199 lines
17 KiB
XML
199 lines
17 KiB
XML
<document ID-DOI="10.1206/0003-0090.457.1.1" ID-ISSN="0003-0090" ID-Zenodo-Dep="6971356" approvalRequired="59" approvalRequired_for_taxonomicNames="59" checkinTime="1659882917898" checkinUser="felipe" docAuthor="Beck, Robin M. D., Voss, Robert S. & Jansa, Sharon A." docDate="2022" docId="03EFDD5DF7066917D96BFDD11BCBFA25" docLanguage="en" docName="BulAmeMusNatHist.2022.457.1-350.pdf" docOrigin="Bulletin of the American Museum of Natural History 2022 (457)" docSource="https://bioone.org/journals/bulletin-of-the-american-museum-of-natural-history/volume-457/issue-1/0003-0090.457.1.1/Craniodental-Morphology-and-Phylogeny-of-Marsupials/10.1206/0003-0090.457.1.1.full" docStyle="DocumentStyle:915933466F796C9C739DF4DB6B8DCFA6.7:BulAmeMusNatHis.2011-.journal_article.1cover.type1" docStyleId="915933466F796C9C739DF4DB6B8DCFA6" docStyleName="BulAmeMusNatHis.2011-.journal_article.1cover.type1" docStyleVersion="7" docTitle="Hesperocynus dolgopolae" docType="treatment" docVersion="7" lastPageNumber="318" masterDocId="FFD6A525F63B6829DA0BFFAE1A62FFC7" masterDocTitle="Craniodental Morphology And Phylogeny Of Marsupials" masterLastPageNumber="353" masterPageNumber="1" pageNumber="317" updateTime="1661891901480" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:titleInfo>
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<mods:title>Craniodental Morphology And Phylogeny Of Marsupials</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Beck, Robin M. D.</mods:namePart>
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<mods:affiliation>School of Science, Engineering and Environment University of Salford, U. K. & School of Biological, Earth & Environmental Sciences University of New South Wales, Australia & Division of Vertebrate Zoology (Mammalogy) American Museum of Natural History</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Voss, Robert S.</mods:namePart>
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<mods:affiliation>Division of Vertebrate Zoology (Mammalogy) American Museum of Natural History</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Jansa, Sharon A.</mods:namePart>
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<mods:affiliation>Bell Museum and Department of Ecology, Evolution, and Behavior University of Minnesota</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:title>Bulletin of the American Museum of Natural History</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2022</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2022-06-28</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>2022</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>457</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>353</mods:end>
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<mods:location>
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<mods:url>https://bioone.org/journals/bulletin-of-the-american-museum-of-natural-history/volume-457/issue-1/0003-0090.457.1.1/Craniodental-Morphology-and-Phylogeny-of-Marsupials/10.1206/0003-0090.457.1.1.full</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.1206/0003-0090.457.1.1</mods:identifier>
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<mods:identifier type="ISSN">0003-0090</mods:identifier>
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<mods:identifier type="Zenodo-Dep">6971356</mods:identifier>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.6974436" ID-Zenodo-Dep="6974436" LSID="urn:lsid:plazi:treatment:03EFDD5DF7066917D96BFDD11BCBFA25" httpUri="http://treatment.plazi.org/id/03EFDD5DF7066917D96BFDD11BCBFA25" lastPageId="318" lastPageNumber="318" pageId="317" pageNumber="317">
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<paragraph blockId="317.[864,1032,639,663]" box="[864,1032,639,663]" pageId="317" pageNumber="317">
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<heading box="[864,1032,639,663]" centered="true" fontSize="10" level="2" pageId="317" pageNumber="317" reason="4">
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<emphasis bold="true" box="[864,1032,639,663]" pageId="317" pageNumber="317">
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†
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<taxonomicName box="[878,1032,639,662]" class="Mammalia" family="Sparassocynidae" genus="Hesperocynus" kingdom="Animalia" order="Didelphimorphia" pageId="317" pageNumber="317" phylum="Chordata" rank="species" species="undetermined">
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<emphasis bold="true" box="[878,1032,639,662]" italics="true" pageId="317" pageNumber="317">Hesperocynus</emphasis>
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</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="317" pageNumber="317" type="nomenclature">
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<paragraph blockId="317.[684,1214,688,1738]" pageId="317" pageNumber="317">
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SPECIES SCORED: †
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<taxonomicName baseAuthorityName="Reig" baseAuthorityYear="1952" box="[935,1212,688,712]" class="Mammalia" family="Sparassocynidae" genus="Hesperocynus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Didelphimorphia" pageId="317" pageNumber="317" phylum="Chordata" rank="species" species="dolgopolae">
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<emphasis box="[935,1212,688,712]" italics="true" pageId="317" pageNumber="317">Hesperocynus dolgopolae</emphasis>
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</taxonomicName>
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(
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<typeStatus box="[692,741,722,746]" pageId="317" pageNumber="317">type</typeStatus>
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and only described species).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="317" pageNumber="317" type="materials_examined">
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<paragraph blockId="317.[684,1214,688,1738]" pageId="317" pageNumber="317">
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GEOLOGICAL PROVENANCE OF SCORED
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<collectionCode box="[684,743,788,812]" pageId="317" pageNumber="317">SPECIMENS</collectionCode>
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: Level 20 (sensu Stahlecker; see Riggs and Patterson, 1939) of the Andalhuala Formation,
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<collectingRegion box="[684,907,854,878]" country="Argentina" name="Catamarca" pageId="317" pageNumber="317">Catamarca Province</collectingRegion>
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,
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<collectingCountry box="[919,1031,854,878]" name="Argentina" pageId="317" pageNumber="317">Argentina</collectingCountry>
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; Arroyo Seco de la Frazada, La Huertita Formation (previously middle section of the Aisol Formation; Forasiepi et al., 2009, 2011; Garrido et al., 2014),
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<collectingRegion country="Argentina" name="Mendoza" pageId="317" pageNumber="317">Mendoza Province</collectingRegion>
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,
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<collectingCountry box="[794,907,986,1010]" name="Argentina" pageId="317" pageNumber="317">Argentina</collectingCountry>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="317" pageNumber="317" type="description">
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<paragraph blockId="317.[684,1214,688,1738]" pageId="317" pageNumber="317">
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AGE OF SCORED SPECIMENS: Esteban et al. (2014: fig. 2) summarized the stratigraphy of the Andalhuala Formation and published radiometric dates for the various Levels sensu Stahlecker (see Riggs and Patterson, 1939). Of these, Butler et al. (1984) gave a radiometric date of 6.70 ± 0.05 Mya for Level 15, while Latorre et al. (1997) presented a radiometric date of 5.64 ± 0.16 Mya for Level 23, which collectively bracket Level 20, from which the Andalhuala
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<taxonomicName box="[995,1141,1318,1341]" class="Mammalia" family="Sparassocynidae" genus="Hesperocynus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Didelphimorphia" pageId="317" pageNumber="317" phylum="Chordata" rank="genus">
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<emphasis box="[995,1141,1318,1341]" italics="true" pageId="317" pageNumber="317">Hesperocynus</emphasis>
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</taxonomicName>
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specimen was collected. This falls within the late Miocene Huayquerian
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<collectionCode box="[884,973,1383,1407]" pageId="317" pageNumber="317">SALMA</collectionCode>
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(9–5.28 Mya; Prevosti and Forasiepi, 2018:
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<tableCitation box="[928,1029,1416,1440]" captionStart="TABLE 1" captionStartId="7.[611,691,226,250]" captionTargetBox="[119,1110,327,1337]" captionTargetPageId="7" captionText="TABLE 1 Higher Classification and Geographic Distribution of Recent Marsupialsa" httpUri="http://table.plazi.org/id/DF393CC3F63C682ED868FF4C1E65FED9" pageId="317" pageNumber="317" tableUuid="DF393CC3F63C682ED868FF4C1E65FED9">table 1.1</tableCitation>
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). Garrido et al. (2014) renamed the “middle section of the Aisol Formation” as the La Huertita Formation and identified it as Montehermosan-Chapadmalalan based on biostratigraphy. The ages of the Montehermosan and Chapadmalalan SALMAs are discussed above (see †
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<taxonomicName baseAuthorityName="Beck and Taglioretti" baseAuthorityYear="2020" box="[898,1051,1615,1638]" class="Mammalia" family="Didelphidae" genus="Sparassocynus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Didelphimorphia" pageId="317" pageNumber="317" phylum="Chordata" rank="genus">
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<emphasis box="[898,1051,1615,1638]" italics="true" pageId="317" pageNumber="317">Sparassocynus</emphasis>
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</taxonomicName>
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).
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</paragraph>
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<paragraph blockId="317.[684,1214,688,1738]" box="[716,1164,1647,1672]" pageId="317" pageNumber="317">
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ASSIGNED AGE
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<collectionCode box="[883,957,1648,1672]" pageId="317" pageNumber="317">RANGE</collectionCode>
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:
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<geoCoordinate box="[971,1030,1647,1671]" degrees="6.750" direction="north" orientation="latitude" pageId="317" pageNumber="317" precision="55" value="6.75">6.750</geoCoordinate>
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<geoCoordinate box="[1030,1102,1647,1671]" degrees="3.100" direction="west" orientation="longitude" pageId="317" pageNumber="317" precision="55" value="-3.1">–3.100</geoCoordinate>
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Mya.
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="318" lastPageNumber="318" pageId="317" pageNumber="317" type="discussion">
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<paragraph blockId="317.[684,1214,688,1738]" lastBlockId="318.[108,638,226,1506]" lastPageId="318" lastPageNumber="318" pageId="317" pageNumber="317">
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REMARKS: Based largely on information provided by a partial skull from the La Huertita Formation (MHNSR-PV 1046), Forasiepi et al. (2009) erected the genus †
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<taxonomicName box="[445,597,259,282]" class="Mammalia" family="Sparassocynidae" genus="Hesperocynus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Didelphimorphia" pageId="318" pageNumber="318" phylum="Chordata" rank="genus">
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<emphasis box="[445,597,259,282]" italics="true" pageId="318" pageNumber="318">Hesperocynus</emphasis>
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</taxonomicName>
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to include specimens originally referred to †
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<taxonomicName class="Mammalia" family="Didelphidae" genus="Thylatheridium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Didelphimorphia" pageId="318" pageNumber="318" phylum="Chordata" rank="species" species="dolgopolae">
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<emphasis italics="true" pageId="318" pageNumber="318">Thylatheridium dolgopolae</emphasis>
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</taxonomicName>
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(Reig, 1958a; Goin and Montalvo, 1988; Goin et al., 2000). Forasiepi et al. (2009) noted that †
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<taxonomicName box="[391,551,391,415]" class="Mammalia" family="Didelphidae" genus="Thylatheridium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Didelphimorphia" pageId="318" pageNumber="318" phylum="Chordata" rank="species" species="dolgopolae">
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<emphasis box="[391,551,391,415]" italics="true" pageId="318" pageNumber="318">H. dolgopolae</emphasis>
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</taxonomicName>
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shares derived craniodental similarities with †
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<taxonomicName baseAuthorityName="Beck and Taglioretti" baseAuthorityYear="2020" class="Mammalia" family="Didelphidae" genus="Sparassocynus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Didelphimorphia" pageId="318" pageNumber="318" phylum="Chordata" rank="genus">
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<emphasis italics="true" pageId="318" pageNumber="318">Sparassocynus</emphasis>
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</taxonomicName>
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(a partial posterior braincase of †
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<taxonomicName authority=", FMNH" authorityName="FMNH" class="Mammalia" family="Didelphidae" genus="Thylatheridium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Didelphimorphia" pageId="318" pageNumber="318" phylum="Chordata" rank="species" species="dolgopolae">
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<emphasis italics="true" pageId="318" pageNumber="318">H. dolgopolae</emphasis>
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, FMNH
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</taxonomicName>
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<collectionCode box="[298,323,490,514]" pageId="318" pageNumber="318">P-</collectionCode>
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15225, had previously been identified as †
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<taxonomicName box="[281,484,524,547]" class="Mammalia" family="Didelphidae" genus="Sparassocynus" kingdom="Animalia" order="Didelphimorphia" pageId="318" pageNumber="318" phylum="Chordata" rank="species" species="undetermined">
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<emphasis box="[281,439,524,547]" italics="true" pageId="318" pageNumber="318">Sparassocynus</emphasis>
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sp.
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</taxonomicName>
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by previous authors; Reig, 1958b; Reig and Simpson, 1972; Simpson, 1974), but is somewhat more plesiomorphic, and they classified both genera within
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<taxonomicName authorityName="Reig" authorityYear="1958" box="[110,301,655,679]" class="Mammalia" family="Sparassocynidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Didelphimorphia" pageId="318" pageNumber="318" phylum="Chordata" rank="family">Sparassocynidae</taxonomicName>
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. As already discussed (see †
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<taxonomicName baseAuthorityName="Beck and Taglioretti" baseAuthorityYear="2020" box="[121,273,689,712]" class="Mammalia" family="Didelphidae" genus="Sparassocynus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Didelphimorphia" pageId="318" pageNumber="318" phylum="Chordata" rank="genus">
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<emphasis box="[121,273,689,712]" italics="true" pageId="318" pageNumber="318">Sparassocynus</emphasis>
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</taxonomicName>
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, above), the total-evidence analyses of Beck and Taglioretti (2020) recovered †
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<taxonomicName box="[122,270,755,778]" class="Mammalia" family="Sparassocynidae" genus="Hesperocynus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Didelphimorphia" pageId="318" pageNumber="318" phylum="Chordata" rank="genus">
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<emphasis box="[122,270,755,778]" italics="true" pageId="318" pageNumber="318">Hesperocynus</emphasis>
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</taxonomicName>
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and †
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<taxonomicName baseAuthorityName="Beck and Taglioretti" baseAuthorityYear="2020" box="[345,500,755,778]" class="Mammalia" family="Didelphidae" genus="Sparassocynus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Didelphimorphia" pageId="318" pageNumber="318" phylum="Chordata" rank="genus">
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<emphasis box="[345,500,755,778]" italics="true" pageId="318" pageNumber="318">Sparassocynus</emphasis>
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</taxonomicName>
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as a triballevel clade within
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<taxonomicName baseAuthorityName="sensu Voss and Jansa" baseAuthorityYear="2009" box="[307,440,788,812]" class="Mammalia" family="Didelphidae" kingdom="Animalia" order="Didelphimorphia" pageId="318" pageNumber="318" phylum="Chordata" rank="family">Didelphidae</taxonomicName>
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(Sparassocynini). Our analyses provide an additional test of the relationship between †
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<taxonomicName box="[349,493,855,878]" class="Mammalia" family="Sparassocynidae" genus="Hesperocynus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Didelphimorphia" pageId="318" pageNumber="318" phylum="Chordata" rank="genus">
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<emphasis box="[349,493,855,878]" italics="true" pageId="318" pageNumber="318">Hesperocynus</emphasis>
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</taxonomicName>
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and †
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<taxonomicName baseAuthorityName="Beck and Taglioretti" baseAuthorityYear="2020" class="Mammalia" family="Didelphidae" genus="Sparassocynus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Didelphimorphia" pageId="318" pageNumber="318" phylum="Chordata" rank="genus">
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<emphasis italics="true" pageId="318" pageNumber="318">Sparassocynus</emphasis>
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</taxonomicName>
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and Recent didelphids.
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</paragraph>
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<paragraph blockId="318.[108,638,226,1506]" pageId="318" pageNumber="318">
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We were able to examine FMNH
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<collectionCode box="[521,545,920,944]" pageId="318" pageNumber="318">P-</collectionCode>
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15225, a partial posterior braincase of †
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<taxonomicName box="[479,636,953,977]" class="Mammalia" family="Didelphidae" genus="Thylatheridium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Didelphimorphia" pageId="318" pageNumber="318" phylum="Chordata" rank="species" species="dolgopolae">
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<emphasis box="[479,636,953,977]" italics="true" pageId="318" pageNumber="318">H. dolgopolae</emphasis>
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</taxonomicName>
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from the Andalhuala Formation (see Simpson, 1974: figs. 7–10), but not the well-preserved skull from the Aisol Formation (MHNSR–PV 1046) described by Forasiepi et al. (2009). A number of character scores (particularly those relating to the dentition) have therefore been taken directly from Forasiepi et al.’s (2009) description and figures. As well as the Andalhuala and Aisol formations, †
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<taxonomicName box="[201,363,1251,1275]" class="Mammalia" family="Didelphidae" genus="Thylatheridium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Didelphimorphia" pageId="318" pageNumber="318" phylum="Chordata" rank="species" species="dolgopolae">
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<emphasis box="[201,363,1251,1275]" italics="true" pageId="318" pageNumber="318">H. dolgopolae</emphasis>
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</taxonomicName>
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specimens have been collected from the Cerro Azul Formation (
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<collectingRegion country="Argentina" name="La Pampa" pageId="318" pageNumber="318">La Pampa Province</collectingRegion>
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,
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<collectingCountry box="[308,421,1317,1341]" name="Argentina" pageId="318" pageNumber="318">Argentina</collectingCountry>
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; Goin et al., 2000; Abello et al., 2002; Forasiepi et al., 2009), but this last material has not been used for scoring purposes here and was not used to inform the assigned age range of our †
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<taxonomicName box="[412,558,1450,1473]" class="Mammalia" family="Sparassocynidae" genus="Hesperocynus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Didelphimorphia" pageId="318" pageNumber="318" phylum="Chordata" rank="genus">
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<emphasis box="[412,558,1450,1473]" italics="true" pageId="318" pageNumber="318">Hesperocynus</emphasis>
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</taxonomicName>
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terminal (see Püschel et al., 2020).
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |