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<mods:title id="9C3EFE7095A8506252967C351B3001CE">Integrating museum collections and molecules reveals genus-level synonymy and new species in red devil spiders (Araneae, Dysderidae) from the Middle East and Central Asia</mods:title>
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<mods:namePart id="82C1471332A9E174B8598B412D3FD432">Bellvert, Adrià</mods:namePart>
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<mods:affiliation id="C1C08DAC79ACEAA512E34B788AE146A9">Departament de Biologia Evolutiva, Ecologia i Ciències Ambientals, Universitat de Barcelona (UB), Av. Diagonal, 643, 08028 Barcelona, Spain. &amp; Institut de Recerca de la Biodiversitat (IRBio), Universitat de Barcelona (UB) Barcelona, Spain.</mods:affiliation>
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<mods:namePart id="0CFB44E6851FC09D65CA6E40D62CE6F5">Dimitrov, Dragomir</mods:namePart>
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<mods:affiliation id="CA711FF9E515C017E6D4D6B56F7ECB0F">Departament de Biologia Evolutiva, Ecologia i Ciències Ambientals, Universitat de Barcelona (UB), Av. Diagonal, 643, 08028 Barcelona, Spain. &amp; Institut de Recerca de la Biodiversitat (IRBio), Universitat de Barcelona (UB) Barcelona, Spain. &amp; National Museum of Natural Sciences, Bulgarian Academy of Sciences, 1 Tsar Osvoboditel Blvd, 1000 Sofia, Bulgaria.</mods:affiliation>
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<mods:namePart id="9D8455ECD16B157D328D4A14B0394588">Zamani, Alireza</mods:namePart>
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<mods:affiliation id="FF862975595C422E5EBAA3BEBC78B0F8">Zoological Museum, Biodiversity Unit, FI- 20014 University of Turku, Finland.</mods:affiliation>
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<mods:namePart id="209980C14B98AD9730B3E03DB6AB9708">Arnedo, Miquel A.</mods:namePart>
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<mods:date id="76DF6C2546F599B0EB778C6C2E1D644D">2024</mods:date>
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<taxonomicName id="4C73B3D6B732D85B72588E924788FC21" authority="Bellvert &amp; Dimitrov" authorityName="Bellvert &amp; Dimitrov &amp; Zamani &amp; Arnedo" authorityYear="2024" box="[513,978,969,996]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="220" phylum="Arthropoda" rank="species" species="naouelae" status="sp. nov.">
<emphasis id="B9071447B732D85B72588E924680FC21" bold="true" box="[513,730,969,996]" italics="true" pageId="10" pageNumber="220">Dysdera naouelae</emphasis>
Bellvert &amp; Dimitrov
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<figureCitation id="1348D4D0B732D85B72868956470EFBE2" box="[735,852,1037,1063]" captionStart="Figs 914. 911" captionStartId="11.[189,241,1848,1874]" captionTargetBox="[189,1398,265,1810]" captionTargetId="figure-16@11.[189,1398,265,1810]" captionTargetPageId="11" captionText="Figs 914. 911. Male Dysdera naouelae Bellvert &amp; Dimitrov sp. nov. (SMF). 9. Prolateral view. 10. Anterior view. 11. Retrolateral view. 1214. Female Dysdera naouelae Bellvert &amp; Dimitrov sp. nov. (SMF), vulva. 12. Ventral view. 13. Lateral view. 14. Dorsal view. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10663888" httpUri="https://zenodo.org/record/10663888/files/figure.png" pageId="10" pageNumber="220">Figs 914</figureCitation>
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<taxonomicName id="4C73B3D6B732D85B70E4891645CAFBA2" ID-CoL="38989" authorityName="Kroneberg" authorityYear="1875" box="[189,400,1101,1127]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="220" phylum="Arthropoda" rank="species" species="tartarica">
<emphasis id="B9071447B732D85B70E4891645CAFBA2" box="[189,400,1101,1127]" italics="true" pageId="10" pageNumber="220">Dysdera tartarica</emphasis>
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<treatmentCitation id="0AD2EE44B732D85B71F4891546D4FBA2" author="Lazarov S." box="[429,654,1101,1128]" page="104" pageId="10" pageNumber="220" year="2009">
<bibRefCitation id="EFE2B5A4B732D85B71F4891546D4FBA2" author="Lazarov" box="[429,654,1101,1128]" firstAuthor="Lazarov" page="104" pageId="10" pageNumber="220" pagination="104 - 106" refId="ref12449" refString="Lazarov S. 2009. A new record of Dysdera tartarica Kroneberg, 1875 with a description of the previously unknown female (Araneae, Dysderidae). Turkish Journal of Arachnology 1 (2, for 2008): 104 - 106." type="journal article" year="2009">Lazarov 2009: 104</bibRefCitation>
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, figs 15 [misidentification].
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<emphasis id="B9071447B732D85B70E489D6456CFB62" bold="true" box="[189,310,1165,1191]" pageId="10" pageNumber="220">Diagnosis</emphasis>
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The new species is morphologically most similar to
<taxonomicName id="4C73B3D6B732D85B731A89E1404BFB11" authority="Bellvert &amp; Dimitrov &amp; Zamani &amp; Arnedo, 2024" authorityName="Bellvert &amp; Dimitrov &amp; Zamani &amp; Arnedo" authorityYear="2024" box="[835,1041,1210,1236]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="220" phylum="Arthropoda" rank="species" species="kourosh" status="sp. nov.">
<emphasis id="B9071447B732D85B731A89E1404BFB11" box="[835,1041,1210,1236]" italics="true" pageId="10" pageNumber="220">Dysdera kourosh</emphasis>
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<taxonomicNameLabel id="A234A93CB732D85B744089E14020FB11" box="[1049,1146,1210,1236]" pageId="10" pageNumber="220" rank="species">sp. nov.</taxonomicNameLabel>
,
<taxonomicName id="4C73B3D6B732D85B74D289E14640FB33" authority="Fomichev &amp; Marusik, 2021" authorityName="Fomichev &amp; Marusik" authorityYear="2021" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="220" phylum="Arthropoda" rank="species" species="mikhailovi">
<emphasis id="B9071447B732D85B74D289E14123FB11" box="[1163,1401,1210,1236]" italics="true" pageId="10" pageNumber="220">Dysdera mikhailovi</emphasis>
<bibRefCitation id="EFE2B5A4B732D85B70E489804640FB33" author="Fomichev &amp; Marusik" box="[189,538,1243,1270]" firstAuthor="Fomichev" pageId="10" pageNumber="220" pagination="73 - 89" refId="ref11966" refString="Fomichev A. A. &amp; Marusik Y. M. 2021. Notes on the spider genus Dysdera Latreille, 1804 (Araneae: Dysderidae) in Central Asia. Zootaxa 5006 (1): 73 - 89. https: // doi. org / 10.11646 / zootaxa. 5006.1.10" type="journal article" year="2021">Fomichev &amp; Marusik, 2021</bibRefCitation>
</taxonomicName>
, and
<taxonomicName id="4C73B3D6B732D85B72398980408EFB33" authority="Zamani, Marusik &amp; Szuts, 2023" authorityName="Zamani, Marusik &amp; Szuts" authorityYear="2023" box="[608,1236,1243,1270]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="220" phylum="Arthropoda" rank="species" species="sagartia">
<emphasis id="B9071447B732D85B72398980476DFB30" box="[608,823,1243,1269]" italics="true" pageId="10" pageNumber="220">Dysdera sagartia</emphasis>
<bibRefCitation id="EFE2B5A4B732D85B731B8980408EFB33" author="Zamani A. &amp; Marusik Y. M. &amp; Szuts T." box="[834,1236,1243,1270]" journalOrPublisher="Zoosystematics and Evolution" pageId="10" pageNumber="220" pagination="337 - 344" part="99" refId="ref13524" refString="Zamani A., Marusik Y. M. &amp; Szuts T. 2023 a. A survey of Dysderella Dunin, 1992 (Araneae, Dysderidae), with a new species from Iran. Zoosystematics and Evolution 99 (2): 337 - 344. https: // doi. org / 10.3897 / zse. 99.104613" title="A survey of Dysderella Dunin, 1992 (Araneae, Dysderidae), with a new species from Iran" type="journal article" year="2023">Zamani, Marusik &amp; Szűts, 2023</bibRefCitation>
</taxonomicName>
by the well-developed, wide and rounded crest (
<figureCitation id="1348D4D0B732D85B722B89A646B7FAD2" box="[626,749,1277,1303]" captionStart="Figs 914. 911" captionStartId="11.[189,241,1848,1874]" captionTargetBox="[189,1398,265,1810]" captionTargetId="figure-16@11.[189,1398,265,1810]" captionTargetPageId="11" captionText="Figs 914. 911. Male Dysdera naouelae Bellvert &amp; Dimitrov sp. nov. (SMF). 9. Prolateral view. 10. Anterior view. 11. Retrolateral view. 1214. Female Dysdera naouelae Bellvert &amp; Dimitrov sp. nov. (SMF), vulva. 12. Ventral view. 13. Lateral view. 14. Dorsal view. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10663888" httpUri="https://zenodo.org/record/10663888/files/figure.png" pageId="10" pageNumber="220">Figs 911</figureCitation>
), internal sclerite significantly wider than the exterior one, and the characteristic lateral margin of the lateral sheet (
<figureCitation id="1348D4D0B732D85B73C9884447B2FAFC" box="[912,1000,1311,1337]" captionStart="Figs 914. 911" captionStartId="11.[189,241,1848,1874]" captionTargetBox="[189,1398,265,1810]" captionTargetId="figure-16@11.[189,1398,265,1810]" captionTargetPageId="11" captionText="Figs 914. 911. Male Dysdera naouelae Bellvert &amp; Dimitrov sp. nov. (SMF). 9. Prolateral view. 10. Anterior view. 11. Retrolateral view. 1214. Female Dysdera naouelae Bellvert &amp; Dimitrov sp. nov. (SMF), vulva. 12. Ventral view. 13. Lateral view. 14. Dorsal view. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10663888" httpUri="https://zenodo.org/record/10663888/files/figure.png" pageId="10" pageNumber="220">Fig. 10</figureCitation>
). The male differs by (1) the thin, well sclerotized processus-like lateral margin of the lateral sheet, almost perpendicular to the embolic division tip (
<figureCitation id="1348D4D0B732D85B710F883945F7FAB9" box="[342,429,1378,1404]" captionStart="Figs 914. 911" captionStartId="11.[189,241,1848,1874]" captionTargetBox="[189,1398,265,1810]" captionTargetId="figure-16@11.[189,1398,265,1810]" captionTargetPageId="11" captionText="Figs 914. 911. Male Dysdera naouelae Bellvert &amp; Dimitrov sp. nov. (SMF). 9. Prolateral view. 10. Anterior view. 11. Retrolateral view. 1214. Female Dysdera naouelae Bellvert &amp; Dimitrov sp. nov. (SMF), vulva. 12. Ventral view. 13. Lateral view. 14. Dorsal view. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10663888" httpUri="https://zenodo.org/record/10663888/files/figure.png" pageId="10" pageNumber="220">Fig. 11</figureCitation>
), vs pointed backward in
<taxonomicName id="4C73B3D6B732D85B72BA8838473FFAB9" authorityName="Bellvert &amp; Dimitrov &amp; Zamani &amp; Arnedo" authorityYear="2024" box="[739,869,1378,1404]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="220" phylum="Arthropoda" rank="species" species="kourosh">
<emphasis id="B9071447B732D85B72BA8838473FFAB9" box="[739,869,1378,1404]" italics="true" pageId="10" pageNumber="220">D. kourosh</emphasis>
</taxonomicName>
(arrowed in
<figureCitation id="1348D4D0B732D85B73A68839400CFAB9" box="[1023,1110,1378,1404]" captionStart="Figs 1820" captionStartId="15.[189,241,1963,1989]" captionTargetBox="[194,1398,1197,1925]" captionTargetId="figure-300@15.[189,1398,1197,1925]" captionTargetPageId="15" captionText="Figs 1820. Male Dysdera kourosh Bellvert, Zamani &amp; Dimitrov sp. nov. (SMF), left palp. 18. Prolateral view. 19. Anterior view. 20. Retrolateral view. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10663892" httpUri="https://zenodo.org/record/10663892/files/figure.png" pageId="10" pageNumber="220">Fig. 20</figureCitation>
) and significantly wider in
<taxonomicName id="4C73B3D6B732D85B708588DE4527FA5B" authorityName="Fomichev &amp; Marusik" authorityYear="2021" box="[220,381,1412,1438]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="220" phylum="Arthropoda" rank="species" species="mikhailovi">
<emphasis id="B9071447B732D85B708588DE4527FA5B" box="[220,381,1412,1438]" italics="true" pageId="10" pageNumber="220">D. mikhailovi</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFE2B5A4B732D85B71D488DF468CFA5B" author="Fomichev &amp; Marusik" box="[397,726,1412,1438]" firstAuthor="Fomichev" pageId="10" pageNumber="220" pagination="73 - 89" refId="ref11966" refString="Fomichev A. A. &amp; Marusik Y. M. 2021. Notes on the spider genus Dysdera Latreille, 1804 (Araneae: Dysderidae) in Central Asia. Zootaxa 5006 (1): 73 - 89. https: // doi. org / 10.11646 / zootaxa. 5006.1.10" type="journal article" year="2021">Fomichev &amp; Marusik 2021</bibRefCitation>
: fig. 46) and
<taxonomicName id="4C73B3D6B732D85B732A88DE47A0FA5B" authorityName="Zamani, Marusik &amp; Szuts" authorityYear="2023" box="[883,1018,1413,1438]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="220" phylum="Arthropoda" rank="species" species="sagartia">
<emphasis id="B9071447B732D85B732A88DE47A0FA5B" box="[883,1018,1413,1438]" italics="true" pageId="10" pageNumber="220">D. sagartia</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFE2B5A4B732D85B745388DF415AFA5B" author="Zamani" box="[1034,1280,1412,1438]" etAl="et al." firstAuthor="Zamani" pageId="10" pageNumber="220" pagination="43 - 86" refId="ref13584" refString="Zamani A., Marusik Y. M. &amp; Szuts T. 2023 b. A survey of the spider genus Dysdera Latreille, 1804 (Araneae, Dysderidae) in Iran, with fourteen new species and notes on two fossil genera. 86: 43 - 86. https: // doi. org / 10.3897 / zookeys. 1146.97517" type="journal article" year="2023" yearSuffix="b">
Zamani
<emphasis id="B9071447B732D85B743688DE40F1FA5B" box="[1135,1195,1412,1438]" italics="true" pageId="10" pageNumber="220">et al.</emphasis>
2023b
</bibRefCitation>
: fig. 19a), (2) the convex crest, with wider apical part (
<figureCitation id="1348D4D0B732D85B728988FE4740FA05" box="[720,794,1445,1472]" captionStart="Figs 914. 911" captionStartId="11.[189,241,1848,1874]" captionTargetBox="[189,1398,265,1810]" captionTargetId="figure-16@11.[189,1398,265,1810]" captionTargetPageId="11" captionText="Figs 914. 911. Male Dysdera naouelae Bellvert &amp; Dimitrov sp. nov. (SMF). 9. Prolateral view. 10. Anterior view. 11. Retrolateral view. 1214. Female Dysdera naouelae Bellvert &amp; Dimitrov sp. nov. (SMF), vulva. 12. Ventral view. 13. Lateral view. 14. Dorsal view. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10663888" httpUri="https://zenodo.org/record/10663888/files/figure.png" pageId="10" pageNumber="220">Fig. 9</figureCitation>
), vs concave at the middle in
<taxonomicName id="4C73B3D6B732D85B74DD88FD415CFA7A" authorityName="Bellvert &amp; Dimitrov &amp; Zamani &amp; Arnedo" authorityYear="2024" box="[1156,1286,1445,1471]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="220" phylum="Arthropoda" rank="species" species="kourosh">
<emphasis id="B9071447B732D85B74DD88FD415CFA7A" box="[1156,1286,1445,1471]" italics="true" pageId="10" pageNumber="220">D. kourosh</emphasis>
</taxonomicName>
(
<figureCitation id="1348D4D0B732D85B754E88FE4137FA05" box="[1303,1389,1445,1472]" captionStart="Figs 1820" captionStartId="15.[189,241,1963,1989]" captionTargetBox="[194,1398,1197,1925]" captionTargetId="figure-300@15.[189,1398,1197,1925]" captionTargetPageId="15" captionText="Figs 1820. Male Dysdera kourosh Bellvert, Zamani &amp; Dimitrov sp. nov. (SMF), left palp. 18. Prolateral view. 19. Anterior view. 20. Retrolateral view. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10663892" httpUri="https://zenodo.org/record/10663892/files/figure.png" pageId="10" pageNumber="220">Fig. 18</figureCitation>
) and wider in the basal part than in the apical part in
<taxonomicName id="4C73B3D6B732D85B7345889347E6FA24" authorityName="Fomichev &amp; Marusik" authorityYear="2021" box="[796,956,1479,1505]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="220" phylum="Arthropoda" rank="species" species="mikhailovi">
<emphasis id="B9071447B732D85B7345889347E6FA24" box="[796,956,1479,1505]" italics="true" pageId="10" pageNumber="220">D. mikhailovi</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFE2B5A4B732D85B7392889C4155FA24" author="Fomichev &amp; Marusik" box="[971,1295,1479,1505]" firstAuthor="Fomichev" pageId="10" pageNumber="220" pagination="73 - 89" refId="ref11966" refString="Fomichev A. A. &amp; Marusik Y. M. 2021. Notes on the spider genus Dysdera Latreille, 1804 (Araneae: Dysderidae) in Central Asia. Zootaxa 5006 (1): 73 - 89. https: // doi. org / 10.11646 / zootaxa. 5006.1.10" type="journal article" year="2021">Fomichev &amp; Marusik 2021</bibRefCitation>
: fig. 44), and (3) proximal border of the additional lateral sheet not fused with distal haematodocha (
<figureCitation id="1348D4D0B732D85B74B688B2413DF9C6" box="[1263,1383,1513,1539]" captionStart="Figs 914. 911" captionStartId="11.[189,241,1848,1874]" captionTargetBox="[189,1398,265,1810]" captionTargetId="figure-16@11.[189,1398,265,1810]" captionTargetPageId="11" captionText="Figs 914. 911. Male Dysdera naouelae Bellvert &amp; Dimitrov sp. nov. (SMF). 9. Prolateral view. 10. Anterior view. 11. Retrolateral view. 1214. Female Dysdera naouelae Bellvert &amp; Dimitrov sp. nov. (SMF), vulva. 12. Ventral view. 13. Lateral view. 14. Dorsal view. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10663888" httpUri="https://zenodo.org/record/10663888/files/figure.png" pageId="10" pageNumber="220">Figs 9, 11</figureCitation>
), vs fused in the other 2 species. Female differs by the curved spermatheca in dorsal view (
<figureCitation id="1348D4D0B732D85B74B48B51411FF9E0" box="[1261,1349,1546,1573]" captionStart="Figs 914. 911" captionStartId="11.[189,241,1848,1874]" captionTargetBox="[189,1398,265,1810]" captionTargetId="figure-16@11.[189,1398,265,1810]" captionTargetPageId="11" captionText="Figs 914. 911. Male Dysdera naouelae Bellvert &amp; Dimitrov sp. nov. (SMF). 9. Prolateral view. 10. Anterior view. 11. Retrolateral view. 1214. Female Dysdera naouelae Bellvert &amp; Dimitrov sp. nov. (SMF), vulva. 12. Ventral view. 13. Lateral view. 14. Dorsal view. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10663888" httpUri="https://zenodo.org/record/10663888/files/figure.png" pageId="10" pageNumber="220">Fig. 12</figureCitation>
), vs straight in
<taxonomicName id="4C73B3D6B732D85B71658B764586F983" authorityName="Fomichev &amp; Marusik" authorityYear="2021" box="[316,476,1580,1606]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="220" phylum="Arthropoda" rank="species" species="mikhailovi">
<emphasis id="B9071447B732D85B71658B764586F983" box="[316,476,1580,1606]" italics="true" pageId="10" pageNumber="220">D. mikhailovi</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFE2B5A4B732D85B71B28B77476BF983" author="Fomichev &amp; Marusik" box="[491,817,1580,1606]" firstAuthor="Fomichev" pageId="10" pageNumber="220" pagination="73 - 89" refId="ref11966" refString="Fomichev A. A. &amp; Marusik Y. M. 2021. Notes on the spider genus Dysdera Latreille, 1804 (Araneae: Dysderidae) in Central Asia. Zootaxa 5006 (1): 73 - 89. https: // doi. org / 10.11646 / zootaxa. 5006.1.10" type="journal article" year="2021">Fomichev &amp; Marusik 2021</bibRefCitation>
: fig. 54) and the presence of a sclerotized neck in the spermatheca of
<taxonomicName id="4C73B3D6B732D85B71FB8B144668F9AD" authority="Bellvert &amp; Dimitrov &amp; Zamani &amp; Arnedo, 2024" authorityName="Bellvert &amp; Dimitrov &amp; Zamani &amp; Arnedo" authorityYear="2024" box="[418,562,1614,1640]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="220" phylum="Arthropoda" rank="species" species="naouelae" status="sp. nov.">
<emphasis id="B9071447B732D85B71FB8B144668F9AD" box="[418,562,1614,1640]" italics="true" pageId="10" pageNumber="220">D. naouelae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A234A93CB732D85B72608B1546C9F9AD" box="[569,659,1614,1640]" pageId="10" pageNumber="220" rank="species">sp. nov.</taxonomicNameLabel>
(
<figureCitation id="1348D4D0B732D85B72FA8B1546A1F9AD" box="[675,763,1614,1640]" captionStart="Figs 914. 911" captionStartId="11.[189,241,1848,1874]" captionTargetBox="[189,1398,265,1810]" captionTargetId="figure-16@11.[189,1398,265,1810]" captionTargetPageId="11" captionText="Figs 914. 911. Male Dysdera naouelae Bellvert &amp; Dimitrov sp. nov. (SMF). 9. Prolateral view. 10. Anterior view. 11. Retrolateral view. 1214. Female Dysdera naouelae Bellvert &amp; Dimitrov sp. nov. (SMF), vulva. 12. Ventral view. 13. Lateral view. 14. Dorsal view. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10663888" httpUri="https://zenodo.org/record/10663888/files/figure.png" pageId="10" pageNumber="220">Fig. 14</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3699BDEB732D85B70E48BD54515F932" pageId="10" pageNumber="220" type="etymology">
<paragraph id="8BCCC855B732D85B70E48BD5451EF96D" blockId="10.[189,324,1678,1704]" box="[189,324,1678,1704]" pageId="10" pageNumber="220">
<heading id="D0847F39B732D85B70E48BD5451EF96D" bold="true" box="[189,324,1678,1704]" fontSize="11" level="3" pageId="10" pageNumber="220" reason="3">
<emphasis id="B9071447B732D85B70E48BD5451EF96D" bold="true" box="[189,324,1678,1704]" pageId="10" pageNumber="220">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8BCCC855B732D85B70E48BE04515F932" blockId="10.[189,1398,1723,1783]" pageId="10" pageNumber="220">The new species is named after Naouel El Jaarani Oualkadi, partner of the first author, for all her patience and support.</paragraph>
</subSubSection>
<subSubSection id="C3699BDEB732D85D70E48A474507FE5D" lastPageId="12" lastPageNumber="222" pageId="10" pageNumber="220" type="materials_examined">
<paragraph id="8BCCC855B732D85B70E48A474536F8F3" blockId="10.[189,364,1820,1846]" box="[189,364,1820,1846]" pageId="10" pageNumber="220">
<heading id="D0847F39B732D85B70E48A474536F8F3" bold="true" box="[189,364,1820,1846]" fontSize="11" level="3" pageId="10" pageNumber="220" reason="3">
<emphasis id="B9071447B732D85B70E48A474536F8F3" bold="true" box="[189,364,1820,1846]" pageId="10" pageNumber="220">Type material</emphasis>
</heading>
</paragraph>
<paragraph id="8BCCC855B732D85B70B38A124506F8A6" blockId="10.[189,1192,1865,1925]" box="[234,348,1865,1891]" pageId="10" pageNumber="220">
<emphasis id="B9071447B732D85B70B38A124506F8A6" bold="true" box="[234,348,1865,1891]" pageId="10" pageNumber="220">
<typeStatus id="54C876F7B732D85B70B38A124506F8A6" box="[234,348,1865,1891]" pageId="10" pageNumber="220" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8BCCC855B732D85B70E48A3040F2F840" blockId="10.[189,1192,1865,1925]" box="[189,1192,1899,1925]" pageId="10" pageNumber="220">
<materialsCitation id="3B1BC208B732D85B709E8A3040F2F840" box="[199,1192,1899,1925]" collectionCode="SMF" country="TURKESTAN" location="TURKESTAN" pageId="10" pageNumber="220" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<collectingCountry id="F36488C5B732D85B709E8A304523F840" box="[199,377,1899,1925]" pageId="10" pageNumber="220">TURKESTAN</collectingCountry>
” •
<specimenCount id="9D7503DCB732D85B71C58A304591F840" box="[412,459,1899,1925]" count="1" pageId="10" pageNumber="220" type="male">1 ♂</specimenCount>
; precise country, collection date and collector unknown;
<collectionCode id="ED625090B732D85B74318A3040FEF840" box="[1128,1188,1899,1925]" collectionName="Germany, Frankfurt-am-Main, Forschungsinstitut und Naturmuseum Senckenberg" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="10" pageNumber="220">SMF</collectionCode>
.
</materialsCitation>
</paragraph>
<paragraph id="8BCCC855B732D85B70B38AF04506F800" blockId="10.[189,1192,1963,2025]" box="[234,348,1963,1989]" pageId="10" pageNumber="220">
<emphasis id="B9071447B732D85B70B38AF04506F800" bold="true" box="[234,348,1963,1989]" pageId="10" pageNumber="220">
<typeStatus id="54C876F7B732D85B70B38AF04506F800" box="[234,348,1963,1989]" pageId="10" pageNumber="220" type="paratype">Paratype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8BCCC855B732D85B70E48A9540F2F82D" blockId="10.[189,1192,1963,2025]" box="[189,1192,1998,2025]" pageId="10" pageNumber="220">
<materialsCitation id="3B1BC208B732D85B709E8A9540FEF82D" box="[199,1188,1998,2025]" collectionCode="SMF" country="TURKESTAN" location="TURKESTAN" pageId="10" pageNumber="220" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<collectingCountry id="F36488C5B732D85B709E8A954523F82D" box="[199,377,1998,2024]" pageId="10" pageNumber="220">TURKESTAN</collectingCountry>
” •
<specimenCount id="9D7503DCB732D85B71C58A954593F82C" box="[412,457,1998,2025]" count="1" pageId="10" pageNumber="220" type="female">1 ♀</specimenCount>
; precise country, collection date and collector unknown;
<collectionCode id="ED625090B732D85B74318A9540FEF82D" box="[1128,1188,1998,2024]" collectionName="Germany, Frankfurt-am-Main, Forschungsinstitut und Naturmuseum Senckenberg" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="10" pageNumber="220">SMF</collectionCode>
</materialsCitation>
.
</paragraph>
<caption id="DF0C98DDB733D85A70E48A634099F85F" ID-DOI="http://doi.org/10.5281/zenodo.10663888" ID-Zenodo-Dep="10663888" httpUri="https://zenodo.org/record/10663888/files/figure.png" pageId="11" pageNumber="221" startId="11.[189,241,1848,1874]" targetBox="[189,1398,265,1810]" targetPageId="11">
<paragraph id="8BCCC855B733D85A70E48A634099F85F" blockId="11.[189,1399,1848,1946]" pageId="11" pageNumber="221">
<emphasis id="B9071447B733D85A70E48A6345DFF897" bold="true" box="[189,389,1848,1874]" pageId="11" pageNumber="221">Figs 914. 911.</emphasis>
Male
<taxonomicName id="4C73B3D6B733D85A718D8A6247FCF896" authority="Bellvert &amp; Dimitrov" authorityName="Bellvert &amp; Dimitrov &amp; Zamani &amp; Arnedo" authorityYear="2024" box="[468,934,1849,1875]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="221" phylum="Arthropoda" rank="species" species="naouelae" status="sp. nov.">
<emphasis id="B9071447B733D85A718D8A6246F6F896" box="[468,684,1849,1875]" italics="true" pageId="11" pageNumber="221">Dysdera naouelae</emphasis>
Bellvert &amp; Dimitrov
</taxonomicName>
<taxonomicNameLabel id="A234A93CB733D85A73F78A624050F896" box="[942,1034,1849,1875]" pageId="11" pageNumber="221" rank="species">sp. nov.</taxonomicNameLabel>
(SMF).
<emphasis id="B9071447B733D85A742B8A634025F897" bold="true" box="[1138,1151,1848,1874]" pageId="11" pageNumber="221">9</emphasis>
. Prolateral view.
<emphasis id="B9071447B733D85A750A8A634135F897" bold="true" box="[1363,1391,1848,1874]" pageId="11" pageNumber="221">10</emphasis>
. Anterior view.
<emphasis id="B9071447B733D85A712D8A0745D4F8B3" bold="true" box="[372,398,1884,1910]" pageId="11" pageNumber="221">11</emphasis>
. Retrolateral view.
<emphasis id="B9071447B733D85A72278A07469DF8B3" bold="true" box="[638,711,1884,1910]" pageId="11" pageNumber="221">1214</emphasis>
. Female
<taxonomicName id="4C73B3D6B733D85A73618A07414BF8B3" authority="Bellvert &amp; Dimitrov" authorityName="Bellvert &amp; Dimitrov &amp; Zamani &amp; Arnedo" authorityYear="2024" box="[824,1297,1884,1910]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="221" phylum="Arthropoda" rank="species" species="naouelae" status="sp. nov.">
<emphasis id="B9071447B733D85A73618A074048F8B3" box="[824,1042,1884,1910]" italics="true" pageId="11" pageNumber="221">Dysdera naouelae</emphasis>
Bellvert &amp; Dimitrov
</taxonomicName>
<taxonomicNameLabel id="A234A93CB733D85A75438A07412DF8B3" box="[1306,1399,1884,1910]" pageId="11" pageNumber="221" rank="species">sp. nov.</taxonomicNameLabel>
(SMF), vulva.
<emphasis id="B9071447B733D85A71338A2445DDF85C" bold="true" box="[362,391,1919,1945]" pageId="11" pageNumber="221">12</emphasis>
. Ventral view.
<emphasis id="B9071447B733D85A726F8A244609F85C" bold="true" box="[566,595,1919,1945]" pageId="11" pageNumber="221">13</emphasis>
. Lateral view.
<emphasis id="B9071447B733D85A735A8A24477AF85C" bold="true" box="[771,800,1919,1945]" pageId="11" pageNumber="221">14</emphasis>
. Dorsal view. Scale bars = 0.5 mm.
</paragraph>
</caption>
<paragraph id="8BCCC855B734D85D70E48C534609FEE7" blockId="12.[189,595,264,290]" box="[189,595,264,290]" pageId="12" pageNumber="222">
<heading id="D0847F39B734D85D70E48C534609FEE7" bold="true" box="[189,595,264,290]" fontSize="11" level="3" pageId="12" pageNumber="222" reason="3">
<emphasis id="B9071447B734D85D70E48C534609FEE7" bold="true" box="[189,595,264,290]" pageId="12" pageNumber="222">Comparative material examined</emphasis>
</heading>
</paragraph>
<paragraph id="8BCCC855B734D85D70B38C6D4507FE5D" blockId="12.[189,1399,310,408]" pageId="12" pageNumber="222">
<materialsCitation id="3B1BC208B734D85D70B38C6D4503FE5D" collectingDate="1983-06-12" collectionCode="NHMW" collectorName="S. L. Zonstein" country="KYRGYZYSTAN" county="Arslanbob" location="Jalal-Abad Region" municipality="Ferganski" pageId="12" pageNumber="222" specimenCount="1" specimenCount-male="1">
<taxonomicName id="4C73B3D6B734D85D70B38C6D46CDFE94" ID-CoL="38989" authority="Kroneberg, 1875" authorityName="Kroneberg" authorityYear="1875" box="[234,663,310,337]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="222" phylum="Arthropoda" rank="species" species="tartarica">
<emphasis id="B9071447B734D85D70B38C6D46CDFE94" bold="true" box="[234,663,310,337]" pageId="12" pageNumber="222">
<originalDetermination id="A86D70A9B734D85D70B38C6D46CDFE94" box="[234,663,310,337]" pageId="12" pageNumber="222">
<emphasis id="B9071447B734D85D70B38C6D45E7FE94" bold="true" box="[234,445,310,337]" italics="true" pageId="12" pageNumber="222">Dysdera tartarica</emphasis>
<bibRefCitation id="EFE2B5A4B734D85D719C8C6C46CDFE94" author="Kroneberg" box="[453,663,311,337]" firstAuthor="Kroneberg" pageId="12" pageNumber="222" pagination="1 - 58" refId="ref12310" refString="Kroneberg A. 1875. Araneae. In: Fedtschenko A. P. (ed.) Puteshestvie v Tourkestan. Reisen in Turkestan. Zoologischer Theil. Nachrichten der Gesellschaft der Freunde der Naturwissenschaften zu Moskau 19 (3): 1 - 58." type="journal article" year="1875">Kroneberg, 1875</bibRefCitation>
</originalDetermination>
</emphasis>
</taxonomicName>
(
<figureCitation id="1348D4D0B734D85D72FF8C6C4776FE97" box="[678,812,311,338]" captionStart="Figs 1517" captionStartId="12.[188,239,1963,1989]" captionTargetBox="[189,1398,1188,1924]" captionTargetId="figure-286@12.[189,1398,1188,1924]" captionTargetPageId="12" captionText="Figs 1517. Male Dysdera tartarica Kroneberg, 1875 (NHMW), left palp. 15. Prolateral view. 16. Anterior view.17. Retrolateral view. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10663890" httpUri="https://zenodo.org/record/10663890/files/figure.png" pageId="12" pageNumber="222">Figs 1517</figureCitation>
)
<collectingCountry id="F36488C5B734D85D70E48C0145CDFEB1" box="[189,407,346,372]" pageId="12" pageNumber="222">KYRGYZYSTAN</collectingCountry>
<specimenCount id="9D7503DCB734D85D71E88C0145BBFEB0" box="[433,481,346,373]" count="1" pageId="12" pageNumber="222" type="male">1 ♂</specimenCount>
;
<location id="8EAC9E8EB734D85D71B78C014697FEB1" LSID="urn:lsid:plazi:treatment:03DA7943B732D85F72588E9240F3FE27:8EAC9E8EB734D85D71B78C014697FEB1" box="[494,717,346,372]" country="KYRGYZYSTAN" county="Arslanbob" municipality="Ferganski" name="Jalal-Abad Region" pageId="12" pageNumber="222">Jalal-Abad Region</location>
,
<collectingMunicipality id="6BA8522FB734D85D72838C014717FEB1" box="[730,845,346,372]" pageId="12" pageNumber="222">Ferganski</collectingMunicipality>
range,
<collectingCounty id="62ADB0D9B734D85D73FD8C014078FEB1" box="[932,1058,346,372]" pageId="12" pageNumber="222">Arslanbob</collectingCounty>
;
<date id="FFCDEE95B734D85D74778C014090FEB1" box="[1070,1226,346,373]" pageId="12" pageNumber="222" value="1983-06-12">
<collectingDate id="EF89177DB734D85D74778C014090FEB1" box="[1070,1226,346,373]" pageId="12" pageNumber="222" value="1983-06-12">12 Jun. 1983</collectingDate>
</date>
;
<collectorName id="2686AD83B734D85D748E8C01412DFEB1" box="[1239,1399,346,372]" pageId="12" pageNumber="222">S.L. Zonstein</collectorName>
leg.;
<collectionCode id="ED625090B734D85D70A18C254503FE5D" box="[248,345,382,408]" collectionName="Austria, Wien, Naturhistorisches Museum Wien" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="12" pageNumber="222" type="Museum">NHMW</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3699BDEB734D85C70E48CE4475AF8BD" lastPageId="13" lastPageNumber="223" pageId="12" pageNumber="222" type="description">
<paragraph id="8BCCC855B734D85D70E48CE44514FE1C" blockId="12.[189,334,447,473]" box="[189,334,447,473]" pageId="12" pageNumber="222">
<heading id="D0847F39B734D85D70E48CE44514FE1C" bold="true" box="[189,334,447,473]" fontSize="11" level="3" pageId="12" pageNumber="222" reason="3">
<emphasis id="B9071447B734D85D70E48CE44514FE1C" bold="true" box="[189,334,447,473]" pageId="12" pageNumber="222">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8BCCC855B734D85D70B38CB545E2FDCD" blockId="12.[187,1400,493,802]" box="[234,440,493,520]" pageId="12" pageNumber="222">
<emphasis id="B9071447B734D85D70B38CB54570FDCD" bold="true" box="[234,298,494,520]" pageId="12" pageNumber="222">Male</emphasis>
(
<figureCitation id="1348D4D0B734D85D71608CB645EAFDCD" box="[313,432,493,520]" captionStart="Figs 914. 911" captionStartId="11.[189,241,1848,1874]" captionTargetBox="[189,1398,265,1810]" captionTargetId="figure-16@11.[189,1398,265,1810]" captionTargetPageId="11" captionText="Figs 914. 911. Male Dysdera naouelae Bellvert &amp; Dimitrov sp. nov. (SMF). 9. Prolateral view. 10. Anterior view. 11. Retrolateral view. 1214. Female Dysdera naouelae Bellvert &amp; Dimitrov sp. nov. (SMF), vulva. 12. Ventral view. 13. Lateral view. 14. Dorsal view. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10663888" httpUri="https://zenodo.org/record/10663888/files/figure.png" pageId="12" pageNumber="222">Figs 911</figureCitation>
)
</paragraph>
<paragraph id="8BCCC855B734D85D70E48F4A404DFCE4" blockId="12.[187,1400,493,802]" pageId="12" pageNumber="222">PROSOMA. 3.55 long; maximum width 2.63; minimum width 1.63. Uniformly orange-red; smooth with some small black grains mainly at front. Frontal border roughly triangular, about ½ of carapace length; anterior lateral borders convergent; rounded at maximum dorsal width, back lateral borders rounded; posterior margin narrow, straight. Eye diameters: AME 0.18; PLE 0.14; PME 0.11; AMEs on edge of the frontal border, separated from one another by about ⅔ diameter, close to PLEs; PMEs very close to one another, about ⅓ of PME diameter from PLEs. Labium trapezoid-shaped, base wider than the distal part; longer than wide at base; semi-circular groove at the tip. Sternum orange, frontally darker, becoming lighter towards the back; wrinkled; covered with setae mainly on margin.</paragraph>
<paragraph id="8BCCC855B734D85D70E48E124045FC08" blockId="12.[189,1399,839,973]" pageId="12" pageNumber="222">CHELICERAE. 1.36 long, about ⅖ of carapace length in dorsal view; fang medium-sized, 0.99 long; paturon dorsal and ventral side completely covered with piligerous granulations. Cheliceral inner groove short, about ⅓ of cheliceral length; armed with three teeth and lamina at base; B&gt; D= M; D triangular, located roughly at the centre of the groove; B close to basal lamina; M close to B.</paragraph>
<paragraph id="8BCCC855B734D85C70E48EAF4122FDF8" blockId="12.[188,1400,1011,1109]" lastBlockId="13.[187,1400,265,573]" lastPageId="13" lastPageNumber="223" pageId="12" pageNumber="222">LEGS. Front legs orange, back legs yellow. Lengths of leg segments: fe1 2.85; pa1 1.81; ti1 2.40; me1 2.65; ta1 0.67; total 10.38; fe2 2.62; pa2 1.65; ti2 2.17; me2 2.48; ta2 0.61; total 9.52; fe3 2.03; pa3 1.12; ti3 1.30; me3 1.98; ta3 0.52; total 6.94; fe4 2.74; pa4 1.40; ti4 1.93; me4 2.57; ta4 0.66; total 9.30; fe Pdp 1.67; pa Pdp 0.94; ti Pdp 0.81; ta Pdp 0.93; total 4.35; leg formula: 1&gt;2&gt;4&gt;3. Leg 1 has two terminal spines on the forward margin; leg 2 two terminal spines on the forward margin. Fe3d spines in two rows; forward 2; backward 1; pa3 spineless; tb3d spines arranged in two bands; proximal 1.0.1; medial-proximal 0; medial-distal 0; distal 1.0.1; tb3v spines arranged in four bands; proximal 0.1.0; medial-proximal 1.1.0; medial-distal 0.1.0; distal 1.0.0; with two terminal spines. Fe4d spines in two rows, forward 2; backward 6-5; pa4 spineless; tb4d spines arranged in two bands; proximal 1.0.1; medial-proximal 0; medial-distal 0; distal 1.0.1; tb4v spines arranged in four bands; proximal 1.2.0; medial-proximal 1.1.0; medial-distal 0.1.0; distal 1.0.0; with two terminal spines. Dorsal side of the frontal legs smooth; ventral side of the pedipalp covered with small piligerous grains. Claws with 8 teeth or less; hardly larger than claw width.</paragraph>
<caption id="DF0C98DDB734D85D70E58AF046B6F82C" ID-DOI="http://doi.org/10.5281/zenodo.10663890" ID-Zenodo-Dep="10663890" httpUri="https://zenodo.org/record/10663890/files/figure.png" pageId="12" pageNumber="222" startId="12.[188,239,1963,1989]" targetBox="[189,1398,1188,1924]" targetPageId="12">
<paragraph id="8BCCC855B734D85D70E58AF046B6F82C" blockId="12.[186,1399,1963,2025]" pageId="12" pageNumber="222">
<emphasis id="B9071447B734D85D70E58AF04519F800" bold="true" box="[188,323,1963,1989]" pageId="12" pageNumber="222">Figs 1517.</emphasis>
Male
<taxonomicName id="4C73B3D6B734D85D71D28AF04747F803" authority="Kroneberg, 1875" authorityName="Kroneberg" authorityYear="1875" box="[395,797,1963,1990]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="222" phylum="Arthropoda" rank="species" species="tartarica">
<emphasis id="B9071447B734D85D71D28AF04602F800" box="[395,600,1963,1989]" italics="true" pageId="12" pageNumber="222">Dysdera tartarica</emphasis>
<bibRefCitation id="EFE2B5A4B734D85D72058AF04747F803" author="Kroneberg" box="[604,797,1963,1990]" firstAuthor="Kroneberg" pageId="12" pageNumber="222" pagination="1 - 58" refId="ref12310" refString="Kroneberg A. 1875. Araneae. In: Fedtschenko A. P. (ed.) Puteshestvie v Tourkestan. Reisen in Turkestan. Zoologischer Theil. Nachrichten der Gesellschaft der Freunde der Naturwissenschaften zu Moskau 19 (3): 1 - 58." type="journal article" year="1875">Kroneberg, 1875</bibRefCitation>
</taxonomicName>
(NHMW), left palp.
<emphasis id="B9071447B734D85D74538AF0407DF800" bold="true" box="[1034,1063,1963,1989]" pageId="12" pageNumber="222">15</emphasis>
. Prolateral view.
<emphasis id="B9071447B734D85D74B58AF04153F800" bold="true" box="[1260,1289,1963,1989]" pageId="12" pageNumber="222">16</emphasis>
. Anterior view.
<emphasis id="B9071447B734D85D70A28A954542F82D" bold="true" box="[251,280,1998,2024]" pageId="12" pageNumber="222">17</emphasis>
. Retrolateral view. Scale bar = 0.5 mm.
</paragraph>
</caption>
<paragraph id="8BCCC855B735D85C70E48F3F4799FD67" blockId="13.[189,1398,612,674]" pageId="13" pageNumber="223">OPISTHOSOMA. 3.76 long; cream-colored; cylindrical. Abdominal dorsal setae 0.02 long; thick, roughly straight, blunt, tip not enlarged; uniformly and thickly distributed.</paragraph>
<paragraph id="8BCCC855B735D85C70E48F9246DDFC38" blockId="13.[188,1400,713,1022]" pageId="13" pageNumber="223">
PALP (
<figureCitation id="1348D4D0B735D85C715F8F924527FD26" box="[262,381,713,739]" captionStart="Figs 914. 911" captionStartId="11.[189,241,1848,1874]" captionTargetBox="[189,1398,265,1810]" captionTargetId="figure-16@11.[189,1398,265,1810]" captionTargetPageId="11" captionText="Figs 914. 911. Male Dysdera naouelae Bellvert &amp; Dimitrov sp. nov. (SMF). 9. Prolateral view. 10. Anterior view. 11. Retrolateral view. 1214. Female Dysdera naouelae Bellvert &amp; Dimitrov sp. nov. (SMF), vulva. 12. Ventral view. 13. Lateral view. 14. Dorsal view. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10663888" httpUri="https://zenodo.org/record/10663888/files/figure.png" pageId="13" pageNumber="223">Figs 911</figureCitation>
). T slightly shorter than ED, external distal border sloped forwards, internal one sloped backward. ED not bent, same T axis in lateral view, internal distal border not expanded. IS wider than ES, both more or less parallel; IS continuous to tip (
<figureCitation id="1348D4D0B735D85C73418E544734FCEF" box="[792,878,783,810]" captionStart="Figs 914. 911" captionStartId="11.[189,241,1848,1874]" captionTargetBox="[189,1398,265,1810]" captionTargetId="figure-16@11.[189,1398,265,1810]" captionTargetPageId="11" captionText="Figs 914. 911. Male Dysdera naouelae Bellvert &amp; Dimitrov sp. nov. (SMF). 9. Prolateral view. 10. Anterior view. 11. Retrolateral view. 1214. Female Dysdera naouelae Bellvert &amp; Dimitrov sp. nov. (SMF), vulva. 12. Ventral view. 13. Lateral view. 14. Dorsal view. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10663888" httpUri="https://zenodo.org/record/10663888/files/figure.png" pageId="13" pageNumber="223">Fig. 10</figureCitation>
). ED tip sloped towards back in lateral view. C present, long; distal end beside ED internal tip; distal border rounded, smooth, markedly expanded, perpendicular to ED (
<figureCitation id="1348D4D0B735D85C71E28E0D4659FCB5" box="[443,515,854,880]" captionStart="Figs 914. 911" captionStartId="11.[189,241,1848,1874]" captionTargetBox="[189,1398,265,1810]" captionTargetId="figure-16@11.[189,1398,265,1810]" captionTargetPageId="11" captionText="Figs 914. 911. Male Dysdera naouelae Bellvert &amp; Dimitrov sp. nov. (SMF). 9. Prolateral view. 10. Anterior view. 11. Retrolateral view. 1214. Female Dysdera naouelae Bellvert &amp; Dimitrov sp. nov. (SMF), vulva. 12. Ventral view. 13. Lateral view. 14. Dorsal view. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10663888" httpUri="https://zenodo.org/record/10663888/files/figure.png" pageId="13" pageNumber="223">Fig. 9</figureCitation>
). AC absent. LF absent. L well developed, with a sclerotized processus-like basal lateral apophysis. LA absent. F absent. AL present, well developed; proximal border in posterior view smooth, not fused with distal haematodocha. P not fused to T (
<figureCitation id="1348D4D0B735D85C73AF8EC74065FC72" box="[1014,1087,924,951]" captionStart="Figs 914. 911" captionStartId="11.[189,241,1848,1874]" captionTargetBox="[189,1398,265,1810]" captionTargetId="figure-16@11.[189,1398,265,1810]" captionTargetPageId="11" captionText="Figs 914. 911. Male Dysdera naouelae Bellvert &amp; Dimitrov sp. nov. (SMF). 9. Prolateral view. 10. Anterior view. 11. Retrolateral view. 1214. Female Dysdera naouelae Bellvert &amp; Dimitrov sp. nov. (SMF), vulva. 12. Ventral view. 13. Lateral view. 14. Dorsal view. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10663888" httpUri="https://zenodo.org/record/10663888/files/figure.png" pageId="13" pageNumber="223">Fig. 9</figureCitation>
); lateral length from ⅔ to as long as T width; ridge present, parallel to T; not expanded, upper margin smooth; distally ridge-like expanded; posterior margin not folded.
</paragraph>
<paragraph id="8BCCC855B735D85C70B3897E45CBFB09" blockId="13.[189,1400,1060,1228]" pageId="13" pageNumber="223">
<emphasis id="B9071447B735D85C70B3897E451FFBFA" bold="true" box="[234,325,1061,1087]" pageId="13" pageNumber="223">Female</emphasis>
(
<figureCitation id="1348D4D0B735D85C710C897F4581FBFA" box="[341,475,1060,1087]" captionStart="Figs 914. 911" captionStartId="11.[189,241,1848,1874]" captionTargetBox="[189,1398,265,1810]" captionTargetId="figure-16@11.[189,1398,265,1810]" captionTargetPageId="11" captionText="Figs 914. 911. Male Dysdera naouelae Bellvert &amp; Dimitrov sp. nov. (SMF). 9. Prolateral view. 10. Anterior view. 11. Retrolateral view. 1214. Female Dysdera naouelae Bellvert &amp; Dimitrov sp. nov. (SMF), vulva. 12. Ventral view. 13. Lateral view. 14. Dorsal view. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10663888" httpUri="https://zenodo.org/record/10663888/files/figure.png" pageId="13" pageNumber="223">Figs 1214</figureCitation>
) PROSOMA. 3.77 long; maximum width 2.73; minimum width 1.75. Orange, anteriorly darker, becoming lighter towards posterior. Anterior border almost round. Eye diameters: AME 0.19; PLE 0.11; PME 0.15, separated from one another by about 1 diameter and about ½ PME diameter from PLEs. Sternum uniformly orange.
</paragraph>
<paragraph id="8BCCC855B735D85C70E489A846F8FAF4" blockId="13.[189,1399,1265,1329]" pageId="13" pageNumber="223">CHELICERAE. 1.53 long; 1.18. Cheliceral inner groove short, about ⅓ of cheliceral length; armed with three teeth and lamina at base; B&gt; D&gt; M.</paragraph>
<paragraph id="8BCCC855B735D85C70E4880345BCF9AD" blockId="13.[188,1400,1367,1641]" pageId="13" pageNumber="223">LEGS. Lengths of leg segments: fe1 2.74; pa1 1.81; ti1 2.17; me1 2.26; ta1 0.61; total 9.61; fe2 2.51; pa2 1.69; ti2 2.02; me2 2.19; ta2 0.55; total 8.96; fe3 1.96; pa3 1.13; ti3 1.32; me3 1.87; ta3 0.49; total 6.78; fe4 2.61; pa4 1.38; ti4 1.93; me4 2.53; ta4 0.56; total 9.01; fe Pdp 1.66; pa Pdp 0.80; ti Pdp 0.77; ta Pdp 0.94; total 4.18; leg formula 1&gt;4&gt; 2&gt;3. Leg 1 one terminal spine on forward margin. Fe3d spines in one row; 3; tb3v spines arranged in three bands; proximal 1.1.1; medial-proximal 1.1.1; medial-distal 0; distal 1.0.0; with two terminal spines. Fe4d spines in two rows; forward 3-2; backward 6-5; tb4v spines arranged in four bands; proximal 1.1.1-0; medial-proximal 1.1.0; medial-distal 0.0-1.0-1; distal 1.1-0.0; with two terminal spines.</paragraph>
<paragraph id="8BCCC855B735D85C70E48BCB410AF96F" blockId="13.[189,1360,1680,1706]" box="[189,1360,1680,1706]" pageId="13" pageNumber="223">OPISTHOSOMA. 4.41 long. Abdominal dorsal hairs 0.08 long. All other somatic characters as in male.</paragraph>
<paragraph id="8BCCC855B735D85C70E48B8A475AF8BD" blockId="13.[187,1400,1745,1913]" pageId="13" pageNumber="223">
VULVA (
<figureCitation id="1348D4D0B735D85C71438B8A45F9F92E" box="[282,419,1745,1771]" captionStart="Figs 914. 911" captionStartId="11.[189,241,1848,1874]" captionTargetBox="[189,1398,265,1810]" captionTargetId="figure-16@11.[189,1398,265,1810]" captionTargetPageId="11" captionText="Figs 914. 911. Male Dysdera naouelae Bellvert &amp; Dimitrov sp. nov. (SMF). 9. Prolateral view. 10. Anterior view. 11. Retrolateral view. 1214. Female Dysdera naouelae Bellvert &amp; Dimitrov sp. nov. (SMF), vulva. 12. Ventral view. 13. Lateral view. 14. Dorsal view. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10663888" httpUri="https://zenodo.org/record/10663888/files/figure.png" pageId="13" pageNumber="223">Figs 1214</figureCitation>
). DA wider than long, fused to VA (
<figureCitation id="1348D4D0B735D85C73098B8A47F2F92E" box="[848,936,1745,1771]" captionStart="Figs 914. 911" captionStartId="11.[189,241,1848,1874]" captionTargetBox="[189,1398,265,1810]" captionTargetId="figure-16@11.[189,1398,265,1810]" captionTargetPageId="11" captionText="Figs 914. 911. Male Dysdera naouelae Bellvert &amp; Dimitrov sp. nov. (SMF). 9. Prolateral view. 10. Anterior view. 11. Retrolateral view. 1214. Female Dysdera naouelae Bellvert &amp; Dimitrov sp. nov. (SMF), vulva. 12. Ventral view. 13. Lateral view. 14. Dorsal view. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10663888" httpUri="https://zenodo.org/record/10663888/files/figure.png" pageId="13" pageNumber="223">Fig. 13</figureCitation>
); DF wide in dorsal view. MF margins fused, sheet-like, well developed, and completely sclerotized. VA rectangular, transparent (
<figureCitation id="1348D4D0B735D85C749A8BAF414DF8CA" box="[1219,1303,1780,1807]" captionStart="Figs 914. 911" captionStartId="11.[189,241,1848,1874]" captionTargetBox="[189,1398,265,1810]" captionTargetId="figure-16@11.[189,1398,265,1810]" captionTargetPageId="11" captionText="Figs 914. 911. Male Dysdera naouelae Bellvert &amp; Dimitrov sp. nov. (SMF). 9. Prolateral view. 10. Anterior view. 11. Retrolateral view. 1214. Female Dysdera naouelae Bellvert &amp; Dimitrov sp. nov. (SMF), vulva. 12. Ventral view. 13. Lateral view. 14. Dorsal view. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10663888" httpUri="https://zenodo.org/record/10663888/files/figure.png" pageId="13" pageNumber="223">Fig. 14</figureCitation>
); frontal region completely sclerotized; posterior region sclerotized except for most internal area; AVD absent. S attachment projected under VA; arms as long as DA (
<figureCitation id="1348D4D0B735D85C737E8A604724F890" box="[807,894,1851,1877]" captionStart="Figs 914. 911" captionStartId="11.[189,241,1848,1874]" captionTargetBox="[189,1398,265,1810]" captionTargetId="figure-16@11.[189,1398,265,1810]" captionTargetPageId="11" captionText="Figs 914. 911. Male Dysdera naouelae Bellvert &amp; Dimitrov sp. nov. (SMF). 9. Prolateral view. 10. Anterior view. 11. Retrolateral view. 1214. Female Dysdera naouelae Bellvert &amp; Dimitrov sp. nov. (SMF), vulva. 12. Ventral view. 13. Lateral view. 14. Dorsal view. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10663888" httpUri="https://zenodo.org/record/10663888/files/figure.png" pageId="13" pageNumber="223">Fig. 12</figureCitation>
), distinctly curved; tips not projected; neck as wide as arms. Laterals of TB directed forward.
</paragraph>
</subSubSection>
<subSubSection id="C3699BDEB735D85C70E48AC4465AF82D" box="[189,512,1951,2024]" pageId="13" pageNumber="223" type="distribution">
<paragraph id="8BCCC855B735D85C70E48AC4450CF87C" blockId="13.[189,342,1951,1977]" box="[189,342,1951,1977]" pageId="13" pageNumber="223">
<heading id="D0847F39B735D85C70E48AC4450CF87C" bold="true" box="[189,342,1951,1977]" fontSize="11" level="3" pageId="13" pageNumber="223" reason="3">
<emphasis id="B9071447B735D85C70E48AC4450CF87C" bold="true" box="[189,342,1951,1977]" pageId="13" pageNumber="223">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8BCCC855B735D85C70E48A95465AF82D" blockId="13.[189,512,1998,2024]" box="[189,512,1998,2024]" pageId="13" pageNumber="223">Central Asia (“Turkestan”).</paragraph>
</subSubSection>
<subSubSection id="C3699BDEB736D85F70E48C5240F3FE27" pageId="14" pageNumber="224" type="discussion">
<paragraph id="8BCCC855B736D85F70E48C524575FEE6" blockId="14.[189,303,265,291]" box="[189,303,265,291]" pageId="14" pageNumber="224">
<heading id="D0847F39B736D85F70E48C524575FEE6" bold="true" box="[189,303,265,291]" fontSize="11" level="3" pageId="14" pageNumber="224" reason="3">
<emphasis id="B9071447B736D85F70E48C524575FEE6" bold="true" box="[189,303,265,291]" pageId="14" pageNumber="224">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8BCCC855B736D85F70E48C6340F3FE27" blockId="14.[189,1399,312,482]" pageId="14" pageNumber="224">
This species belongs to the
<emphasis id="B9071447B736D85F724E8C634625FE97" box="[535,639,312,338]" italics="true" pageId="14" pageNumber="224">aculeata</emphasis>
group (sensu
<bibRefCitation id="EFE2B5A4B736D85F73688C63414EFE97" author="Deeleman-Reinhold &amp; Deeleman" box="[817,1300,312,338]" firstAuthor="Deeleman-Reinhold" pageId="14" pageNumber="224" pagination="141 - 269" refId="ref11600" refString="Deeleman-Reinhold C. L. &amp; Deeleman P. R. 1988. Revision des Dysderinae (Araneae, Dysderidae), les especes mediterraneennes occidentales exceptees. Tijdschrift voor Entomologie 131: 141 - 269." type="journal article" year="1988">Deeleman-Reinhold &amp; Deeleman 1988</bibRefCitation>
). It was erroneously recorded as
<taxonomicName id="4C73B3D6B736D85F71858C074727FEB3" authority="Kroneberg, 1875" authorityName="Kroneberg" authorityYear="1875" box="[476,893,347,374]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="224" phylum="Arthropoda" rank="species" species="tartarica">
<emphasis id="B9071447B736D85F71858C0746F5FEB0" box="[476,687,348,374]" italics="true" pageId="14" pageNumber="224">Dysdera tartarica</emphasis>
<bibRefCitation id="EFE2B5A4B736D85F72EF8C074727FEB3" author="Kroneberg" box="[694,893,347,374]" firstAuthor="Kroneberg" pageId="14" pageNumber="224" pagination="1 - 58" refId="ref12310" refString="Kroneberg A. 1875. Araneae. In: Fedtschenko A. P. (ed.) Puteshestvie v Tourkestan. Reisen in Turkestan. Zoologischer Theil. Nachrichten der Gesellschaft der Freunde der Naturwissenschaften zu Moskau 19 (3): 1 - 58." type="journal article" year="1875">Kroneberg, 1875</bibRefCitation>
</taxonomicName>
by
<bibRefCitation id="EFE2B5A4B736D85F73F18C074038FEB3" author="Lazarov" box="[936,1122,348,374]" firstAuthor="Lazarov" pageId="14" pageNumber="224" pagination="104 - 106" refId="ref12449" refString="Lazarov S. 2009. A new record of Dysdera tartarica Kroneberg, 1875 with a description of the previously unknown female (Araneae, Dysderidae). Turkish Journal of Arachnology 1 (2, for 2008): 104 - 106." type="journal article" year="2009">Lazarov (2009)</bibRefCitation>
. We examined the male specimen of
<taxonomicName id="4C73B3D6B736D85F71008CDB45BCFE5C" authorityName="Kroneberg" authorityYear="1875" box="[345,486,384,409]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="224" phylum="Arthropoda" rank="species" species="tartarica">
<emphasis id="B9071447B736D85F71008CDB45BCFE5C" box="[345,486,384,409]" italics="true" pageId="14" pageNumber="224">D. tartarica</emphasis>
</taxonomicName>
, described and depicted by
<bibRefCitation id="EFE2B5A4B736D85F73618CDB417BFE5F" author="Deeleman-Reinhold &amp; Deeleman" box="[824,1313,384,410]" firstAuthor="Deeleman-Reinhold" pageId="14" pageNumber="224" pagination="141 - 269" refId="ref11600" refString="Deeleman-Reinhold C. L. &amp; Deeleman P. R. 1988. Revision des Dysderinae (Araneae, Dysderidae), les especes mediterraneennes occidentales exceptees. Tijdschrift voor Entomologie 131: 141 - 269." type="journal article" year="1988">Deeleman-Reinhold &amp; Deeleman (1988)</bibRefCitation>
, which fits the original description and the drawing of
<bibRefCitation id="EFE2B5A4B736D85F72AA8CF847ADFE7B" author="Charitonov" box="[755,1015,419,446]" firstAuthor="Charitonov" page="22" pageId="14" pageNumber="224" pagination="17 - 39" refId="ref11354" refString="Charitonov D. E. 1956. Obzor paukov semeistva Dysderidae fauny SSSR. Uchenye Zapiski Molotovskogo Gosudarstvennogo Universiteta Imeni A. M. Gor'kogo 10: 17 - 39." type="journal article" year="1956">Charitonov (1956: 22</bibRefCitation>
, fig. 2). As clearly seen from its photographs (
<figureCitation id="1348D4D0B736D85F71078C9C45BCFE27" box="[350,486,455,482]" captionStart="Figs 1517" captionStartId="12.[188,239,1963,1989]" captionTargetBox="[189,1398,1188,1924]" captionTargetId="figure-286@12.[189,1398,1188,1924]" captionTargetPageId="12" captionText="Figs 1517. Male Dysdera tartarica Kroneberg, 1875 (NHMW), left palp. 15. Prolateral view. 16. Anterior view.17. Retrolateral view. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10663890" httpUri="https://zenodo.org/record/10663890/files/figure.png" pageId="14" pageNumber="224">Figs 1517</figureCitation>
), it is not conspecific with
<taxonomicName id="4C73B3D6B736D85F73788C9347EBFE24" authority="Bellvert &amp; Dimitrov &amp; Zamani &amp; Arnedo, 2024" authorityName="Bellvert &amp; Dimitrov &amp; Zamani &amp; Arnedo" authorityYear="2024" box="[801,945,455,481]" class="Arachnida" family="Dysderidae" genus="Dysdera" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="224" phylum="Arthropoda" rank="species" species="naouelae" status="sp. nov.">
<emphasis id="B9071447B736D85F73788C9347EBFE24" box="[801,945,455,481]" italics="true" pageId="14" pageNumber="224">D. naouelae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A234A93CB736D85F73E18C934048FE27" box="[952,1042,456,482]" pageId="14" pageNumber="224" rank="species">sp. nov.</taxonomicNameLabel>
(
<figureCitation id="1348D4D0B736D85F74788C9C40C1FE27" box="[1057,1179,455,482]" captionStart="Figs 914. 911" captionStartId="11.[189,241,1848,1874]" captionTargetBox="[189,1398,265,1810]" captionTargetId="figure-16@11.[189,1398,265,1810]" captionTargetPageId="11" captionText="Figs 914. 911. Male Dysdera naouelae Bellvert &amp; Dimitrov sp. nov. (SMF). 9. Prolateral view. 10. Anterior view. 11. Retrolateral view. 1214. Female Dysdera naouelae Bellvert &amp; Dimitrov sp. nov. (SMF), vulva. 12. Ventral view. 13. Lateral view. 14. Dorsal view. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10663888" httpUri="https://zenodo.org/record/10663888/files/figure.png" pageId="14" pageNumber="224">Figs 914</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>