600 lines
92 KiB
XML
600 lines
92 KiB
XML
<document id="42BCEFB102FCC48B4951F0230D9308F3" ID-DOI="10.5281/zenodo.195391" ID-GBIF-Dataset="369a933b-24a3-49ce-a98c-7c1bb931a178" ID-ISSN="1175-5326" ID-Zenodo-Dep="195391" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460226407764" checkinUser="plazi" docAuthor="Lindquist, Evert E. & Moraza, Maria L." docDate="2010" docId="03ACF95E27003D52FF6DF9E1FEB69260" docLanguage="en" docName="zt02479p021.pdf" docOrigin="Zootaxa 2479" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Opilioseius Lindquist & Moraza, 2010, new genus" docType="treatment" docVersion="8" lastPageNumber="14" masterDocId="FF95812627033D5FFFFAFFE3FFF69522" masterDocTitle="Revised diagnosis of the family Blattisociidae (Acari: Mesostigmata: Phytoseioidea), with a key to its genera and description of a new fungus- inhabiting genus from Costa Rica" masterLastPageNumber="21" masterPageNumber="1" pageNumber="4" updateTime="1698238420118" updateUser="plazi">
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<mods:mods id="223502987697DC4605EEDBFC45964CDF" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="1C2AFA787BB11498099EDDD8CDC60027">
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<mods:title id="11A493F5E21C4C323C5949C358E19AC6">Revised diagnosis of the family Blattisociidae (Acari: Mesostigmata: Phytoseioidea), with a key to its genera and description of a new fungus- inhabiting genus from Costa Rica</mods:title>
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</mods:titleInfo>
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<mods:name id="FCB491EA89660DD2471DD00664D67AA5" type="personal">
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<mods:role id="B9AF4BCEADD9816F08C1B5C2F22670A3">
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<mods:roleTerm id="0DDCC012EADD393EAA26FDF3099FB441">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="C8B2AA83A68215D5A677B22B14B98D8A">Lindquist, Evert E.</mods:namePart>
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</mods:name>
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<mods:name id="A5FB4F12A4F8C71E39776E668C27B2EB" type="personal">
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<mods:role id="4877A185B63EC1F82B0F8FB0A7558F03">
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<mods:roleTerm id="3DDE32EACB5DDCA6DCA9E98C86EB60A9">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="7E55D5E243E4266ED1F9B24FCB9E6DB4">Moraza, Maria L.</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="42B23B07EF8AF913AC3E6AE0A9D14F32">text</mods:typeOfResource>
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<mods:relatedItem id="5E2BDF151324B4F9B43AE29651D5B42D" type="host">
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<mods:titleInfo id="81488494BB57132B9C401C15F3D6D626">
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<mods:title id="ED2E1B1ECAAB7A961938C475BDB840D6">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="27AB66957BF6501F4E712207F06F7CF6">
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<mods:date id="3188F0D58539CB75CD94DC8D91A3A407">2010</mods:date>
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<mods:detail id="9A0DD0BBA26374CA191BC4AE4D9FE168" type="volume">
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<mods:number id="8BCB998A12A711C72D20CD06D4DC6B39">2479</mods:number>
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<mods:start id="9740CAE8CDE84082CCE0B2EA0A12EA6C">1</mods:start>
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<mods:end id="6F175C36B83BAF7F1842A45767E36193">21</mods:end>
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<mods:classification id="EDFEDA122CF8AAFAF228CD000B19549D">journal article</mods:classification>
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<mods:identifier id="37B628CB50C5EE9060FDF69CB244A82E" type="DOI">10.5281/zenodo.195391</mods:identifier>
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<mods:identifier id="6DDC31136FC4426843C7D52F7ED03471" type="GBIF-Dataset">369a933b-24a3-49ce-a98c-7c1bb931a178</mods:identifier>
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<mods:identifier id="5963C5192862F2ABB849ED20E0E9C7B8" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="DB382F8912FD7A4E624BB565DE4295FA" type="Zenodo-Dep">195391</mods:identifier>
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<treatment id="03ACF95E27003D52FF6DF9E1FEB69260" ID-DOI="http://doi.org/10.5281/zenodo.6198575" ID-GBIF-Taxon="119386430" ID-Zenodo-Dep="6198575" LSID="urn:lsid:plazi:treatment:03ACF95E27003D52FF6DF9E1FEB69260" httpUri="http://treatment.plazi.org/id/03ACF95E27003D52FF6DF9E1FEB69260" lastPageId="13" lastPageNumber="14" pageId="3" pageNumber="4">
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<subSubSection id="C31F1BC327003D5CFF6DF9E1FE38931D" pageId="3" pageNumber="4" type="nomenclature">
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<paragraph id="8BBA484827003D5CFF6DF9E1FE58933E" blockId="3.[151,462,1538,1599]" box="[151,430,1538,1564]" pageId="3" pageNumber="4">
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<heading id="D0F2FF2427003D5CFF6DF9E1FE58933E" bold="true" box="[151,430,1538,1564]" fontSize="11" level="1" pageId="3" pageNumber="4" reason="1">
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<emphasis id="B971945A27003D5CFF6DF9E1FE58933E" bold="true" box="[151,430,1538,1564]" pageId="3" pageNumber="4">
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<taxonomicName id="4C0533CB27003D5CFF6DF9E1FEE8933E" ID-CoL="9BBZN" box="[151,286,1538,1564]" class="Arachnida" family="Blattisociidae" genus="Opilioseius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="B971945A27003D5CFF6DF9E1FEE8933E" bold="true" box="[151,286,1538,1564]" italics="true" pageId="3" pageNumber="4">Opilioseius</emphasis>
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</taxonomicName>
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,
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<taxonomicNameLabel id="A242292127003D5CFED7F9E1FE58933E" box="[301,430,1538,1564]" pageId="3" pageNumber="4" rank="genus">new genus</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BBA484827003D5CFF6DF9C6FE38931D" blockId="3.[151,462,1538,1599]" box="[151,462,1573,1599]" pageId="3" pageNumber="4">
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(
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<figureCitation id="133E54CD27003D5CFF65F9C6FEE1931D" box="[159,279,1573,1599]" captionStart-0="FIGURES 1 – 11" captionStart-1="FIGURES 12 – 13" captionStart-2="FIGURES 14 – 16" captionStart-3="FIGURES 17 – 22" captionStart-4="FIGURES 23 – 31. 23 – 29" captionStartId-0="4.[151,269,1877,1901]" captionStartId-1="5.[151,272,1979,2003]" captionStartId-2="6.[151,270,1764,1788]" captionStartId-3="8.[151,269,1841,1865]" captionStartId-4="9.[151,271,1803,1827]" captionTargetBox-0="[157,1435,197,1824]" captionTargetBox-1="[157,1436,500,1919]" captionTargetBox-2="[154,1435,194,1763]" captionTargetBox-3="[150,1436,197,1772]" captionTargetBox-4="[163,1436,194,1764]" captionTargetId-0="figure@4.[157,1442,197,1853]" captionTargetId-1="figure@5.[153,1438,494,1958]" captionTargetId-2="figure@6.[154,1439,194,1789]" captionTargetId-3="figure@8.[132,1455,197,1935]" captionTargetId-4="figure@9.[163,1448,194,1782]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionTargetPageId-2="6" captionTargetPageId-3="8" captionTargetPageId-4="9" captionText-0="FIGURES 1 – 11. Opilioseius grallator n. sp., adult female: 1, subcapitulum; 2 a – 2 b, corniculi; 3, two views of corniculus and salivary stylet, dorsal view; 4, paralabral processes; 5, tritosternum, showing denticulate chevron at base of laciniae; 6, palpus, dorsolateral view; 7, detail of seta al of palpfemur; 8, chelicera, lateral antiaxial view; 9 a, separate ventrolateral view of movable digit; 9 b, separate paraxial lateral view of movable digit; 10, separate paraxial view of fixed digit; 11, gnathotectum." captionText-1="FIGURES 12 – 13. Opilioseius grallator n. sp., adult female: 12, idiosoma, dorsal aspect with detail of peritrematal shield; 13, enlarged separate views of dorsal setae denoted." captionText-2="FIGURES 14 – 16. Opilioseius grallator n. sp., adult female: 14 a – b, detail of spermatheca showing minor duct arising from embolus at base of calyx; 15, detail of peritrematal shield, free from exopodal strip alongside legs II – IV except where they join at mid-level of leg IV; 16, idiosoma, ventral aspect with detail of glandular pores on coxa I." captionText-3="FIGURES 17 – 22. Opilioseius grallator n. sp., adult female: 17 – 20, legs I-IV, excluding tarsi: 17, leg I, dorsal view; 18, leg II, posterolateral view; 19, leg III, posterolateral view; 20, leg IV, posterolateral view with detail of elongate setae on genu and tibia; 21, tarsus IV, dorsal view; 22 a – b, details of distal structures of tarsus IV: 22 a, ventroapical tarsal process below base of pretarsus, and distal minute tarsal setae ad - 1, pd - 1; 22 b, pretarsal claws, paradactyli and pulvillus." captionText-4="FIGURES 23 – 31. 23 – 29, Opilioseius grallator n. sp., adult: 23, tarsus I, right leg, dorsal view; 24, tarsus I, detail of tarsal seta “ S ” and associated setae; 25 – 29, male: 25, chelicera, paraxial view; 26 a separate, lateroventral view of movable digit; 26 b, separate, lateroventral view of movable digit with scale over dentate region; 27 a spermatodactyl with ventral scale; 27 b, ventral cheliceral scale; 28, gnathotectum; 29, subcapitulum. 30 – 31, Podocinum sp., adult: 30, tarsus I, right leg, dorsal view; 31, tarsus I, detail of tarsal seta “ S ” and associated setae." httpUri-0="https://zenodo.org/record/195392/files/figure.png" httpUri-1="https://zenodo.org/record/195393/files/figure.png" httpUri-2="https://zenodo.org/record/195394/files/figure.png" httpUri-3="https://zenodo.org/record/195395/files/figure.png" httpUri-4="https://zenodo.org/record/195396/files/figure.png" pageId="3" pageNumber="4">Figs 1–29</figureCitation>
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,
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<figureCitation id="133E54CD27003D5CFED9F9C6FE99931D" box="[291,367,1573,1599]" captionStart-0="FIGURES 32 – 33" captionStart-1="FIGURES 34 – 38. 34 – 35" captionStartId-0="10.[151,269,1980,2004]" captionStartId-1="11.[151,270,1817,1841]" captionTargetBox-0="[154,1436,642,1930]" captionTargetBox-1="[157,1436,204,1770]" captionTargetId-0="figure@10.[154,1439,642,1958]" captionTargetId-1="figure@11.[156,1441,197,1794]" captionTargetPageId-0="10" captionTargetPageId-1="11" captionText-0="FIGURES 32 – 33. Opilioseius grallator n. sp., adult male: 32, idiosoma, dorsal aspect, with laterally expanded detail of peritrematal shield fused to podonotal dorsal region; 33, idiosoma, ventral view." captionText-1="FIGURES 34 – 38. 34 – 35, Opilioseius grallator n. sp., deutonymph: 34, idiosoma, dorsal aspect; 35, idiosoma, ventral aspect. 36 – 38, leg III, tarsal apex and pretarsus, showing different forms of ventroapical tarsal processes and conjectured acrotarsal demarcation: 36, Adenoseius sp. (Sejidae), anterolateral view; 37, Euzercon sp. (Euzerconidae), ventral view; 38, Arctacarus rostratus Evans (Arctacaridae), ventral view. Abbreviation: acr mar, acrotarsal margin." httpUri-0="https://zenodo.org/record/195397/files/figure.png" httpUri-1="https://zenodo.org/record/195398/files/figure.png" pageId="3" pageNumber="4">32–35</figureCitation>
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,
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<figureCitation id="133E54CD27003D5CFE81F9C6FE30931D" box="[379,454,1573,1599]" captionStart="FIGURES 39 – 40" captionStartId="12.[151,272,1939,1963]" captionTargetBox="[221,1378,178,1918]" captionTargetId="figure@12.[221,1379,176,1918]" captionTargetPageId="12" captionText="FIGURES 39 – 40. Opilioseius grallator n. sp.: 39, photomicrograph of adult female, dorsoventral habitus view; 40, photomicrograph of gravid adult female, showing relative size of fully developed egg and maternal idiosoma, dorsoventral habitus view." httpUri="https://zenodo.org/record/195399/files/figure.png" pageId="3" pageNumber="4">39, 40</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C31F1BC327003D5CFF6DF989FCD193A3" box="[151,807,1642,1666]" pageId="3" pageNumber="4" type="materials_examined">
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<paragraph id="8BBA484827003D5CFF6DF989FCD193A3" blockId="3.[151,807,1642,1666]" box="[151,807,1642,1666]" pageId="3" pageNumber="4">
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<emphasis id="B971945A27003D5CFF6DF989FEDB93A0" bold="true" box="[151,301,1642,1666]" pageId="3" pageNumber="4">
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<typeStatus id="54BEF6EA27003D5CFF6DF989FF3993A0" box="[151,207,1642,1666]" pageId="3" pageNumber="4">Type</typeStatus>
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species:
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</emphasis>
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<taxonomicName id="4C0533CB27003D5CFEC9F989FDE593A3" box="[307,531,1642,1665]" class="Arachnida" family="Blattisociidae" genus="Opilioseius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="grallator" status="sp. nov.">
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<emphasis id="B971945A27003D5CFEC9F989FDE593A3" box="[307,531,1642,1665]" italics="true" pageId="3" pageNumber="4">Opilioseius grallator</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A242292127003D5CFDE0F989FD6B93A0" box="[538,669,1642,1666]" pageId="3" pageNumber="4" rank="species">
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<emphasis id="B971945A27003D5CFDE0F989FD6B93A0" bold="true" box="[538,669,1642,1666]" pageId="3" pageNumber="4">new species</emphasis>
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</taxonomicNameLabel>
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. Monotypic.
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</paragraph>
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</subSubSection>
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<subSubSection id="C31F1BC327003D5AFF6DF949FB4B94E5" lastPageId="5" lastPageNumber="6" pageId="3" pageNumber="4" type="diagnosis">
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<paragraph id="8BBA484827003D5AFF6DF949FB4B94E5" blockId="3.[151,1437,1706,2031]" lastBlockId="5.[151,1437,152,455]" lastPageId="5" lastPageNumber="6" pageId="3" pageNumber="4">
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<emphasis id="B971945A27003D5CFF6DF949FEE193E6" bold="true" box="[151,279,1706,1732]" pageId="3" pageNumber="4">Diagnosis.</emphasis>
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Among the genera of Blattisociinae, adults of
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<taxonomicName id="4C0533CB27003D5CFCC2F948FC4B93E6" box="[824,957,1707,1732]" class="Arachnida" family="Blattisociidae" genus="Opilioseius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="B971945A27003D5CFCC2F948FC4B93E6" box="[824,957,1707,1732]" italics="true" pageId="3" pageNumber="4">Opilioseius</emphasis>
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</taxonomicName>
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are most similar to those of
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<taxonomicName id="4C0533CB27003D5CFAF1F948FA6B93E6" box="[1291,1437,1707,1732]" class="Arachnida" family="Blattisociidae" genus="Aceodromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
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<emphasis id="B971945A27003D5CFAF1F948FA6B93E6" box="[1291,1437,1707,1732]" italics="true" pageId="3" pageNumber="4">Aceodromus</emphasis>
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</taxonomicName>
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in sharing the following apomorphic attributes: dorsal shield hypotrichous, lacking podonotal setae
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<emphasis id="B971945A27003D5CFAD3F933FAB593CB" box="[1321,1347,1744,1769]" italics="true" pageId="3" pageNumber="4">z1</emphasis>
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,
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<emphasis id="B971945A27003D5CFAA8F933FA9D93CB" box="[1362,1387,1744,1769]" italics="true" pageId="3" pageNumber="4">z3</emphasis>
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,
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<emphasis id="B971945A27003D5CFA81F933FA6293CB" box="[1403,1428,1744,1769]" italics="true" pageId="3" pageNumber="4">z6</emphasis>
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,
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<emphasis id="B971945A27003D5CFF6DF916FF47922C" box="[151,177,1781,1806]" italics="true" pageId="3" pageNumber="4">r2</emphasis>
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and opisthonotal setae
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<emphasis id="B971945A27003D5CFE36F916FDFB922C" box="[460,525,1781,1807]" italics="true" pageId="3" pageNumber="4">J1-J3</emphasis>
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,
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<emphasis id="B971945A27003D5CFDE7F916FDCC922C" box="[541,570,1781,1806]" italics="true" pageId="3" pageNumber="4">S1</emphasis>
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; genua of legs II and III with 10 and 8 setae, respectively, lacking
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<emphasis id="B971945A27003D5CFAA5F916FA62922C" box="[1375,1428,1781,1807]" italics="true" pageId="3" pageNumber="4">pv -1</emphasis>
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; female ventrianal shield weakly sclerotized, with 2, 3, or 4 pairs of opisthogastric setae plus the circumanal setae; female spermatheca with weakly sclerotized calyx. They are distinguished apomorphically from those of
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<taxonomicName id="4C0533CB27003D5CFF40F886FEA3925C" box="[186,341,1893,1918]" class="Arachnida" family="Blattisociidae" genus="Aceodromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
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<emphasis id="B971945A27003D5CFF40F886FEA3925C" box="[186,341,1893,1918]" italics="true" pageId="3" pageNumber="4">Aceodromus</emphasis>
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</taxonomicName>
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and other blattisociine genera by the following attributes: idiosomatic dorsum lacking paravertical lyrifissures
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<emphasis id="B971945A27003D5CFE4BF868FE299286" box="[433,479,1931,1956]" italics="true" pageId="3" pageNumber="4">idj1</emphasis>
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, lacking opisthonotal setae
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<emphasis id="B971945A27003D5CFCD9F868FCB49286" box="[803,834,1931,1956]" italics="true" pageId="3" pageNumber="4">Z2</emphasis>
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,
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<emphasis id="B971945A27003D5CFCAAF868FC9B9286" box="[848,877,1931,1956]" italics="true" pageId="3" pageNumber="4">S1</emphasis>
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,
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<emphasis id="B971945A27003D5CFC86F868FC6F9286" box="[892,921,1931,1956]" italics="true" pageId="3" pageNumber="4">S2</emphasis>
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, and having the
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<emphasis id="B971945A27003D5CFBA0F868FB769286" box="[1114,1152,1930,1956]" italics="true" pageId="3" pageNumber="4">r -R</emphasis>
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marginal series reduced to just four pairs in females, three of which on soft cuticle, and to two pairs in males, none of which on soft cuticle; peritremes shortened, not reaching to the vertex area; legs I to IV greatly elongated, with I and IV two to three times as long as dorsal shield, and with tarsi II to IV over three times as long as the respective tibial segments; movable cheliceral digit with five teeth and, more basally, with a series of indentations to accommodate the basal teeth of the fixed digit; movable digit with two groove-like structures on its ventroparaxial surface. In contrast, adults of
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<taxonomicName id="4C0533CB27063D5AFD98FEE3FD03943B" box="[610,757,256,281]" class="Arachnida" family="Blattisociidae" genus="Aceodromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
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<emphasis id="B971945A27063D5AFD98FEE3FD03943B" box="[610,757,256,281]" italics="true" pageId="5" pageNumber="6">Aceodromus</emphasis>
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</taxonomicName>
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are apomorphic in lacking podonotal setae
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<emphasis id="B971945A27063D5AFB05FEE3FAE0943B" box="[1279,1302,256,281]" italics="true" pageId="5" pageNumber="6">j1</emphasis>
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,
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<emphasis id="B971945A27063D5AFADFFEE3FAC9943B" box="[1317,1343,256,281]" italics="true" pageId="5" pageNumber="6">s1</emphasis>
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,
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<emphasis id="B971945A27063D5AFAB5FEE3FA9C943B" box="[1359,1386,256,281]" italics="true" pageId="5" pageNumber="6">s2</emphasis>
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,
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<emphasis id="B971945A27063D5AFA83FEE3FA65943B" box="[1401,1427,256,281]" italics="true" pageId="5" pageNumber="6">s5</emphasis>
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; plesiomorphic differences include females retaining eight pairs of
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<emphasis id="B971945A27063D5AFC67FEC1FC329419" box="[925,964,290,316]" italics="true" pageId="5" pageNumber="6">r -R</emphasis>
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marginals, including seven pairs on soft cuticle (undescribed males have two pairs, of which one pair on soft cuticle, EEL, personal observation); peritremes fully developed, their apices nearly contiguous in vertex area, legs I to IV are subequal to at most 1.5 as long as dorsal shield, and with tarsi II-IV 1.5 to 2.8 times as long as respective tibial segments; movable cheliceral digit quadridentate, lacking basal indentations and lateral groove-like structures.
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</paragraph>
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</subSubSection>
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<caption id="DF7A18C027073D5BFF6DF8B6FE2592CE" httpUri="https://zenodo.org/record/195392/files/figure.png" pageId="4" pageNumber="5" targetBox="[157,1435,197,1824]" targetPageId="4">
|
||
<paragraph id="8BBA484827073D5BFF6DF8B6FE2592CE" blockId="4.[151,1437,1877,2028]" pageId="4" pageNumber="5">
|
||
<emphasis id="B971945A27073D5BFF6DF8B6FEB8924F" bold="true" box="[151,334,1877,1901]" pageId="4" pageNumber="5">FIGURES 1–11.</emphasis>
|
||
<taxonomicName id="4C0533CB27073D5BFEAEF8B6FDC5924E" box="[340,563,1877,1900]" class="Arachnida" family="Blattisociidae" genus="Opilioseius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="grallator" status="sp. nov.">
|
||
<emphasis id="B971945A27073D5BFEAEF8B6FDC5924E" box="[340,563,1877,1900]" italics="true" pageId="4" pageNumber="5">Opilioseius grallator</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A242292127073D5BFDC0F8B6FD83924F" box="[570,629,1877,1901]" pageId="4" pageNumber="5" rank="species">
|
||
<emphasis id="B971945A27073D5BFDC0F8B6FD83924F" bold="true" box="[570,629,1877,1901]" pageId="4" pageNumber="5">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, adult female: 1, subcapitulum; 2a–2b, corniculi; 3, two views of corniculus and salivary stylet, dorsal view; 4, paralabral processes; 5, tritosternum, showing denticulate chevron at base of laciniae; 6, palpus, dorsolateral view; 7, detail of seta
|
||
<emphasis id="B971945A27073D5BFD51F876FD34928E" box="[683,706,1941,1964]" italics="true" pageId="4" pageNumber="5">al</emphasis>
|
||
of palpfemur; 8, chelicera, lateral antiaxial view; 9a, separate ventrolateral view of movable digit; 9b, separate paraxial lateral view of movable digit; 10, separate paraxial view of fixed digit; 11, gnathotectum.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF7A18C027063D5AFF6DF858FCF892D1" httpUri="https://zenodo.org/record/195393/files/figure.png" pageId="5" pageNumber="6" targetBox="[157,1436,500,1919]" targetPageId="5">
|
||
<paragraph id="8BBA484827063D5AFF6DF858FCF892D1" blockId="5.[151,1436,1979,2035]" pageId="5" pageNumber="6">
|
||
<emphasis id="B971945A27063D5AFF6DF858FE9592F1" bold="true" box="[151,355,1979,2003]" pageId="5" pageNumber="6">FIGURES 12–13.</emphasis>
|
||
<taxonomicName id="4C0533CB27063D5AFE91F85FFDA292F1" box="[363,596,1980,2003]" class="Arachnida" family="Blattisociidae" genus="Opilioseius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="grallator" status="sp. nov.">
|
||
<emphasis id="B971945A27063D5AFE91F85FFDA292F1" box="[363,596,1980,2003]" italics="true" pageId="5" pageNumber="6">Opilioseius grallator</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A242292127063D5AFDA6F858FD6D92F1" box="[604,667,1979,2003]" pageId="5" pageNumber="6" rank="species">
|
||
<emphasis id="B971945A27063D5AFDA6F858FD6D92F1" bold="true" box="[604,667,1979,2003]" pageId="5" pageNumber="6">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, adult female: 12, idiosoma, dorsal aspect with detail of peritrematal shield; 13, enlarged separate views of dorsal setae denoted.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF7A18C027053D59FF6DF907FAE79219" httpUri="https://zenodo.org/record/195394/files/figure.png" pageId="6" pageNumber="7" targetBox="[154,1435,194,1763]" targetPageId="6">
|
||
<paragraph id="8BBA484827053D59FF6DF907FAE79219" blockId="6.[151,1436,1764,1851]" pageId="6" pageNumber="7">
|
||
<emphasis id="B971945A27053D59FF6DF907FEA993DE" bold="true" box="[151,351,1764,1788]" pageId="6" pageNumber="7">FIGURES 14–16.</emphasis>
|
||
<taxonomicName id="4C0533CB27053D59FE9DF906FDBC93DE" box="[359,586,1765,1788]" class="Arachnida" family="Blattisociidae" genus="Opilioseius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="grallator" status="sp. nov.">
|
||
<emphasis id="B971945A27053D59FE9DF906FDBC93DE" box="[359,586,1765,1788]" italics="true" pageId="6" pageNumber="7">Opilioseius grallator</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A242292127053D59FDA8F907FD7993DE" box="[594,655,1764,1788]" pageId="6" pageNumber="7" rank="species">
|
||
<emphasis id="B971945A27053D59FDA8F907FD7993DE" bold="true" box="[594,655,1764,1788]" pageId="6" pageNumber="7">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, adult female: 14a–b, detail of spermatheca showing minor duct arising from embolus at base of calyx; 15, detail of peritrematal shield, free from exopodal strip alongside legs II–IV except where they join at mid-level of leg IV; 16, idiosoma, ventral aspect with detail of glandular pores on coxa I.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C31F1BC327053D55FF3CF887FAC8976E" lastPageId="10" lastPageNumber="11" pageId="6" pageNumber="7" type="description">
|
||
<paragraph id="8BBA484827053D58FF3CF887FCB3944E" blockId="6.[151,1436,1892,2031]" lastBlockId="7.[151,1437,152,1970]" lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7">
|
||
<emphasis id="B971945A27053D59FF3CF887FEA9925C" bold="true" box="[198,351,1892,1918]" pageId="6" pageNumber="7">Description.</emphasis>
|
||
<emphasis id="B971945A27053D59FE93F886FDB6925C" box="[361,576,1893,1918]" italics="true" pageId="6" pageNumber="7">Idiosomal dorsum</emphasis>
|
||
. Adult female (
|
||
<figureCitation id="133E54CD27053D59FCF9F886FCA8925D" box="[771,862,1893,1919]" captionStart="FIGURES 12 – 13" captionStartId="5.[151,272,1979,2003]" captionTargetBox="[157,1436,500,1919]" captionTargetId="figure@5.[153,1438,494,1958]" captionTargetPageId="5" captionText="FIGURES 12 – 13. Opilioseius grallator n. sp., adult female: 12, idiosoma, dorsal aspect with detail of peritrematal shield; 13, enlarged separate views of dorsal setae denoted." httpUri="https://zenodo.org/record/195393/files/figure.png" pageId="6" pageNumber="7">Fig. 12</figureCitation>
|
||
). Dorsal shield entire, without lateral incisions, lightly sclerotized, surrounding soft integument smoothly striate. Dorsal shield with complement of 23 pairs of setae, including 15 podonotal pairs (
|
||
<emphasis id="B971945A27053D59FD9EF853FD6D92EB" box="[612,667,1967,1993]" italics="true" pageId="6" pageNumber="7">j1-j6</emphasis>
|
||
,
|
||
<emphasis id="B971945A27053D59FD51F853FD3392EB" box="[683,709,1968,1993]" italics="true" pageId="6" pageNumber="7">z2</emphasis>
|
||
,
|
||
<emphasis id="B971945A27053D59FD2EF853FD1892EB" box="[724,750,1968,1993]" italics="true" pageId="6" pageNumber="7">z4</emphasis>
|
||
,
|
||
<emphasis id="B971945A27053D59FD04F853FCE192EB" box="[766,791,1968,1993]" italics="true" pageId="6" pageNumber="7">z5</emphasis>
|
||
,
|
||
<emphasis id="B971945A27053D59FCDDF853FCB792EB" box="[807,833,1968,1993]" italics="true" pageId="6" pageNumber="7">s1</emphasis>
|
||
,
|
||
<emphasis id="B971945A27053D59FCAAF853FC9C92EB" box="[848,874,1968,1993]" italics="true" pageId="6" pageNumber="7">s2</emphasis>
|
||
,
|
||
<emphasis id="B971945A27053D59FC80F853FC4192EB" box="[890,951,1967,1993]" italics="true" pageId="6" pageNumber="7">s4-s6</emphasis>
|
||
,
|
||
<emphasis id="B971945A27053D59FC3DF853FC1692EB" box="[967,992,1968,1993]" italics="true" pageId="6" pageNumber="7">r3</emphasis>
|
||
) and eight opisthonotal pairs (
|
||
<emphasis id="B971945A27053D59FAB1F853FA9192EB" box="[1355,1383,1968,1993]" italics="true" pageId="6" pageNumber="7">J5</emphasis>
|
||
,
|
||
<emphasis id="B971945A27053D59FA8FF853FA6292EB" box="[1397,1428,1968,1993]" italics="true" pageId="6" pageNumber="7">Z1</emphasis>
|
||
,
|
||
<emphasis id="B971945A27053D59FF6DF836FF1792CC" box="[151,225,2005,2031]" italics="true" pageId="6" pageNumber="7">Z3-Z5</emphasis>
|
||
,
|
||
<emphasis id="B971945A27053D59FF0BF836FEC192CC" box="[241,311,2005,2031]" italics="true" pageId="6" pageNumber="7">S3-S5</emphasis>
|
||
); setae attenuate, of widely disparate lengths,
|
||
<emphasis id="B971945A27053D59FC8AF836FC7B92CC" box="[880,909,2005,2030]" italics="true" pageId="6" pageNumber="7">J5</emphasis>
|
||
shortest,
|
||
<emphasis id="B971945A27053D59FBFEF836FBED92CC" box="[1028,1051,2005,2030]" italics="true" pageId="6" pageNumber="7">j6</emphasis>
|
||
,
|
||
<emphasis id="B971945A27053D59FBD6F836FBB092CC" box="[1068,1094,2005,2030]" italics="true" pageId="6" pageNumber="7">z2</emphasis>
|
||
,
|
||
<emphasis id="B971945A27053D59FBADF836FB8492CC" box="[1111,1138,2005,2030]" italics="true" pageId="6" pageNumber="7">z5</emphasis>
|
||
short,
|
||
<emphasis id="B971945A27053D59FB3DF836FB1592CC" box="[1223,1251,2005,2030]" italics="true" pageId="6" pageNumber="7">s5</emphasis>
|
||
longest,
|
||
<emphasis id="B971945A27053D59FAA9F836FA6A92CC" box="[1363,1436,2005,2031]" italics="true" pageId="6" pageNumber="7">Z3-Z5</emphasis>
|
||
nearly as long; most setae smooth, but
|
||
<emphasis id="B971945A27043D58FD8FFF7BFD679593" box="[629,657,152,177]" italics="true" pageId="7" pageNumber="8">J5</emphasis>
|
||
minutely barbed. Dorsal shield with complement of 18 pairs of discernible pore-like structures (eight podonotal, ten opisthonotal), of which seven pairs (four podonotal, three opisthonotal) superficially appear secretory (gland pores) and 11 pairs (four podonotal, seven opisthonotal) non-secretory (poroids). Soft surrounding cuticle with only three pairs of
|
||
<emphasis id="B971945A27043D58FB6EFEEBFB4C9403" box="[1172,1210,264,289]" italics="true" pageId="7" pageNumber="8">r-R</emphasis>
|
||
marginal setae and pair of marginal poroids
|
||
<emphasis id="B971945A27043D58FE39FECEFE0D9464" box="[451,507,301,326]" italics="true" pageId="7" pageNumber="8">idRp</emphasis>
|
||
. Peritrematal plates narrowly uniting with dorsal shield anteriorly to level of setae
|
||
<emphasis id="B971945A27043D58FF23FEB1FF049449" box="[217,242,338,363]" italics="true" pageId="7" pageNumber="8">s1</emphasis>
|
||
; peritremes shortened, not reaching to vertex area.
|
||
</paragraph>
|
||
<paragraph id="8BBA484827043D58FF3EFE9BFD9F9720" blockId="7.[151,1437,152,1970]" pageId="7" pageNumber="8">
|
||
Adult male (
|
||
<figureCitation id="133E54CD27043D58FEA2FE9BFE4694B0" box="[344,432,376,402]" captionStart="FIGURES 32 – 33" captionStartId="10.[151,269,1980,2004]" captionTargetBox="[154,1436,642,1930]" captionTargetId="figure@10.[154,1439,642,1958]" captionTargetPageId="10" captionText="FIGURES 32 – 33. Opilioseius grallator n. sp., adult male: 32, idiosoma, dorsal aspect, with laterally expanded detail of peritrematal shield fused to podonotal dorsal region; 33, idiosoma, ventral view." httpUri="https://zenodo.org/record/195397/files/figure.png" pageId="7" pageNumber="8">Fig. 32</figureCitation>
|
||
). Dorsal shield setation, ornamentation, and form as in female except more expanded anterolaterally so as to bear setae
|
||
<emphasis id="B971945A27043D58FDD9FE7EFDCA9494" box="[547,572,413,438]" italics="true" pageId="7" pageNumber="8">r5</emphasis>
|
||
and be more broadly united to peritrematal shields at level between setae
|
||
<emphasis id="B971945A27043D58FF6DFE21FF4794F9" box="[151,177,450,475]" italics="true" pageId="7" pageNumber="8">r3</emphasis>
|
||
and
|
||
<emphasis id="B971945A27043D58FF11FE21FEF094F9" box="[235,262,450,475]" italics="true" pageId="7" pageNumber="8">z4</emphasis>
|
||
. Form and proportionate size of dorsal shield setae as in female. Lateral soft cuticle without any
|
||
<emphasis id="B971945A27043D58FA7AFE21FA6494F9" box="[1408,1426,450,475]" italics="true" pageId="7" pageNumber="8">R</emphasis>
|
||
- marginal setae. Peritremes as in female.
|
||
</paragraph>
|
||
<paragraph id="8BBA484827043D58FF3EFDEEFC5E915E" blockId="7.[151,1437,152,1970]" pageId="7" pageNumber="8">
|
||
<emphasis id="B971945A27043D58FF3EFDEEFE7A9704" box="[196,396,525,550]" italics="true" pageId="7" pageNumber="8">Idiosomal venter</emphasis>
|
||
. Adult female (
|
||
<figureCitation id="133E54CD27043D58FDB0FDEEFD559705" box="[586,675,525,551]" captionStart="FIGURES 14 – 16" captionStartId="6.[151,270,1764,1788]" captionTargetBox="[154,1435,194,1763]" captionTargetId="figure@6.[154,1439,194,1789]" captionTargetPageId="6" captionText="FIGURES 14 – 16. Opilioseius grallator n. sp., adult female: 14 a – b, detail of spermatheca showing minor duct arising from embolus at base of calyx; 15, detail of peritrematal shield, free from exopodal strip alongside legs II – IV except where they join at mid-level of leg IV; 16, idiosoma, ventral aspect with detail of glandular pores on coxa I." httpUri="https://zenodo.org/record/195394/files/figure.png" pageId="7" pageNumber="8">Fig. 16</figureCitation>
|
||
). Tritosternum with laciniae free for most of length, their fused bases with an obliquely lined series of denticles (
|
||
<figureCitation id="133E54CD27043D58FD1EFDD1FCDB976E" box="[740,813,562,588]" captionStart="FIGURES 1 – 11" captionStartId="4.[151,269,1877,1901]" captionTargetBox="[157,1435,197,1824]" captionTargetId="figure@4.[157,1442,197,1853]" captionTargetPageId="4" captionText="FIGURES 1 – 11. Opilioseius grallator n. sp., adult female: 1, subcapitulum; 2 a – 2 b, corniculi; 3, two views of corniculus and salivary stylet, dorsal view; 4, paralabral processes; 5, tritosternum, showing denticulate chevron at base of laciniae; 6, palpus, dorsolateral view; 7, detail of seta al of palpfemur; 8, chelicera, lateral antiaxial view; 9 a, separate ventrolateral view of movable digit; 9 b, separate paraxial lateral view of movable digit; 10, separate paraxial view of fixed digit; 11, gnathotectum." httpUri="https://zenodo.org/record/195392/files/figure.png" pageId="7" pageNumber="8">Fig. 5</figureCitation>
|
||
). Ventral shields unornamented, weakly sclerotized. Presternal region with weakly developed platelets confluent with sternal shield. Sternal shield entire, with weakly developed endopodal extensions between coxae I-II and II-III, and with three pairs of setae and two pairs of lyrifissures; setae
|
||
<emphasis id="B971945A27043D58FE37FD41FE199799" box="[461,495,674,699]" italics="true" pageId="7" pageNumber="8">st4</emphasis>
|
||
on metasternal plates; sternal lyrifissures
|
||
<date id="FFBB6E8827043D58FC1AFD41FBF29799" box="[992,1028,674,699]" pageId="7" pageNumber="8">
|
||
<emphasis id="B971945A27043D58FC1AFD41FBF29799" box="[992,1028,674,699]" italics="true" pageId="7" pageNumber="8">iv3</emphasis>
|
||
</date>
|
||
absent (
|
||
<figureCitation id="133E54CD27043D58FB92FD41FB36979E" box="[1128,1216,674,700]" captionStart="FIGURES 14 – 16" captionStartId="6.[151,270,1764,1788]" captionTargetBox="[154,1435,194,1763]" captionTargetId="figure@6.[154,1439,194,1789]" captionTargetPageId="6" captionText="FIGURES 14 – 16. Opilioseius grallator n. sp., adult female: 14 a – b, detail of spermatheca showing minor duct arising from embolus at base of calyx; 15, detail of peritrematal shield, free from exopodal strip alongside legs II – IV except where they join at mid-level of leg IV; 16, idiosoma, ventral aspect with detail of glandular pores on coxa I." httpUri="https://zenodo.org/record/195394/files/figure.png" pageId="7" pageNumber="8">Fig. 16</figureCitation>
|
||
). Endopodal strips between coxae III and IV free, weakly defined. Epigynal shield with its rounded, hyaline anterior margin between legs III–IV not overlapping posterior edge of sternal shield, its posterior margin truncate; setae
|
||
<emphasis id="B971945A27043D58FAAFFD0EFA8E9624" box="[1365,1400,749,774]" italics="true" pageId="7" pageNumber="8">st5</emphasis>
|
||
on shield’s posterolateral margins, paragenital poroids
|
||
<date id="FFBB6E8827043D58FCDEFCF1FCBD9609" box="[804,843,786,811]" pageId="7" pageNumber="8">
|
||
<emphasis id="B971945A27043D58FCDEFCF1FCBD9609" box="[804,843,786,811]" italics="true" pageId="7" pageNumber="8">iv5</emphasis>
|
||
</date>
|
||
on soft cuticle. Opisthosomatic venter without discernible metapodal and postgenital platelets; soft integument with seven pairs of setae and three pairs of poroids. Ventrianal shield small, with two pairs of opisthogastric setae and the three circumanal setae; paranals inserted at level slightly anterior to posterior edge of anus, and similar in size to postanal seta; ventrianal shield with a pair of gland pores on posterolateral margins, and a well-developed cribrum behind level of postanal seta. Peritrematal shield with two poroids and one gland pore in area behind stigma where connecting with exopodal strip curving behind coxa IV (
|
||
<figureCitation id="133E54CD27043D58FCD2FC11FC58912E" box="[808,942,1010,1036]" captionStart="FIGURES 14 – 16" captionStartId="6.[151,270,1764,1788]" captionTargetBox="[154,1435,194,1763]" captionTargetId="figure@6.[154,1439,194,1789]" captionTargetPageId="6" captionText="FIGURES 14 – 16. Opilioseius grallator n. sp., adult female: 14 a – b, detail of spermatheca showing minor duct arising from embolus at base of calyx; 15, detail of peritrematal shield, free from exopodal strip alongside legs II – IV except where they join at mid-level of leg IV; 16, idiosoma, ventral aspect with detail of glandular pores on coxa I." httpUri="https://zenodo.org/record/195394/files/figure.png" pageId="7" pageNumber="8">Figs 15, 16</figureCitation>
|
||
); peritrematal shield continuous alongside coxae II-IV but without exopodal extensions between bases of coxae II-III and III-IV; shield with weakly developed gland pore anterior to apex of peritreme (
|
||
<figureCitation id="133E54CD27043D58FD02FBDEFCA69175" box="[760,848,1085,1111]" captionStart="FIGURES 12 – 13" captionStartId="5.[151,272,1979,2003]" captionTargetBox="[157,1436,500,1919]" captionTargetId="figure@5.[153,1438,494,1958]" captionTargetPageId="5" captionText="FIGURES 12 – 13. Opilioseius grallator n. sp., adult female: 12, idiosoma, dorsal aspect with detail of peritrematal shield; 13, enlarged separate views of dorsal setae denoted." httpUri="https://zenodo.org/record/195393/files/figure.png" pageId="7" pageNumber="8">Fig. 12</figureCitation>
|
||
). Spermathecal apparatus of phytoseiid-type, with a minor duct emanating from embolus near base of calyx (
|
||
<figureCitation id="133E54CD27043D58FCB9FB81FC6C915E" box="[835,922,1122,1148]" captionStart="FIGURES 14 – 16" captionStartId="6.[151,270,1764,1788]" captionTargetBox="[154,1435,194,1763]" captionTargetId="figure@6.[154,1439,194,1789]" captionTargetPageId="6" captionText="FIGURES 14 – 16. Opilioseius grallator n. sp., adult female: 14 a – b, detail of spermatheca showing minor duct arising from embolus at base of calyx; 15, detail of peritrematal shield, free from exopodal strip alongside legs II – IV except where they join at mid-level of leg IV; 16, idiosoma, ventral aspect with detail of glandular pores on coxa I." httpUri="https://zenodo.org/record/195394/files/figure.png" pageId="7" pageNumber="8">Fig. 14</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BBA484827043D58FF3EFB6BFBB390A0" blockId="7.[151,1437,152,1970]" pageId="7" pageNumber="8">
|
||
Adult male (
|
||
<figureCitation id="133E54CD27043D58FEACFB6BFE5B9180" box="[342,429,1160,1186]" captionStart="FIGURES 32 – 33" captionStartId="10.[151,269,1980,2004]" captionTargetBox="[154,1436,642,1930]" captionTargetId="figure@10.[154,1439,642,1958]" captionTargetPageId="10" captionText="FIGURES 32 – 33. Opilioseius grallator n. sp., adult male: 32, idiosoma, dorsal aspect, with laterally expanded detail of peritrematal shield fused to podonotal dorsal region; 33, idiosoma, ventral view." httpUri="https://zenodo.org/record/195397/files/figure.png" pageId="7" pageNumber="8">Fig. 33</figureCitation>
|
||
). Tritosternum and presternal area as in female. Ventral shields unornamented, weakly sclerotized. Sternitigenital shield with five pairs of setae and two pairs of poroids (
|
||
<date id="FFBB6E8827043D58FB9EFB4EFB7191E4" box="[1124,1159,1197,1222]" pageId="7" pageNumber="8">
|
||
<emphasis id="B971945A27043D58FB9EFB4EFB7191E4" box="[1124,1159,1197,1222]" italics="true" pageId="7" pageNumber="8">iv3</emphasis>
|
||
</date>
|
||
absent), deeply incised between sternal and genital sections, and lacking endopodal extensions between coxae I and II. Endopodal strips between coxae II and III, III and IV free, weakly defined. Opisthosomatic venter without discernible metapodal platelets; soft integument lacking setae and discernible poroids. Ventrianal shield small, with four pairs of opisthogastric setae and the three circumanal setae; attributes of circumanal setae, gland pores, and cribrum as in female. Posteriorly, peritrematal shield and peritreme as in female.
|
||
</paragraph>
|
||
<paragraph id="8BBA484827043D58FF3EFA6EFD2A9290" blockId="7.[151,1437,152,1970]" pageId="7" pageNumber="8">
|
||
<emphasis id="B971945A27043D58FF3EFA6EFE969084" box="[196,352,1421,1446]" italics="true" pageId="7" pageNumber="8">Gnathosoma</emphasis>
|
||
. Female & male. Gnathotectum with convex, denticulate anterior margin (
|
||
<figureCitation id="133E54CD27043D58FB04FA6EFAAB9085" box="[1278,1373,1421,1447]" captionStart="FIGURES 1 – 11" captionStartId="4.[151,269,1877,1901]" captionTargetBox="[157,1435,197,1824]" captionTargetId="figure@4.[157,1442,197,1853]" captionTargetPageId="4" captionText="FIGURES 1 – 11. Opilioseius grallator n. sp., adult female: 1, subcapitulum; 2 a – 2 b, corniculi; 3, two views of corniculus and salivary stylet, dorsal view; 4, paralabral processes; 5, tritosternum, showing denticulate chevron at base of laciniae; 6, palpus, dorsolateral view; 7, detail of seta al of palpfemur; 8, chelicera, lateral antiaxial view; 9 a, separate ventrolateral view of movable digit; 9 b, separate paraxial lateral view of movable digit; 10, separate paraxial view of fixed digit; 11, gnathotectum." httpUri="https://zenodo.org/record/195392/files/figure.png" pageId="7" pageNumber="8">Figs 11</figureCitation>
|
||
,
|
||
<figureCitation id="133E54CD27043D58FA91FA6EFA789085" box="[1387,1422,1421,1447]" captionStart="FIGURES 23 – 31. 23 – 29" captionStartId="9.[151,271,1803,1827]" captionTargetBox="[163,1436,194,1764]" captionTargetId="figure@9.[163,1448,194,1782]" captionTargetPageId="9" captionText="FIGURES 23 – 31. 23 – 29, Opilioseius grallator n. sp., adult: 23, tarsus I, right leg, dorsal view; 24, tarsus I, detail of tarsal seta “ S ” and associated setae; 25 – 29, male: 25, chelicera, paraxial view; 26 a separate, lateroventral view of movable digit; 26 b, separate, lateroventral view of movable digit with scale over dentate region; 27 a spermatodactyl with ventral scale; 27 b, ventral cheliceral scale; 28, gnathotectum; 29, subcapitulum. 30 – 31, Podocinum sp., adult: 30, tarsus I, right leg, dorsal view; 31, tarsus I, detail of tarsal seta “ S ” and associated setae." httpUri="https://zenodo.org/record/195396/files/figure.png" pageId="7" pageNumber="8">28</figureCitation>
|
||
). Chelicerae of moderate length, without any conspicuous process along antiaxial or paraxial lateral surfaces basal to the digits; fixed digit with tiny, setiform pilus dentilis and row of coarse teeth along entire length of masticatory surface and an offset tooth (gabelzahn) subapically (
|
||
<figureCitation id="133E54CD27043D58FC60FA1EFBE09335" box="[922,1046,1533,1559]" captionStart="FIGURES 1 – 11" captionStartId="4.[151,269,1877,1901]" captionTargetBox="[157,1435,197,1824]" captionTargetId="figure@4.[157,1442,197,1853]" captionTargetPageId="4" captionText="FIGURES 1 – 11. Opilioseius grallator n. sp., adult female: 1, subcapitulum; 2 a – 2 b, corniculi; 3, two views of corniculus and salivary stylet, dorsal view; 4, paralabral processes; 5, tritosternum, showing denticulate chevron at base of laciniae; 6, palpus, dorsolateral view; 7, detail of seta al of palpfemur; 8, chelicera, lateral antiaxial view; 9 a, separate ventrolateral view of movable digit; 9 b, separate paraxial lateral view of movable digit; 10, separate paraxial view of fixed digit; 11, gnathotectum." httpUri="https://zenodo.org/record/195392/files/figure.png" pageId="7" pageNumber="8">Figs 8, 10</figureCitation>
|
||
,
|
||
<figureCitation id="133E54CD27043D58FBD9FA1EFBB29335" box="[1059,1092,1533,1559]" captionStart="FIGURES 23 – 31. 23 – 29" captionStartId="9.[151,271,1803,1827]" captionTargetBox="[163,1436,194,1764]" captionTargetId="figure@9.[163,1448,194,1782]" captionTargetPageId="9" captionText="FIGURES 23 – 31. 23 – 29, Opilioseius grallator n. sp., adult: 23, tarsus I, right leg, dorsal view; 24, tarsus I, detail of tarsal seta “ S ” and associated setae; 25 – 29, male: 25, chelicera, paraxial view; 26 a separate, lateroventral view of movable digit; 26 b, separate, lateroventral view of movable digit with scale over dentate region; 27 a spermatodactyl with ventral scale; 27 b, ventral cheliceral scale; 28, gnathotectum; 29, subcapitulum. 30 – 31, Podocinum sp., adult: 30, tarsus I, right leg, dorsal view; 31, tarsus I, detail of tarsal seta “ S ” and associated setae." httpUri="https://zenodo.org/record/195396/files/figure.png" pageId="7" pageNumber="8">25</figureCitation>
|
||
); movable cheliceral digit of female with a row of several coarse teeth, and with two fine paraxial grooves running proximally and ventrally from the bases of subapical teeth; arthrodial envelope fringed (
|
||
<figureCitation id="133E54CD27043D58FBF8F9ABFB8C9340" box="[1026,1146,1608,1634]" captionStart="FIGURES 1 – 11" captionStartId="4.[151,269,1877,1901]" captionTargetBox="[157,1435,197,1824]" captionTargetId="figure@4.[157,1442,197,1853]" captionTargetPageId="4" captionText="FIGURES 1 – 11. Opilioseius grallator n. sp., adult female: 1, subcapitulum; 2 a – 2 b, corniculi; 3, two views of corniculus and salivary stylet, dorsal view; 4, paralabral processes; 5, tritosternum, showing denticulate chevron at base of laciniae; 6, palpus, dorsolateral view; 7, detail of seta al of palpfemur; 8, chelicera, lateral antiaxial view; 9 a, separate ventrolateral view of movable digit; 9 b, separate paraxial lateral view of movable digit; 10, separate paraxial view of fixed digit; 11, gnathotectum." httpUri="https://zenodo.org/record/195392/files/figure.png" pageId="7" pageNumber="8">Figs 8-10</figureCitation>
|
||
); movable digit of male with teeth restricted to basal half of masticatory surface, and with short, bilobate spermatodactyl (
|
||
<figureCitation id="133E54CD27043D58FAFDF98EFA6693A5" box="[1287,1424,1645,1671]" captionStart="FIGURES 23 – 31. 23 – 29" captionStartId="9.[151,271,1803,1827]" captionTargetBox="[163,1436,194,1764]" captionTargetId="figure@9.[163,1448,194,1782]" captionTargetPageId="9" captionText="FIGURES 23 – 31. 23 – 29, Opilioseius grallator n. sp., adult: 23, tarsus I, right leg, dorsal view; 24, tarsus I, detail of tarsal seta “ S ” and associated setae; 25 – 29, male: 25, chelicera, paraxial view; 26 a separate, lateroventral view of movable digit; 26 b, separate, lateroventral view of movable digit with scale over dentate region; 27 a spermatodactyl with ventral scale; 27 b, ventral cheliceral scale; 28, gnathotectum; 29, subcapitulum. 30 – 31, Podocinum sp., adult: 30, tarsus I, right leg, dorsal view; 31, tarsus I, detail of tarsal seta “ S ” and associated setae." httpUri="https://zenodo.org/record/195396/files/figure.png" pageId="7" pageNumber="8">Figs 25–27</figureCitation>
|
||
). Paralabral processes simple, acute (
|
||
<figureCitation id="133E54CD27043D58FDBAF971FD7D938E" box="[576,651,1682,1708]" captionStart="FIGURES 1 – 11" captionStartId="4.[151,269,1877,1901]" captionTargetBox="[157,1435,197,1824]" captionTargetId="figure@4.[157,1442,197,1853]" captionTargetPageId="4" captionText="FIGURES 1 – 11. Opilioseius grallator n. sp., adult female: 1, subcapitulum; 2 a – 2 b, corniculi; 3, two views of corniculus and salivary stylet, dorsal view; 4, paralabral processes; 5, tritosternum, showing denticulate chevron at base of laciniae; 6, palpus, dorsolateral view; 7, detail of seta al of palpfemur; 8, chelicera, lateral antiaxial view; 9 a, separate ventrolateral view of movable digit; 9 b, separate paraxial lateral view of movable digit; 10, separate paraxial view of fixed digit; 11, gnathotectum." httpUri="https://zenodo.org/record/195392/files/figure.png" pageId="7" pageNumber="8">Fig. 4</figureCitation>
|
||
). Corniculi normal in form, well separated, more splayed in male (
|
||
<figureCitation id="133E54CD27043D58FF65F95BFF0393F0" box="[159,245,1720,1746]" captionStart="FIGURES 23 – 31. 23 – 29" captionStartId="9.[151,271,1803,1827]" captionTargetBox="[163,1436,194,1764]" captionTargetId="figure@9.[163,1448,194,1782]" captionTargetPageId="9" captionText="FIGURES 23 – 31. 23 – 29, Opilioseius grallator n. sp., adult: 23, tarsus I, right leg, dorsal view; 24, tarsus I, detail of tarsal seta “ S ” and associated setae; 25 – 29, male: 25, chelicera, paraxial view; 26 a separate, lateroventral view of movable digit; 26 b, separate, lateroventral view of movable digit with scale over dentate region; 27 a spermatodactyl with ventral scale; 27 b, ventral cheliceral scale; 28, gnathotectum; 29, subcapitulum. 30 – 31, Podocinum sp., adult: 30, tarsus I, right leg, dorsal view; 31, tarsus I, detail of tarsal seta “ S ” and associated setae." httpUri="https://zenodo.org/record/195396/files/figure.png" pageId="7" pageNumber="8">Fig. 29</figureCitation>
|
||
) than in female (
|
||
<figureCitation id="133E54CD27043D58FE43F95BFE0B93F0" box="[441,509,1720,1746]" captionStart="FIGURES 1 – 11" captionStartId="4.[151,269,1877,1901]" captionTargetBox="[157,1435,197,1824]" captionTargetId="figure@4.[157,1442,197,1853]" captionTargetPageId="4" captionText="FIGURES 1 – 11. Opilioseius grallator n. sp., adult female: 1, subcapitulum; 2 a – 2 b, corniculi; 3, two views of corniculus and salivary stylet, dorsal view; 4, paralabral processes; 5, tritosternum, showing denticulate chevron at base of laciniae; 6, palpus, dorsolateral view; 7, detail of seta al of palpfemur; 8, chelicera, lateral antiaxial view; 9 a, separate ventrolateral view of movable digit; 9 b, separate paraxial lateral view of movable digit; 10, separate paraxial view of fixed digit; 11, gnathotectum." httpUri="https://zenodo.org/record/195392/files/figure.png" pageId="7" pageNumber="8">Fig. 1</figureCitation>
|
||
), with salivary styli appressed to dorsal cornicular faces and difficult to discern (
|
||
<figureCitation id="133E54CD27043D58FF65F93EFEF193D5" box="[159,263,1757,1783]" captionStart="FIGURES 1 – 11" captionStartId="4.[151,269,1877,1901]" captionTargetBox="[157,1435,197,1824]" captionTargetId="figure@4.[157,1442,197,1853]" captionTargetPageId="4" captionText="FIGURES 1 – 11. Opilioseius grallator n. sp., adult female: 1, subcapitulum; 2 a – 2 b, corniculi; 3, two views of corniculus and salivary stylet, dorsal view; 4, paralabral processes; 5, tritosternum, showing denticulate chevron at base of laciniae; 6, palpus, dorsolateral view; 7, detail of seta al of palpfemur; 8, chelicera, lateral antiaxial view; 9 a, separate ventrolateral view of movable digit; 9 b, separate paraxial lateral view of movable digit; 10, separate paraxial view of fixed digit; 11, gnathotectum." httpUri="https://zenodo.org/record/195392/files/figure.png" pageId="7" pageNumber="8">Figs 2, 3</figureCitation>
|
||
); internal malae fimbriated laterally. Deutosternum with seven transverse rows of denticles, anterior five rows similarly moderately wide and multidenticulate, sixth and seventh rows more widened and denticulate, all rows connected except the seventh (
|
||
<figureCitation id="133E54CD27043D58FD14F8CBFCCE9260" box="[750,824,1832,1858]" captionStart="FIGURES 1 – 11" captionStartId="4.[151,269,1877,1901]" captionTargetBox="[157,1435,197,1824]" captionTargetId="figure@4.[157,1442,197,1853]" captionTargetPageId="4" captionText="FIGURES 1 – 11. Opilioseius grallator n. sp., adult female: 1, subcapitulum; 2 a – 2 b, corniculi; 3, two views of corniculus and salivary stylet, dorsal view; 4, paralabral processes; 5, tritosternum, showing denticulate chevron at base of laciniae; 6, palpus, dorsolateral view; 7, detail of seta al of palpfemur; 8, chelicera, lateral antiaxial view; 9 a, separate ventrolateral view of movable digit; 9 b, separate paraxial lateral view of movable digit; 10, separate paraxial view of fixed digit; 11, gnathotectum." httpUri="https://zenodo.org/record/195392/files/figure.png" pageId="7" pageNumber="8">Figs 1</figureCitation>
|
||
,
|
||
<figureCitation id="133E54CD27043D58FCBFF8CBFC909260" box="[837,870,1832,1858]" captionStart="FIGURES 23 – 31. 23 – 29" captionStartId="9.[151,271,1803,1827]" captionTargetBox="[163,1436,194,1764]" captionTargetId="figure@9.[163,1448,194,1782]" captionTargetPageId="9" captionText="FIGURES 23 – 31. 23 – 29, Opilioseius grallator n. sp., adult: 23, tarsus I, right leg, dorsal view; 24, tarsus I, detail of tarsal seta “ S ” and associated setae; 25 – 29, male: 25, chelicera, paraxial view; 26 a separate, lateroventral view of movable digit; 26 b, separate, lateroventral view of movable digit with scale over dentate region; 27 a spermatodactyl with ventral scale; 27 b, ventral cheliceral scale; 28, gnathotectum; 29, subcapitulum. 30 – 31, Podocinum sp., adult: 30, tarsus I, right leg, dorsal view; 31, tarsus I, detail of tarsal seta “ S ” and associated setae." httpUri="https://zenodo.org/record/195396/files/figure.png" pageId="7" pageNumber="8">29</figureCitation>
|
||
). Subcapitular setae similarly smooth, attenuate, capitular pair
|
||
<emphasis id="B971945A27043D58FEB3F8AEFE919244" box="[329,359,1869,1894]" italics="true" pageId="7" pageNumber="8">pc</emphasis>
|
||
longer than hypostomatic pairs
|
||
<emphasis id="B971945A27043D58FCFBF8AEFC9F9244" box="[769,873,1869,1895]" italics="true" pageId="7" pageNumber="8">hp1-hp3</emphasis>
|
||
. Palpi with normal setation as described for
|
||
<taxonomicName id="4C0533CB27043D58FF6DF891FEE592AE" box="[151,275,1906,1932]" class="Arachnida" higherTaxonomySource="GBIF" infraOrder="Gamasina" kingdom="Animalia" order="Mesostigmata" pageId="7" pageNumber="8" phylum="Arthropoda" rank="infraOrder">Gamasina</taxonomicName>
|
||
by
|
||
<bibRefCitation id="EF9435B927043D58FEBEF891FE0692AE" author="Evans" box="[324,496,1906,1932]" pageId="7" pageNumber="20" refString="Evans, G. O. (1964) Some observations on the chaetotaxy of the pedipalps in the Mesostigmata (Acari). Annals and Magazine of Natural History, Series 13, 6, 513 - 527." type="journal article" year="1964">Evans (1964)</bibRefCitation>
|
||
; palpfemoral seta
|
||
<emphasis id="B971945A27043D58FD22F890FD0692AE" box="[728,752,1907,1932]" italics="true" pageId="7" pageNumber="8">al</emphasis>
|
||
spatulate, palpgenual setae
|
||
<emphasis id="B971945A27043D58FBAEF890FB7192AE" box="[1108,1159,1906,1932]" italics="true" pageId="7" pageNumber="8">al -1</emphasis>
|
||
and
|
||
<emphasis id="B971945A27043D58FB3CF890FB0E92AE" box="[1222,1272,1906,1932]" italics="true" pageId="7" pageNumber="8">al -2</emphasis>
|
||
more or less spatulate; palptarsal apotele two-tined (
|
||
<figureCitation id="133E54CD27043D58FD99F87BFD399290" box="[611,719,1944,1970]" captionStart="FIGURES 1 – 11" captionStartId="4.[151,269,1877,1901]" captionTargetBox="[157,1435,197,1824]" captionTargetId="figure@4.[157,1442,197,1853]" captionTargetPageId="4" captionText="FIGURES 1 – 11. Opilioseius grallator n. sp., adult female: 1, subcapitulum; 2 a – 2 b, corniculi; 3, two views of corniculus and salivary stylet, dorsal view; 4, paralabral processes; 5, tritosternum, showing denticulate chevron at base of laciniae; 6, palpus, dorsolateral view; 7, detail of seta al of palpfemur; 8, chelicera, lateral antiaxial view; 9 a, separate ventrolateral view of movable digit; 9 b, separate paraxial lateral view of movable digit; 10, separate paraxial view of fixed digit; 11, gnathotectum." httpUri="https://zenodo.org/record/195392/files/figure.png" pageId="7" pageNumber="8">Figs 6–7</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="DF7A18C0270B3D57FF6DF8D2FAB1928A" httpUri="https://zenodo.org/record/195395/files/figure.png" pageId="8" pageNumber="9" targetBox="[150,1436,197,1772]" targetPageId="8">
|
||
<paragraph id="8BBA4848270B3D57FF6DF8D2FAB1928A" blockId="8.[151,1436,1841,1960]" pageId="8" pageNumber="9">
|
||
<emphasis id="B971945A270B3D57FF6DF8D2FEAB926B" bold="true" box="[151,349,1841,1865]" pageId="8" pageNumber="9">FIGURES 17–22.</emphasis>
|
||
<taxonomicName id="4C0533CB270B3D57FE9EF8D2FDB2926A" box="[356,580,1841,1864]" class="Arachnida" family="Blattisociidae" genus="Opilioseius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="grallator" status="sp. nov.">
|
||
<emphasis id="B971945A270B3D57FE9EF8D2FDB2926A" box="[356,580,1841,1864]" italics="true" pageId="8" pageNumber="9">Opilioseius grallator</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2422921270B3D57FDB1F8D2FD71926B" box="[587,647,1841,1865]" pageId="8" pageNumber="9" rank="species">
|
||
<emphasis id="B971945A270B3D57FDB1F8D2FD71926B" bold="true" box="[587,647,1841,1865]" pageId="8" pageNumber="9">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, adult female: 17–20, legs I-IV, excluding tarsi: 17, leg I, dorsal view; 18, leg II, posterolateral view; 19, leg III, posterolateral view; 20, leg IV, posterolateral view with detail of elongate setae on genu and tibia; 21, tarsus IV, dorsal view; 22a–b, details of distal structures of tarsus IV: 22a,ventroapical tarsal process below base of pretarsus, and distal minute tarsal setae
|
||
<emphasis id="B971945A270B3D57FD20F872FCFC928A" box="[730,778,1937,1960]" italics="true" pageId="8" pageNumber="9">ad -1</emphasis>
|
||
,
|
||
<emphasis id="B971945A270B3D57FCE2F872FCBE928A" box="[792,840,1937,1960]" italics="true" pageId="8" pageNumber="9">pd -1</emphasis>
|
||
; 22b, pretarsal claws, paradactyli and pulvillus.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF7A18C0270A3D56FF6DF8E8FC0D9281" httpUri="https://zenodo.org/record/195396/files/figure.png" pageId="9" pageNumber="10" targetBox="[163,1436,194,1764]" targetPageId="9">
|
||
<paragraph id="8BBA4848270A3D56FF6DF8E8FC0D9281" blockId="9.[151,1436,1803,1955]" pageId="9" pageNumber="10">
|
||
<emphasis id="B971945A270A3D56FF6DF8E8FE979201" bold="true" box="[151,353,1803,1827]" pageId="9" pageNumber="10">FIGURES 23–31.</emphasis>
|
||
23–29,
|
||
<taxonomicName id="4C0533CB270A3D56FE40F8EFFD569201" box="[442,672,1804,1827]" class="Arachnida" family="Blattisociidae" genus="Opilioseius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="grallator" status="sp. nov.">
|
||
<emphasis id="B971945A270A3D56FE40F8EFFD569201" box="[442,672,1804,1827]" italics="true" pageId="9" pageNumber="10">Opilioseius grallator</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2422921270A3D56FD5DF8E8FD139201" box="[679,741,1803,1827]" pageId="9" pageNumber="10" rank="species">
|
||
<emphasis id="B971945A270A3D56FD5DF8E8FD139201" bold="true" box="[679,741,1803,1827]" pageId="9" pageNumber="10">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, adult: 23, tarsus I, right leg, dorsal view; 24, tarsus I, detail of tarsal seta “S” and associated setae; 25–29, male: 25, chelicera, paraxial view; 26a separate, lateroventral view of movable digit; 26b, separate, lateroventral view of movable digit with scale over dentate region; 27a spermatodactyl with ventral scale; 27b, ventral cheliceral scale; 28, gnathotectum; 29, subcapitulum. 30–31,
|
||
<taxonomicName id="4C0533CB270A3D56FBBEF88FFB4B92A1" box="[1092,1213,1900,1923]" class="Arachnida" family="Podocinidae" genus="Podocinum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B971945A270A3D56FBBEF88FFB4B92A1" box="[1092,1213,1900,1923]" italics="true" pageId="9" pageNumber="10">Podocinum</emphasis>
|
||
</taxonomicName>
|
||
sp., adult: 30, tarsus I, right leg, dorsal view; 31, tarsus I, detail of tarsal seta “S” and associated setae.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BBA484827093D55FF3EFF7BFAC8976E" blockId="10.[151,1436,152,588]" pageId="10" pageNumber="11">
|
||
<emphasis id="B971945A27093D55FF3EFF7BFF0A9593" box="[196,252,152,177]" italics="true" pageId="10" pageNumber="11">Legs</emphasis>
|
||
. Female & male. Legs I to IV with elongated pretarsi bearing paired small claws, paradactyli, and rounded pulvillae (
|
||
<figureCitation id="133E54CD27093D55FE80FF5EFE2295F5" box="[378,468,189,215]" captionStart="FIGURES 17 – 22" captionStartId="8.[151,269,1841,1865]" captionTargetBox="[150,1436,197,1772]" captionTargetId="figure@8.[132,1455,197,1935]" captionTargetPageId="8" captionText="FIGURES 17 – 22. Opilioseius grallator n. sp., adult female: 17 – 20, legs I-IV, excluding tarsi: 17, leg I, dorsal view; 18, leg II, posterolateral view; 19, leg III, posterolateral view; 20, leg IV, posterolateral view with detail of elongate setae on genu and tibia; 21, tarsus IV, dorsal view; 22 a – b, details of distal structures of tarsus IV: 22 a, ventroapical tarsal process below base of pretarsus, and distal minute tarsal setae ad - 1, pd - 1; 22 b, pretarsal claws, paradactyli and pulvillus." httpUri="https://zenodo.org/record/195395/files/figure.png" pageId="10" pageNumber="11">Fig. 22</figureCitation>
|
||
b). Legs elongate, especially I and IV, which two to three times as long as dorsal shield (
|
||
<figureCitation id="133E54CD27093D55FF15FF01FEBE95DE" box="[239,328,226,252]" captionStart="FIGURES 39 – 40" captionStartId="12.[151,272,1939,1963]" captionTargetBox="[221,1378,178,1918]" captionTargetId="figure@12.[221,1379,176,1918]" captionTargetPageId="12" captionText="FIGURES 39 – 40. Opilioseius grallator n. sp.: 39, photomicrograph of adult female, dorsoventral habitus view; 40, photomicrograph of gravid adult female, showing relative size of fully developed egg and maternal idiosoma, dorsoventral habitus view." httpUri="https://zenodo.org/record/195399/files/figure.png" pageId="10" pageNumber="11">Fig. 39</figureCitation>
|
||
); legs II to IV with tarsus three to four times as long as tibia (
|
||
<figureCitation id="133E54CD27093D55FBD6FF01FB7F95DE" box="[1068,1161,226,252]" captionStart="FIGURES 17 – 22" captionStartId="8.[151,269,1841,1865]" captionTargetBox="[150,1436,197,1772]" captionTargetId="figure@8.[132,1455,197,1935]" captionTargetPageId="8" captionText="FIGURES 17 – 22. Opilioseius grallator n. sp., adult female: 17 – 20, legs I-IV, excluding tarsi: 17, leg I, dorsal view; 18, leg II, posterolateral view; 19, leg III, posterolateral view; 20, leg IV, posterolateral view with detail of elongate setae on genu and tibia; 21, tarsus IV, dorsal view; 22 a – b, details of distal structures of tarsus IV: 22 a, ventroapical tarsal process below base of pretarsus, and distal minute tarsal setae ad - 1, pd - 1; 22 b, pretarsal claws, paradactyli and pulvillus." httpUri="https://zenodo.org/record/195395/files/figure.png" pageId="10" pageNumber="11">Figs 21</figureCitation>
|
||
,
|
||
<figureCitation id="133E54CD27093D55FB6CFF01FB4E95DE" box="[1174,1208,226,252]" captionStart="FIGURES 39 – 40" captionStartId="12.[151,272,1939,1963]" captionTargetBox="[221,1378,178,1918]" captionTargetId="figure@12.[221,1379,176,1918]" captionTargetPageId="12" captionText="FIGURES 39 – 40. Opilioseius grallator n. sp.: 39, photomicrograph of adult female, dorsoventral habitus view; 40, photomicrograph of gravid adult female, showing relative size of fully developed egg and maternal idiosoma, dorsoventral habitus view." httpUri="https://zenodo.org/record/195399/files/figure.png" pageId="10" pageNumber="11">39</figureCitation>
|
||
). Distal margins of coxae I-IV lacking prominent serrations or spur-like processes. Tarsus I with sensilla
|
||
<emphasis id="B971945A27093D55FB5DFEEBFB409403" box="[1191,1206,264,289]" italics="true" pageId="10" pageNumber="11">S</emphasis>
|
||
slightly lanceolate apically (
|
||
<figureCitation id="133E54CD27093D55FEF9FECEFEAD9465" box="[259,347,301,327]" captionStart="FIGURES 23 – 31. 23 – 29" captionStartId="9.[151,271,1803,1827]" captionTargetBox="[163,1436,194,1764]" captionTargetId="figure@9.[163,1448,194,1782]" captionTargetPageId="9" captionText="FIGURES 23 – 31. 23 – 29, Opilioseius grallator n. sp., adult: 23, tarsus I, right leg, dorsal view; 24, tarsus I, detail of tarsal seta “ S ” and associated setae; 25 – 29, male: 25, chelicera, paraxial view; 26 a separate, lateroventral view of movable digit; 26 b, separate, lateroventral view of movable digit with scale over dentate region; 27 a spermatodactyl with ventral scale; 27 b, ventral cheliceral scale; 28, gnathotectum; 29, subcapitulum. 30 – 31, Podocinum sp., adult: 30, tarsus I, right leg, dorsal view; 31, tarsus I, detail of tarsal seta “ S ” and associated setae." httpUri="https://zenodo.org/record/195396/files/figure.png" pageId="10" pageNumber="11">Fig. 24</figureCitation>
|
||
), and without markedly elongated setae apically. Tarsi II–IV with apical setal processes
|
||
<emphasis id="B971945A27093D55FAA5FECEFA639464" box="[1375,1429,301,327]" italics="true" pageId="10" pageNumber="11">ad -1</emphasis>
|
||
,
|
||
<emphasis id="B971945A27093D55FF6DFEB1FF3B9449" box="[151,205,338,364]" italics="true" pageId="10" pageNumber="11">pd -1</emphasis>
|
||
short, about 1/10th as long as pretarsi, and with slender, acute apical process ventrally (
|
||
<figureCitation id="133E54CD27093D55FB0EFEB1FA76944E" box="[1268,1408,338,364]" captionStart="FIGURES 17 – 22" captionStartId="8.[151,269,1841,1865]" captionTargetBox="[150,1436,197,1772]" captionTargetId="figure@8.[132,1455,197,1935]" captionTargetPageId="8" captionText="FIGURES 17 – 22. Opilioseius grallator n. sp., adult female: 17 – 20, legs I-IV, excluding tarsi: 17, leg I, dorsal view; 18, leg II, posterolateral view; 19, leg III, posterolateral view; 20, leg IV, posterolateral view with detail of elongate setae on genu and tibia; 21, tarsus IV, dorsal view; 22 a – b, details of distal structures of tarsus IV: 22 a, ventroapical tarsal process below base of pretarsus, and distal minute tarsal setae ad - 1, pd - 1; 22 b, pretarsal claws, paradactyli and pulvillus." httpUri="https://zenodo.org/record/195395/files/figure.png" pageId="10" pageNumber="11">Figs 21, 22</figureCitation>
|
||
a). Complement of setae on segments of legs I-II-III-IV normal for Blattisociinae as presented by
|
||
<bibRefCitation id="EF9435B927093D55FAF1FE9BFECE9495" author="Lindquist" pageId="10" pageNumber="20" refString="Lindquist, E. E. & Evans, G. O. (1965) Taxonomic concepts in the Ascidae, with a modified setal nomenclature for the idiosoma of the Gamasina (Acarina: Mesostigmata). Memoirs of the Entomological Society of Canada, 47, 1 - 64." type="journal article" year="1965">Lindquist & Evans (1965)</bibRefCitation>
|
||
: coxae, 2-2-2-1; trochanters, 6-5-5-5; femora, (2 3/1 2/2 2) - (1 3/1 2/2 1) - (1 2/1 1/0 1) - (1 2/0 1/1 1); genua, (2 3/1 3/1 2) - (2 3/1 2/0 2) - (2 2/1 2/0 1) - (2 2/1 3/0 1); tibiae, (2 3/1 3/1 2) - (2 2/1 2/1 2) - (2 1/1 2/1 1) - (2 1/1 3/1 2); femur II lacking
|
||
<emphasis id="B971945A27093D55FD64FE0BFD389723" box="[670,718,488,514]" italics="true" pageId="10" pageNumber="11">al -2</emphasis>
|
||
; genu I and tibia I each lacking
|
||
<emphasis id="B971945A27093D55FBA3FE0BFB789723" box="[1113,1166,488,514]" italics="true" pageId="10" pageNumber="11">av -2</emphasis>
|
||
; genua II and III each lacking
|
||
<emphasis id="B971945A27093D55FF0CFDEEFEDD9704" box="[246,299,525,551]" italics="true" pageId="10" pageNumber="11">pv -1</emphasis>
|
||
(
|
||
<figureCitation id="133E54CD27093D55FEC6FDEEFE379705" box="[316,449,525,551]" captionStart="FIGURES 17 – 22" captionStartId="8.[151,269,1841,1865]" captionTargetBox="[150,1436,197,1772]" captionTargetId="figure@8.[132,1455,197,1935]" captionTargetPageId="8" captionText="FIGURES 17 – 22. Opilioseius grallator n. sp., adult female: 17 – 20, legs I-IV, excluding tarsi: 17, leg I, dorsal view; 18, leg II, posterolateral view; 19, leg III, posterolateral view; 20, leg IV, posterolateral view with detail of elongate setae on genu and tibia; 21, tarsus IV, dorsal view; 22 a – b, details of distal structures of tarsus IV: 22 a, ventroapical tarsal process below base of pretarsus, and distal minute tarsal setae ad - 1, pd - 1; 22 b, pretarsal claws, paradactyli and pulvillus." httpUri="https://zenodo.org/record/195395/files/figure.png" pageId="10" pageNumber="11">Figs 17-21</figureCitation>
|
||
). Leg setae collectively smooth, attenuated, some elongated on tibia III, femur IV, and especially on genu, tibia and tarsus of leg IV. Legs of male without dimorphically modified setae.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF7A18C027093D55FF6DF85FFC1B92D1" httpUri="https://zenodo.org/record/195397/files/figure.png" pageId="10" pageNumber="11" targetBox="[154,1436,642,1930]" targetPageId="10">
|
||
<paragraph id="8BBA484827093D55FF6DF85FFC1B92D1" blockId="10.[151,1436,1980,2035]" pageId="10" pageNumber="11">
|
||
<emphasis id="B971945A27093D55FF6DF85FFEA892F6" bold="true" box="[151,350,1980,2004]" pageId="10" pageNumber="11">FIGURES 32–33.</emphasis>
|
||
<taxonomicName id="4C0533CB27093D55FE9EF85FFDB292F1" box="[356,580,1980,2003]" class="Arachnida" family="Blattisociidae" genus="Opilioseius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="grallator" status="sp. nov.">
|
||
<emphasis id="B971945A27093D55FE9EF85FFDB292F1" box="[356,580,1980,2003]" italics="true" pageId="10" pageNumber="11">Opilioseius grallator</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A242292127093D55FDB6F85FFD7192F6" box="[588,647,1980,2004]" pageId="10" pageNumber="11" rank="species">
|
||
<emphasis id="B971945A27093D55FDB6F85FFD7192F6" bold="true" box="[588,647,1980,2004]" pageId="10" pageNumber="11">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, adult male: 32, idiosoma, dorsal aspect, with laterally expanded detail of peritrematal shield fused to podonotal dorsal region; 33, idiosoma, ventral view.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF7A18C027083D54FF6DF8FAFB1592B3" httpUri="https://zenodo.org/record/195398/files/figure.png" pageId="11" pageNumber="12" targetBox="[157,1436,204,1770]" targetPageId="11">
|
||
<paragraph id="8BBA484827083D54FF6DF8FAFB1592B3" blockId="11.[151,1436,1817,1937]" pageId="11" pageNumber="12">
|
||
<emphasis id="B971945A27083D54FF6DF8FAFEA99213" bold="true" box="[151,351,1817,1841]" pageId="11" pageNumber="12">FIGURES 34–38.</emphasis>
|
||
34–35,
|
||
<taxonomicName id="4C0533CB27083D54FE42F8F9FD6D9213" box="[440,667,1818,1841]" class="Arachnida" family="Blattisociidae" genus="Opilioseius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="grallator" status="sp. nov.">
|
||
<emphasis id="B971945A27083D54FE42F8F9FD6D9213" box="[440,667,1818,1841]" italics="true" pageId="11" pageNumber="12">Opilioseius grallator</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A242292127083D54FD58F8FAFD299213" box="[674,735,1817,1841]" pageId="11" pageNumber="12" rank="species">
|
||
<emphasis id="B971945A27083D54FD58F8FAFD299213" bold="true" box="[674,735,1817,1841]" pageId="11" pageNumber="12">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, deutonymph: 34, idiosoma, dorsal aspect; 35, idiosoma, ventral aspect. 36–38, leg III, tarsal apex and pretarsus, showing different forms of ventroapical tarsal processes and conjectured acrotarsal demarcation: 36,
|
||
<emphasis id="B971945A27083D54FE3BF8B9FDCC9253" box="[449,570,1882,1905]" italics="true" pageId="11" pageNumber="12">Adenoseius</emphasis>
|
||
sp. (
|
||
<taxonomicName id="4C0533CB27083D54FD94F8B9FD349253" box="[622,706,1882,1905]" class="Arachnida" family="Sejidae" kingdom="Animalia" order="Mesostigmata" pageId="11" pageNumber="12" phylum="Arthropoda" rank="family">Sejidae</taxonomicName>
|
||
), anterolateral view; 37,
|
||
<taxonomicName id="4C0533CB27083D54FC37F8B9FBC69253" box="[973,1072,1882,1905]" class="Arachnida" family="Euzerconidae" genus="Euzercon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B971945A27083D54FC37F8B9FBC69253" box="[973,1072,1882,1905]" italics="true" pageId="11" pageNumber="12">Euzercon</emphasis>
|
||
</taxonomicName>
|
||
sp. (
|
||
<taxonomicName id="4C0533CB27083D54FB9CF8B9FB0B9253" box="[1126,1277,1882,1905]" class="Arachnida" family="Euzerconidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="11" pageNumber="12" phylum="Arthropoda" rank="family">Euzerconidae</taxonomicName>
|
||
), ventral view; 38,
|
||
<taxonomicName id="4C0533CB27083D54FF45F899FD7392B3" authority="Evans" authorityName="Evans" box="[191,645,1914,1937]" class="Arachnida" family="Arctacaridae" genus="Arctacarus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="rostratus">
|
||
<emphasis id="B971945A27083D54FF45F899FE6C92B3" box="[191,410,1914,1937]" italics="true" pageId="11" pageNumber="12">Arctacarus rostratus</emphasis>
|
||
Evans (Arctacaridae)
|
||
</taxonomicName>
|
||
, ventral view. Abbreviation:
|
||
<emphasis id="B971945A27083D54FC41F89AFBE292B3" bold="true" box="[955,1044,1913,1937]" pageId="11" pageNumber="12">acr mar</emphasis>
|
||
, acrotarsal margin.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF7A18C0270F3D53FF6DF870FE4692C9" httpUri="https://zenodo.org/record/195399/files/figure.png" pageId="12" pageNumber="13" targetBox="[221,1378,178,1918]" targetPageId="12">
|
||
<paragraph id="8BBA4848270F3D53FF6DF870FE4692C9" blockId="12.[151,1437,1939,2027]" pageId="12" pageNumber="13">
|
||
<emphasis id="B971945A270F3D53FF6DF870FE959289" bold="true" box="[151,355,1939,1963]" pageId="12" pageNumber="13">FIGURES 39–40.</emphasis>
|
||
<taxonomicName id="4C0533CB270F3D53FE90F877FDA49289" box="[362,594,1940,1963]" class="Arachnida" family="Blattisociidae" genus="Opilioseius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="grallator" status="sp. nov.">
|
||
<emphasis id="B971945A270F3D53FE90F877FDA49289" box="[362,594,1940,1963]" italics="true" pageId="12" pageNumber="13">Opilioseius grallator</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2422921270F3D53FDA0F870FD6F9289" box="[602,665,1939,1963]" pageId="12" pageNumber="13" rank="species">
|
||
<emphasis id="B971945A270F3D53FDA0F870FD6F9289" bold="true" box="[602,665,1939,1963]" pageId="12" pageNumber="13">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
: 39, photomicrograph of adult female, dorsoventral habitus view; 40, photomicrograph of gravid adult female, showing relative size of fully developed egg and maternal idiosoma, dorsoventral habitus view.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C31F1BC3270E3D52FF3EFF74FCDF9465" pageId="13" pageNumber="14" type="etymology">
|
||
<paragraph id="8BBA4848270E3D52FF3EFF74FCDF9465" blockId="13.[151,1437,151,1858]" pageId="13" pageNumber="14">
|
||
<emphasis id="B971945A270E3D52FF3EFF74FEA79593" bold="true" box="[196,337,151,177]" pageId="13" pageNumber="14">Etymology.</emphasis>
|
||
The name of the genus is a Latinized combination of the term
|
||
<emphasis id="B971945A270E3D52FBD3FF7BFB9B9593" box="[1065,1133,152,177]" italics="true" pageId="13" pageNumber="14">opilio</emphasis>
|
||
, meaning “shepherd” and applied to the arachnid order Opiliones whose members are typically long-legged, and
|
||
<emphasis id="B971945A270E3D52FB71FF5EFB3095F4" box="[1163,1222,189,214]" italics="true" pageId="13" pageNumber="14">seius</emphasis>
|
||
or
|
||
<emphasis id="B971945A270E3D52FB11FF5EFAD095F4" box="[1259,1318,189,214]" italics="true" pageId="13" pageNumber="14">sejus</emphasis>
|
||
, a Roman surname (also meaning “divine” or “a wanderer”, according to
|
||
<bibRefCitation id="EF9435B9270E3D52FC5DFF01FB1695DE" author="Chant" box="[935,1248,226,252]" pageId="13" pageNumber="19" refString="Chant, D. A. & McMurtry, J. A. (1994) A review of the subfamilies Phytoseiinae and Typhlodrominae (Acari: Phytoseiidae). International Journal of Acarology, 20, 222 - 310." type="journal article" year="1994">Chant & McMurtry 1994</bibRefCitation>
|
||
) used by many authors to form names for genera of mesostigmatic mites. The name is masculine in gender, and is intended to refer to the remarkably long-legged form of these mites.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31F1BC3270E3D52FF3CFEB1FB3E94B0" pageId="13" pageNumber="14" type="distribution">
|
||
<paragraph id="8BBA4848270E3D52FF3CFEB1FB3E94B0" blockId="13.[151,1437,151,1858]" pageId="13" pageNumber="14">
|
||
<emphasis id="B971945A270E3D52FF3CFEB1FDFA944E" bold="true" box="[198,524,338,364]" pageId="13" pageNumber="14">Distribution and habitats.</emphasis>
|
||
This monobasic genus is based on material associated with perennial fungal growths on decaying logs from just one locality in coastal lowland rainforest of
|
||
<collectingCountry id="F31208D8270E3D52FBBAFE9BFB3294B0" box="[1088,1220,376,402]" name="Costa Rica" pageId="13" pageNumber="14">Costa Rica</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31F1BC3270E3D52FF3CFE7EFEB69260" pageId="13" pageNumber="14" type="discussion">
|
||
<paragraph id="8BBA4848270E3D52FF3CFE7EFCA497B5" blockId="13.[151,1437,151,1858]" pageId="13" pageNumber="14">
|
||
<emphasis id="B971945A270E3D52FF3CFE7EFEB69495" bold="true" box="[198,320,413,439]" pageId="13" pageNumber="14">Remarks.</emphasis>
|
||
The genus
|
||
<taxonomicName id="4C0533CB270E3D52FE2BFE7EFDA19494" box="[465,599,413,438]" class="Arachnida" family="Blattisociidae" genus="Opilioseius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="13" pageNumber="4" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B971945A270E3D52FE2BFE7EFDA19494" box="[465,599,413,438]" italics="true" pageId="13" pageNumber="14">Opilioseius</emphasis>
|
||
</taxonomicName>
|
||
is based on several apomorphies, four of which appear to be unique among the known taxa of
|
||
<taxonomicName id="4C0533CB270E3D52FE26FE21FD7A94FE" box="[476,652,450,476]" class="Arachnida" family="Blattisociidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="13" pageNumber="14" phylum="Arthropoda" rank="family">Blattisociidae</taxonomicName>
|
||
: (1) absence of paravertical lyrifissures
|
||
<emphasis id="B971945A270E3D52FB84FE21FB5994F9" box="[1150,1199,450,475]" italics="true" pageId="13" pageNumber="14">idj1</emphasis>
|
||
on the idiosomatic dorsum; (2) great elongation of legs I to IV, and especially of the tarsal segments of legs II to IV; (3) reduction in size of the male ventrianal shield; and (4) groove-like structures on the paraxial surface of the movable digit of the chelicera. However, the uniqueness of the cheliceral structures at the genus level is problematic, as we have observed bizarrely modified cheliceral digits among various undescribed species of
|
||
<taxonomicName id="4C0533CB270E3D52FB55FDBBFADD9753" box="[1199,1323,600,625]" class="Arachnida" family="Ascidae" genus="Lasioseius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B971945A270E3D52FB55FDBBFADD9753" box="[1199,1323,600,625]" italics="true" pageId="13" pageNumber="14">Lasioseius</emphasis>
|
||
</taxonomicName>
|
||
found on bracket fungi, including some that coexist with
|
||
<taxonomicName id="4C0533CB270E3D52FD3FFD9EFCBC97B4" box="[709,842,637,662]" class="Arachnida" family="Blattisociidae" genus="Opilioseius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="13" pageNumber="4" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B971945A270E3D52FD3FFD9EFCBC97B4" box="[709,842,637,662]" italics="true" pageId="13" pageNumber="14">Opilioseius</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BBA4848270E3D52FF3CFD41FDA896E0" blockId="13.[151,1437,151,1858]" pageId="13" pageNumber="14">
|
||
The apomorphic similarities shared between
|
||
<taxonomicName id="4C0533CB270E3D52FD27FD41FC959799" box="[733,867,674,699]" class="Arachnida" family="Blattisociidae" genus="Opilioseius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="13" pageNumber="4" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B971945A270E3D52FD27FD41FC959799" box="[733,867,674,699]" italics="true" pageId="13" pageNumber="14">Opilioseius</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C0533CB270E3D52FC65FD41FBC79799" box="[927,1073,674,699]" class="Arachnida" family="Blattisociidae" genus="Aceodromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B971945A270E3D52FC65FD41FBC79799" box="[927,1073,674,699]" italics="true" pageId="13" pageNumber="14">Aceodromus</emphasis>
|
||
</taxonomicName>
|
||
, as given in the diagnosis and key, are all subject to homoplasy and may be convergences, as members of these two genera are adapted to very different habitats and substrates. While those of
|
||
<taxonomicName id="4C0533CB270E3D52FCF3FD0EFC789624" box="[777,910,749,774]" class="Arachnida" family="Blattisociidae" genus="Opilioseius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="13" pageNumber="4" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B971945A270E3D52FCF3FD0EFC789624" box="[777,910,749,774]" italics="true" pageId="13" pageNumber="14">Opilioseius</emphasis>
|
||
</taxonomicName>
|
||
are restricted to perennial fungi on decaying logs, members of the four species of
|
||
<taxonomicName id="4C0533CB270E3D52FDB1FCF1FD2A9609" box="[587,732,786,811]" class="Arachnida" family="Blattisociidae" genus="Aceodromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B971945A270E3D52FDB1FCF1FD2A9609" box="[587,732,786,811]" italics="true" pageId="13" pageNumber="14">Aceodromus</emphasis>
|
||
</taxonomicName>
|
||
known to us (two of them undescribed) occupy herbaceous and grassy habitats, where they may be associated with copiphorine tettigoniids, on which they are phoretic as adult females (
|
||
<bibRefCitation id="EF9435B9270E3D52FEBDFCBEFD999655" author="Lindquist" box="[327,623,861,887]" pageId="13" pageNumber="20" refString="Lindquist, E. E. & Chant, D. A. (1964) A redescription of Aceodromus Muma and its transfer to the Blattisociinae (Acarina: Blattosociidae). Canadian Entomologist, 96, 500 - 507." type="journal article" year="1964">Lindquist & Chant 1964</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EF9435B9270E3D52FD86FCBEFC509655" author="Lindquist" box="[636,934,861,887]" pageId="13" pageNumber="20" refString="Lindquist, E. E. & Evans, G. O. (1965) Taxonomic concepts in the Ascidae, with a modified setal nomenclature for the idiosoma of the Gamasina (Acarina: Mesostigmata). Memoirs of the Entomological Society of Canada, 47, 1 - 64." type="journal article" year="1965">Lindquist & Evans 1965</bibRefCitation>
|
||
).
|
||
<taxonomicName id="4C0533CB270E3D52FC41FCBEFB439654" box="[955,1205,861,886]" class="Arachnida" family="Blattisociidae" genus="Opilioseius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="grallator">
|
||
<emphasis id="B971945A270E3D52FC41FCBEFB439654" box="[955,1205,861,886]" italics="true" pageId="13" pageNumber="14">Opilioseius grallator</emphasis>
|
||
</taxonomicName>
|
||
is not known to be phoretic, and it is difficult to envisage phoretic behavior among mites with such ungainly long legs and relatively slow movement (see below).
|
||
</paragraph>
|
||
<paragraph id="8BBA4848270E3D52FF3CFC2EFBA691E5" blockId="13.[151,1437,151,1858]" pageId="13" pageNumber="14">
|
||
Adults of the
|
||
<emphasis id="B971945A270E3D52FE97FC2EFDFB96C4" box="[365,525,973,998]" italics="true" pageId="13" pageNumber="14">phytoseioides</emphasis>
|
||
species group
|
||
<emphasis id="B971945A270E3D52FD3BFC2EFCF496C4" box="[705,770,973,998]" italics="true" pageId="13" pageNumber="14">sensu</emphasis>
|
||
Lindquist (1964) in the genus
|
||
<taxonomicName id="4C0533CB270E3D52FB82FC2EFB0296C4" box="[1144,1268,973,998]" class="Arachnida" family="Ascidae" genus="Lasioseius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B971945A270E3D52FB82FC2EFB0296C4" box="[1144,1268,973,998]" italics="true" pageId="13" pageNumber="14">Lasioseius</emphasis>
|
||
</taxonomicName>
|
||
are similar to those of
|
||
<taxonomicName id="4C0533CB270E3D52FEFCFC11FE549129" box="[262,418,1010,1035]" class="Arachnida" family="Blattisociidae" genus="Aceodromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B971945A270E3D52FEFCFC11FE549129" box="[262,418,1010,1035]" italics="true" pageId="13" pageNumber="14">Aceodromus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C0533CB270E3D52FE18FC11FD859129" box="[482,627,1010,1035]" class="Arachnida" family="Blattisociidae" genus="Opilioseius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="13" pageNumber="4" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B971945A270E3D52FE18FC11FD859129" box="[482,627,1010,1035]" italics="true" pageId="13" pageNumber="14">Opilioseius</emphasis>
|
||
</taxonomicName>
|
||
in the hypotrichous condition of the dorsal shield, which lacks podonotal setae
|
||
<emphasis id="B971945A270E3D52FE9CFBFBFE759113" box="[358,387,1048,1073]" italics="true" pageId="13" pageNumber="14">z1</emphasis>
|
||
,
|
||
<emphasis id="B971945A270E3D52FE6EFBFBFE469113" box="[404,432,1048,1073]" italics="true" pageId="13" pageNumber="14">z3</emphasis>
|
||
,
|
||
<emphasis id="B971945A270E3D52FE3BFBFBFE2B9113" box="[449,477,1048,1073]" italics="true" pageId="13" pageNumber="14">z6</emphasis>
|
||
,
|
||
<emphasis id="B971945A270E3D52FE14FBFBFDFC9113" box="[494,522,1048,1073]" italics="true" pageId="13" pageNumber="14">s3</emphasis>
|
||
,
|
||
<emphasis id="B971945A270E3D52FDE6FBFBFDCE9113" box="[540,568,1048,1073]" italics="true" pageId="13" pageNumber="14">r2</emphasis>
|
||
,
|
||
<emphasis id="B971945A270E3D52FDB3FBFBFD909113" box="[585,614,1048,1073]" italics="true" pageId="13" pageNumber="14">r4</emphasis>
|
||
and opisthonotal setae
|
||
<emphasis id="B971945A270E3D52FC69FBFBFC469113" box="[915,944,1048,1073]" italics="true" pageId="13" pageNumber="14">J1</emphasis>
|
||
,
|
||
<emphasis id="B971945A270E3D52FC38FBFBFC169113" box="[962,992,1048,1073]" italics="true" pageId="13" pageNumber="14">J3</emphasis>
|
||
,
|
||
<emphasis id="B971945A270E3D52FC0BFBFBFBE69113" box="[1009,1040,1048,1073]" italics="true" pageId="13" pageNumber="14">S1</emphasis>
|
||
. They are especially similar to
|
||
<taxonomicName id="4C0533CB270E3D52FF6DFBDEFEDF9174" box="[151,297,1085,1110]" class="Arachnida" family="Blattisociidae" genus="Aceodromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B971945A270E3D52FF6DFBDEFEDF9174" box="[151,297,1085,1110]" italics="true" pageId="13" pageNumber="14">Aceodromus</emphasis>
|
||
</taxonomicName>
|
||
in lacking the true vertical setae,
|
||
<emphasis id="B971945A270E3D52FD49FBDEFD3C9174" box="[691,714,1085,1110]" italics="true" pageId="13" pageNumber="14">j1</emphasis>
|
||
, and also
|
||
<emphasis id="B971945A270E3D52FCBAFBDEFCAF9174" box="[832,857,1085,1110]" italics="true" pageId="13" pageNumber="14">s1</emphasis>
|
||
,
|
||
<emphasis id="B971945A270E3D52FC92FBDEFC759174" box="[872,899,1085,1110]" italics="true" pageId="13" pageNumber="14">s2</emphasis>
|
||
. However, these similarities are thought to be convergences, with the
|
||
<emphasis id="B971945A270E3D52FE51FB80FDBC915E" box="[427,586,1123,1148]" italics="true" pageId="13" pageNumber="14">phytoseioides</emphasis>
|
||
group being derived from a lineage in
|
||
<taxonomicName id="4C0533CB270E3D52FBEEFB80FB66915E" box="[1044,1168,1123,1148]" class="Arachnida" family="Ascidae" genus="Lasioseius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B971945A270E3D52FBEEFB80FB66915E" box="[1044,1168,1123,1148]" italics="true" pageId="13" pageNumber="14">Lasioseius</emphasis>
|
||
</taxonomicName>
|
||
which is characterised in part by having heavily sclerotized dorsal and ventral shields, the female ventrianal shield expansive and bearing six pairs of opisthogastric setae, and the female movable chela tridentate.
|
||
</paragraph>
|
||
<paragraph id="8BBA4848270E3D52FF3CFB31FEB69260" blockId="13.[151,1437,151,1858]" pageId="13" pageNumber="14">
|
||
Adults of the family
|
||
<taxonomicName id="4C0533CB270E3D52FE40FB31FDBD91CE" box="[442,587,1234,1260]" class="Arachnida" family="Podocinidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="13" pageNumber="14" phylum="Arthropoda" rank="family">Podocinidae</taxonomicName>
|
||
resemble those of
|
||
<taxonomicName id="4C0533CB270E3D52FCD2FB30FC5B91CE" box="[808,941,1235,1260]" class="Arachnida" family="Blattisociidae" genus="Opilioseius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="13" pageNumber="4" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B971945A270E3D52FCD2FB30FC5B91CE" box="[808,941,1235,1260]" italics="true" pageId="13" pageNumber="14">Opilioseius</emphasis>
|
||
</taxonomicName>
|
||
in having long legs, and especially legs II to IV having elongated tarsi that may be three to five times as long as each of the respective tibiae and genua. Legs I of podocinids, however, are disproportionately long relative to legs II to IV, lack claws, and have one or two greatly elongated tactile setae apically, such that their function appears to be strictly tactile, antenna-like, rather than optionally ambulatory. And, while elongation of tarsus I in
|
||
<taxonomicName id="4C0533CB270E3D52FC01FA8BFB7090A3" box="[1019,1158,1384,1409]" class="Arachnida" family="Blattisociidae" genus="Opilioseius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="13" pageNumber="4" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B971945A270E3D52FC01FA8BFB7090A3" box="[1019,1158,1384,1409]" italics="true" pageId="13" pageNumber="14">Opilioseius</emphasis>
|
||
</taxonomicName>
|
||
occurs along its entire length (
|
||
<figureCitation id="133E54CD270E3D52FF0BFA6EFEBE9085" box="[241,328,1421,1447]" captionStart="FIGURES 23 – 31. 23 – 29" captionStartId="9.[151,271,1803,1827]" captionTargetBox="[163,1436,194,1764]" captionTargetId="figure@9.[163,1448,194,1782]" captionTargetPageId="9" captionText="FIGURES 23 – 31. 23 – 29, Opilioseius grallator n. sp., adult: 23, tarsus I, right leg, dorsal view; 24, tarsus I, detail of tarsal seta “ S ” and associated setae; 25 – 29, male: 25, chelicera, paraxial view; 26 a separate, lateroventral view of movable digit; 26 b, separate, lateroventral view of movable digit with scale over dentate region; 27 a spermatodactyl with ventral scale; 27 b, ventral cheliceral scale; 28, gnathotectum; 29, subcapitulum. 30 – 31, Podocinum sp., adult: 30, tarsus I, right leg, dorsal view; 31, tarsus I, detail of tarsal seta “ S ” and associated setae." httpUri="https://zenodo.org/record/195396/files/figure.png" pageId="13" pageNumber="14">Fig. 23</figureCitation>
|
||
), that in podocinids differs by the elongation being restricted primarily to the region proximal to the sensory receptor complex, i.e., the region that bears only attenuated solid-core setae (delimited by the angled line in
|
||
<figureCitation id="133E54CD270E3D52FEBFFA3BFE6990D0" box="[325,415,1496,1522]" captionStart="FIGURES 23 – 31. 23 – 29" captionStartId="9.[151,271,1803,1827]" captionTargetBox="[163,1436,194,1764]" captionTargetId="figure@9.[163,1448,194,1782]" captionTargetPageId="9" captionText="FIGURES 23 – 31. 23 – 29, Opilioseius grallator n. sp., adult: 23, tarsus I, right leg, dorsal view; 24, tarsus I, detail of tarsal seta “ S ” and associated setae; 25 – 29, male: 25, chelicera, paraxial view; 26 a separate, lateroventral view of movable digit; 26 b, separate, lateroventral view of movable digit with scale over dentate region; 27 a spermatodactyl with ventral scale; 27 b, ventral cheliceral scale; 28, gnathotectum; 29, subcapitulum. 30 – 31, Podocinum sp., adult: 30, tarsus I, right leg, dorsal view; 31, tarsus I, detail of tarsal seta “ S ” and associated setae." httpUri="https://zenodo.org/record/195396/files/figure.png" pageId="13" pageNumber="14">Fig. 30</figureCitation>
|
||
). Also, in contrast to tarsus I of
|
||
<taxonomicName id="4C0533CB270E3D52FCD3FA3BFC4490D3" box="[809,946,1496,1521]" class="Arachnida" family="Blattisociidae" genus="Opilioseius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="13" pageNumber="4" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B971945A270E3D52FCD3FA3BFC4490D3" box="[809,946,1496,1521]" italics="true" pageId="13" pageNumber="14">Opilioseius</emphasis>
|
||
</taxonomicName>
|
||
having a complete complement of setae typical of the Dermanyssina (6-28/9-6), that of podocinids is setal-deficient in both regions, having a larval complement of setae (along with one protonymphal seta added) in the proximal region and a few less ventral setae in the sensory receptor complex region. Legs II to IV of podocinids have short pretarsi relative to those of
|
||
<taxonomicName id="4C0533CB270E3D52FF43F98EFEB393A4" box="[185,325,1645,1670]" class="Arachnida" family="Blattisociidae" genus="Opilioseius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="13" pageNumber="4" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B971945A270E3D52FF43F98EFEB393A4" box="[185,325,1645,1670]" italics="true" pageId="13" pageNumber="14">Opilioseius</emphasis>
|
||
</taxonomicName>
|
||
. Also, unlike
|
||
<taxonomicName id="4C0533CB270E3D52FE0CF98EFD7493A4" box="[502,642,1645,1670]" class="Arachnida" family="Blattisociidae" genus="Opilioseius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="13" pageNumber="4" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B971945A270E3D52FE0CF98EFD7493A4" box="[502,642,1645,1670]" italics="true" pageId="13" pageNumber="14">Opilioseius</emphasis>
|
||
</taxonomicName>
|
||
and other blattisociids, podocinids have a three-tined palptarsal apotele, a serrated transverse ridge across the dorsal face of the gnathotectum, a more reduced dorsal body setation that lacks clunal setae
|
||
<emphasis id="B971945A270E3D52FDEEF95BFDC793F3" box="[532,561,1720,1745]" italics="true" pageId="13" pageNumber="14">J5</emphasis>
|
||
and all
|
||
<emphasis id="B971945A270E3D52FD6EF95BFD5093F3" box="[660,678,1720,1745]" italics="true" pageId="13" pageNumber="14">R</emphasis>
|
||
- marginal setae, and their males have a holoventral shield. In contrast, legs I of
|
||
<taxonomicName id="4C0533CB270E3D52FE96F93EFE0493D4" box="[364,498,1757,1782]" class="Arachnida" family="Blattisociidae" genus="Opilioseius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="13" pageNumber="4" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B971945A270E3D52FE96F93EFE0493D4" box="[364,498,1757,1782]" italics="true" pageId="13" pageNumber="14">Opilioseius</emphasis>
|
||
</taxonomicName>
|
||
are similar in elongated length to legs IV, lack greatly elongated setae apically, are equipped with small paired claws on long pretarsi like legs II to IV, and are thought to be ambulatory as well as tactile.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |