagus in dorsal, ventral, and lateral view of Pharaonus (Pharaonus): 15: P. adelphus, paratype (a-c) and holotype (d) (Kerman, Iran); 16: P. caucasicus (Gorovan, Armenia); 17: P. fasciculatus (Egypt); 18: P. farsensis, holotype (Darab, Iran); 19: P. saharicus, paratype (Tamrit, Tassili n’Ajjer, Algeria); 20: P. varicoloreus (Halfeti, Birecik, Adiyaman, Turkey). 21: female genital sclerites of P. adelphus (Kerman, Bakram-djord [= Bahramjerd], Iran). 22 – 23: aedeagus in dorsal, ventral, and lateral view of Pharaonus (Tamerlanius): 22: P. lederi (Koron 15 km Ekoc. Dzejran, Buckara, Uzbekistan); 23: P. semenowi (Kul'dzhuktau, Kizilkum desert, Uzbekistan)." httpUri="https://zenodo.org/record/244638/files/figure.png" pageId="11" pageNumber="178">20</figureCitation>
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)...............................
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<taxonomicName id="4C734D1D4C34FFEBFBD2B5AAFA146F85" authority="Burmeister, 1844" authorityName="Burmeister" authorityYear="1844" box="[1098,1430,1985,2006]" class="Insecta" family="Rutelidae" genus="Pharaonus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="178" phylum="Arthropoda" rank="species" species="varicoloreus">
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<emphasis id="B907EA8C4C34FFEBFBD2B5AAFB5B6F85" box="[1098,1241,1985,2006]" italics="true" pageId="11" pageNumber="178">P. varicoloreus</emphasis>
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(
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<bibRefCitation id="EFE24B6F4C34FFEBFB7FB5AAFA0D6F85" author="Burmeister" box="[1255,1423,1985,2006]" pageId="11" pageNumber="178" refString="Burmeister, H. C. C. (1844) Handbuch der Entomologie. Vierter Band, Erste Abtheilung. Coleoptera Lamellicornia Anthobia et Phyllophaga systellochela. T. C. F. Enslin, Berlin, 584 pp." type="book" year="1844">Burmeister, 1844</bibRefCitation>
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)
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</taxonomicName>
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</keyLead>
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</paragraph>
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<paragraph id="8BCC369E4C34FFEBFF0FB5B5FA1E6FA0" blockId="11.[151,1437,1780,2035]" box="[151,1436,2014,2035]" pageId="11" pageNumber="178">
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<keyLead id="308295AB4C34FFEBFF0FB5B5FA1E6FA0" box="[151,1436,2014,2035]" pageId="11" pageNumber="178">- Pronotum with long pilosity on most or entire surface. Protibia of male bidentate. Elytra yellowish brown to light brown.... 3</keyLead>
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</paragraph>
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</keyStep>
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||
<keyStep id="30872E3B4C33FFECFF0FB2FCFA1468B4" pageId="12" pageNumber="179">
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<paragraph id="8BCC369E4C33FFECFF0FB2FCFA1E68FF" blockId="12.[151,1437,151,994]" box="[151,1436,151,172]" pageId="12" pageNumber="179">
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<keyLead id="308295AB4C33FFECFF0FB2FCFA1E68FF" box="[151,1436,151,172]" pageId="12" pageNumber="179">3. (2) Pronotum with dense and strong punctation, punctures separated by a distance of less than one puncture diameter......... 4</keyLead>
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</paragraph>
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<paragraph id="8BCC369E4C33FFECFF0FB2DEFA1468B4" blockId="12.[151,1437,151,994]" pageId="12" pageNumber="179">
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<keyLead id="308295AB4C33FFECFF0FB2DEFA1468B4" pageId="12" pageNumber="179">
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- Pronotum with scattered and fine punctation, punctures separated by a distance of more than one puncture diameter.
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<collectingCountry id="F364760E4C33FFECFAD9B2DEFA156899" box="[1345,1431,181,202]" name="Armenia" pageId="12" pageNumber="179">Armenia</collectingCountry>
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, Caucasus,
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<collectingCountry id="F364760E4C33FFECFEDFB2B9FE0968B4" box="[327,395,210,231]" name="Turkey" pageId="12" pageNumber="179">Turkey</collectingCountry>
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||
(
|
||
<figureCitation id="13482A1B4C33FFECFE01B2B9FE5E68B4" box="[409,476,210,231]" captionStart="FIGURES 3 – 6" captionStartId="3.[151,264,1953,1975]" captionTargetBox="[212,1376,196,1915]" captionTargetId="figure@3.[201,1386,193,1932]" captionTargetPageId="3" captionText="FIGURES 3 – 6. Habitus of Pharaonus (Pharaonus) species. 3: P. adelphus, male paratype (Kerman, Iran). 4: P. adelphus, female paratype (Kerman, Bakram-djord [= Bahramjerd], Iran). 5: P. caucasicus, male (Gorovan sands, Armenia). 6: P. farsensis, male paratype (Darab, Iran)." httpUri="https://zenodo.org/record/244634/files/figure.png" pageId="12" pageNumber="179">Figs. 5</figureCitation>
|
||
,
|
||
<figureCitation id="13482A1B4C33FFECFE7FB2B9FE7D68B4" box="[487,511,210,231]" captionStart="FIGURES 15 – 23" captionStartId="8.[151,264,1847,1869]" captionTargetBox="[174,1402,241,1792]" captionTargetId="figure@8.[174,1405,193,1813]" captionTargetPageId="8" captionText="FIGURES 15 – 23. Genitalia of Pharaonus. 15 – 19: aedeagus in dorsal, ventral, and lateral view of Pharaonus (Pharaonus): 15: P. adelphus, paratype (a-c) and holotype (d) (Kerman, Iran); 16: P. caucasicus (Gorovan, Armenia); 17: P. fasciculatus (Egypt); 18: P. farsensis, holotype (Darab, Iran); 19: P. saharicus, paratype (Tamrit, Tassili n’Ajjer, Algeria); 20: P. varicoloreus (Halfeti, Birecik, Adiyaman, Turkey). 21: female genital sclerites of P. adelphus (Kerman, Bakram-djord [= Bahramjerd], Iran). 22 – 23: aedeagus in dorsal, ventral, and lateral view of Pharaonus (Tamerlanius): 22: P. lederi (Koron 15 km Ekoc. Dzejran, Buckara, Uzbekistan); 23: P. semenowi (Kul'dzhuktau, Kizilkum desert, Uzbekistan)." httpUri="https://zenodo.org/record/244638/files/figure.png" pageId="12" pageNumber="179">16</figureCitation>
|
||
)....................................................
|
||
<taxonomicName id="4C734D1D4C33FFECFB1AB2B9FA1468B4" authority="Reitter, 1888" authorityName="Reitter" authorityYear="1888" box="[1154,1430,210,231]" class="Insecta" family="Rutelidae" genus="Pharaonus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="179" phylum="Arthropoda" rank="species" species="caucasicus">
|
||
<emphasis id="B907EA8C4C33FFECFB1AB2B9FA8068B4" box="[1154,1282,210,231]" italics="true" pageId="12" pageNumber="179">P. caucasicus</emphasis>
|
||
(
|
||
<bibRefCitation id="EFE24B6F4C33FFECFA88B2B9FA0D68B4" author="Reitter" box="[1296,1423,210,231]" pageId="12" pageNumber="179" refString="Reitter, E. (1888) Ubersicht der Arten der Coleopteren-Gattung Phyllopertha Kirby aus Europa, den angrenzenden Landern, Sibirien mit Central-Asien. Entomologische Nachrichten, 14, 289 - 294." type="journal article" year="1888">Reitter, 1888</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30872E3B4C33FFECFF0FB284FA19690F" pageId="12" pageNumber="179">
|
||
<paragraph id="8BCC369E4C33FFECFF0FB284FA196971" blockId="12.[151,1437,151,994]" pageId="12" pageNumber="179">
|
||
<keyLead id="308295AB4C33FFECFF0FB284FA196971" pageId="12" pageNumber="179">
|
||
4. (3) Pygidium entirely punctate and striolate. Clypeus 1.8–2.0 times as wide as long. Pronotum without impressed median line. North Africa and Levant:
|
||
<collectingCountry id="F364760E4C33FFECFE42B366FDA66971" box="[474,548,269,290]" name="Algeria" pageId="12" pageNumber="179">Algeria</collectingCountry>
|
||
,
|
||
<collectingCountry id="F364760E4C33FFECFDB5B366FDEB6971" box="[557,617,269,290]" name="Egypt" pageId="12" pageNumber="179">Egypt</collectingCountry>
|
||
,
|
||
<collectingCountry id="F364760E4C33FFECFDEBB366FD296971" box="[627,683,269,290]" name="Israel" pageId="12" pageNumber="179">Israel</collectingCountry>
|
||
,
|
||
<collectingCountry id="F364760E4C33FFECFD2CB366FD6C6971" box="[692,750,269,290]" name="Libya" pageId="12" pageNumber="179">Libya</collectingCountry>
|
||
,
|
||
<collectingCountry id="F364760E4C33FFECFD60B366FCB46971" box="[760,822,269,290]" name="Sudan" pageId="12" pageNumber="179">Sudan</collectingCountry>
|
||
,
|
||
<collectingCountry id="F364760E4C33FFECFCA7B366FC046971" box="[831,902,269,290]" name="Tunisia" pageId="12" pageNumber="179">Tunisia</collectingCountry>
|
||
........................................... 5
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BCC369E4C33FFECFF0FB341FA19690F" blockId="12.[151,1437,151,994]" pageId="12" pageNumber="179">
|
||
<keyLead id="308295AB4C33FFECFF0FB341FA19690F" pageId="12" pageNumber="179">
|
||
- Pygidium with a smooth area between the tufts of setae. Clypeus 2.1–2.5 times as wide as long. Pronotum with a more-or-less distinct longitudinal impression along the midline.
|
||
<collectingCountry id="F364760E4C33FFECFD52B32CFD73690F" box="[714,753,327,348]" name="Iran" pageId="12" pageNumber="179">Iran</collectingCountry>
|
||
........................................................ 6
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30872E3B4C33FFECFF0FB30EFA1E6A14" pageId="12" pageNumber="179">
|
||
<paragraph id="8BCC369E4C33FFECFF0FB30EFA1E6981" blockId="12.[151,1437,151,994]" pageId="12" pageNumber="179">
|
||
<keyLead id="308295AB4C33FFECFF0FB30EFA1E6981" pageId="12" pageNumber="179">
|
||
5. (4) Dorsal surface of head, pronotum, and lateral parts of abdominal ventrites with denser, markedly long, erect or semierect setae. Pygidium covered with long and dense, almost recumbent setae forming two tufts near the base. Aedeagus stout, broader, especially in lateral view; lateral outline of parameres shallowly sinuate (
|
||
<figureCitation id="13482A1B4C33FFECFCE8B3F4FC3569E7" box="[880,951,415,436]" captionStart="FIGURES 15 – 23" captionStartId="8.[151,264,1847,1869]" captionTargetBox="[174,1402,241,1792]" captionTargetId="figure@8.[174,1405,193,1813]" captionTargetPageId="8" captionText="FIGURES 15 – 23. Genitalia of Pharaonus. 15 – 19: aedeagus in dorsal, ventral, and lateral view of Pharaonus (Pharaonus): 15: P. adelphus, paratype (a-c) and holotype (d) (Kerman, Iran); 16: P. caucasicus (Gorovan, Armenia); 17: P. fasciculatus (Egypt); 18: P. farsensis, holotype (Darab, Iran); 19: P. saharicus, paratype (Tamrit, Tassili n’Ajjer, Algeria); 20: P. varicoloreus (Halfeti, Birecik, Adiyaman, Turkey). 21: female genital sclerites of P. adelphus (Kerman, Bakram-djord [= Bahramjerd], Iran). 22 – 23: aedeagus in dorsal, ventral, and lateral view of Pharaonus (Tamerlanius): 22: P. lederi (Koron 15 km Ekoc. Dzejran, Buckara, Uzbekistan); 23: P. semenowi (Kul'dzhuktau, Kizilkum desert, Uzbekistan)." httpUri="https://zenodo.org/record/244638/files/figure.png" pageId="12" pageNumber="179">Fig. 17</figureCitation>
|
||
):
|
||
<collectingCountry id="F364760E4C33FFECFC51B3F4FB8669E7" box="[969,1028,415,436]" name="Egypt" pageId="12" pageNumber="179">Egypt</collectingCountry>
|
||
,
|
||
<collectingCountry id="F364760E4C33FFECFB96B3F4FBC469E7" box="[1038,1094,415,436]" name="Israel" pageId="12" pageNumber="179">Israel</collectingCountry>
|
||
,
|
||
<collectingCountry id="F364760E4C33FFECFBD7B3F4FB0B69E7" box="[1103,1161,415,436]" name="Libya" pageId="12" pageNumber="179">Libya</collectingCountry>
|
||
,
|
||
<collectingCountry id="F364760E4C33FFECFB0AB3F4FB5269E7" box="[1170,1232,415,436]" name="Sudan" pageId="12" pageNumber="179">Sudan</collectingCountry>
|
||
,
|
||
<collectingCountry id="F364760E4C33FFECFB42B3F4FAA369E7" box="[1242,1313,415,436]" name="Tunisia" pageId="12" pageNumber="179">Tunisia</collectingCountry>
|
||
(
|
||
<figureCitation id="13482A1B4C33FFECFAB6B3F4FAF269E7" box="[1326,1392,415,436]" captionStart="FIGURES 7 – 10" captionStartId="5.[151,264,1955,1977]" captionTargetBox="[211,1377,193,1914]" captionTargetId="figure@5.[196,1390,193,1933]" captionTargetPageId="5" captionText="FIGURES 7 – 10. Habitus of Pharaonus (Pharaonus) species. 7: P. fasciculatus, male (Egypt). 8: P. saharicus, male paratype (Tamrit, Tassili n’Ajjer, Algeria). 9: P. varicoloreus, male (environs of Palymra, Syria). 10: P. varicoloreus, female (Birecik, Turkey)." httpUri="https://zenodo.org/record/244635/files/figure.png" pageId="12" pageNumber="179">Figs. 7</figureCitation>
|
||
,
|
||
<figureCitation id="13482A1B4C33FFECFAE4B3F4FA1669E7" box="[1404,1428,415,436]" captionStart="FIGURES 15 – 23" captionStartId="8.[151,264,1847,1869]" captionTargetBox="[174,1402,241,1792]" captionTargetId="figure@8.[174,1405,193,1813]" captionTargetPageId="8" captionText="FIGURES 15 – 23. Genitalia of Pharaonus. 15 – 19: aedeagus in dorsal, ventral, and lateral view of Pharaonus (Pharaonus): 15: P. adelphus, paratype (a-c) and holotype (d) (Kerman, Iran); 16: P. caucasicus (Gorovan, Armenia); 17: P. fasciculatus (Egypt); 18: P. farsensis, holotype (Darab, Iran); 19: P. saharicus, paratype (Tamrit, Tassili n’Ajjer, Algeria); 20: P. varicoloreus (Halfeti, Birecik, Adiyaman, Turkey). 21: female genital sclerites of P. adelphus (Kerman, Bakram-djord [= Bahramjerd], Iran). 22 – 23: aedeagus in dorsal, ventral, and lateral view of Pharaonus (Tamerlanius): 22: P. lederi (Koron 15 km Ekoc. Dzejran, Buckara, Uzbekistan); 23: P. semenowi (Kul'dzhuktau, Kizilkum desert, Uzbekistan)." httpUri="https://zenodo.org/record/244638/files/figure.png" pageId="12" pageNumber="179">17</figureCitation>
|
||
)..........................................................................
|
||
<taxonomicName id="4C734D1D4C33FFECFBCEB3D6FA1E6981" authority="Burmeister, 1844" authorityName="Burmeister" authorityYear="1844" box="[1110,1436,445,466]" class="Insecta" family="Rutelidae" genus="Pharaonus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="179" phylum="Arthropoda" rank="species" species="fasciculatus">
|
||
<emphasis id="B907EA8C4C33FFECFBCEB3D6FB636981" box="[1110,1249,445,466]" italics="true" pageId="12" pageNumber="179">P. fasciculatus</emphasis>
|
||
(
|
||
<bibRefCitation id="EFE24B6F4C33FFECFB74B3D6FA176981" author="Burmeister" box="[1260,1429,445,466]" pageId="12" pageNumber="179" refString="Burmeister, H. C. C. (1844) Handbuch der Entomologie. Vierter Band, Erste Abtheilung. Coleoptera Lamellicornia Anthobia et Phyllophaga systellochela. T. C. F. Enslin, Berlin, 584 pp." type="book" year="1844">Burmeister, 1844</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BCC369E4C33FFECFF0FB3B1FA1E6A14" blockId="12.[151,1437,151,994]" pageId="12" pageNumber="179">
|
||
<keyLead id="308295AB4C33FFECFF0FB3B1FA1E6A14" pageId="12" pageNumber="179">
|
||
<emphasis id="B907EA8C4C33FFECFF0FB3B1FF1D69BC" box="[151,159,474,495]" italics="true" pageId="12" pageNumber="179">-</emphasis>
|
||
Dorsal surface of head, pronotum, and lateral parts of abdominal ventrites with less dense, not considerably long, erect, pale setae. Pygidium covered with considerably long, sparse, erect macrosetae concentrated laterobasally but not forming two tufts. Aedeagus slender, relatively narrow, especially in lateral view; lateral outline of parameres broadly rounded (
|
||
<figureCitation id="13482A1B4C33FFECFB62B07EFAC36A79" box="[1274,1345,533,554]" captionStart="FIGURES 15 – 23" captionStartId="8.[151,264,1847,1869]" captionTargetBox="[174,1402,241,1792]" captionTargetId="figure@8.[174,1405,193,1813]" captionTargetPageId="8" captionText="FIGURES 15 – 23. Genitalia of Pharaonus. 15 – 19: aedeagus in dorsal, ventral, and lateral view of Pharaonus (Pharaonus): 15: P. adelphus, paratype (a-c) and holotype (d) (Kerman, Iran); 16: P. caucasicus (Gorovan, Armenia); 17: P. fasciculatus (Egypt); 18: P. farsensis, holotype (Darab, Iran); 19: P. saharicus, paratype (Tamrit, Tassili n’Ajjer, Algeria); 20: P. varicoloreus (Halfeti, Birecik, Adiyaman, Turkey). 21: female genital sclerites of P. adelphus (Kerman, Bakram-djord [= Bahramjerd], Iran). 22 – 23: aedeagus in dorsal, ventral, and lateral view of Pharaonus (Tamerlanius): 22: P. lederi (Koron 15 km Ekoc. Dzejran, Buckara, Uzbekistan); 23: P. semenowi (Kul'dzhuktau, Kizilkum desert, Uzbekistan)." httpUri="https://zenodo.org/record/244638/files/figure.png" pageId="12" pageNumber="179">Fig. 19</figureCitation>
|
||
).
|
||
<collectingCountry id="F364760E4C33FFECFACBB07EFA1E6A79" box="[1363,1436,533,554]" name="Algeria" pageId="12" pageNumber="179">Algeria</collectingCountry>
|
||
(
|
||
<figureCitation id="13482A1B4C33FFECFF70B059FEA96A14" box="[232,299,562,583]" captionStart="FIGURES 7 – 10" captionStartId="5.[151,264,1955,1977]" captionTargetBox="[211,1377,193,1914]" captionTargetId="figure@5.[196,1390,193,1933]" captionTargetPageId="5" captionText="FIGURES 7 – 10. Habitus of Pharaonus (Pharaonus) species. 7: P. fasciculatus, male (Egypt). 8: P. saharicus, male paratype (Tamrit, Tassili n’Ajjer, Algeria). 9: P. varicoloreus, male (environs of Palymra, Syria). 10: P. varicoloreus, female (Birecik, Turkey)." httpUri="https://zenodo.org/record/244635/files/figure.png" pageId="12" pageNumber="179">Figs. 8</figureCitation>
|
||
,
|
||
<figureCitation id="13482A1B4C33FFECFEAEB059FECD6A14" box="[310,335,562,583]" captionStart="FIGURES 15 – 23" captionStartId="8.[151,264,1847,1869]" captionTargetBox="[174,1402,241,1792]" captionTargetId="figure@8.[174,1405,193,1813]" captionTargetPageId="8" captionText="FIGURES 15 – 23. Genitalia of Pharaonus. 15 – 19: aedeagus in dorsal, ventral, and lateral view of Pharaonus (Pharaonus): 15: P. adelphus, paratype (a-c) and holotype (d) (Kerman, Iran); 16: P. caucasicus (Gorovan, Armenia); 17: P. fasciculatus (Egypt); 18: P. farsensis, holotype (Darab, Iran); 19: P. saharicus, paratype (Tamrit, Tassili n’Ajjer, Algeria); 20: P. varicoloreus (Halfeti, Birecik, Adiyaman, Turkey). 21: female genital sclerites of P. adelphus (Kerman, Bakram-djord [= Bahramjerd], Iran). 22 – 23: aedeagus in dorsal, ventral, and lateral view of Pharaonus (Tamerlanius): 22: P. lederi (Koron 15 km Ekoc. Dzejran, Buckara, Uzbekistan); 23: P. semenowi (Kul'dzhuktau, Kizilkum desert, Uzbekistan)." httpUri="https://zenodo.org/record/244638/files/figure.png" pageId="12" pageNumber="179">19</figureCitation>
|
||
)........................................................................
|
||
<taxonomicName id="4C734D1D4C33FFECFB58B059FA1E6A14" authority="Kral 2012" authorityName="Kral" authorityYear="2012" box="[1216,1436,562,583]" class="Insecta" family="Rutelidae" genus="Pharaonus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="179" phylum="Arthropoda" rank="species" species="saharicus">
|
||
<emphasis id="B907EA8C4C33FFECFB58B059FAB76A14" box="[1216,1333,562,583]" italics="true" pageId="12" pageNumber="179">P. saharicus</emphasis>
|
||
<bibRefCitation id="EFE24B6F4C33FFECFAA3B059FA1E6A14" author="Kral" box="[1339,1436,562,583]" pageId="12" pageNumber="179" refString="Kral, D. (2012) Pharaonus saharicus sp. nov. (Coleoptera: Scarabaeidae: Rutelinae) from the Central Sahara (Algeria). Klapalekiana, 48, 127 - 136." type="journal article" year="2012">Král 2012</bibRefCitation>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30872E3B4C33FFECFF0FB024FA1E6AA4" pageId="12" pageNumber="179">
|
||
<paragraph id="8BCC369E4C33FFECFF0FB024FA1E6ACC" blockId="12.[151,1437,151,994]" pageId="12" pageNumber="179">
|
||
<keyLead id="308295AB4C33FFECFF0FB024FA1E6ACC" pageId="12" pageNumber="179">
|
||
6. (4) Elytra with sparse pubescence arranged in loose stripes; denser on intervals 3, 5, and 7; setae reaching at least the middle of the elytra (
|
||
<figureCitation id="13482A1B4C33FFECFEBDB006FEEE6AD1" box="[293,364,621,642]" captionStart="FIGURES 13 – 14" captionStartId="7.[151,264,1550,1572]" captionTargetBox="[169,1432,195,1512]" captionTargetId="figure@7.[155,1432,195,1528]" captionTargetPageId="7" captionText="FIGURES 13 – 14. Different elytral setation in evidence. 13: P. adelphus. 14: P. farsensis." httpUri="https://zenodo.org/record/244637/files/figure.png" pageId="12" pageNumber="179">Fig. 14</figureCitation>
|
||
), shorter, but clearly visible (
|
||
<date id="FFCD105E4C33FFECFD10B006FD316AD1" box="[648,691,621,642]" pageId="12" pageNumber="179" value="1920-10">20X</date>
|
||
) setae reaching the apical region. In males: antennal club 1.13–1.16 times as long as the clypeal width.
|
||
<collectingCountry id="F364760E4C33FFECFE44B0E1FD806ACC" box="[476,514,650,671]" name="Iran" pageId="12" pageNumber="179">Iran</collectingCountry>
|
||
(
|
||
<figureCitation id="13482A1B4C33FFECFD88B0E1FDD66ACC" box="[528,596,650,671]" captionStart="FIGURES 3 – 6" captionStartId="3.[151,264,1953,1975]" captionTargetBox="[212,1376,196,1915]" captionTargetId="figure@3.[201,1386,193,1932]" captionTargetPageId="3" captionText="FIGURES 3 – 6. Habitus of Pharaonus (Pharaonus) species. 3: P. adelphus, male paratype (Kerman, Iran). 4: P. adelphus, female paratype (Kerman, Bakram-djord [= Bahramjerd], Iran). 5: P. caucasicus, male (Gorovan sands, Armenia). 6: P. farsensis, male paratype (Darab, Iran)." httpUri="https://zenodo.org/record/244634/files/figure.png" pageId="12" pageNumber="179">Figs. 6</figureCitation>
|
||
,
|
||
<figureCitation id="13482A1B4C33FFECFDC6B0E1FDFA6ACC" box="[606,632,650,671]" captionStart="FIGURES 13 – 14" captionStartId="7.[151,264,1550,1572]" captionTargetBox="[169,1432,195,1512]" captionTargetId="figure@7.[155,1432,195,1528]" captionTargetPageId="7" captionText="FIGURES 13 – 14. Different elytral setation in evidence. 13: P. adelphus. 14: P. farsensis." httpUri="https://zenodo.org/record/244637/files/figure.png" pageId="12" pageNumber="179">14</figureCitation>
|
||
,
|
||
<figureCitation id="13482A1B4C33FFECFD1AB0E1FD186ACC" box="[642,666,650,671]" captionStart="FIGURES 15 – 23" captionStartId="8.[151,264,1847,1869]" captionTargetBox="[174,1402,241,1792]" captionTargetId="figure@8.[174,1405,193,1813]" captionTargetPageId="8" captionText="FIGURES 15 – 23. Genitalia of Pharaonus. 15 – 19: aedeagus in dorsal, ventral, and lateral view of Pharaonus (Pharaonus): 15: P. adelphus, paratype (a-c) and holotype (d) (Kerman, Iran); 16: P. caucasicus (Gorovan, Armenia); 17: P. fasciculatus (Egypt); 18: P. farsensis, holotype (Darab, Iran); 19: P. saharicus, paratype (Tamrit, Tassili n’Ajjer, Algeria); 20: P. varicoloreus (Halfeti, Birecik, Adiyaman, Turkey). 21: female genital sclerites of P. adelphus (Kerman, Bakram-djord [= Bahramjerd], Iran). 22 – 23: aedeagus in dorsal, ventral, and lateral view of Pharaonus (Tamerlanius): 22: P. lederi (Koron 15 km Ekoc. Dzejran, Buckara, Uzbekistan); 23: P. semenowi (Kul'dzhuktau, Kizilkum desert, Uzbekistan)." httpUri="https://zenodo.org/record/244638/files/figure.png" pageId="12" pageNumber="179">18</figureCitation>
|
||
)..............................
|
||
<taxonomicName id="4C734D1D4C33FFECFB90B0E1FAA86ACC" authority="Keith, Sabatinelli" authorityName="Keith, Sabatinelli" box="[1032,1322,650,671]" class="Insecta" family="Rutelidae" genus="Pharaonus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="179" phylum="Arthropoda" rank="species" species="farsensis">
|
||
<emphasis id="B907EA8C4C33FFECFB90B0E1FBFA6ACC" box="[1032,1144,650,671]" italics="true" pageId="12" pageNumber="179">P. farsensis</emphasis>
|
||
Keith, Sabatinelli
|
||
</taxonomicName>
|
||
, and Uliana
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BCC369E4C33FFECFF0FB0CCFA1E6AA4" blockId="12.[151,1437,151,994]" pageId="12" pageNumber="179">
|
||
<keyLead id="308295AB4C33FFECFF0FB0CCFA1E6AA4" pageId="12" pageNumber="179">
|
||
- Elytra with long pubescence limited to the humeral region and to a very narrow basal area, not going beyond the tip of the scutellum (
|
||
<figureCitation id="13482A1B4C33FFECFED3B0AEFE136A89" box="[331,401,709,730]" captionStart="FIGURES 13 – 14" captionStartId="7.[151,264,1550,1572]" captionTargetBox="[169,1432,195,1512]" captionTargetId="figure@7.[155,1432,195,1528]" captionTargetPageId="7" captionText="FIGURES 13 – 14. Different elytral setation in evidence. 13: P. adelphus. 14: P. farsensis." httpUri="https://zenodo.org/record/244637/files/figure.png" pageId="12" pageNumber="179">Fig. 13</figureCitation>
|
||
); the rest of the elytra glabrous or with some microsetae barely visible at
|
||
<date id="FFCD105E4C33FFECFBD5B0AEFBFF6A89" box="[1101,1149,709,730]" pageId="12" pageNumber="179" value="1940-10">40X.</date>
|
||
In males: antennal club 0.91– 0.94 times as long as the clypeal width.
|
||
<collectingCountry id="F364760E4C33FFECFDF8B089FD046AA4" box="[608,646,738,759]" name="Iran" pageId="12" pageNumber="179">Iran</collectingCountry>
|
||
(
|
||
<figureCitation id="13482A1B4C33FFECFD0CB089FD726AA4" box="[660,752,738,759]" captionStart="FIGURES 3 – 6" captionStartId="3.[151,264,1953,1975]" captionTargetBox="[212,1376,196,1915]" captionTargetId="figure@3.[201,1386,193,1932]" captionTargetPageId="3" captionText="FIGURES 3 – 6. Habitus of Pharaonus (Pharaonus) species. 3: P. adelphus, male paratype (Kerman, Iran). 4: P. adelphus, female paratype (Kerman, Bakram-djord [= Bahramjerd], Iran). 5: P. caucasicus, male (Gorovan sands, Armenia). 6: P. farsensis, male paratype (Darab, Iran)." httpUri="https://zenodo.org/record/244634/files/figure.png" pageId="12" pageNumber="179">Figs. 3, 4</figureCitation>
|
||
,
|
||
<figureCitation id="13482A1B4C33FFECFD62B089FC966AA4" box="[762,788,738,759]" captionStart="FIGURES 13 – 14" captionStartId="7.[151,264,1550,1572]" captionTargetBox="[169,1432,195,1512]" captionTargetId="figure@7.[155,1432,195,1528]" captionTargetPageId="7" captionText="FIGURES 13 – 14. Different elytral setation in evidence. 13: P. adelphus. 14: P. farsensis." httpUri="https://zenodo.org/record/244637/files/figure.png" pageId="12" pageNumber="179">13</figureCitation>
|
||
,
|
||
<figureCitation id="13482A1B4C33FFECFC86B089FCB46AA4" box="[798,822,738,759]" captionStart="FIGURES 15 – 23" captionStartId="8.[151,264,1847,1869]" captionTargetBox="[174,1402,241,1792]" captionTargetId="figure@8.[174,1405,193,1813]" captionTargetPageId="8" captionText="FIGURES 15 – 23. Genitalia of Pharaonus. 15 – 19: aedeagus in dorsal, ventral, and lateral view of Pharaonus (Pharaonus): 15: P. adelphus, paratype (a-c) and holotype (d) (Kerman, Iran); 16: P. caucasicus (Gorovan, Armenia); 17: P. fasciculatus (Egypt); 18: P. farsensis, holotype (Darab, Iran); 19: P. saharicus, paratype (Tamrit, Tassili n’Ajjer, Algeria); 20: P. varicoloreus (Halfeti, Birecik, Adiyaman, Turkey). 21: female genital sclerites of P. adelphus (Kerman, Bakram-djord [= Bahramjerd], Iran). 22 – 23: aedeagus in dorsal, ventral, and lateral view of Pharaonus (Tamerlanius): 22: P. lederi (Koron 15 km Ekoc. Dzejran, Buckara, Uzbekistan); 23: P. semenowi (Kul'dzhuktau, Kizilkum desert, Uzbekistan)." httpUri="https://zenodo.org/record/244638/files/figure.png" pageId="12" pageNumber="179">15</figureCitation>
|
||
)................
|
||
<taxonomicName id="4C734D1D4C33FFECFB93B089FAA86AA4" authority="Keith, Sabatinelli" authorityName="Keith, Sabatinelli" box="[1035,1322,738,759]" class="Insecta" family="Rutelidae" genus="Pharaonus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="179" phylum="Arthropoda" rank="species" species="adelphus">
|
||
<emphasis id="B907EA8C4C33FFECFB93B089FBFB6AA4" box="[1035,1145,738,759]" italics="true" pageId="12" pageNumber="179">P. adelphus</emphasis>
|
||
Keith, Sabatinelli
|
||
</taxonomicName>
|
||
, and Uliana
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30872E3B4C33FFECFF0FB094FA1E6BB1" pageId="12" pageNumber="179">
|
||
<paragraph id="8BCC369E4C33FFECFF0FB094FA1E6B3F" blockId="12.[151,1437,151,994]" pageId="12" pageNumber="179">
|
||
<keyLead id="308295AB4C33FFECFF0FB094FA1E6B3F" pageId="12" pageNumber="179">
|
||
7. (1) Body, except elytra and appendages (but including femora), covered with very dense, strongly adpressed, short, thick, white setae; setae almost entirely obscuring the integument. Midline of the pronotum impunctate, glabrous. Pygidium completely and densely covered by uniformly thick setae, except for the narrow midline.
|
||
<collectingCountry id="F364760E4C33FFECFC4DB151FBC56B1C" box="[981,1095,826,847]" name="Kazakhstan" pageId="12" pageNumber="179">Kazakhstan</collectingCountry>
|
||
,
|
||
<collectingCountry id="F364760E4C33FFECFBCAB151FB316B1C" box="[1106,1203,826,847]" name="Tajikistan" pageId="12" pageNumber="179">Tajikistan</collectingCountry>
|
||
,
|
||
<collectingCountry id="F364760E4C33FFECFB25B151FAC66B1C" box="[1213,1348,826,847]" name="Turkmenistan" pageId="12" pageNumber="179">Turkmenistan</collectingCountry>
|
||
,
|
||
<collectingCountry id="F364760E4C33FFECFAD6B151FE8A6B3F" name="Uzbekistan" pageId="12" pageNumber="179">Uzbekistan</collectingCountry>
|
||
(
|
||
<figureCitation id="13482A1B4C33FFECFE8DB13CFEE76B3F" box="[277,357,855,876]" captionStart="FIGURES 11 – 12" captionStartId="6.[151,264,1129,1151]" captionTargetBox="[159,1427,207,1105]" captionTargetId="figure@6.[150,1437,195,1118]" captionTargetPageId="6" captionText="FIGURES 11 – 12. Habitus of Pharaonus (Tamerlanius) species. 11: P. l e de r i, male (Koron 15 km Ekoc. Dzejran, Bukhara, Uzbekistan). 12: P. semenowi, male (Katta-kum, Turkmenkuduk, Uzbekistan)." httpUri="https://zenodo.org/record/244636/files/figure.png" pageId="12" pageNumber="179">Figs. 12</figureCitation>
|
||
,
|
||
<figureCitation id="13482A1B4C33FFECFEF7B13CFE0A6B3F" box="[367,392,855,876]" captionStart="FIGURES 15 – 23" captionStartId="8.[151,264,1847,1869]" captionTargetBox="[174,1402,241,1792]" captionTargetId="figure@8.[174,1405,193,1813]" captionTargetPageId="8" captionText="FIGURES 15 – 23. Genitalia of Pharaonus. 15 – 19: aedeagus in dorsal, ventral, and lateral view of Pharaonus (Pharaonus): 15: P. adelphus, paratype (a-c) and holotype (d) (Kerman, Iran); 16: P. caucasicus (Gorovan, Armenia); 17: P. fasciculatus (Egypt); 18: P. farsensis, holotype (Darab, Iran); 19: P. saharicus, paratype (Tamrit, Tassili n’Ajjer, Algeria); 20: P. varicoloreus (Halfeti, Birecik, Adiyaman, Turkey). 21: female genital sclerites of P. adelphus (Kerman, Bakram-djord [= Bahramjerd], Iran). 22 – 23: aedeagus in dorsal, ventral, and lateral view of Pharaonus (Tamerlanius): 22: P. lederi (Koron 15 km Ekoc. Dzejran, Buckara, Uzbekistan); 23: P. semenowi (Kul'dzhuktau, Kizilkum desert, Uzbekistan)." httpUri="https://zenodo.org/record/244638/files/figure.png" pageId="12" pageNumber="179">23</figureCitation>
|
||
)................................................................
|
||
<taxonomicName id="4C734D1D4C33FFECFB0CB13CFA1E6B3F" authority="Reitter, 1887" authorityName="Reitter" authorityYear="1887" box="[1172,1436,855,876]" class="Insecta" family="Rutelidae" genus="Pharaonus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="179" phylum="Arthropoda" rank="species" species="semenowi">
|
||
<emphasis id="B907EA8C4C33FFECFB0CB13CFA886B3F" box="[1172,1290,855,876]" italics="true" pageId="12" pageNumber="179">P. semenowi</emphasis>
|
||
(
|
||
<bibRefCitation id="EFE24B6F4C33FFECFA8EB13CFA176B3F" author="Reitter" box="[1302,1429,855,876]" pageId="12" pageNumber="179" refString="Reitter, E. (1887) Neue Coleopteren aus Europa, den angrenzenden Landern und Sibirien, mit Bemerkungen uber bekannte Arten. Vierter Theil. Deutsche Entomologische Zeitschrift, 31, 497 - 528." type="journal article" year="1887">Reitter, 1887</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BCC369E4C33FFECFF0FB11EFA1E6BB1" blockId="12.[151,1437,151,994]" pageId="12" pageNumber="179">
|
||
<keyLead id="308295AB4C33FFECFF0FB11EFA1E6BB1" pageId="12" pageNumber="179">
|
||
- Body, except elytra and appendages (but including femora) bearing recumbent or semierect long, thin setae; setae not obscuring the integument. Midline of the pronotum punctate, setose. Pygidium bearing two basal tufts of dense setae, the rest of the surface is clearly visible through a loose pilosity.
|
||
<collectingCountry id="F364760E4C33FFECFD2EB1C4FCAC6B97" box="[694,814,943,964]" name="Afghanistan" pageId="12" pageNumber="179">Afghanistan</collectingCountry>
|
||
,
|
||
<collectingCountry id="F364760E4C33FFECFCAFB1C4FC2B6B97" box="[823,937,943,964]" name="Kazakhstan" pageId="12" pageNumber="179">Kazakhstan</collectingCountry>
|
||
,
|
||
<collectingCountry id="F364760E4C33FFECFC2AB1C4FB916B97" box="[946,1043,943,964]" name="Tajikistan" pageId="12" pageNumber="179">Tajikistan</collectingCountry>
|
||
,
|
||
<collectingCountry id="F364760E4C33FFECFB84B1C4FB216B97" box="[1052,1187,943,964]" name="Turkmenistan" pageId="12" pageNumber="179">Turkmenistan</collectingCountry>
|
||
,
|
||
<collectingCountry id="F364760E4C33FFECFB33B1C4FA956B97" box="[1195,1303,943,964]" name="Uzbekistan" pageId="12" pageNumber="179">Uzbekistan</collectingCountry>
|
||
(
|
||
<figureCitation id="13482A1B4C33FFECFABCB1C4FAF36B97" box="[1316,1393,943,964]" captionStart="FIGURES 11 – 12" captionStartId="6.[151,264,1129,1151]" captionTargetBox="[159,1427,207,1105]" captionTargetId="figure@6.[150,1437,195,1118]" captionTargetPageId="6" captionText="FIGURES 11 – 12. Habitus of Pharaonus (Tamerlanius) species. 11: P. l e de r i, male (Koron 15 km Ekoc. Dzejran, Bukhara, Uzbekistan). 12: P. semenowi, male (Katta-kum, Turkmenkuduk, Uzbekistan)." httpUri="https://zenodo.org/record/244636/files/figure.png" pageId="12" pageNumber="179">Figs. 11</figureCitation>
|
||
,
|
||
<figureCitation id="13482A1B4C33FFECFAE3B1C4FA166B97" box="[1403,1428,943,964]" captionStart="FIGURES 15 – 23" captionStartId="8.[151,264,1847,1869]" captionTargetBox="[174,1402,241,1792]" captionTargetId="figure@8.[174,1405,193,1813]" captionTargetPageId="8" captionText="FIGURES 15 – 23. Genitalia of Pharaonus. 15 – 19: aedeagus in dorsal, ventral, and lateral view of Pharaonus (Pharaonus): 15: P. adelphus, paratype (a-c) and holotype (d) (Kerman, Iran); 16: P. caucasicus (Gorovan, Armenia); 17: P. fasciculatus (Egypt); 18: P. farsensis, holotype (Darab, Iran); 19: P. saharicus, paratype (Tamrit, Tassili n’Ajjer, Algeria); 20: P. varicoloreus (Halfeti, Birecik, Adiyaman, Turkey). 21: female genital sclerites of P. adelphus (Kerman, Bakram-djord [= Bahramjerd], Iran). 22 – 23: aedeagus in dorsal, ventral, and lateral view of Pharaonus (Tamerlanius): 22: P. lederi (Koron 15 km Ekoc. Dzejran, Buckara, Uzbekistan); 23: P. semenowi (Kul'dzhuktau, Kizilkum desert, Uzbekistan)." httpUri="https://zenodo.org/record/244638/files/figure.png" pageId="12" pageNumber="179">22</figureCitation>
|
||
)..................................................................................
|
||
<taxonomicName id="4C734D1D4C33FFECFB22B1A6FA1E6BB1" authority="Reitter, 1888" authorityName="Reitter" authorityYear="1888" box="[1210,1436,973,994]" class="Insecta" family="Rutelidae" genus="Pharaonus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="179" phylum="Arthropoda" rank="species" species="lederi">
|
||
<emphasis id="B907EA8C4C33FFECFB22B1A6FA8B6BB1" box="[1210,1289,973,994]" italics="true" pageId="12" pageNumber="179">P. lederi</emphasis>
|
||
(
|
||
<bibRefCitation id="EFE24B6F4C33FFECFA8EB1A6FA176BB1" author="Reitter" box="[1302,1429,973,994]" pageId="12" pageNumber="179" refString="Reitter, E. (1888) Ubersicht der Arten der Coleopteren-Gattung Phyllopertha Kirby aus Europa, den angrenzenden Landern, Sibirien mit Central-Asien. Entomologische Nachrichten, 14, 289 - 294." type="journal article" year="1888">Reitter, 1888</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |