352 lines
39 KiB
XML
352 lines
39 KiB
XML
<document id="8776B6DEAD081C34540BF36DD24091B9" ID-CLB-Dataset="45702" ID-DOI="10.5281/zenodo.214356" ID-GBIF-Dataset="2ccb32a4-9476-4a35-9231-e9849f5ed473" ID-ISSN="1175-5326" ID-Zenodo-Dep="214356" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460424207514" checkinUser="plazi" docAuthor="Michalczyk, Łukasz, Wełnicz, Weronika, Frohme, Marcus & Kaczmarek, Łukasz" docDate="2012" docId="03A987E9733FFFCC1AB77325FED2E76D" docLanguage="en" docName="zt03154p020.pdf" docOrigin="Zootaxa 3154" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Milnesium granulatum Ramazzotti 1962" docType="treatment" docVersion="9" lastPageNumber="11" masterDocId="FF90FF917337FFC61A207425FFD1E261" masterDocTitle="Redescriptions of three Milnesium Doyère, 1840 taxa (Tardigrada: Eutardigrada: Milnesiidae), including the nominal species for the genus" masterLastPageNumber="20" masterPageNumber="1" pageNumber="9" updateTime="1698330435667" updateUser="ExternalLinkService">
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<mods:title id="3BA625AEB1322B204B97696A34524CE4">Redescriptions of three Milnesium Doyère, 1840 taxa (Tardigrada: Eutardigrada: Milnesiidae), including the nominal species for the genus</mods:title>
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<mods:namePart id="F79C4490166AC75D610B5CDD0A08A228">Michalczyk, Łukasz</mods:namePart>
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<mods:namePart id="BDAC7F4A08381C7130B98A4F23D84561">Wełnicz, Weronika</mods:namePart>
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<mods:namePart id="B5F26ABB1925BFEFAAC15AA3D0E3A506">Frohme, Marcus</mods:namePart>
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<mods:namePart id="38EAC37D08BF5C15CB35C11E1EB92413">Kaczmarek, Łukasz</mods:namePart>
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<mods:date id="85C2A4F65E71D5CC4ABA2E5341B506B1">2012</mods:date>
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<mods:number id="BF6A7C6D273068DF65A8529912FB8609">3154</mods:number>
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<treatment id="03A987E9733FFFCC1AB77325FED2E76D" ID-DOI="http://doi.org/10.5281/zenodo.5612863" ID-GBIF-Taxon="119428117" ID-Zenodo-Dep="5612863" LSID="urn:lsid:plazi:treatment:03A987E9733FFFCC1AB77325FED2E76D" httpUri="http://treatment.plazi.org/id/03A987E9733FFFCC1AB77325FED2E76D" lastPageId="10" lastPageNumber="11" pageId="8" pageNumber="9">
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<subSubSection id="C31A6574733FFFCE1AB77325FEA8E55A" pageId="8" pageNumber="9" type="nomenclature">
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<paragraph id="8BBF36FF733FFFCE1AB77325FCE8E57B" blockId="8.[151,825,1792,1851]" box="[151,825,1792,1818]" pageId="8" pageNumber="9">
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<heading id="D0F78193733FFFCE1AB77325FCE8E57B" bold="true" box="[151,825,1792,1818]" fontSize="11" level="1" pageId="8" pageNumber="9" reason="1">
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<emphasis id="B974EAED733FFFCE1AB77325FCE8E57B" bold="true" box="[151,825,1792,1818]" pageId="8" pageNumber="9">
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<taxonomicName id="4C004D7C733FFFCE1AB77325FD40E57B" authority="Ramazzotti, 1962" authorityName="Ramazzotti" authorityYear="1962" box="[151,657,1792,1818]" class="Polychaeta" family="Aphroditidae" genus="Milnesium" kingdom="Animalia" order="Phyllodocida" pageId="8" pageNumber="9" phylum="Annelida" rank="species" species="granulatum">
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<emphasis id="B974EAED733FFFCE1AB77325FE7EE57B" bold="true" box="[151,431,1792,1818]" italics="true" pageId="8" pageNumber="9">Milnesium granulatum</emphasis>
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<bibRefCitation id="EF914B0E733FFFCE1B977325FD40E57B" author="Ramazzotti" box="[439,657,1792,1818]" pageId="8" pageNumber="9" refString="Ramazzotti, G. (1962) Tardigradi del Cile, con descrizione di quattro nuove specie e di una nuova variet. Atti della Societa ltaliana di Scienze Naturali e del Museo Civico di Storia Naturale in Milano, 101, 275 - 287." type="journal article" year="1962">Ramazzotti, 1962</bibRefCitation>
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</taxonomicName>
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, bona species
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BBF36FF733FFFCE1AB77306FEA8E55A" blockId="8.[151,825,1792,1851]" box="[151,377,1827,1851]" pageId="8" pageNumber="9">
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(
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<figureCitation id="133B2A7A733FFFCE1ABF7306FEC0E55A" box="[159,273,1827,1851]" captionStart-0="FIGURES 7 – 9" captionStart-1="FIGURES 10 – 11" captionStartId-0="9.[151,264,1398,1421]" captionStartId-1="9.[151,264,1985,2008]" captionTargetBox-0="[151,1436,401,1376]" captionTargetBox-1="[151,1436,1501,1964]" captionTargetId-0="figure@9.[151,1436,401,1377]" captionTargetId-1="figure@9.[151,1436,1501,1964]" captionTargetPageId-0="9" captionTargetPageId-1="9" captionText-0="FIGURES 7 – 9. Milnesium granulatum Ramazzotti, 1962. 7 — habitus (dorso-ventral view, paratype); 8 — dorso-caudal cuticle (paratype); 9 — buccal tube (dorso-ventral section, paratype). [All PCM]." captionText-1="FIGURES 10 – 11. Milnesium granulatum Ramazzotti, 1962. 10 — claws III (paratype); 11 — claws IV (paratype). [All PCM]." httpUri-0="https://zenodo.org/record/214360/files/figure.png" httpUri-1="https://zenodo.org/record/214361/files/figure.png" pageId="8" pageNumber="9">Figs 7–11</figureCitation>
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,
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<tableCitation id="C6820344733FFFCE1B3C7306FEA1E55A" box="[284,368,1827,1851]" captionStart="TABLE 2" captionStartId="5.[151,235,289,310]" captionTargetBox="[151,1436,485,1997]" captionTargetPageId="5" captionText="TABLE 2. Measurements [in µm] and pt values of selected morphological structures of the holotype and a specimen from" httpUri="http://table.plazi.org/id/DF7F66777332FFC31AB77504FA4DE356" pageId="8" pageNumber="9" tableUuid="DF7F66777332FFC31AB77504FA4DE356">Table 2</tableCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C31A6574733FFFCE1AB7734EFB44E5C6" pageId="8" pageNumber="9" type="materials_examined">
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<paragraph id="8BBF36FF733FFFCE1AB7734EFB44E5C6" blockId="8.[151,1436,1899,2031]" pageId="8" pageNumber="9">
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<emphasis id="B974EAED733FFFCE1AB7734EFE51E5E5" bold="true" box="[151,384,1899,1924]" pageId="8" pageNumber="9">Material examined.</emphasis>
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One
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<typeStatus id="54BB885D733FFFCE1B9D7349FDF1E5E5" box="[445,544,1900,1924]" pageId="8" pageNumber="9" type="paratype">paratype</typeStatus>
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, female, slide labelled as follows: “Tipo 145. 1
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<emphasis id="B974EAED733FFFCE1E157349FB16E5E2" box="[1077,1223,1899,1923]" italics="true" pageId="8" pageNumber="9">H. baumanni</emphasis>
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,
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<specimenCount id="9D06FD76733FFFCE1EF5734EFAD8E5E2" box="[1237,1289,1899,1923]" pageId="8" pageNumber="9" type="generic">
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1
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<taxonomicName id="4C004D7C733FFFCE1ECA7349FAD8E5E2" authority="Ramazzotti" authorityName="Ramazzotti" box="[1258,1289,1900,1923]" class="Polychaeta" family="Aphroditidae" genus="Milnesium" kingdom="Animalia" order="Phyllodocida" pageId="8" pageNumber="9" phylum="Annelida" rank="subSpecies" species="tardigradum" subSpecies="granulatum">
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<emphasis id="B974EAED733FFFCE1ECA7349FAD8E5E2" box="[1258,1289,1900,1923]" italics="true" pageId="8" pageNumber="9">M.</emphasis>
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</taxonomicName>
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</specimenCount>
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tardigradum granulatum Ramazzotti; Fray Jorge (
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<collectingCountry id="F317766F733FFFCE181473AAFDB7E5C6" box="[564,614,1935,1959]" name="Chile" pageId="8" pageNumber="9">Cile</collectingCountry>
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) a m 620,
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<date id="FFBE103F733FFFCE18FC73AAFCE2E5C6" box="[732,819,1935,1959]" pageId="8" pageNumber="9" value="62-05-22">22.5.62</date>
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, Lichene su parmo (Polivinil).”
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</paragraph>
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</subSubSection>
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<subSubSection id="C31A6574733FFFCC1AE77396FC57E305" lastPageId="10" lastPageNumber="11" pageId="8" pageNumber="9" type="description">
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<paragraph id="8BBF36FF733FFFCF1AE77396FCE2E296" blockId="8.[151,1436,1899,2031]" lastBlockId="9.[151,1436,151,356]" lastPageId="9" lastPageNumber="10" pageId="8" pageNumber="9">
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<emphasis id="B974EAED733FFFCE1AE77396FD62E5AD" bold="true" box="[199,691,1971,1996]" pageId="8" pageNumber="9">
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Description (measurements in
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<tableCitation id="C6820344733FFFCE18657391FD75E5AD" box="[581,676,1971,1996]" captionStart="TABLE 2" captionStartId="5.[151,235,289,310]" captionTargetBox="[151,1436,485,1997]" captionTargetPageId="5" captionText="TABLE 2. Measurements [in µm] and pt values of selected morphological structures of the holotype and a specimen from" httpUri="http://table.plazi.org/id/DF7F66777332FFC31AB77504FA4DE356" pageId="8" pageNumber="9" tableUuid="DF7F66777332FFC31AB77504FA4DE356">Table 2</tableCitation>
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).
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</emphasis>
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Body colour unknown, eyes present according to the original description (
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<bibRefCitation id="EF914B0E733FFFCE1B0473F2FE3BE58E" author="Ramazzotti" box="[292,490,2007,2031]" pageId="8" pageNumber="9" refString="Ramazzotti, G. (1962) Tardigradi del Cile, con descrizione di quattro nuove specie e di una nuova variet. Atti della Societa ltaliana di Scienze Naturali e del Museo Civico di Storia Naturale in Milano, 101, 275 - 287." type="journal article" year="1962">Ramazzotti 1962</bibRefCitation>
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), but now are no longer visible (
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<figureCitation id="133B2A7A733FFFCE194273F2FC79E58E" box="[866,936,2007,2031]" captionStart="FIGURES 7 – 9" captionStartId="9.[151,264,1398,1421]" captionTargetBox="[151,1436,401,1376]" captionTargetId="figure@9.[151,1436,401,1377]" captionTargetPageId="9" captionText="FIGURES 7 – 9. Milnesium granulatum Ramazzotti, 1962. 7 — habitus (dorso-ventral view, paratype); 8 — dorso-caudal cuticle (paratype); 9 — buccal tube (dorso-ventral section, paratype). [All PCM]." httpUri="https://zenodo.org/record/214360/files/figure.png" pageId="8" pageNumber="9">Fig. 7</figureCitation>
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). Dorso-lateral cuticle or only caudo-dorsal cuticle covered with a reticular design (diameter of polygons 0.5–1.5 μm) (
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<figureCitation id="133B2A7A733EFFCF19C274B2FBF7E2CE" box="[994,1062,151,175]" captionStart="FIGURES 7 – 9" captionStartId="9.[151,264,1398,1421]" captionTargetBox="[151,1436,401,1376]" captionTargetId="figure@9.[151,1436,401,1377]" captionTargetPageId="9" captionText="FIGURES 7 – 9. Milnesium granulatum Ramazzotti, 1962. 7 — habitus (dorso-ventral view, paratype); 8 — dorso-caudal cuticle (paratype); 9 — buccal tube (dorso-ventral section, paratype). [All PCM]." httpUri="https://zenodo.org/record/214360/files/figure.png" pageId="9" pageNumber="10">Fig. 8</figureCitation>
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). The design better developed on the caudal cuticle. Ventral cuticle smooth. Granulation or pores in cuticle absent. Two lateral and six peribuccal papillae present(ventral papilla smaller than other papillae).
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</paragraph>
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<paragraph id="8BBF36FF733EFFCF1AE57521FC84E305" blockId="9.[151,1436,151,356]" pageId="9" pageNumber="10">
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Buccal apparatus of the
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<taxonomicName id="4C004D7C733EFFCF1BF97521FD9EE37D" box="[473,591,260,284]" class="Polychaeta" family="Aphroditidae" genus="Milnesium" kingdom="Animalia" order="Phyllodocida" pageId="9" pageNumber="10" phylum="Annelida" rank="genus">
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<emphasis id="B974EAED733EFFCF1BF97521FD9EE37D" box="[473,591,260,284]" italics="true" pageId="9" pageNumber="10">Milnesium</emphasis>
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</taxonomicName>
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<typeStatus id="54BB885D733EFFCF18787520FD59E37C" box="[600,648,261,285]" pageId="9" pageNumber="10">type</typeStatus>
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(
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<figureCitation id="133B2A7A733EFFCF18B97521FD0EE37D" box="[665,735,260,284]" captionStart="FIGURES 7 – 9" captionStartId="9.[151,264,1398,1421]" captionTargetBox="[151,1436,401,1376]" captionTargetId="figure@9.[151,1436,401,1377]" captionTargetPageId="9" captionText="FIGURES 7 – 9. Milnesium granulatum Ramazzotti, 1962. 7 — habitus (dorso-ventral view, paratype); 8 — dorso-caudal cuticle (paratype); 9 — buccal tube (dorso-ventral section, paratype). [All PCM]." httpUri="https://zenodo.org/record/214360/files/figure.png" pageId="9" pageNumber="10">Fig. 9</figureCitation>
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). Six peribuccal lamellae around the mouth opening present. Buccal tube slightly funnel-shaped, wider anteriorly (posterior diameter 87% of the anterior diameter). Pharyngeal bulb elongated, pear-shaped and without placoids or septulum.
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</paragraph>
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<caption id="DF7F6677733EFFCF1AB77153FCA6E7CA" httpUri="https://zenodo.org/record/214360/files/figure.png" pageId="9" pageNumber="10" targetBox="[151,1436,401,1376]" targetPageId="9">
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<paragraph id="8BBF36FF733EFFCF1AB77153FCA6E7CA" blockId="9.[151,1436,1398,1452]" pageId="9" pageNumber="10">
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<emphasis id="B974EAED733EFFCF1AB77153FEEBE7ED" bold="true" box="[151,314,1398,1421]" pageId="9" pageNumber="10">FIGURES 7–9.</emphasis>
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<taxonomicName id="4C004D7C733EFFCF1B627152FD39E7ED" authority="Ramazzotti, 1962" authorityName="Ramazzotti" authorityYear="1962" box="[322,744,1398,1421]" class="Polychaeta" family="Aphroditidae" genus="Milnesium" kingdom="Animalia" order="Phyllodocida" pageId="9" pageNumber="10" phylum="Annelida" rank="species" species="granulatum">
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<emphasis id="B974EAED733EFFCF1B627152FDFBE7EC" box="[322,554,1399,1421]" italics="true" pageId="9" pageNumber="10">Milnesium granulatum</emphasis>
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Ramazzotti, 1962
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</taxonomicName>
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.
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<emphasis id="B974EAED733EFFCF18D17152FD2FE7ED" bold="true" box="[753,766,1399,1420]" pageId="9" pageNumber="10">7</emphasis>
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—habitus (dorso-ventral view, paratype);
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<emphasis id="B974EAED733EFFCF1E897152FB64E7ED" bold="true" box="[1193,1205,1399,1420]" pageId="9" pageNumber="10">8</emphasis>
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—dorso-caudal cuticle (paratype);
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<emphasis id="B974EAED733EFFCF1B2C71B0FEC9E7CB" bold="true" box="[268,280,1429,1450]" pageId="9" pageNumber="10">9</emphasis>
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—buccal tube (dorso-ventral section, paratype). [All PCM].
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</paragraph>
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</caption>
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<caption id="DF7F6677733EFFCF1AB773E4FA59E5B9" box="[151,1416,1985,2008]" httpUri="https://zenodo.org/record/214361/files/figure.png" pageId="9" pageNumber="10" targetBox="[151,1436,1501,1964]" targetPageId="9">
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||
<paragraph id="8BBF36FF733EFFCF1AB773E4FA59E5B9" blockId="9.[151,1416,1985,2008]" box="[151,1416,1985,2008]" pageId="9" pageNumber="10">
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<emphasis id="B974EAED733EFFCF1AB773E4FE80E5B6" bold="true" box="[151,337,1985,2008]" pageId="9" pageNumber="10">FIGURES 10–11.</emphasis>
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<taxonomicName id="4C004D7C733EFFCF1B7873E7FD2DE5B9" authority="Ramazzotti, 1962" authorityName="Ramazzotti" authorityYear="1962" box="[344,764,1986,2008]" class="Polychaeta" family="Aphroditidae" genus="Milnesium" kingdom="Animalia" order="Phyllodocida" pageId="9" pageNumber="10" phylum="Annelida" rank="species" species="granulatum">
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<emphasis id="B974EAED733EFFCF1B7873E7FDEEE5B9" box="[344,575,1986,2008]" italics="true" pageId="9" pageNumber="10">Milnesium granulatum</emphasis>
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Ramazzotti, 1962
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</taxonomicName>
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.
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<emphasis id="B974EAED733EFFCF192573E7FCCFE5B6" bold="true" box="[773,798,1986,2007]" pageId="9" pageNumber="10">10</emphasis>
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—claws III (paratype);
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<emphasis id="B974EAED733EFFCF1E2C73E7FBF3E5B6" bold="true" box="[1036,1058,1986,2007]" pageId="9" pageNumber="10">11</emphasis>
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—claws IV (paratype). [All PCM].
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</paragraph>
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</caption>
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<paragraph id="8BBF36FF733DFFCC1AE574B2FD6FE37D" blockId="10.[151,1436,151,896]" pageId="10" pageNumber="11">
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Claws of the
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<taxonomicName id="4C004D7C733DFFCC1B7974BDFE1EE2D1" box="[345,463,152,176]" class="Polychaeta" family="Aphroditidae" genus="Milnesium" kingdom="Animalia" order="Phyllodocida" pageId="10" pageNumber="11" phylum="Annelida" rank="genus">
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<emphasis id="B974EAED733DFFCC1B7974BDFE1EE2D1" box="[345,463,152,176]" italics="true" pageId="10" pageNumber="11">Milnesium</emphasis>
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</taxonomicName>
|
||
<typeStatus id="54BB885D733DFFCC1BF674BDFDD8E2D1" box="[470,521,152,176]" pageId="10" pageNumber="11">type</typeStatus>
|
||
, slender (
|
||
<figureCitation id="133B2A7A733DFFCC185474B2FD24E2CE" box="[628,757,151,175]" captionStart="FIGURES 10 – 11" captionStartId="9.[151,264,1985,2008]" captionTargetBox="[151,1436,1501,1964]" captionTargetId="figure@9.[151,1436,1501,1964]" captionTargetPageId="9" captionText="FIGURES 10 – 11. Milnesium granulatum Ramazzotti, 1962. 10 — claws III (paratype); 11 — claws IV (paratype). [All PCM]." httpUri="https://zenodo.org/record/214361/files/figure.png" pageId="10" pageNumber="11">Figs 10–11</figureCitation>
|
||
). Primary branches on all legs with small, but distinct accessory points detaching from the branch near its end. Secondary branches with rounded basal thickenings. All secondary branches on all legs with three points (i.e. claw configuration: [3-3]-[3-3]). Single, long transversal, cuticular bars under claws I–III present (
|
||
<figureCitation id="133B2A7A733DFFCC187D7521FD60E37D" box="[605,689,260,284]" captionStart="FIGURES 10 – 11" captionStartId="9.[151,264,1985,2008]" captionTargetBox="[151,1436,1501,1964]" captionTargetId="figure@9.[151,1436,1501,1964]" captionTargetPageId="9" captionText="FIGURES 10 – 11. Milnesium granulatum Ramazzotti, 1962. 10 — claws III (paratype); 11 — claws IV (paratype). [All PCM]." httpUri="https://zenodo.org/record/214361/files/figure.png" pageId="10" pageNumber="11">Fig. 10</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BBF36FF733DFFCC1AE57502FC57E305" blockId="10.[151,1436,151,896]" pageId="10" pageNumber="11">
|
||
Although eggs were not described by
|
||
<bibRefCitation id="EF914B0E733DFFCC18537502FC96E35E" author="Ramazzotti" box="[627,839,295,319]" pageId="10" pageNumber="11" refString="Ramazzotti, G. (1962) Tardigradi del Cile, con descrizione di quattro nuove specie e di una nuova variet. Atti della Societa ltaliana di Scienze Naturali e del Museo Civico di Storia Naturale in Milano, 101, 275 - 287." type="journal article" year="1962">Ramazzotti (1962)</bibRefCitation>
|
||
, we should probably expect them to be oval, smooth and deposited in exuvium as in all other known
|
||
<taxonomicName id="4C004D7C733DFFCC18907569FCF6E305" box="[688,807,332,356]" class="Polychaeta" family="Aphroditidae" genus="Milnesium" kingdom="Animalia" order="Phyllodocida" pageId="10" pageNumber="11" phylum="Annelida" rank="genus">
|
||
<emphasis id="B974EAED733DFFCC18907569FCF6E305" box="[688,807,332,356]" italics="true" pageId="10" pageNumber="11">Milnesium</emphasis>
|
||
</taxonomicName>
|
||
species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31A6574733DFFCC1AE77555FD9DE0E5" pageId="10" pageNumber="11" type="discussion">
|
||
<paragraph id="8BBF36FF733DFFCC1AE77555FEFFE076" blockId="10.[151,1436,151,896]" pageId="10" pageNumber="11">
|
||
<emphasis id="B974EAED733DFFCC1AE77555FEEAE3E9" bold="true" box="[199,315,368,392]" pageId="10" pageNumber="11">Remarks.</emphasis>
|
||
<bibRefCitation id="EF914B0E733DFFCC1B61754AFDC1E3E6" author="Maucci" box="[321,528,367,391]" pageId="10" pageNumber="11" refString="Maucci, W. (1973 - 4) Tardigradi muscicoli del Carso Triestino. Bollettino Della Societa' Adriatica di Scienze - Trieste, 59, 107 - 150." type="journal article" year="1973">
|
||
Maucci (
|
||
<date id="FFBE103F733DFFCC1B84754AFDD9E3E6" box="[420,520,367,391]" pageId="10" pageNumber="11" value="1973" valueMax="1974">1973–74</date>
|
||
)
|
||
</bibRefCitation>
|
||
reported
|
||
<taxonomicName id="4C004D7C733DFFCC185B7554FC67E3E9" box="[635,950,368,392]" class="Polychaeta" family="Aphroditidae" genus="Milnesium" kingdom="Animalia" order="Phyllodocida" pageId="10" pageNumber="11" phylum="Annelida" rank="subSpecies" species="tardigradum" subSpecies="granulatum">
|
||
<emphasis id="B974EAED733DFFCC185B7554FC67E3E9" box="[635,950,368,392]" italics="true" pageId="10" pageNumber="11">M. tardigradum granulatum</emphasis>
|
||
</taxonomicName>
|
||
also from the Trieste Kras, but we have not had an occasion to examine those animals. However, given the
|
||
<emphasis id="B974EAED733DFFCC194875B1FC2BE3CD" box="[872,1018,404,428]" italics="true" pageId="10" pageNumber="11">locus typicus</emphasis>
|
||
of
|
||
<taxonomicName id="4C004D7C733DFFCC1E0375B0FB1DE3CD" box="[1059,1228,404,428]" class="Polychaeta" family="Aphroditidae" genus="Milnesium" kingdom="Animalia" order="Phyllodocida" pageId="10" pageNumber="11" phylum="Annelida" rank="species" species="granulatum">
|
||
<emphasis id="B974EAED733DFFCC1E0375B0FB1DE3CD" box="[1059,1228,404,428]" italics="true" pageId="10" pageNumber="11">M. granulatum</emphasis>
|
||
</taxonomicName>
|
||
lies on a different continent and in a different zoogeographical region, we hypothesise that Maucci could have found a different species of the
|
||
<taxonomicName id="4C004D7C733DFFCC1B3275F9FDC2E395" box="[274,531,476,500]" class="Polychaeta" family="Aphroditidae" genus="Milnesium" kingdom="Animalia" order="Phyllodocida" pageId="10" pageNumber="11" phylum="Annelida" rank="species" species="granulatum">
|
||
<emphasis id="B974EAED733DFFCC1B3275F9FDC2E395" box="[274,531,476,500]" italics="true" pageId="10" pageNumber="11">Milnesium granulatum</emphasis>
|
||
</taxonomicName>
|
||
group. Until the record is re-examined, we suggest this report should be considered dubious.
|
||
</paragraph>
|
||
<paragraph id="8BBF36FF733DFFCC1AE77601FD9DE0E5" blockId="10.[151,1436,151,896]" pageId="10" pageNumber="11">
|
||
<emphasis id="B974EAED733DFFCC1AE77601FEA3E05D" bold="true" box="[199,370,547,572]" pageId="10" pageNumber="11">Locus typicus.</emphasis>
|
||
Coquimbo Region, Bosque de Fray Jorge National Park, Ovalle,
|
||
<emphasis id="B974EAED733DFFCC1E727600FBA5E05D" box="[1106,1140,549,572]" italics="true" pageId="10" pageNumber="11">ca.</emphasis>
|
||
<quantity id="4CF89B1A733DFFCC1E5B7601FB01E05D" box="[1147,1232,548,572]" metricMagnitude="5" metricUnit="m" metricValue="4.2" pageId="10" pageNumber="11" unit="km" value="420.0">420 km</quantity>
|
||
to the North from Santiago de
|
||
<collectingCountry id="F317766F733DFFCC1B057662FEB4E03E" box="[293,357,583,607]" name="Chile" pageId="10" pageNumber="11">Chile</collectingCountry>
|
||
,
|
||
<emphasis id="B974EAED733DFFCC1B51766CFE43E001" box="[369,402,585,608]" italics="true" pageId="10" pageNumber="11">ca.</emphasis>
|
||
<geoCoordinate id="EE345038733DFFCC1BBB7662FE22E03E" box="[411,499,583,607]" direction="south" orientation="latitude" pageId="10" pageNumber="11" precision="925" value="-30.666666">30°40'S</geoCoordinate>
|
||
and
|
||
<geoCoordinate id="EE345038733DFFCC180C7662FD5EE03E" box="[556,655,583,607]" direction="west" orientation="longitude" pageId="10" pageNumber="11" precision="925" value="-71.666664">71°40'W</geoCoordinate>
|
||
,
|
||
<emphasis id="B974EAED733DFFCC18BC766CFD6FE001" box="[668,702,585,608]" italics="true" pageId="10" pageNumber="11">ca.</emphasis>
|
||
<quantity id="4CF89B1A733DFFCC18E67662FCDEE03E" box="[710,783,583,607]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="10" pageNumber="11" unit="m" value="600.0">600 m</quantity>
|
||
asl, temperate hygrophilous forest surrounded by steppe, mosses and lichens growing on shrubs.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31A6574733DFFCC1AE776AAFAC0E0C6" box="[199,1297,655,680]" pageId="10" pageNumber="11" type="distribution">
|
||
<paragraph id="8BBF36FF733DFFCC1AE776AAFAC0E0C6" blockId="10.[151,1436,151,896]" box="[199,1297,655,680]" pageId="10" pageNumber="11">
|
||
<emphasis id="B974EAED733DFFCC1AE776AAFEB1E0C9" bold="true" box="[199,352,655,680]" pageId="10" pageNumber="11">Distribution.</emphasis>
|
||
Currently confirmed is only the
|
||
<typeStatus id="54BB885D733DFFCC18EF76B5FD2EE0C9" box="[719,767,656,680]" pageId="10" pageNumber="11">type</typeStatus>
|
||
locality in
|
||
<collectingCountry id="F317766F733DFFCC195E76AAFC6AE0C6" box="[894,955,655,679]" name="Chile" pageId="10" pageNumber="11">Chile</collectingCountry>
|
||
(but see also
|
||
<emphasis id="B974EAED733DFFCC1E7476B5FB67E0C9" box="[1108,1206,656,680]" italics="true" pageId="10" pageNumber="11">Remarks</emphasis>
|
||
above).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31A6574733DFFCC1AE77691FE5BE175" pageId="10" pageNumber="11" type="etymology">
|
||
<paragraph id="8BBF36FF733DFFCC1AE77691FE5BE175" blockId="10.[151,1436,151,896]" pageId="10" pageNumber="11">
|
||
<emphasis id="B974EAED733DFFCC1AE77691FE9FE0AD" bold="true" box="[199,334,692,716]" pageId="10" pageNumber="11">Etymology.</emphasis>
|
||
Giuseppe Ramazzotti named the species ‘
|
||
<taxonomicName id="4C004D7C733DFFCC19647691FC16E0AD" box="[836,967,692,716]" class="Polychaeta" family="Aphroditidae" genus="Milnesium" kingdom="Animalia" order="Phyllodocida" pageId="10" pageNumber="11" phylum="Annelida" rank="species" species="granulatum">
|
||
<emphasis id="B974EAED733DFFCC19647691FC16E0AD" box="[836,967,692,716]" italics="true" pageId="10" pageNumber="11">granulatum</emphasis>
|
||
</taxonomicName>
|
||
’ most likely because he interpreted the reticular design on the cuticle as granulation. In fact, ‘granules’ are shallow depression in the shape of rounded polygons (see
|
||
<figureCitation id="133B2A7A733DFFCC1B1976D9FEADE175" box="[313,380,764,788]" captionStart="FIGURES 7 – 9" captionStartId="9.[151,264,1398,1421]" captionTargetBox="[151,1436,401,1376]" captionTargetId="figure@9.[151,1436,401,1377]" captionTargetPageId="9" captionText="FIGURES 7 – 9. Milnesium granulatum Ramazzotti, 1962. 7 — habitus (dorso-ventral view, paratype); 8 — dorso-caudal cuticle (paratype); 9 — buccal tube (dorso-ventral section, paratype). [All PCM]." httpUri="https://zenodo.org/record/214360/files/figure.png" pageId="10" pageNumber="11">Fig. 8</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31A6574733DFFCC1AE77705FC19E11E" pageId="10" pageNumber="11" type="materials_examined">
|
||
<paragraph id="8BBF36FF733DFFCC1AE77705FAC9E156" blockId="10.[151,1436,151,896]" box="[199,1304,799,825]" pageId="10" pageNumber="11">
|
||
<emphasis id="B974EAED733DFFCC1AE77705FE4EE159" bold="true" box="[199,415,799,824]" pageId="10" pageNumber="11">
|
||
<typeStatus id="54BB885D733DFFCC1AE77705FED3E159" box="[199,258,800,824]" pageId="10" pageNumber="11">Type</typeStatus>
|
||
depositories.
|
||
</emphasis>
|
||
The
|
||
<typeStatus id="54BB885D733DFFCC1BF87704FDD9E158" box="[472,520,801,825]" pageId="10" pageNumber="11">type</typeStatus>
|
||
material is preserved at the Natural History Museum of Verona,
|
||
<collectingCountry id="F317766F733DFFCC1EFE773AFAC2E156" box="[1246,1299,799,823]" name="Italy" pageId="10" pageNumber="11">Italy</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BBF36FF733DFFCC1AE77766FC19E11E" blockId="10.[151,1436,151,896]" pageId="10" pageNumber="11">
|
||
<emphasis id="B974EAED733DFFCC1AE77766FE03E13D" bold="true" box="[199,466,835,860]" pageId="10" pageNumber="11">Differential diagnosis.</emphasis>
|
||
Apart from
|
||
<taxonomicName id="4C004D7C733DFFCC184B7760FCC9E13D" box="[619,792,836,860]" class="Polychaeta" family="Aphroditidae" genus="Milnesium" kingdom="Animalia" order="Phyllodocida" pageId="10" pageNumber="11" phylum="Annelida" rank="species" species="granulatum">
|
||
<emphasis id="B974EAED733DFFCC184B7760FCC9E13D" box="[619,792,836,860]" italics="true" pageId="10" pageNumber="11">M. granulatum</emphasis>
|
||
</taxonomicName>
|
||
there are four other described
|
||
<taxonomicName id="4C004D7C733DFFCC1EAC7761FAD3E13D" box="[1164,1282,836,860]" class="Polychaeta" family="Aphroditidae" genus="Milnesium" kingdom="Animalia" order="Phyllodocida" pageId="10" pageNumber="11" phylum="Annelida" rank="genus">
|
||
<emphasis id="B974EAED733DFFCC1EAC7761FAD3E13D" box="[1164,1282,836,860]" italics="true" pageId="10" pageNumber="11">Milnesium</emphasis>
|
||
</taxonomicName>
|
||
species with sculptured cuticle (see
|
||
<tableCitation id="C6820344733DFFCC1BB87742FE3BE11E" box="[408,490,871,895]" captionStart="TABLE 1. A" captionStartId="4.[151,235,152,173]" captionTargetBox="[151,1428,201,1611]" captionTargetPageId="4" captionText="TABLE 1. A list of taxonomically important qualitative traits of all known Milnesium species and subspecies." httpUri="http://table.plazi.org/id/DF7F66777333FFC21AB774BDFB22E2CF" pageId="10" pageNumber="11" tableUuid="DF7F66777333FFC21AB774BDFB22E2CF">Table 1</tableCitation>
|
||
).
|
||
<taxonomicName id="4C004D7C733DFFCC1822774CFD7AE1E1" box="[514,683,872,896]" class="Polychaeta" family="Aphroditidae" genus="Milnesium" kingdom="Animalia" order="Phyllodocida" pageId="10" pageNumber="11" phylum="Annelida" rank="species" species="granulatum">
|
||
<emphasis id="B974EAED733DFFCC1822774CFD7AE1E1" box="[514,683,872,896]" italics="true" pageId="10" pageNumber="11">M. granulatum</emphasis>
|
||
</taxonomicName>
|
||
differs specifically from:
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31A6574733DFFCC1AB77794FED2E76D" pageId="10" pageNumber="11" type="reference_group">
|
||
<paragraph id="8BBF36FF733DFFCC1AB77794FB60E1A6" blockId="10.[151,1436,943,1292]" box="[151,1201,943,969]" pageId="10" pageNumber="11">
|
||
<taxonomicName id="4C004D7C733DFFCC1AB77794FD8FE1A6" authority="Wallendorf & Miller, 2009" authorityName="Wallendorf & Miller" authorityYear="2009" box="[151,606,943,968]" class="Polychaeta" family="Aphroditidae" genus="Milnesium" kingdom="Animalia" order="Phyllodocida" pageId="10" pageNumber="11" phylum="Annelida" rank="species" species="alabamae">
|
||
<emphasis id="B974EAED733DFFCC1AB77794FEFBE1A9" box="[151,298,944,968]" italics="true" pageId="10" pageNumber="11">M. alabamae</emphasis>
|
||
<bibRefCitation id="EF914B0E733DFFCC1B12778AFD8FE1A6" author="Wallendorf" box="[306,606,943,967]" pageId="10" pageNumber="11" refString="Wallendorf, M. & Miller, W. R. (2009) Tardigrades of North America: Milnesium alabamae nov. sp. (Eutardigrada: Apochela: Milnesiidae) a New Species from Alabama. Transactions of the Kansas Academy of Science, 112 (3 - 4), 181 - 186." type="journal article" year="2009">Wallendorf & Miller, 2009</bibRefCitation>
|
||
</taxonomicName>
|
||
by having accessory points on the primary branches.
|
||
</paragraph>
|
||
<paragraph id="8BBF36FF733DFFCC1AB777F0FD97E636" blockId="10.[151,1436,943,1292]" pageId="10" pageNumber="11">
|
||
<taxonomicName id="4C004D7C733DFFCC1AB777F0FCC0E18D" authority="Kaczmarek, Michalczyk & Beasley, 2004" authorityName="Kaczmarek, Michalczyk & Beasley" authorityYear="2004" box="[151,785,980,1004]" class="Polychaeta" family="Aphroditidae" genus="Milnesium" kingdom="Animalia" order="Phyllodocida" pageId="10" pageNumber="11" phylum="Annelida" rank="species" species="katarzynae">
|
||
<emphasis id="B974EAED733DFFCC1AB777F0FEE6E18D" box="[151,311,980,1004]" italics="true" pageId="10" pageNumber="11">M. katarzynae</emphasis>
|
||
<bibRefCitation id="EF914B0E733DFFCC1B1E77F1FCC0E18D" author="Kaczmarek" box="[318,785,980,1004]" pageId="10" pageNumber="11" refString="Kaczmarek, L., Michalczyk, L. & Beasley, C. W. (2004) Milnesium katarzynae sp. nov., a new species of eutardigrade (Milnesiidae) from China. Zootaxa, 743, 1 - 5." type="journal article" year="2004">Kaczmarek, Michalczyk & Beasley, 2004</bibRefCitation>
|
||
</taxonomicName>
|
||
by a different claw configuration ([3-3]-[3-3] in
|
||
<taxonomicName id="4C004D7C733DFFCC1F1877F0FEC6E671" class="Polychaeta" family="Aphroditidae" genus="Milnesium" kingdom="Animalia" order="Phyllodocida" pageId="10" pageNumber="11" phylum="Annelida" rank="species" species="granulatum">
|
||
<emphasis id="B974EAED733DFFCC1F1877F0FEC6E671" italics="true" pageId="10" pageNumber="11">M. granulatum</emphasis>
|
||
</taxonomicName>
|
||
and [2-2]-[2-2] in
|
||
<taxonomicName id="4C004D7C733DFFCC1BD277DCFD45E671" box="[498,660,1016,1040]" class="Polychaeta" family="Aphroditidae" genus="Milnesium" kingdom="Animalia" order="Phyllodocida" pageId="10" pageNumber="11" phylum="Annelida" rank="species" species="katarzynae">
|
||
<emphasis id="B974EAED733DFFCC1BD277DCFD45E671" box="[498,660,1016,1040]" italics="true" pageId="10" pageNumber="11">M. katarzynae</emphasis>
|
||
</taxonomicName>
|
||
), a wider buccal tube (
|
||
<emphasis id="B974EAED733DFFCC198077DCFBD4E671" box="[928,1029,1016,1040]" italics="true" pageId="10" pageNumber="11">pt ca. 39</emphasis>
|
||
% in
|
||
<taxonomicName id="4C004D7C733DFFCC1E6477DCFB3FE671" box="[1092,1262,1016,1040]" class="Polychaeta" family="Aphroditidae" genus="Milnesium" kingdom="Animalia" order="Phyllodocida" pageId="10" pageNumber="11" phylum="Annelida" rank="species" species="granulatum">
|
||
<emphasis id="B974EAED733DFFCC1E6477DCFB3FE671" box="[1092,1262,1016,1040]" italics="true" pageId="10" pageNumber="11">M. granulatum</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<emphasis id="B974EAED733DFFCC1F0977DCFF2DE655" italics="true" pageId="10" pageNumber="11">pt ca. 21– 26%</emphasis>
|
||
in
|
||
<taxonomicName id="4C004D7C733DFFCC1B3F7038FE6EE655" box="[287,447,1052,1076]" class="Polychaeta" family="Aphroditidae" genus="Milnesium" kingdom="Animalia" order="Phyllodocida" pageId="10" pageNumber="11" phylum="Annelida" rank="species" species="katarzynae">
|
||
<emphasis id="B974EAED733DFFCC1B3F7038FE6EE655" box="[287,447,1052,1076]" italics="true" pageId="10" pageNumber="11">M. katarzynae</emphasis>
|
||
</taxonomicName>
|
||
) and stylet supports inserted in more anterior position (
|
||
<emphasis id="B974EAED733DFFCC1E0A7038FB72E655" box="[1066,1187,1052,1076]" italics="true" pageId="10" pageNumber="11">pt ca. 66%</emphasis>
|
||
in
|
||
<taxonomicName id="4C004D7C733DFFCC1EE57038FABCE655" box="[1221,1389,1052,1076]" class="Polychaeta" family="Aphroditidae" genus="Milnesium" kingdom="Animalia" order="Phyllodocida" pageId="10" pageNumber="11" phylum="Annelida" rank="species" species="granulatum">
|
||
<emphasis id="B974EAED733DFFCC1EE57038FABCE655" box="[1221,1389,1052,1076]" italics="true" pageId="10" pageNumber="11">M. granulatum</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<emphasis id="B974EAED733DFFCC1AE97064FEA3E639" box="[201,370,1088,1112]" italics="true" pageId="10" pageNumber="11">pt. ca. 73–78%</emphasis>
|
||
in
|
||
<taxonomicName id="4C004D7C733DFFCC1BB67064FDE7E639" box="[406,566,1088,1112]" class="Polychaeta" family="Aphroditidae" genus="Milnesium" kingdom="Animalia" order="Phyllodocida" pageId="10" pageNumber="11" phylum="Annelida" rank="species" species="katarzynae">
|
||
<emphasis id="B974EAED733DFFCC1BB67064FDE7E639" box="[406,566,1088,1112]" italics="true" pageId="10" pageNumber="11">M. katarzynae</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BBF36FF733DFFCC1AB77040FDD4E6FE" blockId="10.[151,1436,943,1292]" pageId="10" pageNumber="11">
|
||
<taxonomicName id="4C004D7C733DFFCC1AB77040FD4DE61D" authority="Kaczmarek & Michalczyk, 2007" authorityName="Kaczmarek & Michalczyk" authorityYear="2007" box="[151,668,1124,1148]" class="Polychaeta" family="Aphroditidae" genus="Milnesium" kingdom="Animalia" order="Phyllodocida" pageId="10" pageNumber="11" phylum="Annelida" rank="species" species="krzysztofi">
|
||
<emphasis id="B974EAED733DFFCC1AB77040FEF7E61D" box="[151,294,1124,1148]" italics="true" pageId="10" pageNumber="11">M. krzysztofi</emphasis>
|
||
<bibRefCitation id="EF914B0E733DFFCC1B0D7041FD4DE61D" author="Kaczmarek" box="[301,668,1124,1148]" pageId="10" pageNumber="11" refString="Kaczmarek, L. & Michalczyk, L. (2007) A new species of Tardigrada (Eutardigrada: Milnesiidae): Milnesium krzysztofi from Costa Rica (Central America). New Zealand Journal of Zoology, 34, 297 - 302." type="journal article" year="2007">Kaczmarek & Michalczyk, 2007</bibRefCitation>
|
||
</taxonomicName>
|
||
by a different claw configuration ([3-3]-[3-3] in
|
||
<taxonomicName id="4C004D7C733DFFCC1EE47040FABDE61D" box="[1220,1388,1124,1148]" class="Polychaeta" family="Aphroditidae" genus="Milnesium" kingdom="Animalia" order="Phyllodocida" pageId="10" pageNumber="11" phylum="Annelida" rank="species" species="granulatum">
|
||
<emphasis id="B974EAED733DFFCC1EE47040FABDE61D" box="[1220,1388,1124,1148]" italics="true" pageId="10" pageNumber="11">M. granulatum</emphasis>
|
||
</taxonomicName>
|
||
and [2-3]-[3-2] in
|
||
<taxonomicName id="4C004D7C733DFFCC1B4670ACFE24E6C1" box="[358,501,1160,1184]" class="Polychaeta" family="Aphroditidae" genus="Milnesium" kingdom="Animalia" order="Phyllodocida" pageId="10" pageNumber="11" phylum="Annelida" rank="species" species="krzysztofi">
|
||
<emphasis id="B974EAED733DFFCC1B4670ACFE24E6C1" box="[358,501,1160,1184]" italics="true" pageId="10" pageNumber="11">M. krzysztofi</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BBF36FF733DFFCC1AB77088FED2E76D" blockId="10.[151,1436,943,1292]" pageId="10" pageNumber="11">
|
||
<taxonomicName id="4C004D7C733DFFCC1AB77088FDBDE6A5" authority="Pilato, Binda & Lisi, 2002" authorityName="Pilato, Binda & Lisi" authorityYear="2002" box="[151,620,1196,1220]" class="Polychaeta" family="Aphroditidae" genus="Milnesium" kingdom="Animalia" order="Phyllodocida" pageId="10" pageNumber="11" phylum="Annelida" rank="species" species="reticulatum">
|
||
<emphasis id="B974EAED733DFFCC1AB77088FEEDE6A5" box="[151,316,1196,1220]" italics="true" pageId="10" pageNumber="11">M. reticulatum</emphasis>
|
||
<bibRefCitation id="EF914B0E733DFFCC1B647089FDBDE6A5" author="Pilato" box="[324,620,1196,1220]" pageId="10" pageNumber="11" refString="Pilato, G., Binda, M. G. & Lisi, O. (2002) Notes on tardigrades of the Seychelles with the description of two new species. Bollettino dell'Accademia Gioenia Scienze Naturali, 35, 503 - 517." type="journal article" year="2002">Pilato, Binda & Lisi, 2002</bibRefCitation>
|
||
</taxonomicName>
|
||
by the absence of gibbosities, having six peribuccal lamellae (four in
|
||
<taxonomicName id="4C004D7C733DFFCC1F5D7088FE98E689" class="Polychaeta" family="Aphroditidae" genus="Milnesium" kingdom="Animalia" order="Phyllodocida" pageId="10" pageNumber="11" phylum="Annelida" rank="species" species="reticulatum">
|
||
<emphasis id="B974EAED733DFFCC1F5D7088FE98E689" italics="true" pageId="10" pageNumber="11">M. reticulatum</emphasis>
|
||
</taxonomicName>
|
||
) and a different claw configuration ([3-3]-[3-3] in
|
||
<taxonomicName id="4C004D7C733DFFCC19B170F4FBEAE689" box="[913,1083,1232,1256]" class="Polychaeta" family="Aphroditidae" genus="Milnesium" kingdom="Animalia" order="Phyllodocida" pageId="10" pageNumber="11" phylum="Annelida" rank="species" species="granulatum">
|
||
<emphasis id="B974EAED733DFFCC19B170F4FBEAE689" box="[913,1083,1232,1256]" italics="true" pageId="10" pageNumber="11">M. granulatum</emphasis>
|
||
</taxonomicName>
|
||
and [2-3]-[3-2] in
|
||
<taxonomicName id="4C004D7C733DFFCC1F3570F4FF22E76D" class="Polychaeta" family="Aphroditidae" genus="Milnesium" kingdom="Animalia" order="Phyllodocida" pageId="10" pageNumber="11" phylum="Annelida" rank="species" species="reticulatum">
|
||
<emphasis id="B974EAED733DFFCC1F3570F4FF22E76D" italics="true" pageId="10" pageNumber="11">M. reticulatum</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |