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<document ID-DOI="10.1206/0003-0090.457.1.1" ID-ISSN="0003-0090" ID-Zenodo-Dep="6971356" approvalRequired="59" approvalRequired_for_taxonomicNames="59" checkinTime="1659882917898" checkinUser="felipe" docAuthor="Beck, Robin M. D., Voss, Robert S. &amp; Jansa, Sharon A." docDate="2022" docId="03EFDD5DF778696ADB14FCAD1E38FC48" docLanguage="en" docName="BulAmeMusNatHist.2022.457.1-350.pdf" docOrigin="Bulletin of the American Museum of Natural History 2022 (457)" docSource="https://bioone.org/journals/bulletin-of-the-american-museum-of-natural-history/volume-457/issue-1/0003-0090.457.1.1/Craniodental-Morphology-and-Phylogeny-of-Marsupials/10.1206/0003-0090.457.1.1.full" docStyle="DocumentStyle:915933466F796C9C739DF4DB6B8DCFA6.7:BulAmeMusNatHis.2011-.journal_article.1cover.type1" docStyleId="915933466F796C9C739DF4DB6B8DCFA6" docStyleName="BulAmeMusNatHis.2011-.journal_article.1cover.type1" docStyleVersion="7" docTitle="Mutpuracinus archibaldi Murray &amp; Megirian 2000" docType="treatment" docVersion="7" lastPageNumber="323" masterDocId="FFD6A525F63B6829DA0BFFAE1A62FFC7" masterDocTitle="Craniodental Morphology And Phylogeny Of Marsupials" masterLastPageNumber="353" masterPageNumber="1" pageNumber="323" updateTime="1661891901480" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title>Craniodental Morphology And Phylogeny Of Marsupials</mods:title>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Beck, Robin M. D.</mods:namePart>
<mods:affiliation>School of Science, Engineering and Environment University of Salford, U. K. &amp; School of Biological, Earth &amp; Environmental Sciences University of New South Wales, Australia &amp; Division of Vertebrate Zoology (Mammalogy) American Museum of Natural History</mods:affiliation>
</mods:name>
<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Voss, Robert S.</mods:namePart>
<mods:affiliation>Division of Vertebrate Zoology (Mammalogy) American Museum of Natural History</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Jansa, Sharon A.</mods:namePart>
<mods:affiliation>Bell Museum and Department of Ecology, Evolution, and Behavior University of Minnesota</mods:affiliation>
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<mods:title>Bulletin of the American Museum of Natural History</mods:title>
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<mods:date>2022</mods:date>
<mods:detail type="pubDate">
<mods:number>2022-06-28</mods:number>
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<mods:detail type="volume">
<mods:number>2022</mods:number>
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<mods:detail type="issue">
<mods:number>457</mods:number>
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<mods:start>1</mods:start>
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<mods:url>https://bioone.org/journals/bulletin-of-the-american-museum-of-natural-history/volume-457/issue-1/0003-0090.457.1.1/Craniodental-Morphology-and-Phylogeny-of-Marsupials/10.1206/0003-0090.457.1.1.full</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.1206/0003-0090.457.1.1</mods:identifier>
<mods:identifier type="ISSN">0003-0090</mods:identifier>
<mods:identifier type="Zenodo-Dep">6971356</mods:identifier>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.6974459" ID-Zenodo-Dep="6974459" LSID="urn:lsid:plazi:treatment:03EFDD5DF778696ADB14FCAD1E38FC48" httpUri="http://treatment.plazi.org/id/03EFDD5DF778696ADB14FCAD1E38FC48" lastPageNumber="323" pageId="323" pageNumber="323">
<subSubSection box="[287,458,771,795]" pageId="323" pageNumber="323" type="multiple">
<paragraph blockId="323.[287,458,771,795]" box="[287,458,771,795]" pageId="323" pageNumber="323">
<heading box="[287,458,771,795]" centered="true" fontSize="10" level="2" pageId="323" pageNumber="323" reason="4">
<taxonomicName box="[300,458,771,795]" class="Mammalia" family="Thylacinidae" genus="Mutpuracinus" kingdom="Animalia" order="Dasyuromorphia" pageId="323" pageNumber="323" phylum="Chordata" rank="species" species="undetermined">
<emphasis bold="true" box="[300,458,771,795]" italics="true" pageId="323" pageNumber="323">Mutpuracinus</emphasis>
</taxonomicName>
</heading>
</paragraph>
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<subSubSection pageId="323" pageNumber="323" type="nomenclature">
<paragraph blockId="323.[108,637,821,1738]" pageId="323" pageNumber="323">
SPECIES SCORED: †
<taxonomicName authorityName="Murray &amp; Megirian" authorityYear="2000" box="[361,636,821,845]" class="Mammalia" family="Thylacinidae" genus="Mutpuracinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dasyuromorphia" pageId="323" pageNumber="323" phylum="Chordata" rank="species" species="archibaldi">
<emphasis box="[361,636,821,845]" italics="true" pageId="323" pageNumber="323">Mutpuracinus archibaldi</emphasis>
</taxonomicName>
(
<typeStatus box="[116,165,854,878]" pageId="323" pageNumber="323">type</typeStatus>
and only described species).
</paragraph>
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<subSubSection pageId="323" pageNumber="323" type="materials_examined">
<paragraph blockId="323.[108,637,821,1738]" pageId="323" pageNumber="323">
GEOLOGICAL PROVENANCE OF SCORED
<collectionCode box="[108,168,920,944]" pageId="323" pageNumber="323">SPECIMENS</collectionCode>
: Bullock Creek Local Fauna, Camfield Beds,
<collectingRegion box="[172,379,953,977]" country="Australia" name="Northern Territory" pageId="323" pageNumber="323">Northern Territory</collectingRegion>
,
<collectingCountry box="[389,489,953,977]" name="Australia" pageId="323" pageNumber="323">Australia</collectingCountry>
; Jaw Junction Site (Riversleigh Faunal Zone D), Riversleigh World Heritage Area,
<collectingRegion box="[350,482,1019,1043]" country="Australia" name="Queensland" pageId="323" pageNumber="323">Queensland</collectingRegion>
,
<collectingCountry box="[494,596,1019,1043]" name="Australia" pageId="323" pageNumber="323">Australia</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection pageId="323" pageNumber="323" type="description">
<paragraph blockId="323.[108,637,821,1738]" pageId="323" pageNumber="323">AGE OF SCORED SPECIMENS: based on biostratigraphy, the Bullock Creek Local Fauna is interpreted to be middle Miocene (Murray and Megirian, 1992; Megirian et al., 2010; Black et al., 2013; Schwartz, 2016; Trusler and Sharp, 2016), whereas Riversleigh Faunal Zone D is considered to be somewhat younger, namely early late Miocene, perhaps 1012 Mya (Archer et al., 1989, 1994, 1995; Myers et al., 2001; Black et al., 2012b, 2013; Woodhead et al., 2014; Arena et al., 2015). In the absence of direct dates, we have conservatively assumed the entire span of the middle to late Miocene (Langhian to Messinian; Cohen et al., 2013 [updated]) for this taxon.</paragraph>
<paragraph blockId="323.[108,637,821,1738]" box="[141,608,1515,1540]" pageId="323" pageNumber="323">
ASSIGNED AGE
<collectionCode box="[312,388,1515,1539]" pageId="323" pageNumber="323">RANGE</collectionCode>
:
<geoCoordinate box="[402,474,1515,1539]" degrees="15.970" direction="north" orientation="latitude" pageId="323" pageNumber="323" precision="55" value="15.97">15.970</geoCoordinate>
<geoCoordinate box="[473,546,1515,1539]" degrees="5.333" direction="west" orientation="longitude" pageId="323" pageNumber="323" precision="55" value="-5.333">5.333</geoCoordinate>
Mya.
</paragraph>
</subSubSection>
<subSubSection pageId="323" pageNumber="323" type="discussion">
<paragraph blockId="323.[108,637,821,1738]" lastBlockId="323.[684,1213,226,911]" pageId="323" pageNumber="323">
REMARKS: †
<taxonomicName authorityName="Murray &amp; Megirian" authorityYear="2000" box="[270,531,1548,1572]" class="Mammalia" family="Thylacinidae" genus="Mutpuracinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dasyuromorphia" pageId="323" pageNumber="323" phylum="Chordata" rank="species" species="archibaldi">
<emphasis box="[270,531,1548,1572]" italics="true" pageId="323" pageNumber="323">Mutpuracinus archibaldi</emphasis>
</taxonomicName>
was originally described by Murray and Megirian (2000) based on fragmentary craniodental material from the Bullock Creek Local Fauna. Subsequently, a relatively complete, albeit somewhat disarticulated, skull (
<collectionCode box="[333,396,1714,1738]" country="Australia" httpUri="http://grbio.org/cool/jg13-7fbc" name="Northern Territory Museum of Arts and Sciences" pageId="323" pageNumber="323">NTM</collectionCode>
<accessionNumber box="[402,489,1713,1737]" httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/P91168" pageId="323" pageNumber="323">P91168</accessionNumber>
-5) was recovered from the Jaw Junction Site at Riversleigh (Murray and Megirian, 2006a). Murray and Megirian (2000, 2006a) identified †
<taxonomicName authorityName="Murray &amp; Megirian" authorityYear="2000" box="[1063,1212,292,315]" class="Mammalia" family="Thylacinidae" genus="Mutpuracinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dasyuromorphia" pageId="323" pageNumber="323" phylum="Chordata" rank="genus">
<emphasis box="[1063,1212,292,315]" italics="true" pageId="323" pageNumber="323">Mutpuracinus</emphasis>
</taxonomicName>
as a thylacinid, but its position was unstable in their phylogenetic analyses (Murray and Megirian, 2006a: appendix 2, figs. 24); notably, it was recovered as closer to Recent dasyurids than to thylacinids when added to the craniodental character matrix of Wroe and Musser (2001; see Murray and Megirian, 2006a: appendix 2 fig. 2B). The phylogenetic analyses of Archer et al. (2016), Kealy and Beck (2017), and Rovinsky et al. (2019) also did not support thylacinid affinities for †
<taxonomicName authorityName="Murray &amp; Megirian" authorityYear="2000" box="[921,1071,656,679]" class="Mammalia" family="Thylacinidae" genus="Mutpuracinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dasyuromorphia" pageId="323" pageNumber="323" phylum="Chordata" rank="genus">
<emphasis box="[921,1071,656,679]" italics="true" pageId="323" pageNumber="323">Mutpuracinus</emphasis>
</taxonomicName>
, and we follow the last two studies in considering this taxon to be
<taxonomicName authorityName="sensu Kealy and Beck" authorityYear="2017" box="[747,933,722,746]" class="Mammalia" kingdom="Animalia" order="Dasyuromorphia" pageId="323" pageNumber="323" phylum="Chordata" rank="order">Dasyuromorphia</taxonomicName>
<emphasis box="[942,1083,722,746]" italics="true" pageId="323" pageNumber="323">incertae sedis</emphasis>
. Regardless of its true phylogenetic relationships, the craniodental morphology of †
<taxonomicName authorityName="Murray &amp; Megirian" authorityYear="2000" box="[965,1119,788,811]" class="Mammalia" family="Thylacinidae" genus="Mutpuracinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dasyuromorphia" pageId="323" pageNumber="323" phylum="Chordata" rank="genus">
<emphasis box="[965,1119,788,811]" italics="true" pageId="323" pageNumber="323">Mutpuracinus</emphasis>
</taxonomicName>
implies previously unsuspected dasyuromorphian homoplasy in either the dentition or auditory region, or both (Murray and Megirian, 2006a).
</paragraph>
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