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<document id="F8BB73E3BED462BEB0C59644C06A3A3B" ID-DOI="10.1111/zoj.12305" ID-ISSN="0024-4082" ID-Zenodo-Dep="4721113" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1619443797950" checkinUser="carolina" docAuthor="Bourguignon, Thomas, Scheffrahn, Rudolf H., Nagy, Zoltán Tamás, Sonet, Gontran, Host, Benoît &amp; Roisin, Yves" docDate="2016" docId="03B687D2F5245C0C5073828CA5D47AEE" docLanguage="en" docName="ZoolJLinnSoc.176.15-35.pdf.imf" docOrigin="Zoological Journal of the Linnean Society 176 (1)" docSource="https://academic.oup.com/zoolinnean/article-lookup/doi/10.1111/zoj.12305" docStyle="DocumentStyle:0DD8C314D74634CE09062A86991413F8.2:ZoolJLinnSoc.2002-2009.journal_article" docStyleId="0DD8C314D74634CE09062A86991413F8" docStyleName="ZoolJLinnSoc.2002-2009.journal_article" docStyleVersion="2" docTitle="Humutermes KRISHNAI BOURGUIGNON &amp; ROISIN 2016, GEN. NOV." docType="treatment" docVersion="4" lastPageNumber="22" masterDocId="FF8FFFAAF5215C0B53398404A52E7855" masterDocTitle="Towards a revision of the Neotropical soldierless termites (Isoptera: Termitidae): redescription of the genus Grigiotermes Mathews and description of five new genera" masterLastPageNumber="35" masterPageNumber="15" pageNumber="20" updateTime="1698910514992" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title id="EAD9E68B25A6C326F3313A69751D331A">Towards a revision of the Neotropical soldierless termites (Isoptera: Termitidae): redescription of the genus Grigiotermes Mathews and description of five new genera</mods:title>
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<mods:namePart id="018746D8FE4244A8210DEA88185B393D">Nagy, Zoltán Tamás</mods:namePart>
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<taxonomicName id="4C1F4D47F5245C0E5073828CA03B7EC9" authority="BOURGUIGNON &amp; ROISIN" authorityName="KRISHNAI BOURGUIGNON &amp; ROISIN" authorityYear="2016" box="[842,1301,1671,1693]" class="Insecta" family="Termitidae" genus="Humutermes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="5" pageNumber="20" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B96BEAD6F5245C0E5073828CA6DE7EC9" bold="true" box="[842,1008,1672,1693]" italics="true" pageId="5" pageNumber="20">HUMUTERMES</emphasis>
BOURGUIGNON &amp; ROISIN
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<taxonomicNameLabel id="A25857ADF5245C0E5624828FA0BF7EC9" box="[1309,1425,1671,1692]" pageId="5" pageNumber="20" rank="genus">GEN. NOV.</taxonomicNameLabel>
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<paragraph id="8BA036C4F5245C0E507E82A3A0BB7EE8" blockId="5.[839,1429,1671,1725]" box="[839,1429,1703,1725]" pageId="5" pageNumber="20">
(
<figureCitation id="13242A41F5245C0E507682A3A6E07EE9" box="[847,974,1703,1724]" captionStart="Figure 3" captionStartId="6.[144,222,932,951]" captionTargetBox="[303,1263,196,901]" captionTargetId="figure-224@6.[303,1263,196,902]" captionTargetPageId="6" captionText="Figure 3. Worker. Humutermes krishnai sp. nov.: A, head; B, mandibles; C, foreleg; E, enteric valve (spread on slide). Worker. Humutermes noiroti sp. nov.: D, enteric valve (spread on slide)." figureDoi="http://doi.org/10.5281/zenodo.4721123" httpUri="https://zenodo.org/record/4721123/files/figure.png" pageId="5" pageNumber="20">FIGS 3AE</figureCitation>
,
<figureCitation id="13242A41F5245C0E50E282A3A10E7EE9" box="[987,1056,1703,1724]" captionStart="Figure 7" captionStartId="13.[164,243,1171,1190]" captionTargetBox="[323,1283,194,1141]" captionTargetId="figure-210@13.[323,1283,194,1141]" captionTargetPageId="13" captionText="Figure 7. Worker enteric valve armature, depicted by confocal microscopy. A, Grigiotermes hageni comb. nov.; B, Amplucrutermes inflatus sp. nov.; C, Humutermes krishnai sp. nov.; D, Humutermes noiroti sp. nov.; E, Hydrecotermes arienesho sp. nov.; F, Hydrecotermes kawaii sp. nov.; G, Patawatermes turricola comb. nov.; H, Patawatermes nigripunctatus comb. nov.; I, Rubeotermes jheringi comb. nov. (G, I, reproduced with the permission of Host et al., 2014)." figureDoi="http://doi.org/10.5281/zenodo.4721141" httpUri="https://zenodo.org/record/4721141/files/figure.png" pageId="5" pageNumber="20">7CD</figureCitation>
,
<figureCitation id="13242A41F5245C0E571482A3A1617EE9" box="[1069,1103,1703,1724]" captionStart="Figure 8" captionStartId="14.[144,223,1445,1464]" captionTargetBox="[161,1413,203,1380]" captionTargetId="figure-157@14.[144,1424,195,1416]" captionTargetPageId="14" captionText="Figure 8. Worker digestive tracts. A, Grigiotermes hageni comb. nov.; B, Amplucrutermes inflatus sp. nov.; C, Humutermes krishnai sp. nov.; D, Hydrecotermes arienesho sp. nov.; E, Patawatermes nigripunctatus comb. nov.; F, Rubeotermes jheringi comb. nov.. From left to right: dorsal, right, ventral, and left views." figureDoi="http://doi.org/10.5281/zenodo.4721145" httpUri="https://zenodo.org/record/4721145/files/figure.png" pageId="5" pageNumber="20">8C</figureCitation>
,
<figureCitation id="13242A41F5245C0E576582A3A18F7EE9" box="[1116,1185,1703,1724]" captionStart="Figure 9" captionStartId="16.[144,219,1344,1363]" captionTargetBox="[144,1424,193,1314]" captionTargetId="figure-174@16.[144,1424,193,1314]" captionTargetPageId="16" captionText="Figure 9. Head, pronotum, mesonotum, and metanotum of alates. A, B, Grigiotermes hageni comb. nov.; C, D, Humutermes krishnai sp. nov.; E, F, Hydrecotermes arienesho sp. nov.; G, H, Hydrecotermes kawaii sp. nov.; I, J, Patawatermes turricola comb. nov.; K, L, Patawatermes nigripunctatus comb. nov.; M, N, Rubeotermes jheringi comb. nov. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4721149" httpUri="https://zenodo.org/record/4721149/files/figure.png" pageId="5" pageNumber="20">9CD</figureCitation>
,
<figureCitation id="13242A41F5245C0E579782A3A1E27EE9" box="[1198,1228,1703,1724]" captionStart="Figure 10" captionStartId="17.[164,243,1158,1177]" captionTargetBox="[164,1444,195,1128]" captionTargetId="figure-263@17.[164,1444,195,1128]" captionTargetPageId="17" captionText="Figure 10. Distribution maps of samples examined in this study." figureDoi="http://doi.org/10.5281/zenodo.4721153" httpUri="https://zenodo.org/record/4721153/files/figure.png" pageId="5" pageNumber="20">10</figureCitation>
,
<figureCitation id="13242A41F5245C0E57E082A3A1D87EE9" box="[1241,1270,1703,1724]" captionStart="Figure 11" captionStartId="18.[144,221,1472,1491]" captionTargetBox="[180,1386,200,1441]" captionTargetId="graphics-133@18.[180,1123,205,1433]" captionTargetPageId="18" captionText="Figure 11. Majority rule consensus phylogenetic tree including 13 focal species constructed by Bayesian inference. Numbers on branches are posterior probabilities. Species from South America are in blue font. The scale bar represents the number of mutation per site." figureDoi="http://doi.org/10.5281/zenodo.4721162" httpUri="https://zenodo.org/record/4721162/files/figure.png" pageId="5" pageNumber="20">11</figureCitation>
;
<tableCitation id="C69D037FF5245C0E563D82ACA05F7EE9" box="[1284,1393,1703,1725]" captionStart="Table 4" captionStartId="6.[144,208,1030,1049]" captionText="Table 4. Measurements (in mm) of 12 workers from ten colonies, ten male imagos from one colony, and six female imagos from one colony of Humutermes krishnai sp. nov." pageId="5" pageNumber="20">TABLES 4</tableCitation>
,
<tableCitation id="C69D037FF5245C0E564782A3A0A27EE8" box="[1406,1420,1703,1725]" captionStart="Table 5" captionStartId="7.[164,228,197,216]" captionText="Table 5. Measurements (in mm) of 12 workers from six colonies, and six male and six female imagos from three colonies of Humutermes noiroti sp. nov." pageId="5" pageNumber="20">5</tableCitation>
)
</paragraph>
<paragraph id="8BA036C4F5245C0E500382CAA1617F57" blockId="5.[826,1441,1742,1794]" pageId="5" pageNumber="20">
<emphasis id="B96BEAD6F5245C0E500382CAA6E57EB6" box="[826,971,1742,1763]" italics="true" pageId="5" pageNumber="20">
<typeStatus id="54A48866F5245C0E500382CAA6407EB6" box="[826,878,1742,1763]" pageId="5" pageNumber="20">Type</typeStatus>
species:
</emphasis>
<taxonomicName id="4C1F4D47F5245C0E50ED82CAA6AB7F57" authority="Bourguignon and Roisin" authorityName="Bourguignon and Roisin" class="Insecta" family="Termitidae" genus="Humutermes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="5" pageNumber="20" phylum="Arthropoda" rank="species" species="krishnai">
<emphasis id="B96BEAD6F5245C0E50ED82CAA1FE7EB6" box="[980,1232,1742,1763]" italics="true" pageId="5" pageNumber="20">Humutermes krishnai</emphasis>
Bourguignon and Roisin
</taxonomicName>
, described below.
</paragraph>
<paragraph id="8BA036C4F5245C0E5003831FA6957F65" blockId="5.[826,1441,1819,1902]" box="[826,955,1819,1840]" pageId="5" pageNumber="20">
<emphasis id="B96BEAD6F5245C0E5003831FA6957F65" box="[826,955,1819,1840]" italics="true" pageId="5" pageNumber="20">Description</emphasis>
</paragraph>
<paragraph id="8BA036C4F5245C0D5003833DA79A7EE3" blockId="5.[826,1441,1819,1902]" lastBlockId="6.[144,759,1451,1902]" lastPageId="6" lastPageNumber="21" pageId="5" pageNumber="20">
<emphasis id="B96BEAD6F5245C0E5003833DA6BE7F1B" box="[826,912,1849,1870]" italics="true" pageId="5" pageNumber="20">Worker:</emphasis>
Very small- to small-sized, head width between 0.49 and
<quantity id="4CE79B21F5245C0E509B835CA1267F38" box="[930,1032,1880,1902]" metricMagnitude="-4" metricUnit="m" metricValue="6.2" pageId="5" pageNumber="20" unit="mm" value="0.62">0.62 mm</quantity>
. Head whitish, fully covered by over 100 small setae. Fontanelle almost indistinct. Fore tibia enlarged near middle. Digestive tract as in
<figureCitation id="13242A41F5275C0D514781CEA7DF7D8A" box="[638,753,1482,1504]" captionStart="Figure 8" captionStartId="14.[144,223,1445,1464]" captionTargetBox="[161,1413,203,1380]" captionTargetId="figure-157@14.[144,1424,195,1416]" captionTargetPageId="14" captionText="Figure 8. Worker digestive tracts. A, Grigiotermes hageni comb. nov.; B, Amplucrutermes inflatus sp. nov.; C, Humutermes krishnai sp. nov.; D, Hydrecotermes arienesho sp. nov.; E, Patawatermes nigripunctatus comb. nov.; F, Rubeotermes jheringi comb. nov.. From left to right: dorsal, right, ventral, and left views." figureDoi="http://doi.org/10.5281/zenodo.4721145" httpUri="https://zenodo.org/record/4721145/files/figure.png" pageId="6" pageNumber="21">Figure 8C</figureCitation>
, closely resembling
<emphasis id="B96BEAD6F5275C0D525181EDA7B87DAB" box="[360,662,1513,1534]" italics="true" pageId="6" pageNumber="21">
<taxonomicName id="4C1F4D47F5275C0D525181EDA72C7DAB" authorityName="Bourguignon &amp; Scheffrahn &amp; Nagy &amp; Sonet &amp; Host &amp; Roisin" authorityYear="2016" box="[360,514,1513,1534]" class="Insecta" family="Termitidae" genus="Anoplotermes" kingdom="Animalia" order="Isoptera" pageId="6" pageNumber="21" phylum="Arthropoda" rank="genus" sensu="stricto">Anoplotermes</taxonomicName>
<taxonomicNameLabel id="A25857ADF5275C0D513081EDA7B87DAB" box="[521,662,1513,1534]" pageId="6" pageNumber="21" sensu="stricto">sensu stricto</taxonomicNameLabel>
</emphasis>
. Mesenteron overlapping with proctodeum to form a distinct, elongated mixed segment. Mesenteric part of mixed segment on internal side apically inflated into a whitish, spherical diverticulum. Enteric valve inserted into an extension of the paunch, composed of six sclerotized pads with well-developed sclerotized armature.
</paragraph>
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<paragraph id="8BA036C4F5275C0D53A987A0A69A7B81" blockId="6.[144,1423,931,980]" pageId="6" pageNumber="21">
<emphasis id="B96BEAD6F5275C0D53A987A0A5D97BE2" bold="true" box="[144,247,932,951]" pageId="6" pageNumber="21">Figure 3.</emphasis>
Worker.
<emphasis id="B96BEAD6F5275C0D526287A0A7947BE2" bold="true" box="[347,698,931,951]" pageId="6" pageNumber="21">
<taxonomicName id="4C1F4D47F5275C0D526287A0A7727BE3" authority="Bourguignon &amp; Scheffrahn &amp; Nagy &amp; Sonet &amp; Host &amp; Roisin, 2016" authorityName="Bourguignon &amp; Scheffrahn &amp; Nagy &amp; Sonet &amp; Host &amp; Roisin" authorityYear="2016" box="[347,604,931,951]" class="Insecta" family="Termitidae" genus="Humutermes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="6" pageNumber="21" phylum="Arthropoda" rank="species" species="krishnai" status="sp. nov.">
<emphasis id="B96BEAD6F5275C0D526287A0A7727BE3" bold="true" box="[347,604,931,951]" italics="true" pageId="6" pageNumber="21">Humutermes krishnai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25857ADF5275C0D515A87A0A7947BE2" box="[611,698,932,951]" pageId="6" pageNumber="21" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
: A, head; B, mandibles; C, foreleg; E, enteric valve (spread on slide). Worker.
<emphasis id="B96BEAD6F5275C0D53D387C5A7147B81" bold="true" box="[234,570,961,980]" pageId="6" pageNumber="21">
<taxonomicName id="4C1F4D47F5275C0D53D387C5A4F67B81" authority="Bourguignon &amp; Scheffrahn &amp; Nagy &amp; Sonet &amp; Host &amp; Roisin, 2016" authorityName="Bourguignon &amp; Scheffrahn &amp; Nagy &amp; Sonet &amp; Host &amp; Roisin" authorityYear="2016" box="[234,472,961,980]" class="Insecta" family="Termitidae" genus="Humutermes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="6" pageNumber="21" phylum="Arthropoda" rank="species" species="noiroti" status="sp. nov.">
<emphasis id="B96BEAD6F5275C0D53D387C5A4F67B81" bold="true" box="[234,472,961,980]" italics="true" pageId="6" pageNumber="21">Humutermes noiroti</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25857ADF5275C0D52D887C5A7147B81" box="[481,570,961,980]" pageId="6" pageNumber="21" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
: D, enteric valve (spread on slide).
</paragraph>
</caption>
<caption id="DF60664CF5275C0D53A98002A7EC7C63" pageId="6" pageNumber="21" startId="6.[144,208,1030,1049]">
<paragraph id="8BA036C4F5275C0D53A98002A7EC7C63" blockId="6.[144,1422,1029,1078]" pageId="6" pageNumber="21">
<emphasis id="B96BEAD6F5275C0D53A98002A5C67C4D" bold="true" box="[144,232,1029,1049]" pageId="6" pageNumber="21">Table 4.</emphasis>
Measurements (in mm) of 12 workers from ten colonies, ten male imagos from one colony, and six female imagos from one colony of
<taxonomicName id="4C1F4D47F5275C0D52678027A74E7C60" authority="Bourguignon &amp; Scheffrahn &amp; Nagy &amp; Sonet &amp; Host &amp; Roisin, 2016" authorityName="Bourguignon &amp; Scheffrahn &amp; Nagy &amp; Sonet &amp; Host &amp; Roisin" authorityYear="2016" box="[350,608,1058,1078]" class="Insecta" family="Termitidae" genus="Humutermes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="6" pageNumber="21" phylum="Arthropoda" rank="species" species="krishnai" status="sp. nov.">
<emphasis id="B96BEAD6F5275C0D52678027A74E7C60" bold="true" box="[350,608,1058,1078]" italics="true" pageId="6" pageNumber="21">Humutermes krishnai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25857ADF5275C0D51508027A7EC7C63" box="[617,706,1059,1078]" pageId="6" pageNumber="21" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="8BA036C4F5275C0D51B78059A0327D0F" pageId="6" pageNumber="21">
<table id="F91FC464F527A3F453A98059A0A37D0F" box="[144,1421,1117,1370]" gridcols="4" gridrows="8" pageId="6" pageNumber="21" rowsContinueIn="7.[164,1441,284,537]">
<tr id="352F3486F527A3F453A98059A0A37C25" box="[144,1421,1117,1136]" gridrow="0" pageId="6" pageNumber="21" rowspan-0="1">
<th id="76FE5DFAF527A3F451B78059A6227C25" box="[654,780,1117,1136]" gridcol="1" gridrow="0" pageId="6" pageNumber="21">Worker</th>
<th id="76FE5DFAF527A3F450F78059A1637C25" box="[974,1101,1117,1136]" gridcol="2" gridrow="0" pageId="6" pageNumber="21">Imago ♂</th>
<th id="76FE5DFAF527A3F456368059A0A37C25" box="[1295,1421,1117,1136]" gridcol="3" gridrow="0" pageId="6" pageNumber="21">Imago ♀</th>
</tr>
<tr id="352F3486F527A3F453A98092A0A37CFF" box="[144,1421,1174,1194]" gridrow="1" pageId="6" pageNumber="21">
<th id="76FE5DFAF527A3F453A98092A4E57CFF" box="[144,459,1174,1194]" gridcol="0" gridrow="1" pageId="6" pageNumber="21">Head length to end of clypeus</th>
<td id="76FE5DFAF527A3F451B78092A6227CFF" box="[654,780,1174,1194]" gridcol="1" gridrow="1" pageId="6" pageNumber="21">0.4910.589</td>
<td id="76FE5DFAF527A3F450F78092A1637CFF" box="[974,1101,1174,1194]" gridcol="2" gridrow="1" pageId="6" pageNumber="21">0.4460.577</td>
<td id="76FE5DFAF527A3F456368092A0A37CFF" box="[1295,1421,1174,1194]" gridcol="3" gridrow="1" pageId="6" pageNumber="21">0.4390.567</td>
</tr>
<tr id="352F3486F527A3F453A980B0A0A37C92" box="[144,1421,1204,1223]" gridrow="2" pageId="6" pageNumber="21">
<th id="76FE5DFAF527A3F453A980B0A4E57C92" box="[144,459,1204,1223]" gridcol="0" gridrow="2" pageId="6" pageNumber="21">Head width</th>
<td id="76FE5DFAF527A3F451B780B0A6227C92" box="[654,780,1204,1223]" gridcol="1" gridrow="2" pageId="6" pageNumber="21">0.5690.636</td>
<td id="76FE5DFAF527A3F450F780B0A1637C92" box="[974,1101,1204,1223]" gridcol="2" gridrow="2" pageId="6" pageNumber="21">0.6820.746</td>
<td id="76FE5DFAF527A3F4563680B0A0A37C92" box="[1295,1421,1204,1223]" gridcol="3" gridrow="2" pageId="6" pageNumber="21">0.6940.748</td>
</tr>
<tr id="352F3486F527A3F453A980D5A0A37CB1" box="[144,1421,1233,1252]" gridrow="3" pageId="6" pageNumber="21">
<th id="76FE5DFAF527A3F453A980D5A4E57CB1" box="[144,459,1233,1252]" gridcol="0" gridrow="3" pageId="6" pageNumber="21">Eye diameter</th>
<td id="76FE5DFAF527A3F451B780D5A6227CB1" box="[654,780,1233,1252]" gridcol="1" gridrow="3" pageId="6" pageNumber="21"></td>
<td id="76FE5DFAF527A3F450F780D5A1637CB1" box="[974,1101,1233,1252]" gridcol="2" gridrow="3" pageId="6" pageNumber="21">0.1760.211</td>
<td id="76FE5DFAF527A3F4563680D5A0A37CB1" box="[1295,1421,1233,1252]" gridcol="3" gridrow="3" pageId="6" pageNumber="21">0.1860.216</td>
</tr>
<tr id="352F3486F527A3F453A980EAA0A37D57" box="[144,1421,1262,1282]" gridrow="4" pageId="6" pageNumber="21">
<th id="76FE5DFAF527A3F453A980EAA4E57D57" box="[144,459,1262,1282]" gridcol="0" gridrow="4" pageId="6" pageNumber="21">Pronotum length</th>
<td id="76FE5DFAF527A3F451B780EAA6227D57" box="[654,780,1262,1282]" gridcol="1" gridrow="4" pageId="6" pageNumber="21"></td>
<td id="76FE5DFAF527A3F450F780EAA1637D57" box="[974,1101,1262,1282]" gridcol="2" gridrow="4" pageId="6" pageNumber="21">0.3430.387</td>
<td id="76FE5DFAF527A3F4563680EAA0A37D57" box="[1295,1421,1262,1282]" gridcol="3" gridrow="4" pageId="6" pageNumber="21">0.3580.397</td>
</tr>
<tr id="352F3486F527A3F453A9810FA0A37D4A" box="[144,1421,1291,1311]" gridrow="5" pageId="6" pageNumber="21">
<th id="76FE5DFAF527A3F453A9810FA4E57D4A" box="[144,459,1291,1311]" gridcol="0" gridrow="5" pageId="6" pageNumber="21">Pronotum width</th>
<td id="76FE5DFAF527A3F451B7810FA6227D4A" box="[654,780,1291,1311]" gridcol="1" gridrow="5" pageId="6" pageNumber="21">0.3160.415</td>
<td id="76FE5DFAF527A3F450F7810FA1637D4A" box="[974,1101,1291,1311]" gridcol="2" gridrow="5" pageId="6" pageNumber="21">0.5200.577</td>
<td id="76FE5DFAF527A3F45636810FA0A37D4A" box="[1295,1421,1291,1311]" gridcol="3" gridrow="5" pageId="6" pageNumber="21">0.5500.604</td>
</tr>
<tr id="352F3486F527A3F453A9812DA0A37D69" box="[144,1421,1321,1340]" gridrow="6" pageId="6" pageNumber="21">
<th id="76FE5DFAF527A3F453A9812DA4E57D69" box="[144,459,1321,1340]" gridcol="0" gridrow="6" pageId="6" pageNumber="21">Hind tibia length</th>
<td id="76FE5DFAF527A3F451B7812DA6227D69" box="[654,780,1321,1340]" gridcol="1" gridrow="6" pageId="6" pageNumber="21">0.4240.532</td>
<td id="76FE5DFAF527A3F450F7812DA1637D69" box="[974,1101,1321,1340]" gridcol="2" gridrow="6" pageId="6" pageNumber="21">0.5590.638</td>
<td id="76FE5DFAF527A3F45636812DA0A37D69" box="[1295,1421,1321,1340]" gridcol="3" gridrow="6" pageId="6" pageNumber="21">0.5910.697</td>
</tr>
<tr id="352F3486F527A3F453A98142A0A37D0F" box="[144,1421,1350,1370]" gridrow="7" pageId="6" pageNumber="21">
<th id="76FE5DFAF527A3F453A98142A4E57D0F" box="[144,459,1350,1370]" gridcol="0" gridrow="7" pageId="6" pageNumber="21">Fore tibia width/length</th>
<td id="76FE5DFAF527A3F451B78142A6227D0F" box="[654,780,1350,1370]" gridcol="1" gridrow="7" pageId="6" pageNumber="21">0.2620.372</td>
<td id="76FE5DFAF527A3F450F78142A1637D0F" box="[974,1101,1350,1370]" gridcol="2" gridrow="7" pageId="6" pageNumber="21"></td>
<td id="76FE5DFAF527A3F456368142A0A37D0F" box="[1295,1421,1350,1370]" gridcol="3" gridrow="7" pageId="6" pageNumber="21"></td>
</tr>
</table>
</paragraph>
<paragraph id="8BA036C4F5275C0D539182BBA7A87F3B" blockId="6.[144,759,1451,1902]" pageId="6" pageNumber="21">
<emphasis id="B96BEAD6F5275C0D539182BBA5D97E81" box="[168,247,1727,1748]" italics="true" pageId="6" pageNumber="21">Imago:</emphasis>
Eyes and ocelli of medium size. Fontanelle of medium size, appearing as an elongated white dot located between eyes. Pronotum semicircular from above, covered by hundreds of small bristles and several dozen long bristles. Posterior margins of meso- and metanotum slightly emarginated, with rounded angles.
</paragraph>
<paragraph id="8BA036C4F5275C0D501F81A8A6B87D94" blockId="6.[806,1422,1452,1902]" box="[806,918,1452,1473]" pageId="6" pageNumber="21">
<emphasis id="B96BEAD6F5275C0D501F81A8A6B87D94" box="[806,918,1452,1473]" italics="true" pageId="6" pageNumber="21">Diagnosis</emphasis>
</paragraph>
<paragraph id="8BA036C4F5275C0D501F81CFA6C67F3B" blockId="6.[806,1422,1452,1902]" pageId="6" pageNumber="21">
<taxonomicName id="4C1F4D47F5275C0D501F81CFA69D7DB5" authorityName="KRISHNAI BOURGUIGNON &amp; ROISIN" authorityYear="2016" box="[806,947,1483,1504]" class="Insecta" family="Termitidae" genus="Humutermes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="6" pageNumber="21" phylum="Arthropoda" rank="genus">
<emphasis id="B96BEAD6F5275C0D501F81CFA69D7DB5" box="[806,947,1483,1504]" italics="true" pageId="6" pageNumber="21">Humutermes</emphasis>
</taxonomicName>
workers have an elongated mixed segment with a whitish, spherical mesenteric diverticulum. These features distinguish them from other South American soldierless termites, except for
<emphasis id="B96BEAD6F5275C0D578B8222A6437E0F" italics="true" pageId="6" pageNumber="21">
<taxonomicName id="4C1F4D47F5275C0D578B8222A0667E6E" authorityName="Bourguignon &amp; Scheffrahn &amp; Nagy &amp; Sonet &amp; Host &amp; Roisin" authorityYear="2016" box="[1202,1352,1574,1595]" class="Insecta" family="Termitidae" genus="Anoplotermes" kingdom="Animalia" order="Isoptera" pageId="6" pageNumber="21" phylum="Arthropoda" rank="genus" sensu="stricto">Anoplotermes</taxonomicName>
<taxonomicNameLabel id="A25857ADF5275C0D56698222A6437E0F" pageId="6" pageNumber="21" sensu="stricto">sensu stricto</taxonomicNameLabel>
</emphasis>
. The overall appearance of the workers is very similar to
<emphasis id="B96BEAD6F5275C0D50A68267A1FE7E2C" box="[927,1232,1635,1657]" italics="true" pageId="6" pageNumber="21">
<taxonomicName id="4C1F4D47F5275C0D50A68267A1177E2D" authorityName="Bourguignon &amp; Scheffrahn &amp; Nagy &amp; Sonet &amp; Host &amp; Roisin" authorityYear="2016" box="[927,1081,1635,1656]" class="Insecta" family="Termitidae" genus="Anoplotermes" kingdom="Animalia" order="Isoptera" pageId="6" pageNumber="21" phylum="Arthropoda" rank="genus" sensu="stricto">Anoplotermes</taxonomicName>
<taxonomicNameLabel id="A25857ADF5275C0D57788260A1FE7E2C" box="[1089,1232,1636,1657]" pageId="6" pageNumber="21" sensu="stricto">sensu stricto</taxonomicNameLabel>
</emphasis>
, with which the genus may form a monophyletic group, but the molecular phylogenetic tree is not sufficiently resolved to support this. The genus mainly differs from
<emphasis id="B96BEAD6F5275C0D501F82D9A1797EA6" box="[806,1111,1757,1779]" italics="true" pageId="6" pageNumber="21">
<taxonomicName id="4C1F4D47F5275C0D501F82D9A6EE7EA7" authorityName="Bourguignon &amp; Scheffrahn &amp; Nagy &amp; Sonet &amp; Host &amp; Roisin" authorityYear="2016" box="[806,960,1757,1778]" class="Insecta" family="Termitidae" genus="Anoplotermes" kingdom="Animalia" order="Isoptera" pageId="6" pageNumber="21" phylum="Arthropoda" rank="genus" sensu="stricto">Anoplotermes</taxonomicName>
<taxonomicNameLabel id="A25857ADF5275C0D50F182DAA1797EA6" box="[968,1111,1758,1779]" pageId="6" pageNumber="21" sensu="stricto">sensu stricto</taxonomicNameLabel>
</emphasis>
by the shape of its enteric valve, composed of six pads, each with ten to 20 long spines protruding into the paunch. Imagos differ from
<emphasis id="B96BEAD6F5275C0D501F833DA1787F1A" box="[806,1110,1849,1871]" italics="true" pageId="6" pageNumber="21">
<taxonomicName id="4C1F4D47F5275C0D501F833DA6EE7F1B" authorityName="Bourguignon &amp; Scheffrahn &amp; Nagy &amp; Sonet &amp; Host &amp; Roisin" authorityYear="2016" box="[806,960,1849,1870]" class="Insecta" family="Termitidae" genus="Anoplotermes" kingdom="Animalia" order="Isoptera" pageId="6" pageNumber="21" phylum="Arthropoda" rank="genus" sensu="stricto">Anoplotermes</taxonomicName>
<taxonomicNameLabel id="A25857ADF5275C0D50F1833EA1787F1A" box="[968,1110,1850,1871]" pageId="6" pageNumber="21" sensu="stricto">sensu stricto</taxonomicNameLabel>
</emphasis>
by their larger, more elongated fontanelle.
</paragraph>
<caption id="DF60664CF5265C0C539D84C1A76D78A3" pageId="7" pageNumber="22" startId="7.[164,228,197,216]">
<paragraph id="8BA036C4F5265C0C539D84C1A76D78A3" blockId="7.[164,1441,197,246]" pageId="7" pageNumber="22">
<emphasis id="B96BEAD6F5265C0C539D84C1A5D3788D" bold="true" box="[164,253,197,216]" pageId="7" pageNumber="22">Table 5.</emphasis>
Measurements (in mm) of 12 workers from six colonies, and six male and six female imagos from three colonies of
<taxonomicName id="4C1F4D47F5265C0C53CA84E7A4CF78A3" authority="Bourguignon &amp; Scheffrahn &amp; Nagy &amp; Sonet &amp; Host &amp; Roisin, 2016" authorityName="Bourguignon &amp; Scheffrahn &amp; Nagy &amp; Sonet &amp; Host &amp; Roisin" authorityYear="2016" box="[243,481,227,246]" class="Insecta" family="Termitidae" genus="Humutermes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="7" pageNumber="22" phylum="Arthropoda" rank="species" species="noiroti" status="sp. nov.">
<emphasis id="B96BEAD6F5265C0C53CA84E7A4CF78A3" bold="true" box="[243,481,227,246]" italics="true" pageId="7" pageNumber="22">Humutermes noiroti</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25857ADF5265C0C52D384E7A76D78A3" box="[490,579,227,246]" pageId="7" pageNumber="22" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="8BA036C4F5265C0C519B8518A01E7A4C" pageId="7" pageNumber="22">
<table id="F91FC464F526A3F4539D8518A08F7A4C" box="[164,1441,284,537]" gridcols="4" gridrows="8" pageId="7" pageNumber="22" rowsContinueFrom="6.[144,1421,1117,1370]" rowsContinueIn="9.[164,1441,1627,1880]">
<tr id="352F3486F526A3F4539D8518A08F7965" box="[164,1441,284,304]" gridrow="0" pageId="7" pageNumber="22" rowspan-0="1">
<th id="76FE5DFAF526A3F4519B8518A60E7965" box="[674,800,284,304]" gridcol="1" gridrow="0" pageId="7" pageNumber="22">Worker</th>
<th id="76FE5DFAF526A3F450DA8518A14E7965" box="[995,1120,284,304]" gridcol="2" gridrow="0" pageId="7" pageNumber="22">Imago ♂</th>
<th id="76FE5DFAF526A3F4561A8518A08F7965" box="[1315,1441,284,304]" gridcol="3" gridrow="0" pageId="7" pageNumber="22">Imago ♀</th>
</tr>
<tr id="352F3486F526A3F4539D8552A08F793F" box="[164,1441,342,362]" gridrow="1" pageId="7" pageNumber="22">
<th id="76FE5DFAF526A3F4539D8552A4F1793F" box="[164,479,342,362]" gridcol="0" gridrow="1" pageId="7" pageNumber="22">Head length to end of clypeus</th>
<td id="76FE5DFAF526A3F4519B8552A60E793F" box="[674,800,342,362]" gridcol="1" gridrow="1" pageId="7" pageNumber="22">0.5580.622</td>
<td id="76FE5DFAF526A3F450DA8552A14E793F" box="[995,1120,342,362]" gridcol="2" gridrow="1" pageId="7" pageNumber="22">0.5730.615</td>
<td id="76FE5DFAF526A3F4561A8552A08F793F" box="[1315,1441,342,362]" gridcol="3" gridrow="1" pageId="7" pageNumber="22">0.6050.649</td>
</tr>
<tr id="352F3486F526A3F4539D8570A08F79D2" box="[164,1441,372,391]" gridrow="2" pageId="7" pageNumber="22">
<th id="76FE5DFAF526A3F4539D8570A4F179D2" box="[164,479,372,391]" gridcol="0" gridrow="2" pageId="7" pageNumber="22">Head width</th>
<td id="76FE5DFAF526A3F4519B8570A60E79D2" box="[674,800,372,391]" gridcol="1" gridrow="2" pageId="7" pageNumber="22">0.5980.638</td>
<td id="76FE5DFAF526A3F450DA8570A14E79D2" box="[995,1120,372,391]" gridcol="2" gridrow="2" pageId="7" pageNumber="22">0.7180.742</td>
<td id="76FE5DFAF526A3F4561A8570A08F79D2" box="[1315,1441,372,391]" gridcol="3" gridrow="2" pageId="7" pageNumber="22">0.7120.778</td>
</tr>
<tr id="352F3486F526A3F4539D8595A08F79F1" box="[164,1441,401,420]" gridrow="3" pageId="7" pageNumber="22">
<th id="76FE5DFAF526A3F4539D8595A4F179F1" box="[164,479,401,420]" gridcol="0" gridrow="3" pageId="7" pageNumber="22">Eye diameter</th>
<td id="76FE5DFAF526A3F4519B8595A60E79F1" box="[674,800,401,420]" gridcol="1" gridrow="3" pageId="7" pageNumber="22"></td>
<td id="76FE5DFAF526A3F450DA8595A14E79F1" box="[995,1120,401,420]" gridcol="2" gridrow="3" pageId="7" pageNumber="22">0.1910.211</td>
<td id="76FE5DFAF526A3F4561A8595A08F79F1" box="[1315,1441,401,420]" gridcol="3" gridrow="3" pageId="7" pageNumber="22">0.2030.213</td>
</tr>
<tr id="352F3486F526A3F4539D85AAA08F7997" box="[164,1441,430,450]" gridrow="4" pageId="7" pageNumber="22">
<th id="76FE5DFAF526A3F4539D85AAA4F17997" box="[164,479,430,450]" gridcol="0" gridrow="4" pageId="7" pageNumber="22">Pronotum length</th>
<td id="76FE5DFAF526A3F4519B85AAA60E7997" box="[674,800,430,450]" gridcol="1" gridrow="4" pageId="7" pageNumber="22"></td>
<td id="76FE5DFAF526A3F450DA85AAA14E7997" box="[995,1120,430,450]" gridcol="2" gridrow="4" pageId="7" pageNumber="22">0.3700.426</td>
<td id="76FE5DFAF526A3F4561A85AAA08F7997" box="[1315,1441,430,450]" gridcol="3" gridrow="4" pageId="7" pageNumber="22">0.3800.416</td>
</tr>
<tr id="352F3486F526A3F4539D85CFA08F798A" box="[164,1441,459,479]" gridrow="5" pageId="7" pageNumber="22">
<th id="76FE5DFAF526A3F4539D85CFA4F1798A" box="[164,479,459,479]" gridcol="0" gridrow="5" pageId="7" pageNumber="22">Pronotum width</th>
<td id="76FE5DFAF526A3F4519B85CFA60E798A" box="[674,800,459,479]" gridcol="1" gridrow="5" pageId="7" pageNumber="22">0.3450.375</td>
<td id="76FE5DFAF526A3F450DA85CFA14E798A" box="[995,1120,459,479]" gridcol="2" gridrow="5" pageId="7" pageNumber="22">0.5540.617</td>
<td id="76FE5DFAF526A3F4561A85CFA08F798A" box="[1315,1441,459,479]" gridcol="3" gridrow="5" pageId="7" pageNumber="22">0.5660.609</td>
</tr>
<tr id="352F3486F526A3F4539D85EDA08F79A9" box="[164,1441,489,508]" gridrow="6" pageId="7" pageNumber="22">
<th id="76FE5DFAF526A3F4539D85EDA4F179A9" box="[164,479,489,508]" gridcol="0" gridrow="6" pageId="7" pageNumber="22">Hind tibia length</th>
<td id="76FE5DFAF526A3F4519B85EDA60E79A9" box="[674,800,489,508]" gridcol="1" gridrow="6" pageId="7" pageNumber="22">0.4510.536</td>
<td id="76FE5DFAF526A3F450DA85EDA14E79A9" box="[995,1120,489,508]" gridcol="2" gridrow="6" pageId="7" pageNumber="22">0.6760.713</td>
<td id="76FE5DFAF526A3F4561A85EDA08F79A9" box="[1315,1441,489,508]" gridcol="3" gridrow="6" pageId="7" pageNumber="22">0.6350.706</td>
</tr>
<tr id="352F3486F526A3F4539D8602A08F7A4C" box="[164,1441,518,537]" gridrow="7" pageId="7" pageNumber="22">
<th id="76FE5DFAF526A3F4539D8602A4F17A4C" box="[164,479,518,537]" gridcol="0" gridrow="7" pageId="7" pageNumber="22">Fore tibia width/length</th>
<td id="76FE5DFAF526A3F4519B8602A60E7A4C" box="[674,800,518,537]" gridcol="1" gridrow="7" pageId="7" pageNumber="22">0.2580.330</td>
<td id="76FE5DFAF526A3F450DA8602A14E7A4C" box="[995,1120,518,537]" gridcol="2" gridrow="7" pageId="7" pageNumber="22"></td>
<td id="76FE5DFAF526A3F4561A8602A08F7A4C" box="[1315,1441,518,537]" gridcol="3" gridrow="7" pageId="7" pageNumber="22"></td>
</tr>
</table>
</paragraph>
<paragraph id="8BA036C4F5265C0C539D866CA4357A28" blockId="7.[164,778,616,699]" box="[164,283,616,637]" pageId="7" pageNumber="22">
<emphasis id="B96BEAD6F5265C0C539D866CA4357A28" box="[164,283,616,637]" italics="true" pageId="7" pageNumber="22">Etymology</emphasis>
</paragraph>
<paragraph id="8BA036C4F5265C0C539D8683A5D47AEE" blockId="7.[164,778,616,699]" pageId="7" pageNumber="22">
We named this genus
<taxonomicName id="4C1F4D47F5265C0C529A8683A71D7AC9" authorityName="KRISHNAI BOURGUIGNON &amp; ROISIN" authorityYear="2016" box="[419,563,647,668]" class="Insecta" family="Termitidae" genus="Humutermes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="7" pageNumber="22" phylum="Arthropoda" rank="genus">
<emphasis id="B96BEAD6F5265C0C529A8683A71D7AC9" box="[419,563,647,668]" italics="true" pageId="7" pageNumber="22">Humutermes</emphasis>
</taxonomicName>
to refer to its diet, humus.
</paragraph>
</subSubSection>
</treatment>
</document>