312 lines
35 KiB
XML
312 lines
35 KiB
XML
<document id="A00060DBDF69E5B319895F43C3B30DBE" ID-CLB-Dataset="225750" ID-DOI="10.11646/zootaxa.5227.4.7" ID-GBIF-Dataset="d3d2e01b-fa8f-49cc-a911-6742ef77aaa2" ID-ISSN="1175-5326" ID-Zenodo-Dep="7518982" ID-ZooBank="97C5B5A7-6A62-45FB-BA5A-A8B761D6A01C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1673312564086" checkinUser="plazi" docAuthor="Bollino, Maurizio" docDate="2023" docId="03B687BCDB5FFFDDFF59FBC0FFF78F30" docLanguage="en" docName="zootaxa.5227.4.7.pdf" docOrigin="Zootaxa 5227 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pachyrhynchus viridis" docType="treatment" docVersion="7" lastPageNumber="500" masterDocId="FF8FFFC4DB5CFFD8FFCEFFF3FD3A8E55" masterDocTitle="About some Indonesian Pachyrhynchus Germar 1824 with description of a new species (Coleoptera, Curculionidae, Entiminae, Pachyrhynchini)" masterLastPageNumber="500" masterPageNumber="495" pageNumber="498" updateTime="1699473680678" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="ADF5A6605C550170A2CB1A93395C6528">About some Indonesian Pachyrhynchus Germar 1824 with description of a new species (Coleoptera, Curculionidae, Entiminae, Pachyrhynchini)</mods:title>
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<mods:namePart id="EC3F6E5C896DD9009D83E45F8FEDB3C5">Bollino, Maurizio</mods:namePart>
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<mods:title id="47DC6738D95F438745EC01D73C85269E">Zootaxa</mods:title>
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<mods:date id="C9F5956D05666945FB9B959F0849A1C3">2023</mods:date>
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<mods:number id="6F9FB129F871C6B56DDFB8D7C7C3A88C">2023-01-09</mods:number>
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<treatment id="03B687BCDB5FFFDDFF59FBC0FFF78F30" ID-DOI="http://doi.org/10.5281/zenodo.7540312" ID-GBIF-Taxon="208973352" ID-Zenodo-Dep="7540312" LSID="urn:lsid:plazi:treatment:03B687BCDB5FFFDDFF59FBC0FFF78F30" httpUri="http://treatment.plazi.org/id/03B687BCDB5FFFDDFF59FBC0FFF78F30" lastPageId="5" lastPageNumber="500" pageId="3" pageNumber="498">
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<subSubSection id="C3056521DB5FFFDBFF59FBC0FFBD8A1B" box="[151,647,1075,1102]" pageId="3" pageNumber="498" type="nomenclature">
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<paragraph id="8BA036AADB5FFFDBFF59FBC0FFBD8A1B" blockId="3.[151,647,1075,1138]" box="[151,647,1075,1102]" pageId="3" pageNumber="498">
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<heading id="D0E881C6DB5FFFDBFF59FBC0FFBD8A1B" bold="true" box="[151,647,1075,1102]" fontSize="11" level="1" pageId="3" pageNumber="498" reason="1">
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<taxonomicName id="4C1F4D29DB5FFFDBFF59FBC0FFBD8A1B" ID-CoL="75HR5" authority="(Chevrolat, 1879)" baseAuthorityName="Chevrolat" baseAuthorityYear="1879" box="[151,647,1075,1102]" class="Insecta" family="Curculionidae" genus="Pachyrhynchus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="498" phylum="Arthropoda" rank="species" species="viridis">
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<emphasis id="B96BEAB8DB5FFFDBFF59FBC0FFBD8A1B" bold="true" box="[151,647,1075,1102]" pageId="3" pageNumber="498">
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<emphasis id="B96BEAB8DB5FFFDBFF59FBC0FC9D8A1B" bold="true" box="[151,423,1075,1102]" italics="true" pageId="3" pageNumber="498">Pachyrhynchus viridis</emphasis>
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(Chevrolat, 1879)
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3056521DB5FFFDBFF59FBABFC0C8A27" box="[151,310,1112,1138]" pageId="3" pageNumber="498" type="description">
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<paragraph id="8BA036AADB5FFFDBFF59FBABFC0C8A27" blockId="3.[151,647,1075,1138]" box="[151,310,1112,1138]" pageId="3" pageNumber="498">
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(
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<figureCitation id="13242A2FDB5FFFDBFF51FBABFC178A27" box="[159,301,1112,1138]" captionStart="FIGURE 2" captionStartId="4.[151,250,1887,1911]" captionTargetBox="[211,1385,196,1862]" captionTargetId="figure-31@4.[202,1386,181,1863]" captionTargetPageId="4" captionText="FIGURE 2. Pachyrhynchus species, dorsal habitus. A: Pachyrhynchus forsteni male (Morotai); B: Pachyrhynchus viridis (West Papua). Pachyrhynchus species, endophallus in lateral view. C: Pachyrhynchus forsteni (Halmahera); D: Pachyrhynchus morotaiensis (Morotai); E: Pachyrhynchus viridis (West Papua); F: Pantorhytes biplagiatus (Papua New Guinea). Red arrow indicates the dorsal median diverticulum in Pachyrhynchus, and the ventral median one in Pantorhytes." figureDoi="http://doi.org/10.5281/zenodo.7518986" httpUri="https://zenodo.org/record/7518986/files/figure.png" pageId="3" pageNumber="498">Figs. 2B, 2E</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C3056521DB5FFFDDFF59FB53FFF78F30" lastPageId="5" lastPageNumber="500" pageId="3" pageNumber="498" type="discussion">
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<paragraph id="8BA036AADB5FFFDBFF59FB53FC1A8B1F" blockId="3.[151,1437,1184,2039]" pageId="3" pageNumber="498">
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The species was originally placed in the genus
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<taxonomicName id="4C1F4D29DB5FFFDBFD78FB53FEEF8AEF" authority="Erichson 1834" authorityName="Erichson" authorityYear="1834" box="[694,981,1184,1210]" class="Insecta" family="Curculionidae" genus="Apocyrtus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="498" phylum="Arthropoda" rank="genus">
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<emphasis id="B96BEAB8DB5FFFDBFD78FB53FE128AEF" box="[694,808,1184,1210]" italics="true" pageId="3" pageNumber="498">Apocyrtus</emphasis>
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Erichson 1834
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</taxonomicName>
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by Chevrolat (1879), with type locality “Nova-Guinea (Dorey) [= Manokwari]”. Later, Dalla Torre
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<emphasis id="B96BEAB8DB5FFFDBFC9BFB36FEB18A8B" box="[853,907,1220,1246]" italics="true" pageId="3" pageNumber="498">et al</emphasis>
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. (1931) treated it as a member of the genus
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<taxonomicName id="4C1F4D29DB5FFFDBFF59FB1BFCCA8B57" authority="Behrens 1887" authorityName="Behrens" authorityYear="1887" box="[151,496,1256,1282]" class="Insecta" family="Curculionidae" genus="Sphenomorpha" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="498" phylum="Arthropoda" rank="genus">
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<emphasis id="B96BEAB8DB5FFFDBFF59FB1BFC7B8B57" box="[151,321,1256,1282]" italics="true" pageId="3" pageNumber="498">Sphenomorpha</emphasis>
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Behrens 1887
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</taxonomicName>
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, a treatment followed by Setliff (2007).
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<bibRefCitation id="EF8E4B5BDB5FFFDBFC18FB1BF9A48B57" author="Yoshitake, H." box="[982,1182,1256,1282]" pageId="3" pageNumber="498" pagination="241 - 246" refId="ref2703" refString="Yoshitake, H. (2017) Notes on Pachyrhynchus Jewel Weevils (Coleoptera, Curculionidae, Entiminae) from the Papuan Region. Elytra, Tokyo, New Series, 7 (1), 241 - 246." type="journal article" year="2017">Yoshitake (2017)</bibRefCitation>
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, after examination of the
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<typeStatus id="54A48808DB5FFFDBFF08FAFFFC128B73" box="[198,296,1292,1318]" pageId="3" pageNumber="498" type="holotype">holotype</typeStatus>
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in Naturhistoriska riksmuseet, Stockholm, proposed the new combination
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<taxonomicName id="4C1F4D29DB5FFFDBFB52FAFFFC268B1F" authority="(Chevrolat)" baseAuthorityName="Chevrolat" baseAuthorityYear="1879" class="Insecta" family="Curculionidae" genus="Pachyrhynchus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="498" phylum="Arthropoda" rank="species" species="viridis">
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<emphasis id="B96BEAB8DB5FFFDBFB52FAFFF8A78B73" box="[1180,1437,1292,1318]" italics="true" pageId="3" pageNumber="498">Pachyrhynchus viridis</emphasis>
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(Chevrolat)
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</taxonomicName>
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.
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</paragraph>
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<paragraph id="8BA036AADB5FFFDBFF09FAA7F8A68BAA" blockId="3.[151,1437,1184,2039]" pageId="3" pageNumber="498">
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Recently I obtained a single male of
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<taxonomicName id="4C1F4D29DB5FFFDBFD96FAA7FE3E8B3B" authorityName="Germar" authorityYear="1824" box="[600,772,1364,1390]" class="Insecta" family="Curculionidae" genus="Pachyrhynchus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="498" phylum="Arthropoda" rank="genus">
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<emphasis id="B96BEAB8DB5FFFDBFD96FAA7FE3E8B3B" box="[600,772,1364,1390]" italics="true" pageId="3" pageNumber="498">Pachyrhynchus</emphasis>
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</taxonomicName>
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from Neney village (Ransiki District, Manokwari Province,
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<collectingRegion id="49DBF848DB5FFFDBFF59FA8AFC218BC7" box="[151,283,1400,1427]" country="Indonesia" name="Papua Barat" pageId="3" pageNumber="498">West Papua</collectingRegion>
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) which matches the diagnosis of
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<taxonomicName id="4C1F4D29DB5FFFDBFD5EFA8AFFCD8BC7" baseAuthorityName="Chevrolat" baseAuthorityYear="1879" box="[656,759,1400,1427]" class="Insecta" family="Curculionidae" genus="Pachyrhynchus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="498" phylum="Arthropoda" rank="species" species="viridis">
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<emphasis id="B96BEAB8DB5FFFDBFD5EFA8AFF9B8BC7" box="[656,673,1401,1426]" italics="true" pageId="3" pageNumber="498">P</emphasis>
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.
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<emphasis id="B96BEAB8DB5FFFDBFD61FA8BFFCD8BC7" box="[687,759,1400,1426]" italics="true" pageId="3" pageNumber="498">viridis</emphasis>
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</taxonomicName>
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provided by
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<bibRefCitation id="EF8E4B5BDB5FFFDBFC44FA8BF9778BC7" author="Yoshitake, H." box="[906,1101,1400,1426]" pageId="3" pageNumber="498" pagination="241 - 246" refId="ref2703" refString="Yoshitake, H. (2017) Notes on Pachyrhynchus Jewel Weevils (Coleoptera, Curculionidae, Entiminae) from the Papuan Region. Elytra, Tokyo, New Series, 7 (1), 241 - 246." type="journal article" year="2017">Yoshitake (2017)</bibRefCitation>
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, but the color of the tegument is shiny black. In the same paper, Yoshitake described
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<taxonomicName id="4C1F4D29DB5FFFDBFD23FA6FF9208BE3" authorityName="Yoshitake" authorityYear="2017" box="[749,1050,1436,1462]" class="Insecta" family="Curculionidae" genus="Pachyrhynchus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="498" phylum="Arthropoda" rank="species" species="ohbayashii">
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<emphasis id="B96BEAB8DB5FFFDBFD23FA6FF9208BE3" box="[749,1050,1436,1462]" italics="true" pageId="3" pageNumber="498">Pachyrhynchus ohbayashii</emphasis>
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</taxonomicName>
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from Biak Island. Comparing both the diagnosis and images of the two species provided by Yoshitake, I have not been able to attribute my specimen with certainty to one or the other species. Based on the simple topological data I consider it
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<taxonomicName id="4C1F4D29DB5FFFDBFB55FA17F8AC8BAB" baseAuthorityName="Chevrolat" baseAuthorityYear="1879" box="[1179,1430,1508,1534]" class="Insecta" family="Curculionidae" genus="Pachyrhynchus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="498" phylum="Arthropoda" rank="species" species="viridis">
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<emphasis id="B96BEAB8DB5FFFDBFB55FA17F8AC8BAB" box="[1179,1430,1508,1534]" italics="true" pageId="3" pageNumber="498">Pachyrhynchus viridis</emphasis>
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</taxonomicName>
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.
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</paragraph>
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<paragraph id="8BA036AADB5FFFDBFF09F9FBFE028882" blockId="3.[151,1437,1184,2039]" pageId="3" pageNumber="498">
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Taking advantage of the availability of specimens of four of the five Indonesian species of
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<taxonomicName id="4C1F4D29DB5FFFDBFB3EF9FBF8A68877" authorityName="Germar" authorityYear="1824" box="[1264,1436,1544,1570]" class="Insecta" family="Curculionidae" genus="Pachyrhynchus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="498" phylum="Arthropoda" rank="genus">
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<emphasis id="B96BEAB8DB5FFFDBFB3EF9FBF8A68877" box="[1264,1436,1544,1570]" italics="true" pageId="3" pageNumber="498">Pachyrhynchus</emphasis>
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</taxonomicName>
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most phenetically similar to each other, I also wanted to compare the morphology of the male genitalia, as well as attempt the evertion of the endophallus. As already pointed out (Bollino & Sandel, 2017;
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<bibRefCitation id="EF8E4B5BDB5FFFDBFB46F9A3F85A883F" author="Cabras, A. A. & Bollino, M. & Medina, M. N." box="[1160,1376,1616,1643]" pageId="3" pageNumber="498" pagination="39 - 46" refId="ref2640" refString="Cabras, A. A., Bollino, M. & Medina, M. N. (2018) A new species of the subgenus Orthocyrtus, genus Metapocyrtus (Coleoptera, Curculionidae, Entiminae, Pachyrhynchini) from Mindanao, with notes on its ecology. Baltic Journal of Coleopterology, 18 (1), 39 - 46." type="journal article" year="2018">
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Cabras
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<emphasis id="B96BEAB8DB5FFFDBFB13F9A2F835883F" box="[1245,1295,1616,1642]" italics="true" pageId="3" pageNumber="498">et al</emphasis>
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., 2018
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</bibRefCitation>
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), this latter procedure is not simple, and usually requires several attempts with different specimens. I was able to obtain a complete evertion only in
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<taxonomicName id="4C1F4D29DB5FFFDBFE1BF96AFFBF88E7" authorityName="Snellen van Vollenhoven" authorityYear="1864" box="[469,645,1689,1715]" class="Insecta" family="Curculionidae" genus="Pachyrhynchus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="498" phylum="Arthropoda" rank="species" species="morotaiensis">
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<emphasis id="B96BEAB8DB5FFFDBFE1BF96AFCDC88E7" box="[469,486,1689,1714]" italics="true" pageId="3" pageNumber="498">P</emphasis>
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.
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<emphasis id="B96BEAB8DB5FFFDBFE3BF96AFFBF88E7" box="[501,645,1689,1714]" italics="true" pageId="3" pageNumber="498">morotaiensis</emphasis>
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</taxonomicName>
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(
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<figureCitation id="13242A2FDB5FFFDBFD5BF96BFFD788E7" box="[661,749,1688,1714]" captionStart="FIGURE 2" captionStartId="4.[151,250,1887,1911]" captionTargetBox="[211,1385,196,1862]" captionTargetId="figure-31@4.[202,1386,181,1863]" captionTargetPageId="4" captionText="FIGURE 2. Pachyrhynchus species, dorsal habitus. A: Pachyrhynchus forsteni male (Morotai); B: Pachyrhynchus viridis (West Papua). Pachyrhynchus species, endophallus in lateral view. C: Pachyrhynchus forsteni (Halmahera); D: Pachyrhynchus morotaiensis (Morotai); E: Pachyrhynchus viridis (West Papua); F: Pantorhytes biplagiatus (Papua New Guinea). Red arrow indicates the dorsal median diverticulum in Pachyrhynchus, and the ventral median one in Pantorhytes." figureDoi="http://doi.org/10.5281/zenodo.7518986" httpUri="https://zenodo.org/record/7518986/files/figure.png" pageId="3" pageNumber="498">Fig. 2D</figureCitation>
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) and
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<taxonomicName id="4C1F4D29DB5FFFDBFCE0F96AFEA888E7" baseAuthorityName="Chevrolat" baseAuthorityYear="1879" box="[814,914,1688,1714]" class="Insecta" family="Curculionidae" genus="Pachyrhynchus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="498" phylum="Arthropoda" rank="species" species="viridis">
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<emphasis id="B96BEAB8DB5FFFDBFCE0F96AFEA888E7" box="[814,914,1688,1714]" italics="true" pageId="3" pageNumber="498">P. viridis</emphasis>
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</taxonomicName>
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(
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<figureCitation id="13242A2FDB5FFFDBFC6CF96BFEC388E7" box="[930,1017,1688,1714]" captionStart="FIGURE 2" captionStartId="4.[151,250,1887,1911]" captionTargetBox="[211,1385,196,1862]" captionTargetId="figure-31@4.[202,1386,181,1863]" captionTargetPageId="4" captionText="FIGURE 2. Pachyrhynchus species, dorsal habitus. A: Pachyrhynchus forsteni male (Morotai); B: Pachyrhynchus viridis (West Papua). Pachyrhynchus species, endophallus in lateral view. C: Pachyrhynchus forsteni (Halmahera); D: Pachyrhynchus morotaiensis (Morotai); E: Pachyrhynchus viridis (West Papua); F: Pantorhytes biplagiatus (Papua New Guinea). Red arrow indicates the dorsal median diverticulum in Pachyrhynchus, and the ventral median one in Pantorhytes." figureDoi="http://doi.org/10.5281/zenodo.7518986" httpUri="https://zenodo.org/record/7518986/files/figure.png" pageId="3" pageNumber="498">Fig. 2E</figureCitation>
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), and a partial eversion in
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<taxonomicName id="4C1F4D29DB5FFFDBFAE9F96AF8A788E7" box="[1319,1437,1688,1715]" class="Insecta" family="Curculionidae" genus="Pachyrhynchus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="498" phylum="Arthropoda" rank="species" species="forsteni">
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<emphasis id="B96BEAB8DB5FFFDBFAE9F96AF80288E7" box="[1319,1336,1689,1714]" italics="true" pageId="3" pageNumber="498">P</emphasis>
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.
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<emphasis id="B96BEAB8DB5FFFDBFA89F96BF8A788E7" box="[1351,1437,1688,1714]" italics="true" pageId="3" pageNumber="498">forsteni</emphasis>
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</taxonomicName>
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(
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<figureCitation id="13242A2FDB5FFFDBFF51F94FFDC28883" box="[159,248,1724,1750]" captionStart="FIGURE 2" captionStartId="4.[151,250,1887,1911]" captionTargetBox="[211,1385,196,1862]" captionTargetId="figure-31@4.[202,1386,181,1863]" captionTargetPageId="4" captionText="FIGURE 2. Pachyrhynchus species, dorsal habitus. A: Pachyrhynchus forsteni male (Morotai); B: Pachyrhynchus viridis (West Papua). Pachyrhynchus species, endophallus in lateral view. C: Pachyrhynchus forsteni (Halmahera); D: Pachyrhynchus morotaiensis (Morotai); E: Pachyrhynchus viridis (West Papua); F: Pantorhytes biplagiatus (Papua New Guinea). Red arrow indicates the dorsal median diverticulum in Pachyrhynchus, and the ventral median one in Pantorhytes." figureDoi="http://doi.org/10.5281/zenodo.7518986" httpUri="https://zenodo.org/record/7518986/files/figure.png" pageId="3" pageNumber="498">Fig. 2C</figureCitation>
|
|
), while I have not obtained any results in
|
|
<taxonomicName id="4C1F4D29DB5FFFDBFD02F94EFE0B8883" authorityName="Bollino" authorityYear="2023" box="[716,817,1724,1751]" class="Insecta" family="Curculionidae" genus="Pachyrhynchus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="498" phylum="Arthropoda" rank="species" species="faisali">
|
|
<emphasis id="B96BEAB8DB5FFFDBFD02F94EFFE78883" box="[716,733,1725,1750]" italics="true" pageId="3" pageNumber="498">P</emphasis>
|
|
.
|
|
<emphasis id="B96BEAB8DB5FFFDBFD25F94FFE0B8883" box="[747,817,1724,1750]" italics="true" pageId="3" pageNumber="498">faisali</emphasis>
|
|
</taxonomicName>
|
|
.
|
|
</paragraph>
|
|
<paragraph id="8BA036AADB5FFFDBFF09F913F8A689A2" blockId="3.[151,1437,1184,2039]" pageId="3" pageNumber="498">
|
|
The morphology of the aedeagus and/or endophallus of the genus
|
|
<taxonomicName id="4C1F4D29DB5FFFDBFC0BF913F94B88AF" authorityName="Germar" authorityYear="1824" box="[965,1137,1760,1786]" class="Insecta" family="Curculionidae" genus="Pachyrhynchus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="498" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="B96BEAB8DB5FFFDBFC0BF913F94B88AF" box="[965,1137,1760,1786]" italics="true" pageId="3" pageNumber="498">Pachyrhynchus</emphasis>
|
|
</taxonomicName>
|
|
has sometimes been used to propose phyletic relationships between species (Bollino
|
|
<emphasis id="B96BEAB8DB5FFFDBFCFCF8F6FE5E894B" box="[818,868,1796,1822]" italics="true" pageId="3" pageNumber="498">et al</emphasis>
|
|
., 2017; Cabras
|
|
<emphasis id="B96BEAB8DB5FFFDBFBD6F8F6F970894B" box="[1048,1098,1796,1822]" italics="true" pageId="3" pageNumber="498">et al</emphasis>
|
|
., 2017;
|
|
<bibRefCitation id="EF8E4B5BDB5FFFDBFB69F8F7F87D894B" author="Bollino, M." box="[1191,1351,1796,1822]" pageId="3" pageNumber="498" pagination="504 - 516" refId="ref2479" refString="Bollino, M. (2022) Three new species of Pachyrhynchus Germar 1824, (Coleoptera, Curculionidae, Entiminae) from Mindanao Island, The Philippines, with taxonomical notes. Zootaxa, 5116 (4), 504 - 516. https: // doi. org / 10.11646 / zootaxa. 5116.4.2" type="journal article" year="2022">Bollino, 2022</bibRefCitation>
|
|
). In the species treated here and on the basis of the morphology of the endophallus, it would seem that
|
|
<taxonomicName id="4C1F4D29DB5FFFDBFB59F8DAF83C8917" box="[1175,1286,1832,1858]" class="Insecta" family="Curculionidae" genus="Pachyrhynchus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="498" phylum="Arthropoda" rank="species" species="forsteni">
|
|
<emphasis id="B96BEAB8DB5FFFDBFB59F8DAF83C8917" box="[1175,1286,1832,1858]" italics="true" pageId="3" pageNumber="498">P. forsteni</emphasis>
|
|
</taxonomicName>
|
|
and
|
|
<taxonomicName id="4C1F4D29DB5FFFDBFAF6F8DAF8A78917" baseAuthorityName="Chevrolat" baseAuthorityYear="1879" box="[1336,1437,1832,1859]" class="Insecta" family="Curculionidae" genus="Pachyrhynchus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="498" phylum="Arthropoda" rank="species" species="viridis">
|
|
<emphasis id="B96BEAB8DB5FFFDBFAF6F8DAF8738917" box="[1336,1353,1833,1858]" italics="true" pageId="3" pageNumber="498">P</emphasis>
|
|
.
|
|
<emphasis id="B96BEAB8DB5FFFDBFA9BF8DBF8A78917" box="[1365,1437,1832,1858]" italics="true" pageId="3" pageNumber="498">viridis</emphasis>
|
|
</taxonomicName>
|
|
are more differentiated from the other species of the genus
|
|
<taxonomicName id="4C1F4D29DB5FFFDBFCF6F8BFFEDE8933" authorityName="Germar" authorityYear="1824" box="[824,996,1868,1894]" class="Insecta" family="Curculionidae" genus="Pachyrhynchus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="498" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="B96BEAB8DB5FFFDBFCF6F8BFFEDE8933" box="[824,996,1868,1894]" italics="true" pageId="3" pageNumber="498">Pachyrhynchus</emphasis>
|
|
</taxonomicName>
|
|
as no other taxon has the long median dorsal diverticulum found in the two Indonesian species (see red arrows in figs. 2C and 2E). Something similar is also present in some
|
|
<taxonomicName id="4C1F4D29DB5FFFDBFE48F867FF0C89FB" authorityName="Schoenherr" authorityYear="1826" box="[390,566,1940,1966]" class="Insecta" family="Curculionidae" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="498" phylum="Arthropoda" rank="tribe" tribe="Pachyrhynchini">Pachyrhynchini</taxonomicName>
|
|
belonging to the genus
|
|
<taxonomicName id="4C1F4D29DB5FFFDBFC88F867F96E89FB" authority="Faust 1892" authorityName="Faust" authorityYear="1892" box="[838,1108,1940,1967]" class="Insecta" family="Curculionidae" genus="Pantorhytes" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="498" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="B96BEAB8DB5FFFDBFC88F867FEF789FB" box="[838,973,1940,1966]" italics="true" pageId="3" pageNumber="498">Pantorhytes</emphasis>
|
|
Faust 1892
|
|
</taxonomicName>
|
|
, even if the general structure of the endophallus of this genus is simpler and the smaller median diverticulum is ventral (fig. 2F).
|
|
<taxonomicName id="4C1F4D29DB5FFFDBFB3EF84BFC1D89A3" authorityName="Snellen van Vollenhoven" authorityYear="1864" class="Insecta" family="Curculionidae" genus="Pachyrhynchus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="498" phylum="Arthropoda" rank="species" species="morotaiensis">
|
|
<emphasis id="B96BEAB8DB5FFFDBFB3EF84BFC1D89A3" italics="true" pageId="3" pageNumber="498">Pachyrhynchus morotaiensis</emphasis>
|
|
</taxonomicName>
|
|
,on the contrary, seems in this respect more similar to the rest of the members of the genus
|
|
<taxonomicName id="4C1F4D29DB5FFFDBFB27F82FF8AF89A3" authorityName="Germar" authorityYear="1824" box="[1257,1429,2012,2038]" class="Insecta" family="Curculionidae" genus="Pachyrhynchus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="498" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="B96BEAB8DB5FFFDBFB27F82FF8AF89A3" box="[1257,1429,2012,2038]" italics="true" pageId="3" pageNumber="498">Pachyrhynchus</emphasis>
|
|
</taxonomicName>
|
|
.
|
|
</paragraph>
|
|
<caption id="DF606622DB58FFDCFF59F8ACF98A89B6" ID-DOI="http://doi.org/10.5281/zenodo.7518986" ID-Zenodo-Dep="7518986" httpUri="https://zenodo.org/record/7518986/files/figure.png" pageId="4" pageNumber="499" startId="4.[151,250,1887,1911]" targetBox="[211,1385,196,1862]" targetPageId="4">
|
|
<paragraph id="8BA036AADB58FFDCFF59F8ACF98A89B6" blockId="4.[151,1437,1887,2019]" pageId="4" pageNumber="499">
|
|
<emphasis id="B96BEAB8DB58FFDCFF59F8ACFC2D8922" bold="true" box="[151,279,1887,1911]" pageId="4" pageNumber="499">FIGURE 2.</emphasis>
|
|
<taxonomicName id="4C1F4D29DB58FFDCFEEFF8ACFF298922" box="[289,531,1887,1911]" pageId="4" pageNumber="499">
|
|
<emphasis id="B96BEAB8DB58FFDCFEEFF8ACFC878922" box="[289,445,1887,1911]" italics="true" pageId="4" pageNumber="499">Pachyrhynchus</emphasis>
|
|
species
|
|
</taxonomicName>
|
|
, dorsal habitus. A:
|
|
<taxonomicName id="4C1F4D29DB58FFDCFD28F8ACFEE38922" box="[742,985,1887,1911]" class="Insecta" family="Curculionidae" genus="Pachyrhynchus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="499" phylum="Arthropoda" rank="species" species="forsteni">
|
|
<emphasis id="B96BEAB8DB58FFDCFD28F8ACFEE38922" box="[742,985,1887,1911]" italics="true" pageId="4" pageNumber="499">Pachyrhynchus forsteni</emphasis>
|
|
</taxonomicName>
|
|
male (Morotai); B:
|
|
<taxonomicName id="4C1F4D29DB58FFDCFB7BF8ACF8A68922" baseAuthorityName="Chevrolat" baseAuthorityYear="1879" box="[1205,1436,1887,1911]" class="Insecta" family="Curculionidae" genus="Pachyrhynchus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="499" phylum="Arthropoda" rank="species" species="viridis">
|
|
<emphasis id="B96BEAB8DB58FFDCFB7BF8ACF8A68922" box="[1205,1436,1887,1911]" italics="true" pageId="4" pageNumber="499">Pachyrhynchus viridis</emphasis>
|
|
</taxonomicName>
|
|
(West Papua).
|
|
<taxonomicName id="4C1F4D29DB58FFDCFEE5F870FF2389CE" box="[299,537,1923,1947]" pageId="4" pageNumber="499">
|
|
<emphasis id="B96BEAB8DB58FFDCFEE5F870FCFD89CE" box="[299,455,1923,1947]" italics="true" pageId="4" pageNumber="499">Pachyrhynchus</emphasis>
|
|
species
|
|
</taxonomicName>
|
|
, endophallus in lateral view. C:
|
|
<taxonomicName id="4C1F4D29DB58FFDCFC90F870F9ED89CE" authority="(Halmahera)" baseAuthorityName="Halmahera" box="[862,1239,1923,1947]" class="Insecta" family="Curculionidae" genus="Pachyrhynchus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="499" phylum="Arthropoda" rank="species" species="forsteni">
|
|
<emphasis id="B96BEAB8DB58FFDCFC90F870F97789CE" box="[862,1101,1923,1947]" italics="true" pageId="4" pageNumber="499">Pachyrhynchus forsteni</emphasis>
|
|
(Halmahera)
|
|
</taxonomicName>
|
|
; D:
|
|
<taxonomicName id="4C1F4D29DB58FFDCFACFF870FCBC89EA" authority="(Morotai)" baseAuthorityName="Morotai" class="Insecta" family="Curculionidae" genus="Pachyrhynchus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="499" phylum="Arthropoda" rank="species" species="morotaiensis">
|
|
<emphasis id="B96BEAB8DB58FFDCFACFF870FC2389EA" italics="true" pageId="4" pageNumber="499">Pachyrhynchus morotaiensis</emphasis>
|
|
(Morotai)
|
|
</taxonomicName>
|
|
; E:
|
|
<taxonomicName id="4C1F4D29DB58FFDCFE7EF854FFAF89EA" baseAuthorityName="Chevrolat" baseAuthorityYear="1879" box="[432,661,1959,1983]" class="Insecta" family="Curculionidae" genus="Pachyrhynchus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="499" phylum="Arthropoda" rank="species" species="viridis">
|
|
<emphasis id="B96BEAB8DB58FFDCFE7EF854FFAF89EA" box="[432,661,1959,1983]" italics="true" pageId="4" pageNumber="499">Pachyrhynchus viridis</emphasis>
|
|
</taxonomicName>
|
|
(West Papua); F:
|
|
<taxonomicName id="4C1F4D29DB58FFDCFC9EF854F97E89EA" authorityName="Guerin-Meneville" authorityYear="1841" box="[848,1092,1959,1983]" class="Insecta" family="Curculionidae" genus="Pantorhytes" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="499" phylum="Arthropoda" rank="species" species="biplagiatus">
|
|
<emphasis id="B96BEAB8DB58FFDCFC9EF854F97E89EA" box="[848,1092,1959,1983]" italics="true" pageId="4" pageNumber="499">Pantorhytes biplagiatus</emphasis>
|
|
</taxonomicName>
|
|
(Papua New Guinea). Red arrow indicates the dorsal median diverticulum in
|
|
<taxonomicName id="4C1F4D29DB58FFDCFD9AF838FFCA89B6" authorityName="Germar" authorityYear="1824" box="[596,752,1995,2019]" class="Insecta" family="Curculionidae" genus="Pachyrhynchus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="499" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="B96BEAB8DB58FFDCFD9AF838FFCA89B6" box="[596,752,1995,2019]" italics="true" pageId="4" pageNumber="499">Pachyrhynchus</emphasis>
|
|
</taxonomicName>
|
|
, and the ventral median one in
|
|
<taxonomicName id="4C1F4D29DB58FFDCFBE1F838F99689B6" authorityName="Faust" authorityYear="1892" box="[1071,1196,1995,2019]" class="Insecta" family="Curculionidae" genus="Pantorhytes" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="499" phylum="Arthropoda" rank="genus">Pantorhytes</taxonomicName>
|
|
.
|
|
</paragraph>
|
|
</caption>
|
|
<paragraph id="8BA036AADB59FFDDFF09FF64FFF78F30" blockId="5.[151,1437,151,357]" pageId="5" pageNumber="500">
|
|
As regards the morphology of the aedeagus, the only relevant data is the shape of the apex, rounded in
|
|
<taxonomicName id="4C1F4D29DB59FFDDFA4AFF64FDD78E80" class="Insecta" family="Curculionidae" genus="Pachyrhynchus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="500" phylum="Arthropoda" rank="species" species="forsteni">
|
|
<emphasis id="B96BEAB8DB59FFDDFA4AFF64F8AF8EE5" box="[1412,1429,151,176]" italics="true" pageId="5" pageNumber="500">P</emphasis>
|
|
.
|
|
<emphasis id="B96BEAB8DB59FFDDFF59FF48FDD78E80" box="[151,237,187,213]" italics="true" pageId="5" pageNumber="500">forsteni</emphasis>
|
|
</taxonomicName>
|
|
,
|
|
<taxonomicName id="4C1F4D29DB59FFDDFF30FF48FC8B8E81" box="[254,433,187,213]" class="Insecta" family="Curculionidae" genus="Pachyrhynchus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="500" phylum="Arthropoda" rank="species" species="morotaiensis">
|
|
<emphasis id="B96BEAB8DB59FFDDFF30FF48FC358E81" box="[254,271,187,212]" italics="true" pageId="5" pageNumber="500">P</emphasis>
|
|
.
|
|
<emphasis id="B96BEAB8DB59FFDDFEEFFF48FC8B8E81" box="[289,433,187,212]" italics="true" pageId="5" pageNumber="500">morotaiensis</emphasis>
|
|
</taxonomicName>
|
|
and
|
|
<taxonomicName id="4C1F4D29DB59FFDDFE21FF48FF6D8E80" authorityName="Bollino" authorityYear="2023" box="[495,599,187,213]" class="Insecta" family="Curculionidae" genus="Pachyrhynchus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="500" phylum="Arthropoda" rank="species" species="faisali">
|
|
<emphasis id="B96BEAB8DB59FFDDFE21FF48FF3A8E81" box="[495,512,187,212]" italics="true" pageId="5" pageNumber="500">P</emphasis>
|
|
.
|
|
<emphasis id="B96BEAB8DB59FFDDFDDFFF48FF6D8E80" box="[529,599,187,213]" italics="true" pageId="5" pageNumber="500">faisali</emphasis>
|
|
</taxonomicName>
|
|
, and slightly pointed in
|
|
<taxonomicName id="4C1F4D29DB59FFDDFCB4FF48FEDF8E80" baseAuthorityName="Chevrolat" baseAuthorityYear="1879" box="[890,997,187,213]" class="Insecta" family="Curculionidae" genus="Pachyrhynchus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="500" phylum="Arthropoda" rank="species" species="viridis">
|
|
<emphasis id="B96BEAB8DB59FFDDFCB4FF48FEB18E81" box="[890,907,187,212]" italics="true" pageId="5" pageNumber="500">P</emphasis>
|
|
.
|
|
<emphasis id="B96BEAB8DB59FFDDFC53FF48FEDF8E80" box="[925,997,187,213]" italics="true" pageId="5" pageNumber="500">viridis</emphasis>
|
|
</taxonomicName>
|
|
and
|
|
<taxonomicName id="4C1F4D29DB59FFDDFBECFF48F9FA8E80" authorityName="Yoshitake" authorityYear="2017" box="[1058,1216,187,213]" class="Insecta" family="Curculionidae" genus="Pachyrhynchus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="500" phylum="Arthropoda" rank="species" species="ohbayashii">
|
|
<emphasis id="B96BEAB8DB59FFDDFBECFF48F9098E81" box="[1058,1075,187,212]" italics="true" pageId="5" pageNumber="500">P</emphasis>
|
|
.
|
|
<emphasis id="B96BEAB8DB59FFDDFB8BFF48F9FA8E80" box="[1093,1216,187,213]" italics="true" pageId="5" pageNumber="500">ohbayashii</emphasis>
|
|
</taxonomicName>
|
|
. The length of the apodemes compared to that of the aedeagus is also different (
|
|
<taxonomicName id="4C1F4D29DB59FFDDFC89FF2CFED28EAC" authority="LA" authorityName="LA" box="[839,1000,223,249]" class="Insecta" family="Curculionidae" genus="Pachyrhynchus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="500" phylum="Arthropoda" rank="species" species="forsteni">
|
|
<emphasis id="B96BEAB8DB59FFDDFC89FF2CFE628EAD" box="[839,856,223,248]" italics="true" pageId="5" pageNumber="500">P</emphasis>
|
|
.
|
|
<emphasis id="B96BEAB8DB59FFDDFCA9FF2CFE878EAC" box="[871,957,223,249]" italics="true" pageId="5" pageNumber="500">forsteni</emphasis>
|
|
LA
|
|
</taxonomicName>
|
|
/ LEd: 2.10;
|
|
<taxonomicName id="4C1F4D29DB59FFDDFBB3FF2CF8348EAC" authority="LA" authorityName="LA" box="[1149,1294,223,249]" class="Insecta" family="Curculionidae" genus="Pachyrhynchus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="500" phylum="Arthropoda" rank="species" species="faisali">
|
|
<emphasis id="B96BEAB8DB59FFDDFBB3FF2CF9B48EAD" box="[1149,1166,223,248]" italics="true" pageId="5" pageNumber="500">P</emphasis>
|
|
.
|
|
<emphasis id="B96BEAB8DB59FFDDFB53FF2CF9D98EAC" box="[1181,1251,223,249]" italics="true" pageId="5" pageNumber="500">faisali</emphasis>
|
|
LA
|
|
</taxonomicName>
|
|
/ LEd: 1.62;
|
|
<taxonomicName id="4C1F4D29DB59FFDDFF59FEF0FC468F48" authority=": LA" authorityName="LA" box="[151,380,259,285]" class="Insecta" family="Curculionidae" genus="Pachyrhynchus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="500" phylum="Arthropoda" rank="species" species="morotaiensis">
|
|
<emphasis id="B96BEAB8DB59FFDDFF59FEF0FD928F49" box="[151,168,259,284]" italics="true" pageId="5" pageNumber="500">P</emphasis>
|
|
.
|
|
<emphasis id="B96BEAB8DB59FFDDFF76FEF0FC728F49" box="[184,328,259,284]" italics="true" pageId="5" pageNumber="500">morotaiensis</emphasis>
|
|
: LA
|
|
</taxonomicName>
|
|
/ LEd: 1.41;
|
|
<taxonomicName id="4C1F4D29DB59FFDDFDD6FEF0FF8F8F48" authority=": LA" authorityName="LA" box="[536,693,259,285]" class="Insecta" family="Curculionidae" genus="Pachyrhynchus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="500" phylum="Arthropoda" rank="species" species="viridis">
|
|
<emphasis id="B96BEAB8DB59FFDDFDD6FEF0FF138F49" box="[536,553,259,284]" italics="true" pageId="5" pageNumber="500">P</emphasis>
|
|
.
|
|
<emphasis id="B96BEAB8DB59FFDDFDF7FEF0FFBB8F48" box="[569,641,259,285]" italics="true" pageId="5" pageNumber="500">viridis</emphasis>
|
|
: LA
|
|
</taxonomicName>
|
|
/ LEd: 1.55;
|
|
<taxonomicName id="4C1F4D29DB59FFDDFC9FFEF0F9278F48" authority=": LA" authorityName="LA" box="[849,1053,259,285]" class="Insecta" family="Curculionidae" genus="Pachyrhynchus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="500" phylum="Arthropoda" rank="species" species="ohbayashii">
|
|
<emphasis id="B96BEAB8DB59FFDDFC9FFEF0FED38F48" box="[849,1001,259,285]" italics="true" pageId="5" pageNumber="500">P. ohbayashii</emphasis>
|
|
: LA
|
|
</taxonomicName>
|
|
/ LEd: 1.79). On the basis of my experience, the length of the apodemes of the
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|
<taxonomicName id="4C1F4D29DB59FFDDFD5CFED4FE788F14" authorityName="Schoenherr" authorityYear="1826" box="[658,834,295,321]" class="Insecta" family="Curculionidae" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="500" phylum="Arthropoda" rank="tribe" tribe="Pachyrhynchini">Pachyrhynchini</taxonomicName>
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is directly proportional to the length of the introflexed endophallus, and therefore of the everted one also.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |