318 lines
38 KiB
XML
318 lines
38 KiB
XML
<document id="F783C83526DA7127B1E784F27D98E1F4" ID-DOI="10.5281/zenodo.5458892" ID-ISSN="1942-1354" ID-Zenodo-Dep="5458892" ID-ZooBank="95AEB9E3-09D8-4BD2-9834-1FA8751B244F" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630211278244" checkinUser="felipe" docAuthor="Huchet, Jean-Bernard" docDate="2020" docId="03CF9B7D9E75FFCFFF283D43FEEC7147" docLanguage="en" docName="0778.pdf" docOrigin="Insecta Mundi 2020 (778)" docStyle="DocumentStyle:6D85C66E4B52897F25DA165649BC06DC.3:InsectaMundi.2009-2020.journal_article" docStyleId="6D85C66E4B52897F25DA165649BC06DC" docStyleName="InsectaMundi.2009-2020.journal_article" docStyleVersion="3" docTitle="Nothochodaeus yeti Huchet 2020, new species" docType="treatment" docVersion="4" lastPageNumber="3" masterDocId="FFF6E3059E76FFCBFFE83E3EFFF07728" masterDocTitle="Two new species of Nothochodaeus Nikolajev, 2005 from the Himalayan region (Coleoptera: Scarabaeoidea: Ochodaeidae)" masterLastPageNumber="11" masterPageNumber="1" pageNumber="2" updateTime="1699187496844" updateUser="plazi" zenodo-license-document="CC-BY-NC-3.0">
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<mods:titleInfo id="30B26B69F5F81AE9EA8F69F2D37B9F8E">
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<mods:title id="F4099EAC9DD7FDB2D20C8C016545E02E">Two new species of Nothochodaeus Nikolajev, 2005 from the Himalayan region (Coleoptera: Scarabaeoidea: Ochodaeidae)</mods:title>
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<mods:name id="00FDBC6776D2B64E4816F081C190DFC1" type="personal">
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<mods:roleTerm id="32DF4A6FECF5E66608E3DBBF7E7CD20C">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="2ED66A220688EB6E8BF2B98DB1CE0034">Huchet, Jean-Bernard</mods:namePart>
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<mods:affiliation id="60C6BDD379C2CCAF78DA798E271C3A84">Muséum National d’Histoire Naturelle, ISYEB UMR 7205, MNHN, CNRS, EPHE, UPMC, Paris-Sorbonne CP 50, Entomologie, 45, rue Buffon F- 75005 Paris, France</mods:affiliation>
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<mods:typeOfResource id="C385EE2515F29AF23F7056E6A0858A9F">text</mods:typeOfResource>
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<mods:title id="859E4B553404DD31983B89B61139A405">Insecta Mundi</mods:title>
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</mods:titleInfo>
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<mods:part id="662A7B82FF7F62911ACF25838143701A">
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<mods:date id="441B34C76A8B286AD175890F7BEEF513">2020</mods:date>
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<mods:detail id="DDE82302254AA6AD321B37B7548078A9" type="pubDate">
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<mods:number id="B138E95755797B90055222648D46B7BA">2020-06-26</mods:number>
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<mods:detail id="F9E642B7C0D4D478B699AAB8D43EBEF8" type="volume">
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<mods:number id="FE93F74CA947E759534AAF1055258C01">2020</mods:number>
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<mods:detail id="495ABCB61EE7F0EA044AE2651B99AD26" type="issue">
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<mods:number id="003D2345430EB21F77E143682B2DC0A7">778</mods:number>
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<mods:identifier id="162F6CB1C1B24C3CFD737EDBA41149C1" type="DOI">10.5281/zenodo.5458892</mods:identifier>
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<mods:identifier id="531271E5E22C517EACECD8B53FC2572E" type="ISSN">1942-1354</mods:identifier>
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<treatment id="03CF9B7D9E75FFCFFF283D43FEEC7147" ID-DOI="http://doi.org/10.5281/zenodo.5451347" ID-Zenodo-Dep="5451347" LSID="urn:lsid:plazi:treatment:03CF9B7D9E75FFCFFF283D43FEEC7147" httpUri="http://treatment.plazi.org/id/03CF9B7D9E75FFCFFF283D43FEEC7147" lastPageId="4" lastPageNumber="3" pageId="3" pageNumber="2">
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<subSubSection id="C37C79E09E75FFC8FF283D43FC8D74BD" box="[192,893,893,917]" pageId="3" pageNumber="2" type="nomenclature">
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<paragraph id="8BD92A6B9E75FFC8FF283D43FC8D74BD" blockId="3.[192,893,893,917]" box="[192,893,893,917]" pageId="3" pageNumber="2">
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<heading id="D0919D079E75FFC8FF283D43FC8D74BD" bold="true" box="[192,893,893,917]" fontSize="10" level="2" pageId="3" pageNumber="2" reason="3">
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<emphasis id="B912F6799E75FFC8FF283D43FC8D74BD" bold="true" box="[192,893,893,917]" pageId="3" pageNumber="2">
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<taxonomicName id="4C6651E89E75FFC8FF283D43FDC774BD" ID-CoL="47SSD" authority="Huchet" authorityName="Huchet" authorityYear="2020" box="[192,567,893,917]" class="Insecta" family="Ochodaeidae" genus="Nothochodaeus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="yeti" status="sp. nov.">
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<emphasis id="B912F6799E75FFC8FF283D43FE3A74BD" bold="true" box="[192,458,893,917]" italics="true" pageId="3" pageNumber="2">Nothochodaeus yeti</emphasis>
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Huchet
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</taxonomicName>
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,
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<taxonomicNameLabel id="A2214B029E75FFC8FDAE3D43FD1974BD" box="[582,745,893,917]" pageId="3" pageNumber="2" rank="species">new species</taxonomicNameLabel>
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(
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<figureCitation id="135D36EE9E75FFC8FD123D43FC8374BD" box="[762,883,893,917]" captionStart-0="Figures 1–3" captionStart-1="Figures 4–6" captionStart-2="Figures 7–9" captionStart-3="Figure 10–11" captionStartId-0="5.[192,287,1787,1809]" captionStartId-1="6.[193,288,1787,1809]" captionStartId-2="7.[192,287,991,1013]" captionStartId-3="7.[192,274,1875,1897]" captionTargetBox-0="[224,1402,273,1757]" captionTargetBox-1="[278,1362,332,1682]" captionTargetBox-2="[341,1295,218,959]" captionTargetBox-3="[503,1114,1079,1849]" captionTargetId-0="figure-20@5.[212,1418,251,1772]" captionTargetId-1="figure-23@6.[256,1376,319,1703]" captionTargetId-2="figure-20@7.[338,1295,213,961]" captionTargetId-3="figure-22@7.[449,1159,1063,1870]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionTargetPageId-2="7" captionTargetPageId-3="7" captionText-0="Figures 1–3. Nothochodaeus yeti Huchet, new species, male holotype. 1) Habitus, dorsal view (missing left antenna figured with shading). 2) Labels. 3) Habitus, laterodorsal view (scale = 5mm)." captionText-1="Figures 4–6. Nothochodaeus yeti Huchet, new species, female allotype. 4) Habitus, dorsal view. 5) Labels. 6) Habitus, lateral-dorsal view (scale = 5mm)." captionText-2="Figures 7–9. Nothochodaeus yeti Huchet, new species. 7) Male (holotype), head, dorsal view, inset illustrating the tripartite subpyramidal horn. 8) Head, ventral view. 9) Female (allotype), head, dorsal view." captionText-3="Figure 10–11. Nothochodaeus yeti Huchet, new species, male genitalia. 10) Genital segment (urite IX). 11) Aedeagus in lateral view, the endophallus not everted (Ep: endophallite)." figureDoi-0="http://doi.org/10.5281/zenodo.5458894" figureDoi-1="http://doi.org/10.5281/zenodo.5458896" figureDoi-2="http://doi.org/10.5281/zenodo.5458898" figureDoi-3="http://doi.org/10.5281/zenodo.5458900" httpUri-0="https://zenodo.org/record/5458894/files/figure.png" httpUri-1="https://zenodo.org/record/5458896/files/figure.png" httpUri-2="https://zenodo.org/record/5458898/files/figure.png" httpUri-3="https://zenodo.org/record/5458900/files/figure.png" pageId="3" pageNumber="2">Fig. 1–11</figureCitation>
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)
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C37C79E09E75FFC8FF283D93FC7972ED" pageId="3" pageNumber="2" type="materials_examined">
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<paragraph id="8BD92A6B9E75FFC8FF283D93FBF6730D" blockId="3.[191,1441,941,1478]" pageId="3" pageNumber="2">
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<emphasis id="B912F6799E75FFC8FF283D93FE7A74ED" bold="true" box="[192,394,941,965]" pageId="3" pageNumber="2">Type material.</emphasis>
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<materialsCitation id="3B0E20369E75FFC8FE7A3D93FBF6730D" collectorName="Genitalia" country="Czech Republic" determinerName="J. - B. Huchet" location="Sikkim" municipality="St. Jakl" pageId="3" pageNumber="2" specimenCount="1" specimenCount-male="1" stateProvince="Praha" typeStatus="holotype">
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<typeStatus id="54DD94C99E75FFC8FE7A3D93FDF074ED" box="[402,512,941,965]" pageId="3" pageNumber="2" type="holotype">Holotype</typeStatus>
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male (CSJP), labeled: a) pale yellow paper, rectangular, printed: “
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<location id="8EB97CB09E75FFC8FADE3D93FA6174ED" LSID="urn:lsid:plazi:treatment:03CF9B7D9E75FFCFFF283D43FEEC7147:8EB97CB09E75FFC8FADE3D93FA6174ED" box="[1334,1425,941,965]" country="Czech Republic" municipality="St. Jakl" name="Sikkim" pageId="3" pageNumber="2" stateProvince="Praha">Sikkim</location>
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”; b) rectangular, orange paper, printed: “
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<collectingMunicipality id="6BBDB0119E75FFC8FD4C3DF3FCF274CD" box="[676,770,973,997]" pageId="3" pageNumber="2">St. Jákl</collectingMunicipality>
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collection /
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<collectingRegion id="49A2E4899E75FFC8FC7B3DF3FBF974CD" box="[915,1033,973,997]" country="Czech Republic" name="Praha" pageId="3" pageNumber="2">PRAGUE</collectingRegion>
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/
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<collectingCountry id="F3716AFB9E75FFC8FBF73DF3FB2674CD" box="[1055,1238,973,997]" name="Czech Republic" pageId="3" pageNumber="2">Czech republic</collectingCountry>
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”; c) rectangular, red paper: “TYPE”; d) red paper: “
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<taxonomicName id="4C6651E89E75FFC8FDB43DD3FCE6732D" authorityName="Nikolajev" authorityYear="2005" box="[604,790,1005,1029]" class="Insecta" family="Ochodaeidae" genus="Nothochodaeus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="genus">
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<emphasis id="B912F6799E75FFC8FDB43DD3FCE6732D" box="[604,790,1005,1029]" italics="true" pageId="3" pageNumber="2">Nothochodaeus</emphasis>
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</taxonomicName>
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/
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<taxonomicName id="4C6651E89E75FFC8FCC33DD0FCA6732D" authority="Huchet, 2020" authorityName="Huchet" authorityYear="2020" box="[811,854,1006,1029]" class="Insecta" family="Ochodaeidae" genus="Nothochodaeus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="yeti" status="sp. nov.">
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<emphasis id="B912F6799E75FFC8FCC33DD0FCA6732D" box="[811,854,1006,1029]" italics="true" pageId="3" pageNumber="2">yeti</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A2214B029E75FFC8FCB53DD0FC6E732D" box="[861,926,1006,1029]" pageId="3" pageNumber="2" rank="species">n. sp.</taxonomicNameLabel>
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/
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<typeStatus id="54DD94C99E75FFC8FC5B3DD3FBBE732D" box="[947,1102,1005,1029]" pageId="3" pageNumber="2" type="holotype">HOLOTYPE</typeStatus>
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<specimenCount id="9D60E1E29E75FFC8FBBD3DD0FB99732D" box="[1109,1129,1006,1029]" pageId="3" pageNumber="2" type="male">♂</specimenCount>
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/
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<determinerName id="7E05C6AF9E75FFC8FB953DD0FAE8732D" box="[1149,1304,1005,1029]" pageId="3" pageNumber="2">J.-B. Huchet</determinerName>
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det. 2020”.
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<collectorName id="26934FBD9E75FFC8FF283A33FEC3730D" box="[192,307,1037,1061]" pageId="3" pageNumber="2">Genitalia</collectorName>
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stored in a small glycerol vial, pinned under the specimen.
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</materialsCitation>
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</paragraph>
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<paragraph id="8BD92A6B9E75FFC8FF003A13FE5F738D" blockId="3.[191,1441,941,1478]" pageId="3" pageNumber="2">
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<materialsCitation id="3B0E20369E75FFC8FF003A13FE5F738D" collectorName="Genitalia & Fig." country="India" county="Chasseurs" determinerName="J. - B. Huchet" location="Anglaise" municipality="Darjeeling" pageId="3" pageNumber="2" specimenCount="1" specimenCount-female="1" typeStatus="allotype">
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||
<typeStatus id="54DD94C99E75FFC8FF003A13FEBD736D" box="[232,333,1069,1093]" pageId="3" pageNumber="2" type="allotype">Allotype</typeStatus>
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female (MNHN) labeled: a) pale yellow paper, rectangular, printed: “
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<collectingCountry id="F3716AFB9E75FFC8FB6B3A13FB4A736D" box="[1155,1210,1069,1093]" name="India" pageId="3" pageNumber="2">Inde</collectingCountry>
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<location id="8EB97CB09E75FFC8FB563A13FAD8736D" LSID="urn:lsid:plazi:treatment:03CF9B7D9E75FFCFFF283D43FEEC7147:8EB97CB09E75FFC8FB563A13FAD8736D" box="[1214,1320,1069,1093]" country="India" county="Chasseurs" municipality="Darjeeling" name="Anglaise" pageId="3" pageNumber="2">Anglaise</location>
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/
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<location id="8EB97CB09E75FFC8FAD23A13FA63736D" LSID="urn:lsid:plazi:treatment:03CF9B7D9E75FFCFFF283D43FEEC7147:8EB97CB09E75FFC8FAD23A13FA63736D" box="[1338,1427,1069,1093]" country="India" county="Chasseurs" municipality="Darjeeling" name="Pedong" pageId="3" pageNumber="2">Pedong</location>
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/
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<location id="8EB97CB09E75FFC8FF283A73FEE5734D" LSID="urn:lsid:plazi:treatment:03CF9B7D9E75FFCFFF283D43FEEC7147:8EB97CB09E75FFC8FF283A73FEE5734D" box="[192,277,1101,1125]" country="India" county="Chasseurs" municipality="Darjeeling" name="Region" pageId="3" pageNumber="2">Région</location>
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de /
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<collectingMunicipality id="6BBDB0119E75FFC8FEA53A73FE3E734D" box="[333,462,1101,1125]" pageId="3" pageNumber="2">Darjeeling</collectingMunicipality>
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/
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<collectingCounty id="62B852E79E75FFC8FE0B3A73FD93734D" box="[483,611,1101,1125]" pageId="3" pageNumber="2">Chasseurs</collectingCounty>
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indigènes / 1934”; b) red paper, printed: “
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<taxonomicName id="4C6651E89E75FFC8FBB53A73FAE7734D" authorityName="Nikolajev" authorityYear="2005" box="[1117,1303,1101,1125]" class="Insecta" family="Ochodaeidae" genus="Nothochodaeus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="genus">
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<emphasis id="B912F6799E75FFC8FBB53A73FAE7734D" box="[1117,1303,1101,1125]" italics="true" pageId="3" pageNumber="2">Nothochodaeus</emphasis>
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</taxonomicName>
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/
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<taxonomicName id="4C6651E89E75FFC8FAC53A70FAA8734D" authority="Huchet, 2020" authorityName="Huchet" authorityYear="2020" box="[1325,1368,1102,1125]" class="Insecta" family="Ochodaeidae" genus="Nothochodaeus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="yeti" status="sp. nov.">
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<emphasis id="B912F6799E75FFC8FAC53A70FAA8734D" box="[1325,1368,1102,1125]" italics="true" pageId="3" pageNumber="2">yeti</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A2214B029E75FFC8FAB73A70FA50734D" box="[1375,1440,1102,1125]" pageId="3" pageNumber="2" rank="species">n. sp.</taxonomicNameLabel>
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/
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<typeStatus id="54DD94C99E75FFC8FF383A53FE9573AD" box="[208,357,1133,1157]" pageId="3" pageNumber="2" type="allotype">ALLOTYPE</typeStatus>
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<specimenCount id="9D60E1E29E75FFC8FE863A50FE7273AD" box="[366,386,1134,1157]" pageId="3" pageNumber="2" type="female">♀</specimenCount>
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/
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<determinerName id="7E05C6AF9E75FFC8FE733A50FDC773AD" box="[411,567,1133,1157]" pageId="3" pageNumber="2">J.-B. Huchet</determinerName>
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det. 2020”.
|
||
<collectorName id="26934FBD9E75FFC8FD253A53FCB073AD" box="[717,832,1133,1157]" pageId="3" pageNumber="2">Genitalia</collectorName>
|
||
stored in a small glycerol vial, pinned under the specimen. (
|
||
<collectorName id="26934FBD9E75FFC8FEA13AB3FE89738D" box="[329,377,1165,1189]" pageId="3" pageNumber="2">Fig.</collectorName>
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x-y)
|
||
</materialsCitation>
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</paragraph>
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<paragraph id="8BD92A6B9E75FFC8FF003A93FC7972ED" blockId="3.[191,1441,941,1478]" pageId="3" pageNumber="2">
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<materialsCitation id="3B0E20369E75FFC8FF003A93FC7A73CD" collectingDate="1994-09-18" collectorName="M. Hreblay & T. Csovan" country="Nepal" elevation="2300" latitude="28.166666" location="One" longLatPrecision="1233" longitude="85.28333" municipality="Ganesh Himal." pageId="3" pageNumber="2" specimenCount="1" typeStatus="paratype">
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<location id="8EB97CB09E75FFC8FF003A93FEEA73ED" LSID="urn:lsid:plazi:treatment:03CF9B7D9E75FFCFFF283D43FEEC7147:8EB97CB09E75FFC8FF003A93FEEA73ED" box="[232,282,1197,1221]" country="Nepal" latitude="28.166666" longLatPrecision="1233" longitude="85.28333" municipality="Ganesh Himal." name="One" pageId="3" pageNumber="2">One</location>
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||
female
|
||
<typeStatus id="54DD94C99E75FFC8FE973A90FE1D73EE" box="[383,493,1198,1222]" pageId="3" pageNumber="2" type="paratype">paratype</typeStatus>
|
||
(NMPC) labeled: a) “
|
||
<collectingCountry id="F3716AFB9E75FFC8FD113A93FCAC73ED" box="[761,860,1197,1221]" name="Nepal" pageId="3" pageNumber="2">NEPAL</collectingCountry>
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.
|
||
<collectingMunicipality id="6BBDB0119E75FFC8FC813A93FBD473ED" box="[873,1060,1197,1221]" pageId="3" pageNumber="2">Ganesh Himal.</collectingMunicipality>
|
||
/
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<locationDeviation id="9A6B33499E75FFC8FBD63A93FAD173ED" box="[1086,1313,1197,1221]" location="Thangjet." pageId="3" pageNumber="2">
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<quantity id="4C9E878E9E75FFC8FBD63A93FB8D73ED" box="[1086,1149,1197,1221]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="3" pageNumber="2" unit="km" value="2.0">2 km</quantity>
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||
W Thangjet.
|
||
</locationDeviation>
|
||
/
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<geoCoordinate id="EE524CAC9E75FFC8FAD23A92FA6B73ED" box="[1338,1435,1196,1221]" degrees="85" direction="east" minutes="17" orientation="longitude" pageId="3" pageNumber="2" precision="925" value="85.28333">85°17′E</geoCoordinate>
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,
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<geoCoordinate id="EE524CAC9E75FFC8FF283AF2FED473CD" box="[192,292,1228,1253]" degrees="28" direction="north" minutes="10" orientation="latitude" pageId="3" pageNumber="2" precision="925" value="28.166666">28°10′N</geoCoordinate>
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. /
|
||
<quantity id="4C9E878E9E75FFC8FEA83AF3FE6F73CD" box="[320,415,1229,1253]" metricMagnitude="3" metricUnit="m" metricValue="2.3" pageId="3" pageNumber="2" unit="m" value="2300.0">
|
||
<elevation id="004BCD589E75FFC8FEA83AF3FE6F73CD" box="[320,415,1229,1253]" metricMagnitude="3" metricUnit="m" metricValue="2.3" pageId="3" pageNumber="2" unit="m" value="2300.0">2300 m</elevation>
|
||
</quantity>
|
||
.
|
||
<date id="FFD80CAB9E75FFC8FE433AF3FDC373CD" box="[427,563,1229,1253]" pageId="3" pageNumber="2" value="1994-09-18">
|
||
<collectingDate id="EF9CF5439E75FFC8FE433AF3FDC373CD" box="[427,563,1229,1253]" pageId="3" pageNumber="2" value="1994-09-18">18.IX.1994</collectingDate>
|
||
</date>
|
||
. /
|
||
<collectorName id="26934FBD9E75FFC8FDA73AF0FD2B73CD" box="[591,731,1229,1253]" pageId="3" pageNumber="2">M. Hreblay</collectorName>
|
||
&
|
||
<collectorName id="26934FBD9E75FFC8FCEA3AF0FC8F73CD" box="[770,895,1229,1253]" pageId="3" pageNumber="2">T. Csőván</collectorName>
|
||
”
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B0E20369E75FFC8FC703AF3FED6726D" collectorName="David Kral" country="Czech Republic" elevation="2200" latitude="27.833334" location="One" longLatPrecision="1235" longitude="85.45" pageId="3" pageNumber="2" specimenCount="1" stateProvince="Praha" typeStatus="paratype">
|
||
b) white label, rectangular: “ex coll.
|
||
<collectorName id="26934FBD9E75FFC8FABF3AF3FF08722D" pageId="3" pageNumber="2">David Král</collectorName>
|
||
/ National Museum /
|
||
<collectingRegion id="49A2E4899E75FFC8FDEC3AD0FDAF722E" box="[516,607,1262,1286]" country="Czech Republic" name="Praha" pageId="3" pageNumber="2">Prague</collectingRegion>
|
||
,
|
||
<collectingCountry id="F3716AFB9E75FFC8FD833AD3FCDB722D" box="[619,811,1261,1285]" name="Czech Republic" pageId="3" pageNumber="2">Czech Republic</collectingCountry>
|
||
”.
|
||
<location id="8EB97CB09E75FFC8FCA83AD3FC82722D" LSID="urn:lsid:plazi:treatment:03CF9B7D9E75FFCFFF283D43FEEC7147:8EB97CB09E75FFC8FCA83AD3FC82722D" box="[832,882,1261,1285]" country="Czech Republic" latitude="27.833334" longLatPrecision="1235" longitude="85.45" name="One" pageId="3" pageNumber="2" stateProvince="Praha">One</location>
|
||
female
|
||
<typeStatus id="54DD94C99E75FFC8FC3C3AD0FBB2722E" box="[980,1090,1262,1286]" pageId="3" pageNumber="2" type="paratype">paratype</typeStatus>
|
||
(NMPC> CJBH) labeled: a) “
|
||
<collectingCountry id="F3716AFB9E75FFC8FF233B33FEE4720D" box="[203,276,1293,1317]" name="Nepal" pageId="3" pageNumber="2">Nepal</collectingCountry>
|
||
Helambu, Mulkharka / - Chisapani
|
||
<geoCoordinate id="EE524CAC9E75FFC8FD273B32FCDD720D" box="[719,813,1292,1317]" degrees="85" direction="east" minutes="27" orientation="longitude" pageId="3" pageNumber="2" precision="925" value="85.45">85°27′E</geoCoordinate>
|
||
/
|
||
<geoCoordinate id="EE524CAC9E75FFC8FCAB3B32FC54720D" box="[835,932,1292,1317]" degrees="27" direction="north" minutes="50" orientation="latitude" pageId="3" pageNumber="2" precision="925" value="27.833334">27°50′N</geoCoordinate>
|
||
1800–2500–
|
||
<quantity id="4C9E878E9E75FFC8FBD63B33FB68720D" box="[1086,1176,1293,1317]" metricMagnitude="3" metricUnit="m" metricValue="2.2" pageId="3" pageNumber="2" unit="m" value="2200.0">
|
||
<elevation id="004BCD589E75FFC8FBD63B33FB68720D" box="[1086,1176,1293,1317]" metricMagnitude="3" metricUnit="m" metricValue="2.2" pageId="3" pageNumber="2" unit="m" value="2200.0">2200 m</elevation>
|
||
</quantity>
|
||
/ 26 8 97 lg Fadrizi & Ahrens”
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B0E20369E75FFC8FED93B13FEF172AD" collectorName="David Kral & One" country="Czech Republic" location="Jochen Martens" pageId="3" pageNumber="2" specimenCount="1" stateProvince="Praha" typeStatus="paratype">
|
||
b) white label, rectangular: “ex coll.
|
||
<collectorName id="26934FBD9E75FFC8FD0B3B13FC9A726D" box="[739,874,1325,1349]" pageId="3" pageNumber="2">David Král</collectorName>
|
||
/ National Museum /
|
||
<collectingRegion id="49A2E4899E75FFC8FB863B10FB39726E" box="[1134,1225,1326,1350]" country="Czech Republic" name="Praha" pageId="3" pageNumber="2">Prague</collectingRegion>
|
||
,
|
||
<collectingCountry id="F3716AFB9E75FFC8FB3C3B13FA62726D" box="[1236,1426,1325,1349]" name="Czech Republic" pageId="3" pageNumber="2">Czech Republic</collectingCountry>
|
||
”.
|
||
<collectorName id="26934FBD9E75FFC8FF283B73FF02724D" box="[192,242,1357,1381]" pageId="3" pageNumber="2">One</collectorName>
|
||
female
|
||
<typeStatus id="54DD94C99E75FFC8FEB83B70FE4E724E" box="[336,446,1358,1382]" pageId="3" pageNumber="2" type="paratype">paratype</typeStatus>
|
||
(SMNS) labeled: a) white label, rectangular: “NEPAL-Expeditionen /
|
||
<location id="8EB97CB09E75FFC8FAE03B73FF0572AD" LSID="urn:lsid:plazi:treatment:03CF9B7D9E75FFCFFF283D43FEEC7147:8EB97CB09E75FFC8FAE03B73FF0572AD" country="Czech Republic" name="Jochen Martens" pageId="3" pageNumber="2" stateProvince="Praha">Jochen Martens</location>
|
||
”
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B0E20369E75FFC8FEE53B53FC7972ED" collectingDate="1999-06-16" collectorName="Ilam Dist. Zw. Mai & Martens & Daams & Patrikeev V." country="Nepal" elevation="1800" location="One" pageId="3" pageNumber="2" specimenCount="1" typeStatus="paratype">
|
||
b) white label, rectangular: “255
|
||
<collectorName id="26934FBD9E75FFC8FD733B53FC7072AD" box="[667,896,1389,1413]" pageId="3" pageNumber="2">Ilam Dist. Zw. Mai</collectorName>
|
||
Pok- / hari, Mai Majuwa und Gitang / Khola 2100–
|
||
<quantity id="4C9E878E9E75FFC8FEE33BB3FE95728D" box="[267,357,1421,1445]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="3" pageNumber="2" unit="m" value="1800.0">
|
||
<elevation id="004BCD589E75FFC8FEE33BB3FE95728D" box="[267,357,1421,1445]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="3" pageNumber="2" unit="m" value="1800.0">1800 m</elevation>
|
||
</quantity>
|
||
Kulturland / 25 Aug 83,
|
||
<collectorName id="26934FBD9E75FFC8FD733BB0FCF3728D" box="[667,771,1422,1445]" pageId="3" pageNumber="2">Martens</collectorName>
|
||
&
|
||
<collectorName id="26934FBD9E75FFC8FCC13BB0FC8F728D" box="[809,895,1422,1445]" pageId="3" pageNumber="2">Daams</collectorName>
|
||
leg.”.
|
||
<location id="8EB97CB09E75FFC8FC253BB3FC0F728D" LSID="urn:lsid:plazi:treatment:03CF9B7D9E75FFCFFF283D43FEEC7147:8EB97CB09E75FFC8FC253BB3FC0F728D" box="[973,1023,1421,1445]" country="Nepal" name="One" pageId="3" pageNumber="2">One</location>
|
||
female
|
||
<typeStatus id="54DD94C99E75FFC8FB893BB0FB3F728E" box="[1121,1231,1422,1446]" pageId="3" pageNumber="2" type="paratype">paratype</typeStatus>
|
||
(CDKC) labeled: “E
|
||
<collectingCountry id="F3716AFB9E75FFC8FF0D3B90FEB872EE" box="[229,328,1454,1478]" name="Nepal" pageId="3" pageNumber="2">NEPAL</collectingCountry>
|
||
, Hilles env. / 10–
|
||
<date id="FFD80CAB9E75FFC8FDFE3B93FD6D72ED" box="[534,669,1453,1477]" pageId="3" pageNumber="2" value="1999-06-16">
|
||
<collectingDate id="EF9CF5439E75FFC8FDFE3B93FD6D72ED" box="[534,669,1453,1477]" pageId="3" pageNumber="2" value="1999-06-16">16.06.1999</collectingDate>
|
||
</date>
|
||
/ leg.
|
||
<collectorName id="26934FBD9E75FFC8FD0D3B93FC8E72ED" box="[741,894,1453,1477]" pageId="3" pageNumber="2">Patrikeev V.</collectorName>
|
||
”
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37C79E09E75FFC8FF283BE3FCD6717D" pageId="3" pageNumber="2" type="diagnosis">
|
||
<paragraph id="8BD92A6B9E75FFC8FF283BE3FCD6717D" blockId="3.[192,1440,1501,1622]" pageId="3" pageNumber="2">
|
||
<emphasis id="B912F6799E75FFC8FF283BE3FEBF72DD" bold="true" box="[192,335,1501,1525]" pageId="3" pageNumber="2">Diagnosis.</emphasis>
|
||
This species has the body robust, strongly convex, densely pubescent, roughly unicolored; the head stretched forward in a pointed direction, with a tripartite subpyramidal horn (male), or with two tubercles (female). The underside is chestnut-brown, the legs orange-brown, and the antennae testaceous yellow. The stridulatory peg is present.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37C79E09E75FFCFFF283853FA507237" lastPageId="4" lastPageNumber="3" pageId="3" pageNumber="2" type="description">
|
||
<paragraph id="8BD92A6B9E75FFCFFF283853FA507387" blockId="3.[192,1441,1645,1925]" lastBlockId="4.[192,1441,215,1199]" lastPageId="4" lastPageNumber="3" pageId="3" pageNumber="2">
|
||
<emphasis id="B912F6799E75FFC8FF283853FE9A71AD" bold="true" box="[192,362,1645,1669]" pageId="3" pageNumber="2">Description.</emphasis>
|
||
<typeStatus id="54DD94C99E75FFC8FE983853FE2E71AD" box="[368,478,1645,1669]" pageId="3" pageNumber="2" type="holotype">Holotype</typeStatus>
|
||
male (
|
||
<figureCitation id="135D36EE9E75FFC8FDD83853FD6471AD" box="[560,660,1645,1669]" captionStart="Figures 1–3" captionStartId="5.[192,287,1787,1809]" captionTargetBox="[224,1402,273,1757]" captionTargetId="figure-20@5.[212,1418,251,1772]" captionTargetPageId="5" captionText="Figures 1–3. Nothochodaeus yeti Huchet, new species, male holotype. 1) Habitus, dorsal view (missing left antenna figured with shading). 2) Labels. 3) Habitus, laterodorsal view (scale = 5mm)." figureDoi="http://doi.org/10.5281/zenodo.5458894" httpUri="https://zenodo.org/record/5458894/files/figure.png" pageId="3" pageNumber="2">Fig. 1, 3</figureCitation>
|
||
,
|
||
<figureCitation id="135D36EE9E75FFC8FD483850FD3D71AD" box="[672,717,1645,1669]" captionStart="Figures 7–9" captionStartId="7.[192,287,991,1013]" captionTargetBox="[341,1295,218,959]" captionTargetId="figure-20@7.[338,1295,213,961]" captionTargetPageId="7" captionText="Figures 7–9. Nothochodaeus yeti Huchet, new species. 7) Male (holotype), head, dorsal view, inset illustrating the tripartite subpyramidal horn. 8) Head, ventral view. 9) Female (allotype), head, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5458898" httpUri="https://zenodo.org/record/5458898/files/figure.png" pageId="3" pageNumber="2">7, 8</figureCitation>
|
||
,
|
||
<figureCitation id="135D36EE9E75FFC8FD323853FCD471AD" box="[730,804,1645,1669]" captionStart="Figure 10–11" captionStartId="7.[192,274,1875,1897]" captionTargetBox="[503,1114,1079,1849]" captionTargetId="figure-22@7.[449,1159,1063,1870]" captionTargetPageId="7" captionText="Figure 10–11. Nothochodaeus yeti Huchet, new species, male genitalia. 10) Genital segment (urite IX). 11) Aedeagus in lateral view, the endophallus not everted (Ep: endophallite)." figureDoi="http://doi.org/10.5281/zenodo.5458900" httpUri="https://zenodo.org/record/5458900/files/figure.png" pageId="3" pageNumber="2">10, 11</figureCitation>
|
||
).
|
||
<taxonomicName id="4C6651E89E75FFC8FCD23853FC3271AD" authorityName="Linnaeus" authorityYear="1758" box="[826,962,1645,1669]" class="Insecta" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="order">Coleoptera</taxonomicName>
|
||
: Scarabaeoidea:
|
||
<taxonomicName id="4C6651E89E75FFC8FB643853FADD71AD" authorityName="Mulsant and Rey" authorityYear="1871" box="[1164,1325,1645,1669]" class="Insecta" family="Ochodaeidae" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="family">Ochodaeidae</taxonomicName>
|
||
.
|
||
<emphasis id="B912F6799E75FFC8FADE3853FA5071AD" bold="true" box="[1334,1440,1645,1669]" pageId="3" pageNumber="2">Length:</emphasis>
|
||
10.5 mm (from the apex of the mandibles to the apical part of the tergite VIII).
|
||
<emphasis id="B912F6799E75FFC8FB6D38B3FB2F718D" bold="true" box="[1157,1247,1677,1701]" pageId="3" pageNumber="2">Width:</emphasis>
|
||
5.4 mm.
|
||
<emphasis id="B912F6799E75FFC8FAB838B3FA50718D" bold="true" box="[1360,1440,1677,1701]" pageId="3" pageNumber="2">Head:</emphasis>
|
||
Transverse, sub-hexagonal in outline, the clypeus projected forward, parabolic; clypeal horn subpyramidal, tripartite, π- shaped, resulting from the fusion of two basal tubercles with a small raised transverse anterior keel (
|
||
<figureCitation id="135D36EE9E75FFC8FE8338D3FE30702D" box="[363,448,1773,1797]" captionStart="Figures 7–9" captionStartId="7.[192,287,991,1013]" captionTargetBox="[341,1295,218,959]" captionTargetId="figure-20@7.[338,1295,213,961]" captionTargetPageId="7" captionText="Figures 7–9. Nothochodaeus yeti Huchet, new species. 7) Male (holotype), head, dorsal view, inset illustrating the tripartite subpyramidal horn. 8) Head, ventral view. 9) Female (allotype), head, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5458898" httpUri="https://zenodo.org/record/5458898/files/figure.png" pageId="3" pageNumber="2">Fig. 7a</figureCitation>
|
||
). Surface matte or barely shiny, shortly pubescent, the setae testaceous, obliquely oriented backwards; surface covered with closely spaced granules of heterogeneous size, distant from each other by 1× their diameter on the disc, the granules smaller and tightened on both sides. Posterior part of the vertex with very distinct transverse ripples. Labrum transverse, strongly emarginate and transversely split into two superimposed laminae in the middle front, the lowest edge and sides pubescent; both sides of the labrum convex, hyaline, long pubescent. Eyes large, prominent, globose, lacking ventral projection of canthus. Mandibles subequal, falciform, slightly concave dorsally, sharpedged near the base, the apex and outer edge distinctly darkened; both mandibles 4-dentate on their inner edge (the apical tooth included), the two median teeth (concealed by the labrum in upper view) strongly developed, obliquely oriented forward. Mentum oblong, subtrapezoidal, slightly broader at its base, the sides lowered, the disc slightly concave, without longitudinal furrow. Antenna 10-segmented, 3-antennomere club, testaceous, shiny, the antennal scape very massive, the outer club segment distinctly brightened and shortly pubescent at the upper edge.
|
||
<emphasis id="B912F6799E72FFCFFCA53F89FC2976E7" bold="true" box="[845,985,439,463]" pageId="4" pageNumber="3">Pronotum</emphasis>
|
||
transverse, strongly convex, entirely margined, the edge forming a gutter on each side and along the base. Anterior margin deeply emarginate behind the head, with a thin hyaline orange membrane in front. Front angles obtuse, projecting forward, the posterior ones regularly rounded. Pronotal surface strongly granulate; granules shiny, setose, the setae obliquely directed anteriorly; surface slightly shiny, black with reddish sides.
|
||
<emphasis id="B912F6799E72FFCFFB513C09FAE07567" bold="true" box="[1209,1296,567,591]" pageId="4" pageNumber="3">Elytra</emphasis>
|
||
transverse, slightly shiny, punctato-striate, the basal third lighter, reddish, entirely margined except for the area in front of the humeral callus; elytral punctation strong and tight consisting of small setose granules on a microreticulate background, the testaceous minute setae oriented backward; elytral striae consisting of sunken medium-sized points, separated by 1× their diameter. Humeral callus present but feebly marked. Scutellum black, triangular elongate, lateral edges finely marginate, surface covered distally with coarse setose punctures turning into small setose granules proximally.
|
||
<emphasis id="B912F6799E72FFCFFBB23CC9FB2C7427" bold="true" box="[1114,1244,759,783]" pageId="4" pageNumber="3">Abdomen</emphasis>
|
||
strongly convex with six visible ventrites (III–VIII), its outline entirely margined (in ventral view), with a distinct median longitudinal depression. Surface very shiny, covered by numerous minute setose punctures, roughly separated by 1× their diameter; the sternites lightened, brownish-orange along their front and base edges. Tergite VIII (pygidium) pubescent, the punctation consisting of small close setose granules embedded on a microreticulate background. Tergite VII (propygidium) with trapezoidal interlocking mechanism. Metasternal process slightly convex with a thin median groove; mesocoxae widely separated. Stridulatory peg (sternite VI) barely perceptible, vestigial, in the form of a thin hyaline plate.
|
||
<emphasis id="B912F6799E72FFCFFB9B3DE9FB4A74C7" bold="true" box="[1139,1210,983,1007]" pageId="4" pageNumber="3">Legs:</emphasis>
|
||
Protibia tridentate externally, the basal tooth very reduced, with short, acute pollex directed anteriorly; a distinct serrated longitudinal carina originating from the base and extending forward just prior to the insertion of the protarsus. Femur without accessory teeth, surface with two parallel rows of setose punctures. Upper spur of metatibia as long as first metatarsomere.
|
||
<emphasis id="B912F6799E72FFCFFCCC3A69FBA27347" bold="true" box="[804,1106,1111,1135]" pageId="4" pageNumber="3">
|
||
Genitalia (
|
||
<figureCitation id="135D36EE9E72FFCFFC5E3A69FBB27347" box="[950,1090,1111,1135]" captionStart="Figure 10–11" captionStartId="7.[192,274,1875,1897]" captionTargetBox="[503,1114,1079,1849]" captionTargetId="figure-22@7.[449,1159,1063,1870]" captionTargetPageId="7" captionText="Figure 10–11. Nothochodaeus yeti Huchet, new species, male genitalia. 10) Genital segment (urite IX). 11) Aedeagus in lateral view, the endophallus not everted (Ep: endophallite)." figureDoi="http://doi.org/10.5281/zenodo.5458900" httpUri="https://zenodo.org/record/5458900/files/figure.png" pageId="4" pageNumber="3">Fig. 10, 11</figureCitation>
|
||
).
|
||
</emphasis>
|
||
Parameres narrow, widely divergent apically. Endophallus including five serrate endophallites of distinct size and shape, the two larger endophallites (Ep. 1, 2) located at the level of the phallobase (the endophallus being not everted).
|
||
</paragraph>
|
||
<paragraph id="8BD92A6B9E72FFCFFF283AF9FA507237" blockId="4.[192,1441,1223,1311]" pageId="4" pageNumber="3">
|
||
<emphasis id="B912F6799E72FFCFFF283AF9FE2473F7" bold="true" box="[192,468,1223,1247]" pageId="4" pageNumber="3">Sexual dimorphism.</emphasis>
|
||
A well distinct tripartite clypeal horn in male, two small transverse tubercles, largely separated, located on the clypeo-frontal line (female); anterior margin of pronotum deeply emarginate (male), evenly convex (female); abdominal sternites in male with a median longitudinal depression.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37C79E09E72FFCFFF283B09FC807247" pageId="4" pageNumber="3" type="etymology">
|
||
<paragraph id="8BD92A6B9E72FFCFFF283B09FC807247" blockId="4.[192,1440,1335,1391]" pageId="4" pageNumber="3">
|
||
<emphasis id="B912F6799E72FFCFFF283B09FEAB7267" bold="true" box="[192,347,1335,1359]" pageId="4" pageNumber="3">Etymology.</emphasis>
|
||
The name is a noun in apposition, in reference to the famous mythical hairy humanoid creature supposed to inhabit the Himalayan mountains.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37C79E09E72FFCFFF283BB9FBDA72B7" box="[192,1066,1415,1439]" pageId="4" pageNumber="3" type="distribution">
|
||
<paragraph id="8BD92A6B9E72FFCFFF283BB9FBDA72B7" blockId="4.[192,1066,1415,1439]" box="[192,1066,1415,1439]" pageId="4" pageNumber="3">
|
||
<emphasis id="B912F6799E72FFCFFF283BB9FE8372B7" bold="true" box="[192,371,1415,1439]" pageId="4" pageNumber="3">Distribution.</emphasis>
|
||
Himalayan region of
|
||
<collectingCountry id="F3716AFB9E72FFCFFD943BB9FD4D72B7" box="[636,701,1415,1439]" name="India" pageId="4" pageNumber="3">India</collectingCountry>
|
||
(
|
||
<collectingRegion id="49A2E4899E72FFCFFD253BB9FCD772B7" box="[717,807,1415,1439]" country="India" name="Sikkim" pageId="4" pageNumber="3">Sikkim</collectingRegion>
|
||
) and
|
||
<collectingCountry id="F3716AFB9E72FFCFFC843BB9FC4572B7" box="[876,949,1415,1439]" name="Nepal" pageId="4" pageNumber="3">Nepal</collectingCountry>
|
||
(
|
||
<figureCitation id="135D36EE9E72FFCFFC2D3BB9FBEC72B7" box="[965,1052,1415,1439]" captionStart="Figure 17" captionStartId="11.[192,275,781,803]" captionTargetBox="[192,1440,213,746]" captionTargetId="figure-294@11.[192,1440,213,746]" captionTargetPageId="11" captionText="Figure 17. Distribution map of Nothochodaeus yeti Huchet, new species (red dots, the type locality with a black spot) and N. martensi Huchet, new species (blue square with a white spot) in Northern India (Sikkim) and Nepal." figureDoi="http://doi.org/10.5281/zenodo.5458906" httpUri="https://zenodo.org/record/5458906/files/figure.png" pageId="4" pageNumber="3">Fig. 17</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37C79E09E72FFCFFF283B89FEEC7147" pageId="4" pageNumber="3" type="discussion">
|
||
<paragraph id="8BD92A6B9E72FFCFFF283B89FEEC7147" blockId="4.[192,1440,1463,1647]" pageId="4" pageNumber="3">
|
||
<emphasis id="B912F6799E72FFCFFF283B89FEB372E7" bold="true" box="[192,323,1463,1487]" pageId="4" pageNumber="3">Remarks.</emphasis>
|
||
This new species is morphologically close to
|
||
<taxonomicName id="4C6651E89E72FFCFFC8A3B89FAD672E7" authority="(Westwood)" baseAuthorityName="Westwood" baseAuthorityYear="1852" box="[866,1318,1463,1487]" class="Insecta" family="Ochodaeidae" genus="Nothochodaeus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="lutescens">
|
||
<emphasis id="B912F6799E72FFCFFC8A3B89FB6072E7" box="[866,1168,1463,1487]" italics="true" pageId="4" pageNumber="3">Nothochodaeus lutescens</emphasis>
|
||
(Westwood)
|
||
</taxonomicName>
|
||
described from “
|
||
<emphasis id="B912F6799E72FFCFFEE13BE9FE4F72C7" box="[265,447,1495,1519]" italics="true" pageId="4" pageNumber="3">
|
||
<collectingCountry id="F3716AFB9E72FFCFFEE13BE9FEBC72C7" box="[265,332,1495,1519]" name="India" pageId="4" pageNumber="3">India</collectingCountry>
|
||
Orientali
|
||
</emphasis>
|
||
”. The latter, presumably only known by the
|
||
<typeStatus id="54DD94C99E72FFCFFC273BE9FBC672C7" box="[975,1078,1495,1519]" pageId="4" pageNumber="3" type="holotype">holotype</typeStatus>
|
||
(HEC), is very likely a female. Both species differ by the cephalic structures, and notably, in
|
||
<taxonomicName id="4C6651E89E72FFCFFC423BC6FBCB7127" baseAuthorityName="Westwood" baseAuthorityYear="1852" box="[938,1083,1527,1551]" class="Insecta" family="Ochodaeidae" genus="Nothochodaeus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="lutescens">
|
||
<emphasis id="B912F6799E72FFCFFC423BC6FBCB7127" box="[938,1083,1527,1551]" italics="true" pageId="4" pageNumber="3">N. lutescens</emphasis>
|
||
</taxonomicName>
|
||
, by the clypeus less produced anteriorly and the presence of a well-marked curved transverse clypeal carina distinctly raised, followed by two elongate tubercles obliquely located backwards; the head granulation is denser and distinctly tighter.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |