treatments-xml/data/03/F4/77/03F47776FFDF5949FF5CBA10317F4480.xml

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<mods:title id="BBAF8BC63E20F3931A343ADBB8299812">Revision of the South American genus Tetragonopterus Cuvier, 1816 (Teleostei: Characidae) with description of four new species</mods:title>
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<mods:namePart id="79BFCE0C4A013A3EDA9103EC79D6023B">Silva, Gabriel S. C.</mods:namePart>
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<mods:namePart id="1AC50DC5C481C3446EAA4AE216988267">Melo, Bruno F.</mods:namePart>
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<mods:namePart id="D9A810CC75706ECB6F7E7DAB4FFD546D">Oliveira, Claudio</mods:namePart>
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<mods:namePart id="3AA6164832564315BB9D5762A54383BD">Benine, Ricardo C.</mods:namePart>
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<treatment id="03F47776FFDF5949FF5CBA10317F4480" ID-DOI="http://doi.org/10.5281/zenodo.4566214" ID-GBIF-Taxon="124864095" ID-Zenodo-Dep="4566214" LSID="urn:lsid:plazi:treatment:03F47776FFDF5949FF5CBA10317F4480" httpUri="http://treatment.plazi.org/id/03F47776FFDF5949FF5CBA10317F4480" lastPageId="25" lastPageNumber="25" pageId="22" pageNumber="23">
<subSubSection id="C34795EBFFDF5946FF5CBA10352E4744" pageId="22" pageNumber="23" type="nomenclature">
<paragraph id="8BE2C660FFDF5946FF5CBA10375B4722" blockId="22.[151,847,545,605]" box="[151,847,545,571]" pageId="22" pageNumber="23">
<heading id="D0AA710CFFDF5946FF5CBA10375B4722" bold="true" box="[151,847,545,571]" fontSize="11" level="1" pageId="22" pageNumber="23" reason="1">
<taxonomicName id="4C5DBDE3FFDF5946FF5CBA10375B4722" authority="Zarske, Gery &amp; Isbrucker, 2004" authorityName="Zarske, Gery &amp; Isbrucker" authorityYear="2004" box="[151,847,545,571]" class="Insecta" family="Anthribidae" genus="Tetragonopterus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="rarus">
<emphasis id="B9291A72FFDF5946FF5CBA10375B4722" bold="true" box="[151,847,545,571]" pageId="22" pageNumber="23">
<emphasis id="B9291A72FFDF5946FF5CBA1035B14722" bold="true" box="[151,421,545,571]" italics="true" pageId="22" pageNumber="23">Tetragonopterus rarus</emphasis>
(
<bibRefCitation id="EFCCBB91FFDF5946FE7FBA1037534722" author="Zarske" box="[436,839,545,571]" pageId="22" pageNumber="23" refString="Zarske, A., Gery, J. &amp; Isbrucker, I. J. (2004) Moenkhausia rara sp. n. - eine neue, bereits bestandsgefahrdete Salmler-Art (Teleostei: Characiformes: Characidae) aus Surinam und Franzosisch Guayana mit einer erganzenden Beschreibung von M. simulata (Eigenmann in Pearson, 1924). Zoologische Abhandlungen, 56, 3 - 14." type="journal article" year="2004">Zarske, Géry &amp; Isbrücker, 2004</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BE2C660FFDF5946FF5CBA75352E4744" blockId="22.[151,847,545,605]" box="[151,314,580,605]" pageId="22" pageNumber="23">
<figureCitation id="1366DAE5FFDF5946FF5CBA7534CE4744" box="[151,218,580,605]" captionStart="FIGURE 6" captionStartId="24.[151,250,1800,1822]" captionTargetBox="[239,1347,267,1778]" captionTargetId="figure@24.[239,1347,267,1778]" captionTargetPageId="24" captionText="FIGURE 6. Tetragonopterus rarus. a) MZUSP 88029, 85.4 mm SL, Brazil, Pará, Rio Paru, Amazon basin. Live specimens from (b) Marowijne and (c) Corantijn river basins, Suriname. Photos (b – c) by R. Covain." httpUri="https://zenodo.org/record/177603/files/figure.png" pageId="22" pageNumber="23">Fig. 6</figureCitation>
, 
<tableCitation id="C6DFF3DBFFDF5946FF2DBA75352E4744" box="[230,314,580,605]" captionStart="TABLE 5" captionStartId="23.[151,239,152,175]" captionTargetBox="[159,1197,203,1184]" captionTargetPageId="23" captionText="TABLE 5. Morphometric data for Tetragonopterus rarus (n = 12). Range includes paratypes." httpUri="http://table.plazi.org/id/DF2296E8FFDE5947FF5CB8A9307A45B6" pageId="22" pageNumber="23" tableUuid="DF2296E8FFDE5947FF5CB8A9307A45B6">Table 5</tableCitation>
</paragraph>
</subSubSection>
<subSubSection id="C34795EBFFDF5946FF5CBABD36004622" pageId="22" pageNumber="23" type="reference_group">
<paragraph id="8BE2C660FFDF5946FF5CBABD378A47D8" blockId="22.[151,1437,652,827]" pageId="22" pageNumber="23">
<treatmentCitationGroup id="AB4DE14EFFDF5946FF5CBABD378A47D8" pageId="22" pageNumber="23">
<taxonomicName id="4C5DBDE3FFDF5946FF5CBABD36D347BB" ID-CoL="73MZ9" authority="Zarske, Gery &amp; Isbrucker, 2004: 19" authorityName="Zarske, Gery &amp; Isbrucker" authorityPageNumber="19" authorityYear="2004" box="[151,711,652,674]" class="Actinopterygii" family="Characidae" genus="Moenkhausia" kingdom="Animalia" order="Characiformes" pageId="22" pageNumber="23" phylum="Chordata" rank="species" species="rara">
<emphasis id="B9291A72FFDF5946FF5CBABD354747BB" box="[151,339,652,674]" italics="true" pageId="22" pageNumber="23">Moenkhausia rara</emphasis>
<treatmentCitation id="0AFCE071FFDF5946FE90BABD36D347BB" author="Zarske" box="[347,711,652,674]" page="19" pageId="22" pageNumber="23" year="2004">
<bibRefCitation id="EFCCBB91FFDF5946FE90BABD36B247BB" author="Zarske" box="[347,678,652,674]" pageId="22" pageNumber="23" refString="Zarske, A., Gery, J. &amp; Isbrucker, I. J. (2004) Moenkhausia rara sp. n. - eine neue, bereits bestandsgefahrdete Salmler-Art (Teleostei: Characiformes: Characidae) aus Surinam und Franzosisch Guayana mit einer erganzenden Beschreibung von M. simulata (Eigenmann in Pearson, 1924). Zoologische Abhandlungen, 56, 3 - 14." type="journal article" year="2004">Zarske, Géry &amp; Isbrücker, 2004</bibRefCitation>
: 19
</treatmentCitation>
</taxonomicName>
–30 (original description; type locality: “Surinam, Marowijne, rapid in Oelemari (Ulemari) River); 
<treatmentCitation id="0AFCE071FFDF5946FE28BA9A36B847D8" author="Vari" box="[483,684,683,705]" page="32" pageId="22" pageNumber="23" year="2009">
<bibRefCitation id="EFCCBB91FFDF5946FE28BA9A369D47D8" author="Vari" box="[483,649,683,705]" pageId="22" pageNumber="23" refString="Vari, R. P., Ferraris, C. J. Jr, Radosavljevic, A. &amp; Funk, V. A. (2009) Checklist of the freshwater fishes of the Guiana Shield. Bulletin of the Biological Society of Washington, 17, 1 - 94. http: // dx. doi. org / 10.2988 / 0097 - 0298 - 17.1. i" type="journal article" year="2009">
Vari 
<emphasis id="B9291A72FFDF5946FDDFBA9D365347D8" box="[532,583,683,705]" italics="true" pageId="22" pageNumber="23">et al.</emphasis>
, 2009
</bibRefCitation>
: 32
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(listed, Guiana Shield).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BE2C660FFDF5946FF5CBAFB37DC47E7" blockId="22.[151,1437,652,827]" pageId="22" pageNumber="23">
<treatmentCitationGroup id="AB4DE14EFFDF5946FF5CBAFB37DC47E7" pageId="22" pageNumber="23">
<taxonomicName id="4C5DBDE3FFDF5946FF5CBAFB373147C6" authority="Benine, Pelicao &amp; Vari, 2004: 339" authorityName="Benine, Pelicao &amp; Vari" authorityPageNumber="339" authorityYear="2004" box="[151,805,713,735]" class="Insecta" family="Anthribidae" genus="Tetragonopterus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="lemniscatus">
<emphasis id="B9291A72FFDF5946FF5CBAFB35A847C6" box="[151,444,713,735]" italics="true" pageId="22" pageNumber="23">Tetragonopterus lemniscatus</emphasis>
<treatmentCitation id="0AFCE071FFDF5946FE0FBAF8373147C6" author="Benine" box="[452,805,713,735]" page="339" pageId="22" pageNumber="23" year="2004">
<bibRefCitation id="EFCCBB91FFDF5946FE0FBAF836E147C6" author="Benine" box="[452,757,713,735]" pageId="22" pageNumber="23" refString="Benine, R. C., Pelicao, G. Z. &amp; Vari, R. P. (2004) Tetragonopterus lemniscatus (Characiformes: Characidae), a new species from the Coratijn River basin in Suriname. Proceedings of the Biological Society of Washington, 117, 339 - 345." type="journal article" year="2004">Benine, Pelição &amp; Vari, 2004</bibRefCitation>
: 339
</treatmentCitation>
</taxonomicName>
–345 (original description; type locality: “Suriname, Nickerie District, tributary to Sisa Creek; 
<treatmentCitation id="0AFCE071FFDF5946FDDABAD936CE47E7" author="Vari" box="[529,730,744,766]" page="35" pageId="22" pageNumber="23" year="2009">
<bibRefCitation id="EFCCBB91FFDF5946FDDABAD936A347E7" author="Vari" box="[529,695,744,766]" pageId="22" pageNumber="23" refString="Vari, R. P., Ferraris, C. J. Jr, Radosavljevic, A. &amp; Funk, V. A. (2009) Checklist of the freshwater fishes of the Guiana Shield. Bulletin of the Biological Society of Washington, 17, 1 - 94. http: // dx. doi. org / 10.2988 / 0097 - 0298 - 17.1. i" type="journal article" year="2009">
Vari 
<emphasis id="B9291A72FFDF5946FD89BAD8366147E7" box="[578,629,744,766]" italics="true" pageId="22" pageNumber="23">et al.</emphasis>
, 2009
</bibRefCitation>
: 35
</treatmentCitation>
(listed, Guiana shield).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BE2C660FFDF5946FF5CBB3936004622" blockId="22.[151,1437,652,827]" pageId="22" pageNumber="23">
<treatmentCitationGroup id="AB4DE14EFFDF5946FF5CBB3936004622" pageId="22" pageNumber="23">
<taxonomicName id="4C5DBDE3FFDF5946FF5CBB3936FF4605" authority="Zarske, Gery &amp; Isbrucker, 2004" authorityName="Zarske, Gery &amp; Isbrucker" authorityYear="2004" box="[151,747,774,797]" class="Insecta" family="Anthribidae" genus="Tetragonopterus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="rarus">
<emphasis id="B9291A72FFDF5946FF5CBB39356A4604" box="[151,382,776,797]" italics="true" pageId="22" pageNumber="23">Tetragonopterus rarus</emphasis>
(
<bibRefCitation id="EFCCBB91FFDF5946FE45BB3736F14605" author="Zarske" box="[398,741,774,796]" pageId="22" pageNumber="23" refString="Zarske, A., Gery, J. &amp; Isbrucker, I. J. (2004) Moenkhausia rara sp. n. - eine neue, bereits bestandsgefahrdete Salmler-Art (Teleostei: Characiformes: Characidae) aus Surinam und Franzosisch Guayana mit einer erganzenden Beschreibung von M. simulata (Eigenmann in Pearson, 1924). Zoologische Abhandlungen, 56, 3 - 14." type="journal article" year="2004">Zarske, Géry &amp; Isbrücker, 2004</bibRefCitation>
)
</taxonomicName>
: 
<treatmentCitation id="0AFCE071FFDF5946FD32BB3737CD4605" author="Melo" box="[761,985,774,797]" page="49" pageId="22" pageNumber="23" year="2011">
<bibRefCitation id="EFCCBB91FFDF5946FD32BB3737A74605" author="Melo" box="[761,947,774,797]" pageId="22" pageNumber="23" refString="Melo, B. F., Benine, R. C., Mariguela, T. C. &amp; Oliveira, C. (2011) A new species of Tetragonopterus Cuvier, 1816 (Characiformes: Characidae: Tetragonopterinae) from the rio Jari, Amapa, northern Brazil. Neotropical Ichthyology, 9, 49 - 56. http: // dx. doi. org / 10.1590 / S 1679 - 62252011000100002" type="journal article" year="2011">
Melo 
<emphasis id="B9291A72FFDF5946FCF3BB39377A4605" box="[824,878,774,797]" italics="true" pageId="22" pageNumber="23">et al.</emphasis>
, 2011
</bibRefCitation>
: 49
</treatmentCitation>
–56 (new combination); 
<treatmentCitation id="0AFCE071FFDF5946FB16BB3734F94622" author="Melo" page="709" pageId="22" pageNumber="23" year="2016">
<bibRefCitation id="EFCCBB91FFDF5946FB16BB37318C4605" author="Melo" box="[1245,1432,774,797]" pageId="22" pageNumber="23" refString="Melo, B. F., Benine, R. C., Silva, G. S. C., Avelino, G. S. &amp; Oliveira, C. (2016) Molecular phylogeny of the Neotropical fish genus Tetragonopterus (Teleostei: Characiformes: Characidae). Molecular Phylogenetics and Evolution, 94, 709 - 717. http: // dx. doi. org / 10.1016 / j. ympev. 2015.10.022" type="journal article" year="2016">
Melo 
<emphasis id="B9291A72FFDF5946FAD7BB3931474605" box="[1308,1363,774,797]" italics="true" pageId="22" pageNumber="23">et al.</emphasis>
, 2016
</bibRefCitation>
: 709
</treatmentCitation>
–717 (molecular phylogeny).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C34795EBFFDF5946FF5CBB5936D1404D" pageId="22" pageNumber="23" type="diagnosis">
<paragraph id="8BE2C660FFDF5946FF5CBB5936D1404D" blockId="22.[151,1437,872,2013]" pageId="22" pageNumber="23">
<emphasis id="B9291A72FFDF5946FF5CBB5935054698" bold="true" box="[151,273,872,897]" pageId="22" pageNumber="23">Diagnosis.</emphasis>
<taxonomicName id="4C5DBDE3FFDF5946FED2BB5836024699" box="[281,534,873,896]" class="Insecta" family="Anthribidae" genus="Tetragonopterus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="rarus">
<emphasis id="B9291A72FFDF5946FED2BB5836024699" box="[281,534,873,896]" italics="true" pageId="22" pageNumber="23">Tetragonopterus rarus</emphasis>
</taxonomicName>
is distinguished from all congeners by the presence of longitudinal dark stripes on the lateral surface of body (
<figureCitation id="1366DAE5FFDF5946FDD6BBBD367C46BC" box="[541,616,908,933]" captionStart="FIGURE 6" captionStartId="24.[151,250,1800,1822]" captionTargetBox="[239,1347,267,1778]" captionTargetId="figure@24.[239,1347,267,1778]" captionTargetPageId="24" captionText="FIGURE 6. Tetragonopterus rarus. a) MZUSP 88029, 85.4 mm SL, Brazil, Pará, Rio Paru, Amazon basin. Live specimens from (b) Marowijne and (c) Corantijn river basins, Suriname. Photos (b – c) by R. Covain." httpUri="https://zenodo.org/record/177603/files/figure.png" pageId="22" pageNumber="23">Fig. 6</figureCitation>
) (
<emphasis id="B9291A72FFDF5946FD43BBBC36B346BD" box="[648,679,909,932]" italics="true" pageId="22" pageNumber="23">vs.</emphasis>
absence of longitudinal dark stripes). In addition, 
<taxonomicName id="4C5DBDE3FFDF5946FAD1BBBC316E46BD" box="[1306,1402,909,932]" class="Insecta" family="Anthribidae" genus="Tetragonopterus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="rarus">
<emphasis id="B9291A72FFDF5946FAD1BBBC316E46BD" box="[1306,1402,909,932]" italics="true" pageId="22" pageNumber="23">T. rarus</emphasis>
</taxonomicName>
is distinguished from all congeners, except 
<taxonomicName id="4C5DBDE3FFDF5946FDA8BB8036F546D1" box="[611,737,945,968]" class="Actinopterygii" family="Characidae" genus="Moenkhausia" kingdom="Animalia" order="Characiformes" pageId="22" pageNumber="23" phylum="Chordata" rank="species" species="georgiae">
<emphasis id="B9291A72FFDF5946FDA8BB8036F546D1" box="[611,737,945,968]" italics="true" pageId="22" pageNumber="23">T. georgiae</emphasis>
</taxonomicName>
, by the number of longitudinal scale rows between lateral line and pelvic-fin origin (4.5–5.5 
<emphasis id="B9291A72FFDF5946FE23BBE4361246F5" box="[488,518,981,1004]" italics="true" pageId="22" pageNumber="23">vs.</emphasis>
3.5). 
<taxonomicName id="4C5DBDE3FFDF5946FD8CBBE4375646F5" box="[583,834,981,1004]" class="Insecta" family="Anthribidae" genus="Tetragonopterus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="rarus">
<emphasis id="B9291A72FFDF5946FD8CBBE4375646F5" box="[583,834,981,1004]" italics="true" pageId="22" pageNumber="23">Tetragonopterus rarus</emphasis>
</taxonomicName>
differs from 
<emphasis id="B9291A72FFDF5946FC1DBBE4310846F4" box="[982,1308,980,1005]" italics="true" pageId="22" pageNumber="23">
<taxonomicName id="4C5DBDE3FFDF5946FC1DBBE4307946F5" box="[982,1133,981,1004]" class="Insecta" family="Anthribidae" genus="Tetragonopterus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="anostomus">T. anostomus</taxonomicName>
, 
<taxonomicName id="4C5DBDE3FFDF5946FBBCBBE4310846F4" box="[1143,1308,980,1005]" class="Insecta" family="Anthribidae" genus="Tetragonopterus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="denticulatus">T. denticulatus</taxonomicName>
</emphasis>
, 
<taxonomicName id="4C5DBDE3FFDF5946FAE2BBE4318846F5" box="[1321,1436,981,1004]" class="Insecta" family="Anthribidae" genus="Tetragonopterus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="juruena">
<emphasis id="B9291A72FFDF5946FAE2BBE4318846F5" box="[1321,1436,981,1004]" italics="true" pageId="22" pageNumber="23">T. juruena</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C5DBDE3FFDF5946FF0DBBC8352D4108" box="[198,313,1016,1041]" class="Insecta" family="Anthribidae" genus="Tetragonopterus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="kuluene">
<emphasis id="B9291A72FFDF5946FF0DBBC8352D4108" box="[198,313,1016,1041]" italics="true" pageId="22" pageNumber="23">T. kuluene</emphasis>
</taxonomicName>
by the presence of four principal dentary teeth (
<emphasis id="B9291A72FFDF5946FC9FBBC837674109" box="[852,883,1017,1040]" italics="true" pageId="22" pageNumber="23">vs.</emphasis>
5–6) and by possessing relatively robust teeth on dentary (
<emphasis id="B9291A72FFDF5946FF30BC2C350D412D" box="[251,281,1053,1076]" italics="true" pageId="22" pageNumber="23">vs.</emphasis>
sharper teeth). 
<taxonomicName id="4C5DBDE3FFDF5946FE0DBC2C36D6412D" box="[454,706,1053,1076]" class="Insecta" family="Anthribidae" genus="Tetragonopterus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="rarus">
<emphasis id="B9291A72FFDF5946FE0DBC2C36D6412D" box="[454,706,1053,1076]" italics="true" pageId="22" pageNumber="23">Tetragonopterus rarus</emphasis>
</taxonomicName>
differs from 
<taxonomicName id="4C5DBDE3FFDF5946FC9EBC2C37FC412D" box="[853,1000,1053,1076]" class="Insecta" family="Anthribidae" genus="Tetragonopterus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="anostomus">
<emphasis id="B9291A72FFDF5946FC9EBC2C37FC412D" box="[853,1000,1053,1076]" italics="true" pageId="22" pageNumber="23">T. anostomus</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C5DBDE3FFDF5946FBD5BC2C30DE412D" box="[1054,1226,1053,1076]" class="Insecta" family="Anthribidae" genus="Tetragonopterus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="araguaiensis">
<emphasis id="B9291A72FFDF5946FBD5BC2C30DE412D" box="[1054,1226,1053,1076]" italics="true" pageId="22" pageNumber="23">T. araguaiensis</emphasis>
</taxonomicName>
by the presence of 12–13 gill-rakers on the lower limb of the first gill arch (
<emphasis id="B9291A72FFDF5946FCE1BC70375D4141" box="[810,841,1089,1112]" italics="true" pageId="22" pageNumber="23">vs.</emphasis>
16–20), and from 
<taxonomicName id="4C5DBDE3FFDF5946FBE9BC7030BF4140" box="[1058,1195,1088,1113]" class="Insecta" family="Anthribidae" genus="Tetragonopterus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="carvalhoi">
<emphasis id="B9291A72FFDF5946FBE9BC7030BF4140" box="[1058,1195,1088,1113]" italics="true" pageId="22" pageNumber="23">T. carvalhoi</emphasis>
</taxonomicName>
by the presence of a rounded dark mark on the caudal peduncle (
<emphasis id="B9291A72FFDF5946FD53BC5436A24165" box="[664,694,1125,1148]" italics="true" pageId="22" pageNumber="23">vs.</emphasis>
lozengular dark mark on the caudal peduncle). 
<taxonomicName id="4C5DBDE3FFDF5946FB2FBC5434C041B9" class="Insecta" family="Anthribidae" genus="Tetragonopterus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="rarus">
<emphasis id="B9291A72FFDF5946FB2FBC5434C041B9" italics="true" pageId="22" pageNumber="23">Tetragonopterus rarus</emphasis>
</taxonomicName>
is further distinguished from 
<taxonomicName id="4C5DBDE3FFDF5946FDEBBCB836B541B9" box="[544,673,1161,1184]" class="Insecta" family="Anthribidae" genus="Tetragonopterus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="ommatus">
<emphasis id="B9291A72FFDF5946FDEBBCB836B541B9" box="[544,673,1161,1184]" italics="true" pageId="22" pageNumber="23">T. ommatus</emphasis>
</taxonomicName>
by having 2–4 teeth on the maxilla (
<emphasis id="B9291A72FFDF5946FBF7BCB8304F41B9" box="[1084,1115,1161,1184]" italics="true" pageId="22" pageNumber="23">vs.</emphasis>
7–8) and by having the dark mark centered on caudal peduncle (
<emphasis id="B9291A72FFDF5946FDE1BC9C365C41DD" box="[554,584,1197,1220]" italics="true" pageId="22" pageNumber="23">vs.</emphasis>
mark limited to the posterior portion of caudal peduncle); it differs from 
<taxonomicName id="4C5DBDE3FFDF5946FA4CBC9C351B41F1" class="Insecta" family="Anthribidae" genus="Tetragonopterus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="anostomus">
<emphasis id="B9291A72FFDF5946FA4CBC9C351B41F1" italics="true" pageId="22" pageNumber="23">T. anostomus</emphasis>
</taxonomicName>
by having terminal mouth (
<emphasis id="B9291A72FFDF5946FD8CBCE0367241F1" box="[583,614,1233,1256]" italics="true" pageId="22" pageNumber="23">vs.</emphasis>
subsuperior mouth) and from 
<taxonomicName id="4C5DBDE3FFDF5946FC71BCE0305941F1" box="[954,1101,1233,1256]" class="Insecta" family="Anthribidae" genus="Tetragonopterus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="anostomus">
<emphasis id="B9291A72FFDF5946FC71BCE0305941F1" box="[954,1101,1233,1256]" italics="true" pageId="22" pageNumber="23">T. anostomus</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C5DBDE3FFDF5946FB48BCE030E141F0" box="[1155,1269,1232,1257]" class="Insecta" family="Anthribidae" genus="Tetragonopterus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="kuluene">
<emphasis id="B9291A72FFDF5946FB48BCE030E141F0" box="[1155,1269,1232,1257]" italics="true" pageId="22" pageNumber="23">T. kuluene</emphasis>
</taxonomicName>
by having two humeral marks (
<emphasis id="B9291A72FFDF5946FE9BBCC4357A4015" box="[336,366,1269,1292]" italics="true" pageId="22" pageNumber="23">vs.</emphasis>
only one humeral mark). 
<taxonomicName id="4C5DBDE3FFDF5946FD5FBCC437854015" box="[660,913,1269,1292]" class="Insecta" family="Anthribidae" genus="Tetragonopterus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="rarus">
<emphasis id="B9291A72FFDF5946FD5FBCC437854015" box="[660,913,1269,1292]" italics="true" pageId="22" pageNumber="23">Tetragonopterus rarus</emphasis>
</taxonomicName>
differs from 
<taxonomicName id="4C5DBDE3FFDF5946FBEDBCC430A44015" box="[1062,1200,1269,1292]" class="Actinopterygii" family="Prochilodontidae" genus="Prochilodus" kingdom="Animalia" order="Characiformes" pageId="22" pageNumber="23" phylum="Chordata" rank="species" species="argenteus">
<emphasis id="B9291A72FFDF5946FBEDBCC430A44015" box="[1062,1200,1269,1292]" italics="true" pageId="22" pageNumber="23">T. argenteus</emphasis>
</taxonomicName>
by the presence of 8 predorsal scales (
<emphasis id="B9291A72FFDF5946FE92BD28356C4029" box="[345,376,1305,1328]" italics="true" pageId="22" pageNumber="23">vs.</emphasis>
11–17) and from 
<taxonomicName id="4C5DBDE3FFDF5946FD89BD2836F24028" box="[578,742,1304,1329]" class="Insecta" family="Anthribidae" genus="Tetragonopterus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="denticulatus">
<emphasis id="B9291A72FFDF5946FD89BD2836F24028" box="[578,742,1304,1329]" italics="true" pageId="22" pageNumber="23">T. denticulatus</emphasis>
</taxonomicName>
by having humeral marks separated by one vertical scale row (
<emphasis id="B9291A72FFDF5946FF6BBD0C34AA404D" box="[160,190,1341,1364]" italics="true" pageId="22" pageNumber="23">vs.</emphasis>
marks separated by three vertical scale rows).
</paragraph>
</subSubSection>
<subSubSection id="C34795EBFFDF5947FF0CBD51367A429C" lastPageId="23" lastPageNumber="24" pageId="22" pageNumber="23" type="description">
<paragraph id="8BE2C660FFDF5946FF0CBD51369C4349" blockId="22.[151,1437,872,2013]" pageId="22" pageNumber="23">
<emphasis id="B9291A72FFDF5946FF0CBD51354D4060" bold="true" box="[199,345,1376,1401]" pageId="22" pageNumber="23">Description.</emphasis>
Morphometric data summarized in 
<tableCitation id="C6DFF3DBFFDF5946FD21BD5137544060" box="[746,832,1376,1401]" captionStart="TABLE 5" captionStartId="23.[151,239,152,175]" captionTargetBox="[159,1197,203,1184]" captionTargetPageId="23" captionText="TABLE 5. Morphometric data for Tetragonopterus rarus (n = 12). Range includes paratypes." httpUri="http://table.plazi.org/id/DF2296E8FFDE5947FF5CB8A9307A45B6" pageId="22" pageNumber="23" tableUuid="DF2296E8FFDE5947FF5CB8A9307A45B6">Table 5</tableCitation>
. Body shape compressed, moderately elongated when compared with congeners. Greatest body depth at dorsal-fin origin. Dorsal profile slightly convex between tip of snout and vertical through middle of orbit; slightly concave from this point to end of occipital process, convex from tip of occipital process to dorsal-fin origin; then slightly convex between dorsal-fin base and adipose-fin end. Prepelvic region transversely flattened with distinct, longitudinally aligned lateral keels. Ventral profile of body convex from lower lip to anal-fin origin; slightly convex or straight along base of anal fin. Caudal peduncle with slightly concave dorsal and ventral profiles.
</paragraph>
<paragraph id="8BE2C660FFDF5946FF0CBE6D36B7421C" blockId="22.[151,1437,872,2013]" pageId="22" pageNumber="23">Mouth terminal. Upper and lower jaws of similar size. Premaxilla in two rows of relatively robust teeth. Outer row with 4 (3), 5 (7) or 6 (1) teeth with three cusps, central cusps longest. Inner row with 5 (11) teeth with three or five cusps, central cusps twice as long as the lateral cusps. Maxilla with 2 (2), 3 (6) or 4 (3) tricuspidate teeth. Dentary with 4 (5) or 5 (6) anteriormost teeth with 3 or 5 cusps, followed by one smaller teeth with three cusps and then several small conical teeth of similar size.</paragraph>
<paragraph id="8BE2C660FFDF5946FF0CBF21374A42C4" blockId="22.[151,1437,872,2013]" pageId="22" pageNumber="23">Dorsal-fin rays ii,9 (11). First unbranched ray shorter than second one. Dorsal-fin origin slightly anterior to middle of body. Anal-fin rays iv,28 (1), iv,30 (4), iv,31 (4) or iv,33 (2); posterior unbranched rays and anterior branched rays usually slightly longer than following rays. Anal-fin origin situated posterior to vertical line through posterior most branched dorsal-fin ray. Pelvic-fin rays i,7 (11); origin situated anterior to vertical line through dorsal-fin origin; tip of adpressed pelvic fin reaching one or two scales before first unbranched anal-fin ray. Pectoral-fin rays i,12 (1), i,13 (7) or i,14 (3). Caudal fin forked.</paragraph>
<caption id="DF2296E8FFDE5947FF5CB8A9307A45B6" ID-Table-UUID="DF2296E8FFDE5947FF5CB8A9307A45B6" box="[151,1134,152,175]" httpUri="http://table.plazi.org/id/DF2296E8FFDE5947FF5CB8A9307A45B6" pageId="23" pageNumber="24" targetBox="[159,1197,203,1184]" targetIsTable="true" targetPageId="23">
<paragraph id="8BE2C660FFDE5947FF5CB8A9307A45B6" blockId="23.[151,1134,152,175]" box="[151,1134,152,175]" pageId="23" pageNumber="24">
<emphasis id="B9291A72FFDE5947FF5CB8A9351E45B6" bold="true" box="[151,266,152,175]" pageId="23" pageNumber="24">TABLE 5.</emphasis>
Morphometric data for 
<taxonomicName id="4C5DBDE3FFDE5947FDC1B8A936EE45B6" box="[522,762,152,175]" class="Insecta" family="Anthribidae" genus="Tetragonopterus" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="rarus">
<emphasis id="B9291A72FFDE5947FDC1B8A936EE45B6" box="[522,762,152,175]" italics="true" pageId="23" pageNumber="24">Tetragonopterus rarus</emphasis>
</taxonomicName>
(n=12). Range includes paratypes.
</paragraph>
</caption>
<paragraph id="8BE2C660FFDE5947FD05B8FA308B41B9" pageId="23" pageNumber="24">
<table id="F95D34C0FFDEA6AFFF54B8FA30B941B9" box="[159,1197,203,1184]" gridcols="3" gridrows="25" pageId="23" pageNumber="24">
<tr id="356DC422FFDEA6AFFF54B8FA30B945F9" box="[159,1197,203,224]" gridrow="0" pageId="23" pageNumber="24" rowspan-0="1">
<th id="76BCAD5EFFDEA6AFFD05B8FA372045F9" box="[718,820,203,224]" gridcol="1" gridrow="0" pageId="23" pageNumber="24">Range</th>
<th id="76BCAD5EFFDEA6AFFBB8B8FA30B945F9" box="[1139,1197,203,224]" gridcol="2" gridrow="0" pageId="23" pageNumber="24">Mean</th>
</tr>
<tr id="356DC422FFDEA6AFFF54B8C330B94411" box="[159,1197,242,264]" gridrow="1" pageId="23" pageNumber="24">
<th id="76BCAD5EFFDEA6AFFF54B8C336814411" box="[159,661,242,264]" gridcol="0" gridrow="1" pageId="23" pageNumber="24">Standard length (mm)</th>
<td id="76BCAD5EFFDEA6AFFD05B8C337204411" box="[718,820,242,264]" gridcol="1" gridrow="1" pageId="23" pageNumber="24">42.1–85.4</td>
<td id="76BCAD5EFFDEA6AFFBB8B8C330B94411" box="[1139,1197,242,264]" gridcol="2" gridrow="1" pageId="23" pageNumber="24">60.7</td>
</tr>
<tr id="356DC422FFDEA6AFFF54B92B30B94429" box="[159,1197,282,304]" gridrow="2" pageId="23" pageNumber="24" rowspan-1="1" rowspan-2="1">
<th id="76BCAD5EFFDEA6AFFF54B92B36814429" box="[159,661,282,304]" gridcol="0" gridrow="2" pageId="23" pageNumber="24">Percentages of standard length</th>
</tr>
<tr id="356DC422FFDEA6AFFF54B97330B94441" box="[159,1197,322,344]" gridrow="3" pageId="23" pageNumber="24">
<th id="76BCAD5EFFDEA6AFFF54B97336814441" box="[159,661,322,344]" gridcol="0" gridrow="3" pageId="23" pageNumber="24">Greatest depth</th>
<td id="76BCAD5EFFDEA6AFFD05B97337204441" box="[718,820,322,344]" gridcol="1" gridrow="3" pageId="23" pageNumber="24">46.0–52.6</td>
<td id="76BCAD5EFFDEA6AFFBB8B97330B94441" box="[1139,1197,322,344]" gridcol="2" gridrow="3" pageId="23" pageNumber="24">49.4</td>
</tr>
<tr id="356DC422FFDEA6AFFF54B95B30B94499" box="[159,1197,362,384]" gridrow="4" pageId="23" pageNumber="24">
<th id="76BCAD5EFFDEA6AFFF54B95B36814499" box="[159,661,362,384]" gridcol="0" gridrow="4" pageId="23" pageNumber="24">Predorsal length</th>
<td id="76BCAD5EFFDEA6AFFD05B95B37204499" box="[718,820,362,384]" gridcol="1" gridrow="4" pageId="23" pageNumber="24">46.9–53.2</td>
<td id="76BCAD5EFFDEA6AFFBB8B95B30B94499" box="[1139,1197,362,384]" gridcol="2" gridrow="4" pageId="23" pageNumber="24">50.8</td>
</tr>
<tr id="356DC422FFDEA6AFFF54B9A330B944B1" box="[159,1197,402,424]" gridrow="5" pageId="23" pageNumber="24">
<th id="76BCAD5EFFDEA6AFFF54B9A3368144B1" box="[159,661,402,424]" gridcol="0" gridrow="5" pageId="23" pageNumber="24">Prepectoral length</th>
<td id="76BCAD5EFFDEA6AFFD05B9A3372044B1" box="[718,820,402,424]" gridcol="1" gridrow="5" pageId="23" pageNumber="24">27.0–29.3</td>
<td id="76BCAD5EFFDEA6AFFBB8B9A330B944B1" box="[1139,1197,402,424]" gridcol="2" gridrow="5" pageId="23" pageNumber="24">27.5</td>
</tr>
<tr id="356DC422FFDEA6AFFF54B98B30B944C9" box="[159,1197,442,464]" gridrow="6" pageId="23" pageNumber="24">
<th id="76BCAD5EFFDEA6AFFF54B98B368144C9" box="[159,661,442,464]" gridcol="0" gridrow="6" pageId="23" pageNumber="24">Prepelvic length</th>
<td id="76BCAD5EFFDEA6AFFD05B98B372044C9" box="[718,820,442,464]" gridcol="1" gridrow="6" pageId="23" pageNumber="24">45.8–50.4</td>
<td id="76BCAD5EFFDEA6AFFBB8B98B30B944C9" box="[1139,1197,442,464]" gridcol="2" gridrow="6" pageId="23" pageNumber="24">48.5</td>
</tr>
<tr id="356DC422FFDEA6AFFF54B9D330B944E1" box="[159,1197,482,504]" gridrow="7" pageId="23" pageNumber="24">
<th id="76BCAD5EFFDEA6AFFF54B9D3368144E1" box="[159,661,482,504]" gridcol="0" gridrow="7" pageId="23" pageNumber="24">Preanal length</th>
<td id="76BCAD5EFFDEA6AFFD05B9D3372044E1" box="[718,820,482,504]" gridcol="1" gridrow="7" pageId="23" pageNumber="24">61.8–68.2</td>
<td id="76BCAD5EFFDEA6AFFBB8B9D330B944E1" box="[1139,1197,482,504]" gridcol="2" gridrow="7" pageId="23" pageNumber="24">66.4</td>
</tr>
<tr id="356DC422FFDEA6AFFF54BA3B30B94739" box="[159,1197,522,544]" gridrow="8" pageId="23" pageNumber="24">
<th id="76BCAD5EFFDEA6AFFF54BA3B36814739" box="[159,661,522,544]" gridcol="0" gridrow="8" pageId="23" pageNumber="24">Caudal peduncle depth</th>
<td id="76BCAD5EFFDEA6AFFD05BA3B37204739" box="[718,820,522,544]" gridcol="1" gridrow="8" pageId="23" pageNumber="24">8.3–11.8</td>
<td id="76BCAD5EFFDEA6AFFBB8BA3B30B94739" box="[1139,1197,522,544]" gridcol="2" gridrow="8" pageId="23" pageNumber="24">10.3</td>
</tr>
<tr id="356DC422FFDEA6AFFF54BA0330B94751" box="[159,1197,562,584]" gridrow="9" pageId="23" pageNumber="24">
<th id="76BCAD5EFFDEA6AFFF54BA0336814751" box="[159,661,562,584]" gridcol="0" gridrow="9" pageId="23" pageNumber="24">Caudal peduncle length</th>
<td id="76BCAD5EFFDEA6AFFD05BA0337204751" box="[718,820,562,584]" gridcol="1" gridrow="9" pageId="23" pageNumber="24">6.3–9.4</td>
<td id="76BCAD5EFFDEA6AFFBB8BA0330B94751" box="[1139,1197,562,584]" gridcol="2" gridrow="9" pageId="23" pageNumber="24">8.5</td>
</tr>
<tr id="356DC422FFDEA6AFFF54BA6B30B94769" box="[159,1197,602,624]" gridrow="10" pageId="23" pageNumber="24">
<th id="76BCAD5EFFDEA6AFFF54BA6B36814769" box="[159,661,602,624]" gridcol="0" gridrow="10" pageId="23" pageNumber="24">Pectoral-fin length</th>
<td id="76BCAD5EFFDEA6AFFD05BA6B37204769" box="[718,820,602,624]" gridcol="1" gridrow="10" pageId="23" pageNumber="24">22.2–23.9</td>
<td id="76BCAD5EFFDEA6AFFBB8BA6B30B94769" box="[1139,1197,602,624]" gridcol="2" gridrow="10" pageId="23" pageNumber="24">23.3</td>
</tr>
<tr id="356DC422FFDEA6AFFF54BAB330B94781" box="[159,1197,642,664]" gridrow="11" pageId="23" pageNumber="24">
<th id="76BCAD5EFFDEA6AFFF54BAB336814781" box="[159,661,642,664]" gridcol="0" gridrow="11" pageId="23" pageNumber="24">Pelvic-fin length</th>
<td id="76BCAD5EFFDEA6AFFD05BAB337204781" box="[718,820,642,664]" gridcol="1" gridrow="11" pageId="23" pageNumber="24">14.6–24.2</td>
<td id="76BCAD5EFFDEA6AFFBB8BAB330B94781" box="[1139,1197,642,664]" gridcol="2" gridrow="11" pageId="23" pageNumber="24">18.7</td>
</tr>
<tr id="356DC422FFDEA6AFFF54BA9B30B947D9" box="[159,1197,682,704]" gridrow="12" pageId="23" pageNumber="24">
<th id="76BCAD5EFFDEA6AFFF54BA9B368147D9" box="[159,661,682,704]" gridcol="0" gridrow="12" pageId="23" pageNumber="24">Dorsal-fin length</th>
<td id="76BCAD5EFFDEA6AFFD05BA9B372047D9" box="[718,820,682,704]" gridcol="1" gridrow="12" pageId="23" pageNumber="24">32.6–42.2</td>
<td id="76BCAD5EFFDEA6AFFBB8BA9B30B947D9" box="[1139,1197,682,704]" gridcol="2" gridrow="12" pageId="23" pageNumber="24">33.2</td>
</tr>
<tr id="356DC422FFDEA6AFFF54BAE330B947F1" box="[159,1197,722,744]" gridrow="13" pageId="23" pageNumber="24">
<th id="76BCAD5EFFDEA6AFFF54BAE3368147F1" box="[159,661,722,744]" gridcol="0" gridrow="13" pageId="23" pageNumber="24">Length of dorsal-fin base</th>
<td id="76BCAD5EFFDEA6AFFD05BAE3372047F1" box="[718,820,722,744]" gridcol="1" gridrow="13" pageId="23" pageNumber="24">14.7–17.6</td>
<td id="76BCAD5EFFDEA6AFFBB8BAE330B947F1" box="[1139,1197,722,744]" gridcol="2" gridrow="13" pageId="23" pageNumber="24">16.7</td>
</tr>
<tr id="356DC422FFDEA6AFFF54BACB30B94609" box="[159,1197,762,784]" gridrow="14" pageId="23" pageNumber="24">
<th id="76BCAD5EFFDEA6AFFF54BACB36814609" box="[159,661,762,784]" gridcol="0" gridrow="14" pageId="23" pageNumber="24">Anal-fin length</th>
<td id="76BCAD5EFFDEA6AFFD05BACB37204609" box="[718,820,762,784]" gridcol="1" gridrow="14" pageId="23" pageNumber="24">16.8–25.4</td>
<td id="76BCAD5EFFDEA6AFFBB8BACB30B94609" box="[1139,1197,762,784]" gridcol="2" gridrow="14" pageId="23" pageNumber="24">17.6</td>
</tr>
<tr id="356DC422FFDEA6AFFF54BB1330B94621" box="[159,1197,802,824]" gridrow="15" pageId="23" pageNumber="24">
<th id="76BCAD5EFFDEA6AFFF54BB1336814621" box="[159,661,802,824]" gridcol="0" gridrow="15" pageId="23" pageNumber="24">Length of anal-fin base</th>
<td id="76BCAD5EFFDEA6AFFD05BB1337204621" box="[718,820,802,824]" gridcol="1" gridrow="15" pageId="23" pageNumber="24">34.2–38.7</td>
<td id="76BCAD5EFFDEA6AFFBB8BB1330B94621" box="[1139,1197,802,824]" gridcol="2" gridrow="15" pageId="23" pageNumber="24">37.6</td>
</tr>
<tr id="356DC422FFDEA6AFFF54BB7B30B94679" box="[159,1197,842,864]" gridrow="16" pageId="23" pageNumber="24">
<th id="76BCAD5EFFDEA6AFFF54BB7B36814679" box="[159,661,842,864]" gridcol="0" gridrow="16" pageId="23" pageNumber="24">Distance from eye to dorsal-fin origin</th>
<td id="76BCAD5EFFDEA6AFFD05BB7B37204679" box="[718,820,842,864]" gridcol="1" gridrow="16" pageId="23" pageNumber="24">35.8–48.6</td>
<td id="76BCAD5EFFDEA6AFFBB8BB7B30B94679" box="[1139,1197,842,864]" gridcol="2" gridrow="16" pageId="23" pageNumber="24">48.1</td>
</tr>
<tr id="356DC422FFDEA6AFFF54BB4330B94691" box="[159,1197,882,904]" gridrow="17" pageId="23" pageNumber="24">
<th id="76BCAD5EFFDEA6AFFF54BB4336814691" box="[159,661,882,904]" gridcol="0" gridrow="17" pageId="23" pageNumber="24">Distance from dorsal-fin origin to caudal-fin base</th>
<td id="76BCAD5EFFDEA6AFFD05BB4337204691" box="[718,820,882,904]" gridcol="1" gridrow="17" pageId="23" pageNumber="24">50.7–57.8</td>
<td id="76BCAD5EFFDEA6AFFBB8BB4330B94691" box="[1139,1197,882,904]" gridcol="2" gridrow="17" pageId="23" pageNumber="24">52.4</td>
</tr>
<tr id="356DC422FFDEA6AFFF54BBAB30B946A9" box="[159,1197,922,944]" gridrow="18" pageId="23" pageNumber="24">
<th id="76BCAD5EFFDEA6AFFF54BBAB368146A9" box="[159,661,922,944]" gridcol="0" gridrow="18" pageId="23" pageNumber="24">Head length</th>
<td id="76BCAD5EFFDEA6AFFD05BBAB372046A9" box="[718,820,922,944]" gridcol="1" gridrow="18" pageId="23" pageNumber="24">23.0–27.6</td>
<td id="76BCAD5EFFDEA6AFFBB8BBAB30B946A9" box="[1139,1197,922,944]" gridcol="2" gridrow="18" pageId="23" pageNumber="24">26.0</td>
</tr>
<tr id="356DC422FFDEA6AFFF54BBF330B946C1" box="[159,1197,962,984]" gridrow="19" pageId="23" pageNumber="24">
<th id="76BCAD5EFFDEA6AFFF54BBF3368146C1" box="[159,661,962,984]" gridcol="0" gridrow="19" pageId="23" pageNumber="24">Head depth</th>
<td id="76BCAD5EFFDEA6AFFD05BBF3372046C1" box="[718,820,962,984]" gridcol="1" gridrow="19" pageId="23" pageNumber="24">21.2–27.6</td>
<td id="76BCAD5EFFDEA6AFFBB8BBF330B946C1" box="[1139,1197,962,984]" gridcol="2" gridrow="19" pageId="23" pageNumber="24">22.0</td>
</tr>
<tr id="356DC422FFDEA6AFFF54BBDB30B94119" box="[159,1197,1002,1024]" gridrow="20" pageId="23" pageNumber="24" rowspan-1="1" rowspan-2="1">
<th id="76BCAD5EFFDEA6AFFF54BBDB36814119" box="[159,661,1002,1024]" gridcol="0" gridrow="20" pageId="23" pageNumber="24">Percentages of head length</th>
</tr>
<tr id="356DC422FFDEA6AFFF54BC2330B94131" box="[159,1197,1042,1064]" gridrow="21" pageId="23" pageNumber="24">
<th id="76BCAD5EFFDEA6AFFF54BC2336814131" box="[159,661,1042,1064]" gridcol="0" gridrow="21" pageId="23" pageNumber="24">Snout length</th>
<td id="76BCAD5EFFDEA6AFFD05BC2337204131" box="[718,820,1042,1064]" gridcol="1" gridrow="21" pageId="23" pageNumber="24">22–24</td>
<td id="76BCAD5EFFDEA6AFFBB8BC2330B94131" box="[1139,1197,1042,1064]" gridcol="2" gridrow="21" pageId="23" pageNumber="24">23.3</td>
</tr>
<tr id="356DC422FFDEA6AFFF54BC0B30B94149" box="[159,1197,1082,1104]" gridrow="22" pageId="23" pageNumber="24">
<th id="76BCAD5EFFDEA6AFFF54BC0B36814149" box="[159,661,1082,1104]" gridcol="0" gridrow="22" pageId="23" pageNumber="24">Maxillary length</th>
<td id="76BCAD5EFFDEA6AFFD05BC0B37204149" box="[718,820,1082,1104]" gridcol="1" gridrow="22" pageId="23" pageNumber="24">41–50</td>
<td id="76BCAD5EFFDEA6AFFBB8BC0B30B94149" box="[1139,1197,1082,1104]" gridcol="2" gridrow="22" pageId="23" pageNumber="24">42.3</td>
</tr>
<tr id="356DC422FFDEA6AFFF54BC5330B94161" box="[159,1197,1122,1144]" gridrow="23" pageId="23" pageNumber="24">
<th id="76BCAD5EFFDEA6AFFF54BC5336814161" box="[159,661,1122,1144]" gridcol="0" gridrow="23" pageId="23" pageNumber="24">Horizontal orbital diameter</th>
<td id="76BCAD5EFFDEA6AFFD05BC5337204161" box="[718,820,1122,1144]" gridcol="1" gridrow="23" pageId="23" pageNumber="24">48–54</td>
<td id="76BCAD5EFFDEA6AFFBB8BC5330B94161" box="[1139,1197,1122,1144]" gridcol="2" gridrow="23" pageId="23" pageNumber="24">48.9</td>
</tr>
<tr id="356DC422FFDEA6AFFF54BCBB30B941B9" box="[159,1197,1162,1184]" gridrow="24" pageId="23" pageNumber="24">
<th id="76BCAD5EFFDEA6AFFF54BCBB368141B9" box="[159,661,1162,1184]" gridcol="0" gridrow="24" pageId="23" pageNumber="24">Least interorbital width</th>
<td id="76BCAD5EFFDEA6AFFD05BCBB372041B9" box="[718,820,1162,1184]" gridcol="1" gridrow="24" pageId="23" pageNumber="24">32–41</td>
<td id="76BCAD5EFFDEA6AFFBB8BCBB30B941B9" box="[1139,1197,1162,1184]" gridcol="2" gridrow="24" pageId="23" pageNumber="24">37.5</td>
</tr>
</table>
</paragraph>
<paragraph id="8BE2C660FFDE5947FF0CBCD536154094" blockId="23.[151,1437,1252,2032]" pageId="23" pageNumber="24">Scales large and cycloid. Lateral line complete and slightly bent downward anteriorly. Longitudinal scales 33 (5), 34 (3), or 35 (3). Scale rows between lateral line and dorsal-fin origin 6 (9) or 7 (2). Scale rows between lateral line and pelvic-fin origin 5 (10). Predorsal scales 7 to 9. Scale rows around caudal peduncle 14 (11). Anal-fin base covered by a single row of small scales. Lobes of caudal-fin base covered by small scales, smaller than those covering lateral surface of body.</paragraph>
<paragraph id="8BE2C660FFDE5947FF0CBDA634C040E1" blockId="23.[151,1437,1252,2032]" pageId="23" pageNumber="24">First gill arch with 7 (2) or 8 (7) gill rakers on upper limb and 12 (3) or 13 (7) gill rakers on lower limb. Total vertebrae 30, precaudal vertebrae 11, intermediate vertebrae 2 and caudal vertebrae 17 (2 c&amp;s). Supraneurals 3 (2 c&amp;s).</paragraph>
<paragraph id="8BE2C660FFDE5947FF0CBE35372E4279" blockId="23.[151,1437,1252,2032]" pageId="23" pageNumber="24">
<emphasis id="B9291A72FFDE5947FF0CBE3535884304" bold="true" box="[199,412,1540,1565]" pageId="23" pageNumber="24">Color in alcohol.</emphasis>
General body color yellowish. Dorsal portions of head and body darkly pigmented. Dorsolateral portion of body with small chromatophores along distal margin of scales; scales of ventrolateral portion of body unpigmented. Opercular and infraorbital bones silvery. Posterior margin of scales densely pigmented resulting in dark, horizontally aligned stripes along the lateral surface of the body. Two conspicuous vertical dark humeral marks separated by one scale, with anterior mark more evident. Anterior humeral mark extending over four horizontal scale rows above and one horizontal scale rows bellow lateral line. Posterior humeral mark located over three horizontal scales rows above lateral line. Caudal peduncle with a rounded dark mark. Midlateral silvery stripe broad, extending from supracleithrum to caudal peduncle. Anal, pelvic, and dorsal fins hyaline, outlined by small chromatophores, more so in distal portions of the rays. Adipose fin mostly hyaline, scattered by small chromatophores at distal border (
<figureCitation id="1366DAE5FFDE5947FD10BF76370A4279" box="[731,798,1863,1888]" captionStart="FIGURE 6" captionStartId="24.[151,250,1800,1822]" captionTargetBox="[239,1347,267,1778]" captionTargetId="figure@24.[239,1347,267,1778]" captionTargetPageId="24" captionText="FIGURE 6. Tetragonopterus rarus. a) MZUSP 88029, 85.4 mm SL, Brazil, Pará, Rio Paru, Amazon basin. Live specimens from (b) Marowijne and (c) Corantijn river basins, Suriname. Photos (b – c) by R. Covain." httpUri="https://zenodo.org/record/177603/files/figure.png" pageId="23" pageNumber="24">Fig. 6</figureCitation>
a).
</paragraph>
<paragraph id="8BE2C660FFDE5947FF0CBF5D367A429C" blockId="23.[151,1437,1252,2032]" box="[199,622,1900,1925]" pageId="23" pageNumber="24">
<emphasis id="B9291A72FFDE5947FF0CBF5D35A3429C" bold="true" box="[199,439,1900,1925]" pageId="23" pageNumber="24">Sexual dimorphism.</emphasis>
None observed.
</paragraph>
</subSubSection>
<subSubSection id="C34795EBFFDE5947FF0CBFBE34EA42E9" pageId="23" pageNumber="24" type="distribution">
<paragraph id="8BE2C660FFDE5947FF0CBFBE34EA42E9" blockId="23.[151,1437,1252,2032]" pageId="23" pageNumber="24">
<emphasis id="B9291A72FFDE5947FF0CBFBE357442B1" bold="true" box="[199,352,1935,1960]" pageId="23" pageNumber="24">Distribution.</emphasis>
<taxonomicName id="4C5DBDE3FFDE5947FEADBFA0367642B1" box="[358,610,1937,1960]" class="Insecta" family="Anthribidae" genus="Tetragonopterus" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="rarus">
<emphasis id="B9291A72FFDE5947FEADBFA0367642B1" box="[358,610,1937,1960]" italics="true" pageId="23" pageNumber="24">Tetragonopterus rarus</emphasis>
</taxonomicName>
is known from Corantijn River basin in western 
<collectingCountry id="F34A86F0FFDE5947FB4BBFBE30FA42B1" box="[1152,1262,1935,1960]" name="Suriname" pageId="23" pageNumber="24">Suriname</collectingCountry>
, the 
<collectingRegion id="49990882FFDE5947FAEABFBE318842B1" box="[1313,1436,1935,1960]" country="Suriname" name="Marowijne" pageId="23" pageNumber="24">Marowijne</collectingRegion>
River basin (
<typeStatus id="54E678C2FFDE5947FEE3BF85354E42D5" box="[296,346,1972,1996]" pageId="23" pageNumber="24">type</typeStatus>
locality of 
<taxonomicName id="4C5DBDE3FFDE5947FE16BF8536BB42D5" box="[477,687,1972,1997]" class="Actinopterygii" family="Characidae" genus="Moenkhausia" kingdom="Animalia" order="Characiformes" pageId="23" pageNumber="24" phylum="Chordata" rank="species" species="rara">
<emphasis id="B9291A72FFDE5947FE16BF8536BB42D5" box="[477,687,1972,1997]" italics="true" pageId="23" pageNumber="24">Moenkhausia rara</emphasis>
</taxonomicName>
), the Rio Jari, and Rio Paru of the Amazon basin, northern 
<collectingCountry id="F34A86F0FFDE5947FA9CBF85318F42D4" box="[1367,1435,1972,1997]" name="Brazil" pageId="23" pageNumber="24">Brazil</collectingCountry>
(
<figureCitation id="1366DAE5FFDE5947FF54BFE634F042E9" box="[159,228,2007,2032]" captionStart="FIGURE 2" captionStartId="4.[151,250,2001,2023]" captionText="FIGURE 2. Distribution of Tetragonopterus species throughout South America." pageId="23" pageNumber="24">Fig. 2</figureCitation>
a).
</paragraph>
</subSubSection>
<subSubSection id="C34795EBFFD15948FF0CB8A635EE45CD" pageId="24" pageNumber="25" type="discussion">
<paragraph id="8BE2C660FFD15948FF0CB8A635EE45CD" blockId="24.[151,1436,151,213]" pageId="24" pageNumber="25">
<emphasis id="B9291A72FFD15948FF0CB8A6352F45A9" bold="true" box="[199,315,151,176]" pageId="24" pageNumber="25">Remarks.</emphasis>
The molecular phylogeny of 
<taxonomicName id="4C5DBDE3FFD15948FD5CB8A8302345A9" authority="Melo et al., 2016" authorityName="Melo et al." authorityYear="2016" box="[663,1079,151,176]" class="Insecta" family="Anthribidae" genus="Tetragonopterus" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="genus">
<emphasis id="B9291A72FFD15948FD5CB8A8375B45A9" box="[663,847,153,176]" italics="true" pageId="24" pageNumber="25">Tetragonopterus</emphasis>
(
<bibRefCitation id="EFCCBB91FFD15948FCA9B8A6303B45A9" author="Melo" box="[866,1071,151,176]" pageId="24" pageNumber="25" refString="Melo, B. F., Benine, R. C., Silva, G. S. C., Avelino, G. S. &amp; Oliveira, C. (2016) Molecular phylogeny of the Neotropical fish genus Tetragonopterus (Teleostei: Characiformes: Characidae). Molecular Phylogenetics and Evolution, 94, 709 - 717. http: // dx. doi. org / 10.1016 / j. ympev. 2015.10.022" type="journal article" year="2016">
Melo 
<emphasis id="B9291A72FFD15948FC62B8A837F045A9" box="[937,996,151,176]" italics="true" pageId="24" pageNumber="25">et al.</emphasis>
, 2016
</bibRefCitation>
)
</taxonomicName>
supports the new combination proposed by 
<bibRefCitation id="EFCCBB91FFD15948FEECB88D35E145CD" box="[295,501,188,213]" pageId="24" pageNumber="25" refString="Melo, B. F., Benine, R. C., Mariguela, T. C. &amp; Oliveira, C. (2011) A new species of Tetragonopterus Cuvier, 1816 (Characiformes: Characidae: Tetragonopterinae) from the rio Jari, Amapa, northern Brazil. Neotropical Ichthyology, 9, 49 - 56. http: // dx. doi. org / 10.1590 / S 1679 - 62252011000100002" type="journal article">
Melo 
<emphasis id="B9291A72FFD15948FEA1B88C35B645CC" box="[362,418,188,213]" italics="true" pageId="24" pageNumber="25">et al.</emphasis>
(2011)
</bibRefCitation>
.
</paragraph>
</subSubSection>
<caption id="DF2296E8FFD15948FF5CBF39300B4224" httpUri="https://zenodo.org/record/177603/files/figure.png" pageId="24" pageNumber="25" targetBox="[239,1347,267,1778]" targetPageId="24">
<paragraph id="8BE2C660FFD15948FF5CBF39300B4224" blockId="24.[151,1435,1800,1853]" pageId="24" pageNumber="25">
<emphasis id="B9291A72FFD15948FF5CBF3935004204" bold="true" box="[151,276,1800,1822]" pageId="24" pageNumber="25">FIGURE 6.</emphasis>
<emphasis id="B9291A72FFD15948FED6BF38361E4207" box="[285,522,1801,1822]" italics="true" pageId="24" pageNumber="25">
<taxonomicName id="4C5DBDE3FFD15948FED6BF3836124207" box="[285,518,1801,1822]" class="Insecta" family="Anthribidae" genus="Tetragonopterus" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="rarus">Tetragonopterus rarus</taxonomicName>
.
</emphasis>
a) MZUSP 88029, 85.4 mm SL, Brazil, Pará, Rio Paru, Amazon basin. Live specimens from (b) Marowijne and (c) Corantijn river basins, Suriname. Photos (b–c) by R. Covain.
</paragraph>
</caption>
<subSubSection id="C34795EBFFD15949FF0CBF55317F4480" lastPageId="25" lastPageNumber="26" pageId="24" pageNumber="25" type="materials_examined">
<paragraph id="8BE2C660FFD15949FF0CBF55317F4480" blockId="24.[151,1436,1892,2017]" lastBlockId="25.[151,1437,151,410]" lastPageId="25" lastPageNumber="26" pageId="24" pageNumber="25">
<materialsCitation id="3B35CC3DFFD15948FF0CBF55377D4287" ID-GBIF-Occurrence="1412767492" collectingDate="1957-11-29" collectionCode="ANSP" collectorName="Tampoc &amp; J. Gery" country="Suriname" location="Marowijne" pageId="24" pageNumber="25" specimenCode="ANSP 94708, 1" specimenCount="1" stateProvince="Marowijne" typeStatus="holotype">
<emphasis id="B9291A72FFD15948FF0CBF5536014264" bold="true" box="[199,533,1892,1917]" pageId="24" pageNumber="25">
Material examined. 
<typeStatus id="54E678C2FFD15948FE0CBF5536044264" box="[455,528,1892,1917]" pageId="24" pageNumber="25">Types</typeStatus>
:
</emphasis>
<specimenCode id="DBFB6E1BFFD15948FDEEBF5436F34264" box="[549,743,1893,1917]" pageId="24" pageNumber="25">
<collectionCode id="ED4C5EA5FFD15948FDEEBF5436784264" box="[549,620,1893,1917]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34784" name="Academy of Natural Sciences of Philadelphia" pageId="24" pageNumber="25">ANSP</collectionCode>
94708, 1
</specimenCode>
paratype, 
<quantity id="4CA56B85FFD15948FCA6BF5537CD4265" box="[877,985,1892,1917]" metricMagnitude="-2" metricUnit="m" metricValue="5.79" pageId="24" pageNumber="25" unit="mm" value="57.9">57.9 mm</quantity>
SL, 
<collectingCountry id="F34A86F0FFD15948FBEABF55309E4264" box="[1057,1162,1892,1917]" name="Suriname" pageId="24" pageNumber="25">Suriname</collectingCountry>
/French 
<collectingCountry id="F34A86F0FFD15948FB3BBF54315D4264" box="[1264,1353,1893,1917]" name="Guyana" pageId="24" pageNumber="25">Guyana</collectingCountry>
, upper 
<collectingRegion id="49990882FFD15948FF5CBFB435004287" box="[151,276,1925,1950]" country="Suriname" name="Marowijne" pageId="24" pageNumber="25">Marowijne</collectingRegion>
, 
<collectorName id="26A8A3B6FFD15948FEE9BFB7356F4284" box="[290,379,1926,1949]" pageId="24" pageNumber="25">Tampoc</collectorName>
(into Litany rivier), 
<collectorName id="26A8A3B6FFD15948FDA7BFB736AB4287" box="[620,703,1926,1950]" pageId="24" pageNumber="25">J. Géry</collectorName>
, 
<collectingDate id="EFA71948FFD15948FD05BFB7377D4287" box="[718,873,1925,1950]" pageId="24" pageNumber="25" value="1957-11-29">
<date id="FFE3E0A0FFD15948FD05BFB7377D4287" box="[718,873,1925,1950]" pageId="24" pageNumber="25" value="1957-11-29">29 Nov 1957</date>
</collectingDate>
</materialsCitation>
. 
<materialsCitation id="3B35CC3DFFD15948FCBCBFB735C642F8" ID-GBIF-Occurrence="1412767510" collectionCode="ANSP" collectorName="Saut-Chien &amp; J. Gery" country="Guyana" location="Mana River" pageId="24" pageNumber="25" specimenCode="ANSP 112246, 1, ANSP 139714, 2" specimenCount="1" stateProvince="Marowijne" typeStatus="holotype">
<specimenCode id="DBFB6E1BFFD15948FCBCBFB7302D4287" box="[887,1081,1925,1950]" pageId="24" pageNumber="25">
<collectionCode id="ED4C5EA5FFD15948FCBCBFB737AA4287" box="[887,958,1926,1950]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34784" name="Academy of Natural Sciences of Philadelphia" pageId="24" pageNumber="25">ANSP</collectionCode>
<date id="FFE3E0A0FFD15948FC03BFB430094287" box="[968,1053,1925,1950]" pageId="24" pageNumber="25" value="46-11-22">112246</date>
, 1
</specimenCode>
paratype, 
<quantity id="4CA56B85FFD15948FB78BFB7310E4284" box="[1203,1306,1926,1950]" metricMagnitude="-2" metricUnit="m" metricValue="3.37" pageId="24" pageNumber="25" unit="mm" value="33.7">33.7 mm</quantity>
SL, 
<specimenCode id="DBFB6E1BFFD15948FA9EBFB7351142A6" pageId="24" pageNumber="25">
<collectionCode id="ED4C5EA5FFD15948FA9EBFB731884287" box="[1365,1436,1926,1950]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34784" name="Academy of Natural Sciences of Philadelphia" pageId="24" pageNumber="25">ANSP</collectionCode>
139714, 2
</specimenCode>
paratypes, 
<quantity id="4CA56B85FFD15948FE4FBF97363142A6" box="[388,549,1958,1983]" metricMagnitude="-2" metricUnit="m" metricValue="4.215" metricValueMax="4.84" metricValueMin="3.59" pageId="24" pageNumber="25" unit="mm" value="42.15" valueMax="48.4" valueMin="35.9">35.9–48.4 mm</quantity>
SL, French 
<collectingCountry id="F34A86F0FFD15948FD66BF96371242A6" box="[685,774,1959,1983]" name="Guyana" pageId="24" pageNumber="25">Guyana</collectingCountry>
, 
<collectingRegion id="49990882FFD15948FCDABF97379F42A6" box="[785,907,1958,1983]" country="Suriname" name="Marowijne" pageId="24" pageNumber="25">Marowijne</collectingRegion>
basin, middle 
<location id="8E8290BBFFD15948FBE6BF9930A642A6" LSID="urn:lsid:plazi:treatment:03F47776FFDF5949FF5CBA10317F4480:8E8290BBFFD15948FBE6BF9930A642A6" box="[1069,1202,1958,1983]" country="Guyana" name="Mana River" pageId="24" pageNumber="25" stateProvince="Marowijne">Mana River</location>
, between 
<collectorName id="26A8A3B6FFD15948FAEBBF97318842A6" box="[1312,1436,1958,1983]" pageId="24" pageNumber="25">Saut-Chien</collectorName>
and Saut-Topi-Topi, 
<collectorName id="26A8A3B6FFD15948FE4BBFF835C642F8" box="[384,466,1993,2017]" pageId="24" pageNumber="25">J. Géry</collectorName>
</materialsCitation>
, 
<materialsCitation id="3B35CC3DFFD15948FE2BBFF9312242F8" ID-GBIF-Occurrence="1412767486" box="[480,1334,1992,2017]" collectingDate="1957-10-15" collectionCode="ANSP" country="Venezuela" location="Venezuela" pageId="24" pageNumber="25" specimenCode="ANSP 168083, 39" specimenCount="1" typeStatus="holotype">
<collectingDate id="EFA71948FFD15948FE2BBFF9366442F8" box="[480,624,1992,2017]" pageId="24" pageNumber="25" value="1957-10-15">
<date id="FFE3E0A0FFD15948FE2BBFF9366442F8" box="[480,624,1992,2017]" pageId="24" pageNumber="25" value="1957-10-15">15 Oct 1957</date>
</collectingDate>
. 
<emphasis id="B9291A72FFD15948FDB7BFF936EA42F8" bold="true" box="[636,766,1992,2017]" pageId="24" pageNumber="25">Non-types:</emphasis>
<collectingCountry id="F34A86F0FFD15948FCCDBFF9376D42F8" box="[774,889,1992,2017]" name="Venezuela" pageId="24" pageNumber="25">Venezuela</collectingCountry>
: 
<specimenCode id="DBFB6E1BFFD15948FC42BFF8304D42F8" box="[905,1113,1992,2017]" pageId="24" pageNumber="25">
<collectionCode id="ED4C5EA5FFD15948FC42BFF837C442F8" box="[905,976,1993,2017]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34784" name="Academy of Natural Sciences of Philadelphia" pageId="24" pageNumber="25">ANSP</collectionCode>
168083, 39
</specimenCode>
, 
<quantity id="4CA56B85FFD15948FBACBFF9311D42F9" box="[1127,1289,1992,2017]" metricMagnitude="-2" metricUnit="m" metricValue="3.565" metricValueMax="5.08" metricValueMin="2.05" pageId="24" pageNumber="25" unit="mm" value="35.65" valueMax="50.8" valueMin="20.5">20.5–50.8 mm</quantity>
SL
</materialsCitation>
, 
<materialsCitation id="3B35CC3DFFD15949FA88BFF9377445A9" ID-GBIF-Occurrence="1412767466" collectionCode="MZUSP" country="Venezuela" lastPageId="25" lastPageNumber="26" location="Rio Hacha." pageId="24" pageNumber="25" specimenCode="MZUSP 77808, 3" specimenCount="1" stateProvince="Bolivar" typeStatus="holotype">
<collectingRegion id="49990882FFD15948FA88BFF9318C42F8" box="[1347,1432,1992,2017]" country="Venezuela" name="Bolivar" pageId="24" pageNumber="25">Bolivar</collectingRegion>
, Cuyuni basin, 
<location id="8E8290BBFFD05949FEF2B8A635AC45A9" LSID="urn:lsid:plazi:treatment:03F47776FFDF5949FF5CBA10317F4480:8E8290BBFFD05949FEF2B8A635AC45A9" box="[313,440,151,176]" country="Venezuela" name="Rio Hacha." pageId="25" pageNumber="26" stateProvince="Bolivar">Rio Hacha.</location>
<specimenCode id="DBFB6E1BFFD05949FE75B8A9369245A9" box="[446,646,152,176]" pageId="25" pageNumber="26">
<collectionCode id="ED4C5EA5FFD05949FE75B8A9360F45A9" box="[446,539,152,176]" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="25" pageNumber="26">MZUSP</collectionCode>
77808, 3
</specimenCode>
, 
<quantity id="4CA56B85FFD05949FD58B8A6372245B6" box="[659,822,151,176]" metricMagnitude="-2" metricUnit="m" metricValue="3.775" metricValueMax="4.09" metricValueMin="3.46" pageId="25" pageNumber="26" unit="mm" value="37.75" valueMax="40.9" valueMin="34.6">34.6–40.9 mm</quantity>
SL
</materialsCitation>
, 
<materialsCitation id="3B35CC3DFFD05949FCA7B8A6351045CB" ID-GBIF-Occurrence="1412767479" collectionCode="ANSP" country="Guyana" location="Bolivar" pageId="25" pageNumber="26" specimenCode="ANSP 177000" specimenCount="1" stateProvince="Bolivar" typeStatus="holotype">
<collectingRegion id="49990882FFD05949FCA7B8A637D545A9" box="[876,961,151,176]" country="Venezuela" name="Bolivar" pageId="25" pageNumber="26">Bolivar</collectingRegion>
, Rio Nichare. 
<collectingCountry id="F34A86F0FFD05949FBA9B8A930A845A9" box="[1122,1212,152,176]" name="Guyana" pageId="25" pageNumber="26">Guyana</collectingCountry>
: 
<specimenCode id="DBFB6E1BFFD05949FB03B8A9317945A9" box="[1224,1389,152,176]" collectionCode="ANSP" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34784" name="Academy of Natural Sciences of Philadelphia" pageId="25" pageNumber="26">
<collectionCode id="ED4C5EA5FFD05949FB03B8A9311B45A9" box="[1224,1295,152,176]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34784" name="Academy of Natural Sciences of Philadelphia" pageId="25" pageNumber="26">ANSP</collectionCode>
177000
</specimenCode>
, not measured
</materialsCitation>
, 
<materialsCitation id="3B35CC3DFFD05949FED9B888378345CB" ID-GBIF-Occurrence="1412767474" box="[274,919,185,210]" country="Guyana" latitude="4.7447224" location="Siparuni river" longLatPrecision="21" longitude="-59.005" pageId="25" pageNumber="26" specimenCount="1" typeStatus="holotype">
<collectingCountry id="F34A86F0FFD05949FED9B888359045CB" box="[274,388,185,210]" name="Guyana" pageId="25" pageNumber="26">Essequibo</collectingCountry>
basin, 
<location id="8E8290BBFFD05949FE12B888366345CB" LSID="urn:lsid:plazi:treatment:03F47776FFDF5949FF5CBA10317F4480:8E8290BBFFD05949FE12B888366345CB" box="[473,631,185,210]" country="Guyana" latitude="4.7447224" longLatPrecision="21" longitude="-59.005" name="Siparuni river" pageId="25" pageNumber="26">Siparuni river</location>
, 
<geoCoordinate id="EE69A0A7FFD05949FD4EB88836EB45C8" box="[645,767,185,209]" direction="north" orientation="latitude" pageId="25" pageNumber="26" precision="15" value="4.7447224">4°44’41”N</geoCoordinate>
<geoCoordinate id="EE69A0A7FFD05949FCC3B888378345CB" box="[776,919,185,210]" direction="west" orientation="longitude" pageId="25" pageNumber="26" precision="15" value="-59.005">59°00’18”W</geoCoordinate>
</materialsCitation>
. 
<materialsCitation id="3B35CC3DFFD05949FC6EB888367845EA" ID-GBIF-Occurrence="1412767483" collectionCode="ANSP" country="Guyana" location="Yukanopito falls" municipality="Rupununi region" pageId="25" pageNumber="26" specimenCode="ANSP 190828, 16" specimenCount="1" typeStatus="holotype">
<specimenCode id="DBFB6E1BFFD05949FC6EB888306C45CB" box="[933,1144,185,210]" pageId="25" pageNumber="26">
<collectionCode id="ED4C5EA5FFD05949FC6EB88837F845C8" box="[933,1004,185,209]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34784" name="Academy of Natural Sciences of Philadelphia" pageId="25" pageNumber="26">ANSP</collectionCode>
<date id="FFE3E0A0FFD05949FC3DB888305845C8" box="[1014,1100,185,209]" pageId="25" pageNumber="26" value="28-08-19">190828</date>
, 16
</specimenCode>
, not measured, 
<collectingMunicipality id="6B865C1AFFD05949FAE4B88834F545EA" pageId="25" pageNumber="26">Rupununi region</collectingMunicipality>
, upper 
<collectingCountry id="F34A86F0FFD05949FEF9B8EB35B345EA" box="[306,423,218,243]" name="Guyana" pageId="25" pageNumber="26">Essequibo</collectingCountry>
, 
<location id="8E8290BBFFD05949FE79B8EB367845EA" LSID="urn:lsid:plazi:treatment:03F47776FFDF5949FF5CBA10317F4480:8E8290BBFFD05949FE79B8EB367845EA" box="[434,620,218,243]" country="Guyana" municipality="Rupununi region" name="Yukanopito falls" pageId="25" pageNumber="26">Yukanopito falls</location>
</materialsCitation>
. 
<materialsCitation id="3B35CC3DFFD05949FDBDB8EA355E440C" ID-GBIF-Occurrence="1412767481" collectionCode="ANSP" country="Guyana" location="Kuyuwini river" municipality="Rupununi region" pageId="25" pageNumber="26" specimenCode="ANSP 190465, 4" specimenCount="1" typeStatus="holotype">
<specimenCode id="DBFB6E1BFFD05949FDBDB8EA372245EA" box="[630,822,218,243]" pageId="25" pageNumber="26">
<collectionCode id="ED4C5EA5FFD05949FDBDB8EA36A945EA" box="[630,701,219,243]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34784" name="Academy of Natural Sciences of Philadelphia" pageId="25" pageNumber="26">ANSP</collectionCode>
<date id="FFE3E0A0FFD05949FD0EB8EB370F45EA" box="[709,795,218,243]" pageId="25" pageNumber="26" value="65-04-19">190465</date>
, 4
</specimenCode>
, 
<quantity id="4CA56B85FFD05949FC89B8EB37F745EB" box="[834,995,218,243]" metricMagnitude="-2" metricUnit="m" metricValue="3.84" metricValueMax="4.15" metricValueMin="3.53" pageId="25" pageNumber="26" unit="mm" value="38.4" valueMax="41.5" valueMin="35.3">35.3–41.5 mm</quantity>
SL, 
<collectingMunicipality id="6B865C1AFFD05949FBD2B8EB30C245EA" box="[1049,1238,218,243]" pageId="25" pageNumber="26">Rupununi region</collectingMunicipality>
, 
<collectingCountry id="F34A86F0FFD05949FB29B8EB314045EA" box="[1250,1364,218,243]" name="Guyana" pageId="25" pageNumber="26">Essequibo</collectingCountry>
basin, 
<location id="8E8290BBFFD05949FF5CB8CD355E440C" LSID="urn:lsid:plazi:treatment:03F47776FFDF5949FF5CBA10317F4480:8E8290BBFFD05949FF5CB8CD355E440C" box="[151,330,252,277]" country="Guyana" municipality="Rupununi region" name="Kuyuwini river" pageId="25" pageNumber="26">Kuyuwini river</location>
</materialsCitation>
. 
<materialsCitation id="3B35CC3DFFD05949FE97B8CD318D440C" ID-GBIF-Occurrence="1412767467" box="[348,1433,252,277]" collectionCode="ANSP" collectorName="Kassi-Atae" country="Guyana" location="Rupununi region" pageId="25" pageNumber="26" specimenCode="ANSP 190525, 7" specimenCount="1" typeStatus="holotype">
<specimenCode id="DBFB6E1BFFD05949FE97B8CD363D440D" box="[348,553,252,277]" pageId="25" pageNumber="26">
<collectionCode id="ED4C5EA5FFD05949FE97B8CD35B7440D" box="[348,419,252,276]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34784" name="Academy of Natural Sciences of Philadelphia" pageId="25" pageNumber="26">ANSP</collectionCode>
<date id="FFE3E0A0FFD05949FE7AB8CD3613440C" box="[433,519,252,277]" pageId="25" pageNumber="26" value="25-05-19">190525</date>
, 7
</specimenCode>
, 
<quantity id="4CA56B85FFD05949FDF7B8CD36F0440D" box="[572,740,252,277]" metricMagnitude="-2" metricUnit="m" metricValue="4.37" metricValueMax="5.62" metricValueMin="3.12" pageId="25" pageNumber="26" unit="mm" value="43.7" valueMax="56.2" valueMin="31.2">31.2–56.2 mm</quantity>
SL, 
<location id="8E8290BBFFD05949FCE2B8CD37F9440C" LSID="urn:lsid:plazi:treatment:03F47776FFDF5949FF5CBA10317F4480:8E8290BBFFD05949FCE2B8CD37F9440C" box="[809,1005,252,277]" country="Guyana" name="Rupununi region" pageId="25" pageNumber="26">Rupununi region</location>
, 
<collectingCountry id="F34A86F0FFD05949FBCBB8CD3066440C" box="[1024,1138,252,277]" name="Guyana" pageId="25" pageNumber="26">Essequibo</collectingCountry>
river, 
<collectorName id="26A8A3B6FFD05949FB03B8CD3157440C" box="[1224,1347,252,277]" pageId="25" pageNumber="26">Kassi-Atae</collectorName>
rapids
</materialsCitation>
. 
<materialsCitation id="3B35CC3DFFD05949FF5CB92C36BD442F" ID-GBIF-Occurrence="1412767493" box="[151,681,285,310]" collectionCode="ANSP" country="Suriname" location="Suriname" pageId="25" pageNumber="26" specimenCode="ANSP 189638, 3" specimenCount="1" typeStatus="holotype">
<collectingCountry id="F34A86F0FFD05949FF5CB92C3512442F" box="[151,262,285,310]" name="Suriname" pageId="25" pageNumber="26">Suriname</collectingCountry>
: 
<specimenCode id="DBFB6E1BFFD05949FEDBB92F35DB442F" box="[272,463,285,310]" pageId="25" pageNumber="26">
<collectionCode id="ED4C5EA5FFD05949FEDBB92F3543442F" box="[272,343,286,310]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34784" name="Academy of Natural Sciences of Philadelphia" pageId="25" pageNumber="26">ANSP</collectionCode>
189638, 3
</specimenCode>
, 
<quantity id="4CA56B85FFD05949FE17B92F366A442C" box="[476,638,286,310]" metricMagnitude="-2" metricUnit="m" metricValue="4.135" metricValueMax="4.43" metricValueMin="3.84" pageId="25" pageNumber="26" unit="mm" value="41.349999999999994" valueMax="44.3" valueMin="38.4">38.4–44.3 mm</quantity>
SL
</materialsCitation>
; 
<materialsCitation id="3B35CC3DFFD05949FD7EB92F310C442F" ID-GBIF-Occurrence="1412767471" box="[693,1304,285,310]" collectionCode="ANSP" country="Suriname" location="Marowijne" pageId="25" pageNumber="26" specimenCode="ANSP 189639, 3" specimenCount="1" stateProvince="Marowijne" typeStatus="holotype">
<specimenCode id="DBFB6E1BFFD05949FD7EB92F3760442F" box="[693,884,285,310]" pageId="25" pageNumber="26">
<collectionCode id="ED4C5EA5FFD05949FD7EB92F36E8442F" box="[693,764,286,310]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34784" name="Academy of Natural Sciences of Philadelphia" pageId="25" pageNumber="26">ANSP</collectionCode>
189639, 3
</specimenCode>
, 
<quantity id="4CA56B85FFD05949FC4AB92F3037442C" box="[897,1059,286,310]" metricMagnitude="-2" metricUnit="m" metricValue="4.135" metricValueMax="4.43" metricValueMin="3.84" pageId="25" pageNumber="26" unit="mm" value="41.349999999999994" valueMax="44.3" valueMin="38.4">38.4–44.3 mm</quantity>
SL; 
<collectingRegion id="49990882FFD05949FB91B92C30C0442F" box="[1114,1236,285,310]" country="Suriname" name="Marowijne" pageId="25" pageNumber="26">Marowijne</collectingRegion>
basin
</materialsCitation>
, 
<materialsCitation id="3B35CC3DFFD05949FAEFB92C373C444E" ID-GBIF-Occurrence="1412767497" collectionCode="USNM" country="Suriname" location="Lawa river" pageId="25" pageNumber="26" specimenCode="USNM 229604, 3" specimenCount="1" stateProvince="Sipaliwini" typeStatus="holotype">
<collectingRegion id="49990882FFD05949FAEFB92C318D442F" box="[1316,1433,285,310]" country="Suriname" name="Sipaliwini" pageId="25" pageNumber="26">Sipaliwini</collectingRegion>
, Anapaike, 
<location id="8E8290BBFFD05949FEDBB90E3598444E" LSID="urn:lsid:plazi:treatment:03F47776FFDF5949FF5CBA10317F4480:8E8290BBFFD05949FEDBB90E3598444E" box="[272,396,318,343]" country="Suriname" name="Lawa river" pageId="25" pageNumber="26" stateProvince="Sipaliwini">Lawa river</location>
. 
<specimenCode id="DBFB6E1BFFD05949FE53B90E3676444E" box="[408,610,318,343]" pageId="25" pageNumber="26">
<collectionCode id="ED4C5EA5FFD05949FE53B90E35FC444E" box="[408,488,319,343]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="25" pageNumber="26">USNM</collectionCode>
229604, 3
</specimenCode>
, 
<quantity id="4CA56B85FFD05949FDBBB90F36EA444F" box="[624,766,318,343]" metricMagnitude="-2" metricUnit="m" metricValue="4.945" metricValueMax="6.1899999999999995" metricValueMin="3.7" pageId="25" pageNumber="26" unit="mm" value="49.45" valueMax="61.9" valueMin="37.0">37–61.9 mm</quantity>
SL
</materialsCitation>
, 
<materialsCitation id="3B35CC3DFFD05949FCFEB90F37D6444E" ID-GBIF-Occurrence="1412767490" box="[821,962,318,343]" country="Suriname" location="Brokopondo" pageId="25" pageNumber="26" specimenCount="1" stateProvince="Brokopondo" typeStatus="holotype">
<collectingRegion id="49990882FFD05949FCFEB90F37D6444E" box="[821,962,318,343]" country="Suriname" name="Brokopondo" pageId="25" pageNumber="26">Brokopondo</collectingRegion>
</materialsCitation>
, 
<materialsCitation id="3B35CC3DFFD05949FC04B90F36724461" ID-GBIF-Occurrence="1412767499" country="Brazil" location="Marowijne" pageId="25" pageNumber="26" specimenCount="1" stateProvince="Marowijne" typeStatus="holotype">
<collectingRegion id="49990882FFD05949FC04B90F305D444E" box="[975,1097,318,343]" country="Suriname" name="Marowijne" pageId="25" pageNumber="26">Marowijne</collectingRegion>
river, 
<quantity id="4CA56B85FFD05949FB58B90F30CF444E" box="[1171,1243,318,343]" metricMagnitude="4" metricUnit="m" metricValue="6.3" pageId="25" pageNumber="26" unit="km" value="63.0">63 km</quantity>
Z. Van Afobaka. 
<collectingCountry id="F34A86F0FFD05949FF5CB96E34CB4461" box="[151,223,351,376]" name="Brazil" pageId="25" pageNumber="26">Brazil</collectingCountry>
: LBP 21093, 1, 
<quantity id="4CA56B85FFD05949FE0AB96E3624446E" box="[449,560,351,376]" metricMagnitude="-2" metricUnit="m" metricValue="2.6399999999999997" pageId="25" pageNumber="26" unit="mm" value="26.4">26.4 mm</quantity>
SL
</materialsCitation>
, 
<materialsCitation id="3B35CC3DFFD05949FDB6B951352E4483" ID-GBIF-Occurrence="1412767509" country="Brazil" county="Oiapoque" latitude="3.8132222" location="Igarape Pantanari" longLatPrecision="1" longitude="-51.808777" municipality="Rio Oyapock" pageId="25" pageNumber="26" specimenCount="1" stateProvince="Amapa" typeStatus="holotype">
<collectingRegion id="49990882FFD05949FDB6B95136C44461" box="[637,720,352,376]" country="Brazil" name="Amapa" pageId="25" pageNumber="26">Amapá</collectingRegion>
, 
<collectingCounty id="6283BEECFFD05949FD2CB96E37424461" box="[743,854,351,376]" pageId="25" pageNumber="26">Oiapoque</collectingCounty>
, 
<collectingMunicipality id="6B865C1AFFD05949FCA6B96E301A4461" box="[877,1038,351,376]" pageId="25" pageNumber="26">Rio Oyapock</collectingMunicipality>
, 
<location id="8E8290BBFFD05949FBEEB95130E24461" LSID="urn:lsid:plazi:treatment:03F47776FFDF5949FF5CBA10317F4480:8E8290BBFFD05949FBEEB95130E24461" box="[1061,1270,351,376]" country="Brazil" county="Oiapoque" latitude="3.8132222" longLatPrecision="1" longitude="-51.808777" municipality="Rio Oyapock" name="Igarape Pantanari" pageId="25" pageNumber="26" stateProvince="Amapa">Igarapé Pantanari</location>
, 
<geoCoordinate id="EE69A0A7FFD05949FAC6B96E31884461" box="[1293,1436,351,376]" direction="north" orientation="latitude" pageId="25" pageNumber="26" precision="1" value="3.8132222">3°48’47.6”N</geoCoordinate>
<geoCoordinate id="EE69A0A7FFD05949FF5CB9B0352E4483" box="[151,314,385,410]" direction="west" orientation="longitude" pageId="25" pageNumber="26" precision="1" value="-51.808777">51°48’31.6”W</geoCoordinate>
</materialsCitation>
. 
<materialsCitation id="3B35CC3DFFD05949FE8FB9B031734480" ID-GBIF-Occurrence="1412767478" box="[324,1383,385,410]" collectionCode="MZUSP" collectorName="Alto Paru d'Oeste &amp; Amazon" country="Brazil" location="upper Rio Paru" pageId="25" pageNumber="26" specimenCode="MZUSP 16758, 18" specimenCount="1" stateProvince="Para" typeStatus="holotype">
<specimenCode id="DBFB6E1BFFD05949FE8FB9B0360D4480" box="[324,537,385,410]" pageId="25" pageNumber="26">
<collectionCode id="ED4C5EA5FFD05949FE8FB9B035B54480" box="[324,417,385,409]" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="25" pageNumber="26">MZUSP</collectionCode>
16758, 18
</specimenCode>
, 50.4–68.0 mm SL, 
<collectingRegion id="49990882FFD05949FD31B9B0373A4480" box="[762,814,385,409]" country="Suriname" name="Para" pageId="25" pageNumber="26">Pará</collectingRegion>
, 
<collectorName id="26A8A3B6FFD05949FCF2B9B030104483" box="[825,1028,385,410]" pageId="25" pageNumber="26">Alto Paru d’Oeste</collectorName>
, 
<collectorName id="26A8A3B6FFD05949FBC4B9B030794480" box="[1039,1133,385,409]" pageId="25" pageNumber="26">Amazon</collectorName>
basin, 
<location id="8E8290BBFFD05949FB77B9B331734480" LSID="urn:lsid:plazi:treatment:03F47776FFDF5949FF5CBA10317F4480:8E8290BBFFD05949FB77B9B331734480" box="[1212,1383,385,410]" country="Brazil" name="upper Rio Paru" pageId="25" pageNumber="26" stateProvince="Para">upper Rio Paru</location>
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>