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<document id="CC9BD8CABB1676B53BEBB8E0B1C1CECC" ID-CLB-Dataset="182444" ID-DOI="10.11646/zootaxa.5202.1.1" ID-GBIF-Dataset="45ba25c1-47a9-4001-93d9-294815bf37e7" ID-ISSN="1175-5326" ID-Zenodo-Dep="7285600" ID-ZooBank="BF4831D2-D98B-4265-9138-03DB8607B826" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" checkinTime="1667554680289" checkinUser="plazi" docAuthor="Sánchez-Vialas, Alberto, Calvo-Revuelta, Marta &amp; Riva, Ignacio De La" docDate="2022" docId="03C287E8FFEEFFDFFF4BFD13FF5BE40E" docLanguage="en" docName="zootaxa.5202.1.1.pdf" docOrigin="Zootaxa 5202 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5202.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Philothamnus carinatus" docType="treatment" docVersion="9" lastPageNumber="77" masterDocId="FFFBFF90FFA2FF93FFDCFF9DFFB5E232" masterDocTitle="Synopsis of the terrestrial Reptiles of Equatorial Guinea" masterLastPageNumber="197" masterPageNumber="1" pageNumber="77" updateTime="1699409742602" updateUser="ExternalLinkService">
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<mods:title id="B31D3267E38A13A6DF797A466DEE74E2">Synopsis of the terrestrial Reptiles of Equatorial Guinea</mods:title>
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<mods:namePart id="2966BCF23BD496D4A3C2B6B87198F9F0">Sánchez-Vialas, Alberto</mods:namePart>
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<mods:namePart id="FF94D67CD1FE6483EE5A1DBFB767FC06">Calvo-Revuelta, Marta</mods:namePart>
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<mods:namePart id="B292F2631EA5C4096D640687AA3B35C4">Riva, Ignacio De La</mods:namePart>
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<mods:title id="52E9923746B334AFFC9E71BB90B82149">Zootaxa</mods:title>
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<treatment id="03C287E8FFEEFFDFFF4BFD13FF5BE40E" ID-DOI="http://doi.org/10.5281/zenodo.7293463" ID-GBIF-Taxon="204305354" ID-Zenodo-Dep="7293463" LSID="urn:lsid:plazi:treatment:03C287E8FFEEFFDFFF4BFD13FF5BE40E" httpUri="http://treatment.plazi.org/id/03C287E8FFEEFFDFFF4BFD13FF5BE40E" lastPageNumber="77" pageId="76" pageNumber="77">
<subSubSection id="C3716575FFEEFFDFFF4BFD13FD2FE09A" box="[151,666,654,681]" pageId="76" pageNumber="77" type="nomenclature">
<paragraph id="8BD436FEFFEEFFDFFF4BFD13FD2FE09A" blockId="76.[151,666,654,717]" box="[151,666,654,681]" pageId="76" pageNumber="77">
<heading id="D09C8192FFEEFFDFFF4BFD13FD2FE09A" bold="true" box="[151,666,654,681]" fontSize="11" level="2" pageId="76" pageNumber="77" reason="4">
<taxonomicName id="4C6B4D7DFFEEFFDFFF4BFD13FD2FE09A" authority="(Anderson, 1901)" baseAuthorityName="Anderson" baseAuthorityYear="1901" box="[151,666,654,681]" class="Reptilia" family="Colubridae" genus="Philothamnus" kingdom="Animalia" order="Squamata" pageId="76" pageNumber="77" phylum="Chordata" rank="species" species="carinatus">
<emphasis id="B91FEAECFFEEFFDFFF4BFD13FD2FE09A" bold="true" box="[151,666,654,681]" pageId="76" pageNumber="77">
<emphasis id="B91FEAECFFEEFFDFFF4BFD13FE09E09B" bold="true" box="[151,444,654,681]" italics="true" pageId="76" pageNumber="77">Philothamnus carinatus</emphasis>
(Anderson, 1901)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3716575FFEEFFDFFF4BFD2EFEA3E0FF" box="[151,278,691,717]" pageId="76" pageNumber="77" type="description">
<paragraph id="8BD436FEFFEEFFDFFF4BFD2EFEA3E0FF" blockId="76.[151,666,654,717]" box="[151,278,691,717]" pageId="76" pageNumber="77">
<figureCitation id="13502A7BFFEEFFDFFF4BFD2EFEA3E0FF" box="[151,278,691,717]" captionStart="FIGURE 18" captionStartId="78.[151,250,1921,1945]" captionTargetBox="[178,1388,193,1893]" captionTargetId="figure-19@78.[150,1437,181,1897]" captionTargetPageId="78" captionText="FIGURE 18. From left to right: lateral, dorsal and ventral views of the head of: Hapsidophrys lineatus (A) (MNCN 23815), Hapsidophrys smaragdina (B) (MNCN 22572), and Philothamnus carinatus (C) (EBD 25022). Photos: AS-V." figureDoi="http://doi.org/10.5281/zenodo.7285675" httpUri="https://zenodo.org/record/7285675/files/figure.png" pageId="76" pageNumber="77">Figure 18C</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3716575FFEEFFDFFF4BFD6AFC2FE159" pageId="76" pageNumber="77" type="reference_group">
<paragraph id="8BD436FEFFEEFFDFFF4BFD6AFEDEE11C" blockId="76.[151,1436,759,875]" pageId="76" pageNumber="77">
<taxonomicName id="4C6B4D7DFFEEFFDFFF4BFD6AFDF0E13D" authority="Andersson, 1901: 9" authorityName="Andersson" authorityPageNumber="9" authorityYear="1901" box="[151,581,759,783]" class="Reptilia" family="Colubridae" genus="Chlorophis" kingdom="Animalia" order="Squamata" pageId="76" pageNumber="77" phylum="Chordata" rank="species" species="carinatus">
<emphasis id="B91FEAECFFEEFFDFFF4BFD6AFEC4E13D" box="[151,369,759,783]" italics="true" pageId="76" pageNumber="77">Chlorophis carinatus</emphasis>
Andersson, 1901: 9
</taxonomicName>
.
<typeStatus id="54D0885CFFEEFFDFFD8DFD65FD36E122" box="[593,643,760,784]" pageId="76" pageNumber="77">Type</typeStatus>
locality: “Mapanja,
<collectingCountry id="F37C766EFFEEFFDFFC87FD6AFC72E13D" box="[859,967,759,783]" name="Cameroon" pageId="76" pageNumber="77">Cameroon</collectingCountry>
” and “
<collectingCountry id="F37C766EFFEEFFDFFBD3FD6AFB35E13D" box="[1039,1152,759,783]" name="Cameroon" pageId="76" pageNumber="77">Cameroon</collectingCountry>
”.
<location id="8EB46025FFEEFFDFFB49FD6AFB49E13D" LSID="urn:lsid:plazi:treatment:03C287E8FFEEFFDFFF4BFD13FF5BE40E:8EB46025FFEEFFDFFB49FD6AFB49E13D" box="[1173,1276,759,783]" country="Cameroon" county="Mertens" municipality="Mapanja" name="Restricted" pageId="76" pageNumber="77">Restricted</location>
to
<collectingMunicipality id="6BB0AC84FFEEFFDFFAFDFD6AFACFE13D" box="[1313,1402,759,783]" pageId="76" pageNumber="77">Mapanja</collectingMunicipality>
by
<collectingCounty id="62B54E72FFEEFFDFFF1BFC8BFEAFE11F" box="[199,282,790,813]" pageId="76" pageNumber="77">Mertens</collectingCounty>
(1964).
</paragraph>
<paragraph id="8BD436FEFFEEFFDFFF4BFCA9FBA0E17F" blockId="76.[151,1436,759,875]" box="[151,1045,820,845]" pageId="76" pageNumber="77">
<treatmentCitationGroup id="AB7B11D0FFEEFFDFFF4BFCA9FBA0E17F" box="[151,1045,820,845]" pageId="76" pageNumber="77">
<treatmentCitation id="0ACA10EFFFEEFFDFFF4BFCA9FD43E17F" author="Peters, W. C. H." box="[151,758,820,845]" page="199" pageId="76" pageNumber="77" year="1875">
<taxonomicName id="4C6B4D7DFFEEFFDFFF4BFCA9FD43E17F" authority="Buchholz &amp; Peters, 1875: 199" authorityName="Buchholz &amp; Peters" authorityPageNumber="199" authorityYear="1875" box="[151,758,820,845]" class="Reptilia" family="Colubridae" genus="Philothamnus" kingdom="Animalia" order="Squamata" pageId="76" pageNumber="77" phylum="Chordata" rank="species" species="nigrofasciatus">
<emphasis id="B91FEAECFFEEFFDFFF4BFCA9FE0FE17F" box="[151,442,820,845]" italics="true" pageId="76" pageNumber="77">Philothamnus nigrofasciatus</emphasis>
Buchholz &amp;
<bibRefCitation id="EFFA4B0FFFEEFFDFFD9EFCA8FD43E17F" author="Peters, W. C. H." box="[578,758,820,845]" pageId="76" pageNumber="77" pagination="196 - 212" refId="ref86332" refString="Peters, W. C. H. (1875) Uber die von Hrn. Professor Dr. R. Buchholz in Westafrika gesammelten Amphibien. Monatsberichte der Koniglich preussischen Akademie der Wissenschaften zu Berlin, 1875, 196 - 212." type="journal article" year="1875">Peters, 1875: 199</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
<typeStatus id="54D0885CFFEEFFDFFD23FCA8FC84E17F" box="[767,817,821,845]" pageId="76" pageNumber="77">Type</typeStatus>
locality: “Cameruns”.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BD436FEFFEEFFDFFF4BFCCEFC2FE159" blockId="76.[151,1436,759,875]" box="[151,922,851,875]" pageId="76" pageNumber="77">
<treatmentCitationGroup id="AB7B11D0FFEEFFDFFF4BFCCEFC2FE159" box="[151,922,851,875]" pageId="76" pageNumber="77">
<taxonomicName id="4C6B4D7DFFEEFFDFFF4BFCCEFD7AE159" authority="Andersson, 1901" authorityName="Andersson" authorityYear="1901" box="[151,719,851,875]" class="Reptilia" family="Colubridae" genus="Philothamnus" kingdom="Animalia" order="Squamata" pageId="76" pageNumber="77" phylum="Chordata" rank="subSpecies" species="heterodermus" subSpecies="carinatus">
<emphasis id="B91FEAECFFEEFFDFFF4BFCCEFDAEE159" box="[151,539,851,875]" italics="true" pageId="76" pageNumber="77">Philothamnus heterodermus carinatus</emphasis>
Andersson, 1901
</taxonomicName>
:
<treatmentCitation id="0ACA10EFFFEEFFDFFD05FCCEFC23E159" author="Loveridge, A." box="[729,918,851,875]" page="6" pageId="76" pageNumber="77" year="1951">
<bibRefCitation id="EFFA4B0FFFEEFFDFFD05FCCEFC23E159" author="Loveridge, A." box="[729,918,851,875]" pageId="76" pageNumber="77" pagination="1 - 12" refId="ref83072" refString="Loveridge, A. (1951) Synopsis of the African green snakes (Philothamnus inc. Chlorophis) with the description of a new form. Bulletin de l'Institut Royal des Sciences Naturelles de Belgique, 27, 1 - 12." type="journal article" year="1951">Loveridge 1951: 6</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3716575FFEEFFDFFF4BFC04FE04E671" pageId="76" pageNumber="77" type="distribution">
<paragraph id="8BD436FEFFEEFFDFFF4BFC04FE04E671" blockId="76.[151,1437,921,1596]" pageId="76" pageNumber="77">
<emphasis id="B91FEAECFFEEFFDFFF4BFC04FE9CE181" bold="true" box="[151,297,921,947]" pageId="76" pageNumber="77">Distribution</emphasis>
. This species is widespread from West Africa (
<collectingCountry id="F37C766EFFEEFFDFFCF3FC04FC37E181" box="[815,898,921,947]" name="Guinea-Bissau" pageId="76" pageNumber="77">Guinea</collectingCountry>
Konakry) to East Africa (
<collectingCountry id="F37C766EFFEEFFDFFB43FC04FB5EE181" box="[1183,1259,921,947]" name="Kenya" pageId="76" pageNumber="77">Kenya</collectingCountry>
) and southward to southern
<collectingCountry id="F37C766EFFEEFFDFFEFEFC20FD0EE1E5" box="[290,699,957,983]" name="Democratic Republic of the Congo" pageId="76" pageNumber="77">Democratic Republic of the Congo</collectingCountry>
. In
<collectingCountry id="F37C766EFFEEFFDFFD36FC20FC0FE1E5" box="[746,954,957,983]" name="Equatorial Guinea" pageId="76" pageNumber="77">Equatorial Guinea</collectingCountry>
it has been recorded in Bioko at Bococo (
<bibRefCitation id="EFFA4B0FFFEEFFDFFF43FC7CFEF5E1C9" author="Mertens, R." box="[159,320,993,1019]" pageId="76" pageNumber="77" pagination="275 - 281" refId="ref84680" refString="Mertens, R. (1941) Zur Kenntnis der Herpetofauna von Fernando Poo. Zoologischer Anzeiger, 135, 275 - 281." type="journal article" year="1941">Mertens 1941</bibRefCitation>
;
<bibRefCitation id="EFFA4B0FFFEEFFDFFE92FC7CFDB0E1CE" author="Loveridge, A." box="[334,517,993,1020]" pageId="76" pageNumber="77" pagination="1 - 198" refId="ref83221" refString="Loveridge, A. (1958) Revision of five african snake genera. Bulletin of the Museum of Comparative Zoology, 119, 1 - 198." type="journal article" year="1958">Loveridge 1958</bibRefCitation>
), Moka (
<bibRefCitation id="EFFA4B0FFFEEFFDFFDB0FC7CFCADE1C9" author="Mertens, R." box="[620,792,993,1019]" pageId="76" pageNumber="77" pagination="211 - 238" refId="ref84765" refString="Mertens, R. (1964 a) Die Reptilien von Fernando Poo. Bonner zoologische Beitrage, 15, 211 - 238." type="journal article" year="1964">Mertens 1964a</bibRefCitation>
) and Bahía de San Carlos (
<bibRefCitation id="EFFA4B0FFFEEFFDFFB88FC7CFABBE1C9" author="Boulenger, G. A." box="[1108,1294,993,1019]" pageId="76" pageNumber="77" pagination="196 - 216" refId="ref72267" refString="Boulenger, G. A. (1906) Report on the reptiles collected by the late L. Fea in West Africa. Annali del Museo Civico di Storia Naturale di Genova, 3 (2), 196 - 216." type="journal article" year="1906">Boulenger 1906</bibRefCitation>
;
<bibRefCitation id="EFFA4B0FFFEEFFDFFAC7FC7CFF6AE62D" author="Capocaccia, L." pageId="76" pageNumber="77" pagination="285 - 309" refId="ref73329" refString="Capocaccia, L. (1961 a) Contributo allo studio dei serpenti delle isole del Golfo di Guinea. Annali del Museo Civico di Storia Naturale Giacomo Doria, 72, 285 - 309." type="journal article" year="1961">Capocaccia 1961a</bibRefCitation>
), and in
<collectingCountry id="F37C766EFFEEFFDFFE9AFB98FE01E62D" box="[326,436,1029,1055]" name="Equatorial Guinea" pageId="76" pageNumber="77">Río Muni</collectingCountry>
at Monte Alén National Park (
<bibRefCitation id="EFFA4B0FFFEEFFDFFCC4FB98FC51E62D" author="Lasso, C. A. &amp; Rial, A. I. &amp; Castroviejo, J. &amp; De la Riva, I." box="[792,996,1029,1055]" pageId="76" pageNumber="77" pagination="21 - 34" refId="ref81678" refString="Lasso, C. A., Rial, A. I., Castroviejo, J. &amp; De la Riva, I. (2002) Herpetofauna del Parque Nacional de Monte Alen (Rio Muni, Guinea Ecuatorial). Graellsia, 58, 21 - 34. https: // doi. org / 10.3989 / graellsia. 2002. v 58. i 2.276" type="journal article" year="2002">
Lasso
<emphasis id="B91FEAECFFEEFFDFFCB8FB9BFC2AE62D" box="[868,927,1029,1055]" italics="true" pageId="76" pageNumber="77">et al.</emphasis>
2002
</bibRefCitation>
), including Monte Mitra (
<bibRefCitation id="EFFA4B0FFFEEFFDFFAD1FB98FE95E676" author="Gonwouo, N. L. &amp; Nsang, A." pageId="76" pageNumber="77" pagination="32 - 49" refId="ref78060" refString="Gonwouo, N. L. &amp; Nsang, A. (2005) An assessment of the reptiles and amphibians of the Monte Mitra forest, Monte Alen National Park. In: Sunderland, T. C. H. (Ed.), A Biodiversity Assessment of the Monte Mitra Forest, Monte Alen National Park, Equatorial Guinea. Smithsonian Institution, Washington, D. C., pp. 32 - 49." type="book chapter" year="2005">Gonwouo &amp; Nsang 2005</bibRefCitation>
) (
<figureCitation id="13502A7BFFEEFFDFFEEAFBB4FE11E671" box="[310,420,1065,1091]" captionStart="MAPS 21" captionStartId="79.[151,223,1943,1967]" captionTargetBox="[202,1385,192,1908]" captionTargetId="figure-18@79.[159,1429,181,1919]" captionTargetPageId="79" captionText="MAPS 21 A–C. Distribution maps for Equatorial Guinean records of (A) Hapsidophrys smaragdina; (B) Philothamnus carinatus; (C) Philothamnus girardi." figureDoi="http://doi.org/10.5281/zenodo.7304727" httpUri="https://zenodo.org/record/7304727/files/figure.png" pageId="76" pageNumber="77">Map 21B</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3716575FFEEFFDFFF1BFBD0FF5BE40E" pageId="76" pageNumber="77" type="discussion">
<paragraph id="8BD436FEFFEEFFDFFF1BFBD0FE02E729" blockId="76.[151,1437,921,1596]" pageId="76" pageNumber="77">
<emphasis id="B91FEAECFFEEFFDFFF1BFBD0FEFAE655" bold="true" box="[199,335,1101,1127]" pageId="76" pageNumber="77">Comments.</emphasis>
All the specimens examined by us have 13 scale rows at mid-body. This species was treated by some authors as a subspecies of
<taxonomicName id="4C6B4D7DFFEEFFDFFE27FBEFFCCEE6BE" authority="(Loveridge 1958)" baseAuthorityName="Loveridge" baseAuthorityYear="1958" box="[507,891,1137,1164]" class="Reptilia" family="Colubridae" genus="Philothamnus" kingdom="Animalia" order="Squamata" pageId="76" pageNumber="77" phylum="Chordata" rank="species" species="heterodermus">
<emphasis id="B91FEAECFFEEFFDFFE27FBEFFD1AE6B9" box="[507,687,1137,1163]" italics="true" pageId="76" pageNumber="77">P. heterodermus</emphasis>
(
<bibRefCitation id="EFFA4B0FFFEEFFDFFD60FBECFCC6E6BE" author="Loveridge, A." box="[700,883,1137,1164]" pageId="76" pageNumber="77" pagination="1 - 198" refId="ref83221" refString="Loveridge, A. (1958) Revision of five african snake genera. Bulletin of the Museum of Comparative Zoology, 119, 1 - 198." type="journal article" year="1958">Loveridge 1958</bibRefCitation>
)
</taxonomicName>
. However, it has been suggested that
<taxonomicName id="4C6B4D7DFFEEFFDFFAC4FBEFFA29E6B9" authorityName="Andersson" authorityYear="1901" box="[1304,1436,1138,1163]" class="Reptilia" family="Colubridae" genus="Chlorophis" kingdom="Animalia" order="Squamata" pageId="76" pageNumber="77" phylum="Chordata" rank="species" species="carinatus">
<emphasis id="B91FEAECFFEEFFDFFAC4FBEFFA29E6B9" box="[1304,1436,1138,1163]" italics="true" pageId="76" pageNumber="77">P. carinatus</emphasis>
</taxonomicName>
could represent a species complex (
<bibRefCitation id="EFFA4B0FFFEEFFDFFDF4FB08FC83E69D" author="Engelbrecht, H. M. &amp; Branch, W. R. &amp; Greenbaum, E. &amp; Alexander, G. J. &amp; Jackson, K. &amp; Burger, M. &amp; Conradie, W. &amp; Kusamba, C. &amp; ZassiBoulou, A-G. &amp; Tolley, K. A." box="[552,822,1173,1199]" pageId="76" pageNumber="77" pagination="357 - 365" refId="ref76556" refString="Engelbrecht, H. M., Branch, W. R., Greenbaum, E., Alexander, G. J., Jackson, K., Burger, M., Conradie, W., Kusamba, C., ZassiBoulou, A-G. &amp; Tolley, K. A. (2019) Diversifying into the branches: species boundaries in African green and bush snakes, Philothamnus (Serpentes: Colubridae). Molecular Phylogenetics and Evolution, 130, 357 - 365. https: // doi. org / 10.1016 / j. ympev. 2018.10.023" type="journal article" year="2019">
Engelbrecht
<emphasis id="B91FEAECFFEEFFDFFD65FB0BFD47E69D" box="[697,754,1173,1199]" italics="true" pageId="76" pageNumber="77">et al.</emphasis>
2019
</bibRefCitation>
). It is formed by at least two main clades, one ranging across Gabon-western
<collectingCountry id="F37C766EFFEEFFDFFE40FB24FD11E6E1" box="[412,676,1209,1235]" name="Republic of the Congo" pageId="76" pageNumber="77">Republic of the Congo</collectingCountry>
and the other occurring at the eastern of
<collectingCountry id="F37C766EFFEEFFDFFB55FB24FEBAE6C5" name="Democratic Republic of the Congo" pageId="76" pageNumber="77">Democratic Republic of the Congo</collectingCountry>
(
<bibRefCitation id="EFFA4B0FFFEEFFDFFEFCFB40FD82E6C5" author="Engelbrecht, H. M. &amp; Branch, W. R. &amp; Greenbaum, E. &amp; Alexander, G. J. &amp; Jackson, K. &amp; Burger, M. &amp; Conradie, W. &amp; Kusamba, C. &amp; ZassiBoulou, A-G. &amp; Tolley, K. A." box="[288,567,1245,1271]" pageId="76" pageNumber="77" pagination="357 - 365" refId="ref76556" refString="Engelbrecht, H. M., Branch, W. R., Greenbaum, E., Alexander, G. J., Jackson, K., Burger, M., Conradie, W., Kusamba, C., ZassiBoulou, A-G. &amp; Tolley, K. A. (2019) Diversifying into the branches: species boundaries in African green and bush snakes, Philothamnus (Serpentes: Colubridae). Molecular Phylogenetics and Evolution, 130, 357 - 365. https: // doi. org / 10.1016 / j. ympev. 2018.10.023" type="journal article" year="2019">
Engelbrecht
<emphasis id="B91FEAECFFEEFFDFFE68FB43FE5AE6C5" box="[436,495,1245,1271]" italics="true" pageId="76" pageNumber="77">et al.</emphasis>
2019
</bibRefCitation>
). The
<taxonomicName id="4C6B4D7DFFEEFFDFFD5EFB40FC27E6C5" baseAuthorityName="Anderson" baseAuthorityYear="1901" box="[642,914,1245,1271]" class="Reptilia" family="Colubridae" genus="Philothamnus" kingdom="Animalia" order="Squamata" pageId="76" pageNumber="77" phylum="Chordata" rank="species" species="carinatus">
<emphasis id="B91FEAECFFEEFFDFFD5EFB40FC27E6C5" box="[642,914,1245,1271]" italics="true" pageId="76" pageNumber="77">Philothamnus carinatus</emphasis>
</taxonomicName>
species complex is sister to
<taxonomicName id="4C6B4D7DFFEEFFDFFB38FB43FE06E729" authority="(Engelbrecht et al. 2019)" baseAuthorityName="Engelbrecht" baseAuthorityYear="2019" class="Reptilia" family="Colubridae" genus="Philothamnus" kingdom="Animalia" order="Squamata" pageId="76" pageNumber="77" phylum="Chordata" rank="species" species="heterodermus">
<emphasis id="B91FEAECFFEEFFDFFB38FB43FA29E6C5" box="[1252,1436,1245,1271]" italics="true" pageId="76" pageNumber="77">P. heterodermus</emphasis>
(
<bibRefCitation id="EFFA4B0FFFEEFFDFFF43FA9CFE1EE729" author="Engelbrecht, H. M. &amp; Branch, W. R. &amp; Greenbaum, E. &amp; Alexander, G. J. &amp; Jackson, K. &amp; Burger, M. &amp; Conradie, W. &amp; Kusamba, C. &amp; ZassiBoulou, A-G. &amp; Tolley, K. A." box="[159,427,1281,1307]" pageId="76" pageNumber="77" pagination="357 - 365" refId="ref76556" refString="Engelbrecht, H. M., Branch, W. R., Greenbaum, E., Alexander, G. J., Jackson, K., Burger, M., Conradie, W., Kusamba, C., ZassiBoulou, A-G. &amp; Tolley, K. A. (2019) Diversifying into the branches: species boundaries in African green and bush snakes, Philothamnus (Serpentes: Colubridae). Molecular Phylogenetics and Evolution, 130, 357 - 365. https: // doi. org / 10.1016 / j. ympev. 2018.10.023" type="journal article" year="2019">
Engelbrecht
<emphasis id="B91FEAECFFEEFFDFFEF3FA9FFED2E729" box="[303,359,1281,1307]" italics="true" pageId="76" pageNumber="77">et al.</emphasis>
2019
</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
<paragraph id="8BD436FEFFEEFFDFFF1BFAB8FC88E79E" blockId="76.[151,1437,921,1596]" pageId="76" pageNumber="77">
The specimen recorded from
<collectingRegion id="49AFF81CFFEEFFDFFDC2FAB8FDEAE70D" box="[542,607,1317,1343]" country="Brazil" name="Bahia" pageId="76" pageNumber="77">Bahía</collectingRegion>
de San Carlos, Bioko, by
<bibRefCitation id="EFFA4B0FFFEEFFDFFC45FAB8FBD6E70D" author="Boulenger, G. A." box="[921,1123,1317,1343]" pageId="76" pageNumber="77" pagination="196 - 216" refId="ref72267" refString="Boulenger, G. A. (1906) Report on the reptiles collected by the late L. Fea in West Africa. Annali del Museo Civico di Storia Naturale di Genova, 3 (2), 196 - 216." type="journal article" year="1906">Boulenger (1906)</bibRefCitation>
was subsequently cited by
<bibRefCitation id="EFFA4B0FFFEEFFDFFF4BFAD4FEEBE756" author="Loveridge, A." box="[151,350,1353,1380]" pageId="76" pageNumber="77" pagination="1 - 198" refId="ref83221" refString="Loveridge, A. (1958) Revision of five african snake genera. Bulletin of the Museum of Comparative Zoology, 119, 1 - 198." type="journal article" year="1958">Loveridge (1958)</bibRefCitation>
as
<taxonomicName id="4C6B4D7DFFEEFFDFFE55FAD7FD56E751" authorityName="Loveridge" authorityYear="1958" box="[393,739,1353,1379]" class="Reptilia" family="Colubridae" genus="Philothamnus" kingdom="Animalia" order="Squamata" pageId="76" pageNumber="77" phylum="Chordata" rank="subSpecies" species="heterodermus" subSpecies="heterodermus">
<emphasis id="B91FEAECFFEEFFDFFE55FAD7FD56E751" box="[393,739,1353,1379]" italics="true" pageId="76" pageNumber="77">P. heterodermus heterodermus</emphasis>
</taxonomicName>
and re-identified by
<bibRefCitation id="EFFA4B0FFFEEFFDFFC05FAD4FB0FE751" author="Capocaccia, L." box="[985,1210,1353,1379]" pageId="76" pageNumber="77" pagination="285 - 309" refId="ref73329" refString="Capocaccia, L. (1961 a) Contributo allo studio dei serpenti delle isole del Golfo di Guinea. Annali del Museo Civico di Storia Naturale Giacomo Doria, 72, 285 - 309." type="journal article" year="1961">Capocaccia (1961a)</bibRefCitation>
as
<taxonomicName id="4C6B4D7DFFEEFFDFFB39FAD7FEB4E7B5" authorityName="Andersson" authorityYear="1901" class="Reptilia" family="Colubridae" genus="Philothamnus" kingdom="Animalia" order="Squamata" pageId="76" pageNumber="77" phylum="Chordata" rank="subSpecies" species="heterodermus" subSpecies="carinatus">
<emphasis id="B91FEAECFFEEFFDFFB39FAD7FEB4E7B5" italics="true" pageId="76" pageNumber="77">P. heterodermus carinatus</emphasis>
</taxonomicName>
. Based on the number of scale rows at mid-body (13 scale rows), we follow
<bibRefCitation id="EFFA4B0FFFEEFFDFFB54FAF0FADFE7B5" author="Capocaccia, L." box="[1160,1386,1389,1415]" pageId="76" pageNumber="77" pagination="285 - 309" refId="ref73329" refString="Capocaccia, L. (1961 a) Contributo allo studio dei serpenti delle isole del Golfo di Guinea. Annali del Museo Civico di Storia Naturale Giacomo Doria, 72, 285 - 309." type="journal article" year="1961">Capocaccia (1961a)</bibRefCitation>
and Mertens (1964) in considering this specimen as
<taxonomicName id="4C6B4D7DFFEEFFDFFD6DFA0FFC83E799" baseAuthorityName="Anderson" baseAuthorityYear="1901" box="[689,822,1426,1451]" class="Reptilia" family="Colubridae" genus="Philothamnus" kingdom="Animalia" order="Squamata" pageId="76" pageNumber="77" phylum="Chordata" rank="species" species="carinatus">
<emphasis id="B91FEAECFFEEFFDFFD6DFA0FFC83E799" box="[689,822,1426,1451]" italics="true" pageId="76" pageNumber="77">P. carinatus</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BD436FEFFEEFFDFFF1BFA28FF5BE40E" blockId="76.[151,1437,921,1596]" pageId="76" pageNumber="77">
<emphasis id="B91FEAECFFEEFFDFFF1BFA28FE7CE7FD" bold="true" box="[199,457,1461,1487]" pageId="76" pageNumber="77">Specimens examined.</emphasis>
<materialsCitation id="3B033CA3FFEEFFDFFE0EFA28FA97E7FD" ID-GBIF-Occurrence="3962324312" accessionNumber="EBD 20927" box="[466,1314,1461,1488]" collectionCode="EBD" country="Equatorial Guinea" location="Eleven" municipality="Miboman" pageId="76" pageNumber="77" specimenCode="EBD 20927-20930" specimenCount="1">
<location id="8EB46025FFEEFFDFFE0EFA28FD95E7FD" LSID="urn:lsid:plazi:treatment:03C287E8FFEEFFDFFF4BFD13FF5BE40E:8EB46025FFEEFFDFFE0EFA28FD95E7FD" box="[466,544,1461,1487]" country="Equatorial Guinea" municipality="Miboman" name="Eleven" pageId="76" pageNumber="77">Eleven</location>
specimens.
<collectingCountry id="F37C766EFFEEFFDFFD71FA28FC95E7FD" box="[685,800,1461,1487]" name="Equatorial Guinea" pageId="76" pageNumber="77">Río Muni</collectingCountry>
:
<collectingMunicipality id="6BB0AC84FFEEFFDFFCF0FA28FC29E7FD" box="[812,924,1461,1487]" pageId="76" pageNumber="77">Miboman</collectingMunicipality>
, 1985–1986 (
<specimenCode id="DBCD9E85FFEEFFDFFBE4FA28FAAEE7FD" box="[1080,1307,1461,1487]" collectionCode="EBD" country="Spain" httpUri="http://grbio.org/cool/ah5f-9iii" name="Estacion Biologica de Donana" pageId="76" pageNumber="77">EBD 20927–20930</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="3B033CA3FFEEFFDFFAF3FA28FC43E7C6" ID-GBIF-Occurrence="3962324432" accessionNumber="EBD 25018, EBD 25020" collectingDate="1987-02-25" collectionCode="EBD" country="Equatorial Guinea" location="Miboman" pageId="76" pageNumber="77" specimenCode="EBD 25020-25021" specimenCount="1">
<location id="8EB46025FFEEFFDFFAF3FA28FA29E7FD" LSID="urn:lsid:plazi:treatment:03C287E8FFEEFFDFFF4BFD13FF5BE40E:8EB46025FFEEFFDFFAF3FA28FA29E7FD" box="[1327,1436,1461,1487]" country="Equatorial Guinea" name="Miboman" pageId="76" pageNumber="77">Miboman</location>
(
<quantity id="4C939B1BFFEEFFDFFF43FA44FF5DE7C1" box="[159,232,1497,1523]" metricMagnitude="4" metricUnit="m" metricValue="3.0" pageId="76" pageNumber="77" unit="km" value="30.0">30 km</quantity>
from Niefang),
<date id="FFD5103EFFEEFFDFFE7DFA44FDDCE7C1" box="[417,617,1497,1524]" pageId="76" pageNumber="77" value="1987-02-25">
<collectingDate id="EF91E9D6FFEEFFDFFE7DFA44FDDCE7C1" box="[417,617,1497,1524]" pageId="76" pageNumber="77" value="1987-02-25">25 February 1987</collectingDate>
</date>
(
<collectionCode id="ED7AAE3BFFEEFFDFFDA5FA44FD06E7C1" box="[633,691,1497,1523]" country="Spain" httpUri="http://grbio.org/cool/ah5f-9iii" name="Estacion Biologica de Donana" pageId="76" pageNumber="77">EBD</collectionCode>
25018,
<specimenCode id="DBCD9E85FFEEFFDFFCD3FA47FC5BE7C6" box="[783,1006,1497,1524]" collectionCode="EBD" country="Spain" httpUri="http://grbio.org/cool/ah5f-9iii" name="Estacion Biologica de Donana" pageId="76" pageNumber="77">EBD 25020–25021</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="3B033CA3FFEEFFDFFBDEFA44FEC1E42A" ID-GBIF-Occurrence="3962324528" accessionNumber="EBD 27548" collectingDate="1987-07-28" collectionCode="EBD" country="Equatorial Guinea" pageId="76" pageNumber="77" specimenCount="1">
Miboman (road to Niefang),
<date id="FFD5103EFFEEFFDFFA96FA44FF7AE425" pageId="76" pageNumber="77" value="1987-07-28">
<collectingDate id="EF91E9D6FFEEFFDFFA96FA44FF7AE425" pageId="76" pageNumber="77" value="1987-07-28">28 July 1987</collectingDate>
</date>
(
<collectionCode id="ED7AAE3BFFEEFFDFFF03FA60FEACE425" box="[223,281,1533,1559]" country="Spain" httpUri="http://grbio.org/cool/ah5f-9iii" name="Estacion Biologica de Donana" pageId="76" pageNumber="77">EBD</collectionCode>
27548)
</materialsCitation>
;
<materialsCitation id="3B033CA3FFEEFFDFFE5CFA60FCC8E425" ID-GBIF-Occurrence="3962324348" accessionNumber="EBD 20792" box="[384,893,1533,1559]" collectingDate="1986-09-06" collectionCode="EBD" country="Equatorial Guinea" pageId="76" pageNumber="77" specimenCount="1">
Miboman,
<date id="FFD5103EFFEEFFDFFE20FA60FD6CE425" box="[508,729,1533,1559]" pageId="76" pageNumber="77" value="1986-09-06">
<collectingDate id="EF91E9D6FFEEFFDFFE20FA60FD6CE425" box="[508,729,1533,1559]" pageId="76" pageNumber="77" value="1986-09-06">06 September 1986</collectingDate>
</date>
(
<collectionCode id="ED7AAE3BFFEEFFDFFD35FA60FC96E425" box="[745,803,1533,1559]" country="Spain" httpUri="http://grbio.org/cool/ah5f-9iii" name="Estacion Biologica de Donana" pageId="76" pageNumber="77">EBD</collectionCode>
20792)
</materialsCitation>
;
<materialsCitation id="3B033CA3FFEEFFDFFC54FA60FF5FE40E" ID-GBIF-Occurrence="3962324759" accessionNumber="EBD 25019, EBD 25022" collectingDate="1986-08-01" country="Equatorial Guinea" pageId="76" pageNumber="77" specimenCount="1">
Ayamiken,
<date id="FFD5103EFFEEFFDFFBD5FA60FB08E425" box="[1033,1213,1533,1560]" pageId="76" pageNumber="77" value="1986-08-01">
<collectingDate id="EF91E9D6FFEEFFDFFBD5FA60FB08E425" box="[1033,1213,1533,1560]" pageId="76" pageNumber="77" value="1986-08-01">01 August 1986</collectingDate>
</date>
(EBD 25019, EBD 25022)
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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