240 lines
35 KiB
XML
240 lines
35 KiB
XML
<document id="13791BA2726086492414FED1609DD0D2" ID-DOI="10.1206/0003-0090.457.1.1" ID-ISSN="0003-0090" ID-Zenodo-Dep="6971356" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1659882917898" checkinUser="felipe" docAuthor="Beck, Robin M. D., Voss, Robert S. & Jansa, Sharon A." docDate="2022" docId="03EFDD5DF6E168F3DAD9FD1A1E17F96E" docLanguage="en" docName="BulAmeMusNatHist.2022.457.1-350.pdf" docOrigin="Bulletin of the American Museum of Natural History 2022 (457)" docSource="https://bioone.org/journals/bulletin-of-the-american-museum-of-natural-history/volume-457/issue-1/0003-0090.457.1.1/Craniodental-Morphology-and-Phylogeny-of-Marsupials/10.1206/0003-0090.457.1.1.full" docStyle="DocumentStyle:915933466F796C9C739DF4DB6B8DCFA6.9:BulAmeMusNatHis.2011-.journal_article.1cover.type1" docStyleId="915933466F796C9C739DF4DB6B8DCFA6" docStyleName="BulAmeMusNatHis.2011-.journal_article.1cover.type1" docStyleVersion="9" docTitle="Thylacinidae Bonaparte 1838" docType="treatment" docVersion="14" lastPageNumber="218" masterDocId="FFD6A525F63B6829DA0BFFAE1A62FFC7" masterDocTitle="Craniodental Morphology And Phylogeny Of Marsupials" masterLastPageNumber="353" masterPageNumber="1" pageNumber="218" updateTime="1699350301078" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="F4D4B90A87B9DF48A6D3AF1BEFEC0D61">Craniodental Morphology And Phylogeny Of Marsupials</mods:title>
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<mods:namePart id="FECBD20A8508966F7170BAA2FBA3E0D3">Beck, Robin M. D.</mods:namePart>
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<mods:affiliation id="2B98083231CA22E550AF4ED476E3D43E">School of Science, Engineering and Environment University of Salford, U. K. & School of Biological, Earth & Environmental Sciences University of New South Wales, Australia & Division of Vertebrate Zoology (Mammalogy) American Museum of Natural History</mods:affiliation>
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<mods:namePart id="0A4D33C2C03F044425975863BD5AF5DB">Voss, Robert S.</mods:namePart>
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<mods:affiliation id="8A3C1592CF2D57733650BD7D736483CA">Division of Vertebrate Zoology (Mammalogy) American Museum of Natural History</mods:affiliation>
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<mods:namePart id="8009501A8AD88CB08FA5E2937F86CD0A">Jansa, Sharon A.</mods:namePart>
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<mods:affiliation id="C150FE1E9EE114BD7EC0F7A091C0ACAB">Bell Museum and Department of Ecology, Evolution, and Behavior University of Minnesota</mods:affiliation>
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<mods:date id="B3DC2030F0AED81248666429C8CD9460">2022</mods:date>
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<mods:number id="773C0AAFEFFFA3576E4CCC2B75EACCFF">2022-06-28</mods:number>
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<mods:number id="F73E6EA966D5C2AF13260DAF5E17B673">457</mods:number>
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<treatment id="03EFDD5DF6E168F3DAD9FD1A1E17F96E" ID-DOI="http://doi.org/10.5281/zenodo.6974914" ID-Zenodo-Dep="6974914" LSID="urn:lsid:plazi:treatment:03EFDD5DF6E168F3DAD9FD1A1E17F96E" httpUri="http://treatment.plazi.org/id/03EFDD5DF6E168F3DAD9FD1A1E17F96E" lastPageNumber="218" pageId="218" pageNumber="218">
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<subSubSection id="C35C3FC0F6E168F3DAD9FD1A1874FD0B" box="[210,534,692,716]" pageId="218" pageNumber="218" type="nomenclature">
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<paragraph id="8BF96C4BF6E168F3DAD9FD1A1874FD0B" blockId="218.[210,534,692,716]" box="[210,534,692,716]" pageId="218" pageNumber="218">
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<heading id="D0B1DB27F6E168F3DAD9FD1A1874FD0B" box="[210,534,692,716]" centered="true" fontSize="9" level="2" pageId="218" pageNumber="218" reason="2">
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<taxonomicName id="4C4617C8F6E168F3DAD9FD1A1874FD0B" ID-CoL="H8N" authority="Bonaparte, 1838" authorityName="Bonaparte" authorityYear="1838" box="[210,534,692,716]" class="Mammalia" family="Thylacinidae" kingdom="Animalia" order="Dasyuromorphia" pageId="218" pageNumber="218" phylum="Chordata" rank="family">
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Thylacinidae
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<bibRefCitation id="EFD711BAF6E168F3DB6FFD1A1874FD0B" author="Bonaparte, C." box="[356,534,692,716]" pageId="218" pageNumber="218" pagination="105 - 133" refId="ref194576" refString="Bonaparte, C. 1838. Synopsis vertebratorum systematis. Annali delle Scienze Naturali, Bologna 2 (1): 105 - 133." type="journal article" year="1838">Bonaparte, 1838</bibRefCitation>
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</taxonomicName>
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||
</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C35C3FC0F6E168F3DA87FD5B1AF4FCE9" pageId="218" pageNumber="218" type="synonymic_list">
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<paragraph id="8BF96C4BF6E168F3DA87FD5B1AF4FCE9" blockId="218.[108,638,757,1707]" pageId="218" pageNumber="218">
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||
CONTENTS: †
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<taxonomicName id="4C4617C8F6E168F3DB16FD5B1BFFFCCA" authorityName="Muirhead & Archer" authorityYear="1990" box="[285,413,757,781]" class="Mammalia" family="Thylacinidae" genus="Nimbacinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dasyuromorphia" pageId="218" pageNumber="218" phylum="Chordata" rank="genus">
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<emphasis id="B932B059F6E168F3DB16FD5B1BFFFCCA" box="[285,413,757,781]" italics="true" pageId="218" pageNumber="218">Nimbacinus</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="4C4617C8F6E168F3DBD3FD5B1828FCCA" authorityName="Temminck" authorityYear="1824" box="[472,586,757,781]" class="Mammalia" family="Thylacinidae" genus="Thylacinus" kingdom="Animalia" order="Dasyuromorphia" pageId="218" pageNumber="218" phylum="Chordata" rank="genus">
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<emphasis id="B932B059F6E168F3DBD3FD5B1828FCCA" box="[472,586,757,781]" italics="true" pageId="218" pageNumber="218">Thylacinus</emphasis>
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</taxonomicName>
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(fig. 40).
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</paragraph>
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</subSubSection>
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<subSubSection id="C35C3FC0F6E168F3DA87FC991BFFFB9F" pageId="218" pageNumber="218" type="description">
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<paragraph id="8BF96C4BF6E168F3DA87FC991ACEFCB7" blockId="218.[108,638,757,1707]" pageId="218" pageNumber="218">STEM AGE: 31.2 Mya (95% HPD: 26.6–36.5 Mya).</paragraph>
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<paragraph id="8BF96C4BF6E168F3DA87FCD71ACFFC74" blockId="218.[108,638,757,1707]" pageId="218" pageNumber="218">CROWN AGE: 17.1 Mya (95% HPD: 11.6–24.0 Mya).</paragraph>
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<paragraph id="8BF96C4BF6E168F3DA87FC121BFFFB9F" blockId="218.[108,638,757,1707]" pageId="218" pageNumber="218">UNAMBIGUOUS CRANIODENTAL SYNAPOMORPHIES: Presphenoid exposed in roof of nasopharyngeal fossa above posterior palate (char. 43: 1→0; ci = 0.091) and p3 distinctly taller than p2 (char. 156: 0→2; ci = 0.118).</paragraph>
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</subSubSection>
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<subSubSection id="C35C3FC0F6E168F3DA87FBCF1E17F96E" pageId="218" pageNumber="218" type="discussion">
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<paragraph id="8BF96C4BF6E168F3DA87FBCF1950FDE4" blockId="218.[108,638,757,1707]" lastBlockId="218.[684,1214,226,1705]" pageId="218" pageNumber="218">
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COMMENTS:
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<taxonomicName id="4C4617C8F6E168F3DB10FBCF1BC4FBBE" authorityName="Bonaparte" authorityYear="1838" box="[283,422,1121,1145]" class="Mammalia" family="Thylacinidae" kingdom="Animalia" order="Dasyuromorphia" pageId="218" pageNumber="218" phylum="Chordata" rank="family">Thylacinidae</taxonomicName>
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—defined by
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<bibRefCitation id="EFD711BAF6E168F3D834FBCF1B43FB5D" author="Kealy, S. & R. M. D. Beck" pageId="218" pageNumber="218" pagination="240" refId="ref210569" refString="Kealy, S., and R. M. D. Beck. 2017. Total evidence phylogeny and evolutionary timescale for Australian faunivorous marsupials (Dasyuromorphia). BMC Evolutionary Biology 17 (1): 240." type="journal article" year="2017">Kealy and Beck (2017</bibRefCitation>
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||
:
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||
<tableCitation id="C6C459F0F6E168F3DB25FB2C1B1BFB5D" box="[302,377,1154,1178]" captionStart="TABLE 1" captionStartId="7.[611,691,226,250]" captionTargetBox="[119,1110,327,1337]" captionTargetPageId="7" captionText="TABLE 1 Higher Classification and Geographic Distribution of Recent Marsupialsa" httpUri="http://table.plazi.org/id/DF393CC3F63C682ED868FF4C1E65FED9" pageId="218" pageNumber="218" tableUuid="DF393CC3F63C682ED868FF4C1E65FED9">table 1</tableCitation>
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||
) as the most inclusive clade including
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<taxonomicName id="4C4617C8F6E168F3DB26FB0D1820FB7C" baseAuthorityName="Harris" baseAuthorityYear="1808" box="[301,578,1187,1211]" class="Mammalia" family="Thylacinidae" genus="Thylacinus" kingdom="Animalia" order="Dasyuromorphia" pageId="218" pageNumber="218" phylum="Chordata" rank="species" species="cynocephalus">
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<emphasis id="B932B059F6E168F3DB26FB0D1820FB7C" box="[301,578,1187,1211]" italics="true" pageId="218" pageNumber="218">Thylacinus cynocephalus</emphasis>
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</taxonomicName>
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, but excluding
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<taxonomicName id="4C4617C8F6E168F3DAD6FB6B1BD4FB1B" baseAuthorityName="Shaw" baseAuthorityYear="1800" box="[221,438,1221,1244]" class="Mammalia" family="Dasyuridae" genus="Dasyurus" kingdom="Animalia" order="Dasyuromorphia" pageId="218" pageNumber="218" phylum="Chordata" rank="species" species="viverrinus">
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<emphasis id="B932B059F6E168F3DAD6FB6B1BD4FB1B" box="[221,438,1221,1244]" italics="true" pageId="218" pageNumber="218">Dasyurus viverrinus</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="4C4617C8F6E168F3DBE6FB6A1AAAFB3A" authorityName="Waterhouse" authorityYear="1836" class="Mammalia" family="Myrmecobiidae" genus="Myrmecobius" kingdom="Animalia" order="Dasyuromorphia" pageId="218" pageNumber="218" phylum="Chordata" rank="species" species="fasciatus">
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<emphasis id="B932B059F6E168F3DBE6FB6A1AAAFB3A" italics="true" pageId="218" pageNumber="218">Myrmecobius fasciatus</emphasis>
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</taxonomicName>
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—comprises only
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<taxonomicName id="4C4617C8F6E168F3DB85FB4B1862FB3A" authorityName="Temminck" authorityYear="1824" box="[398,512,1253,1277]" class="Mammalia" family="Thylacinidae" genus="Thylacinus" kingdom="Animalia" order="Dasyuromorphia" pageId="218" pageNumber="218" phylum="Chordata" rank="genus">
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<emphasis id="B932B059F6E168F3DB85FB4B1862FB3A" box="[398,512,1253,1277]" italics="true" pageId="218" pageNumber="218">Thylacinus</emphasis>
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</taxonomicName>
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and †
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<taxonomicName id="4C4617C8F6E168F3D84FFB481ADFFAD9" authorityName="Muirhead & Archer" authorityYear="1990" class="Mammalia" family="Thylacinidae" genus="Nimbacinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dasyuromorphia" pageId="218" pageNumber="218" phylum="Chordata" rank="genus">
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<emphasis id="B932B059F6E168F3D84FFB481ADFFAD9" italics="true" pageId="218" pageNumber="218">Nimbacinus</emphasis>
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</taxonomicName>
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in our dated total-evidence analysis (
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<figureCitation id="137D70CEF6E168F3D852FAA81AE9FAF8" captionStart="FIG" captionStartId="181.[108,150,938,959]" captionTargetId="figure-12@181.[241,1216,224,1485]" captionTargetPageId="181" captionText="FIG. 33. (opposite page and at right) Fifty-per- cent majority rule consensus of post-burn-in trees that results from dated Bayesian analysis (using combined tip-and-node dating and separate Independent Gamma Rates [IGR] clock models for the molecular and morpho- logical partitions) of our total evidence data- set. Black dots at nodes indicate ≥0.95 Bayesian posterior probability (“strong support”); dark gray dots indicate 0.75–0.94 Bayesian poste- rior probability (“moderate support”); light gray dots indicate 0.50–0.74 Bayesian poste- rior probability (“weak support”). Nodes with- out dots were constrained a priori so that their ages could be calibrated (see tables 6, 13). Orange bars represent 95% Highest Posterior Density (HPD) intervals on the ages of nodes. For clarity, 95% HPD intervals are not shown for the ages of fossil terminals." pageId="218" pageNumber="218">fig. 33</figureCitation>
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). As already discussed (see
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<taxonomicName id="4C4617C8F6E168F3DBC8FA89181FFAF8" authorityName="sensu Kealy and Beck" authorityYear="2017" box="[451,637,1319,1343]" class="Mammalia" kingdom="Animalia" order="Dasyuromorphia" pageId="218" pageNumber="218" phylum="Chordata" rank="order">Dasyuromorphia</taxonomicName>
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above), †
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<taxonomicName id="4C4617C8F6E168F3DADBFAE61B5BFAA7" baseAuthorityName="Muirhead and Wroe" baseAuthorityYear="1998" box="[208,313,1352,1376]" class="Mammalia" family="Thylacinidae" genus="Badjcinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dasyuromorphia" pageId="218" pageNumber="218" phylum="Chordata" rank="genus">
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<emphasis id="B932B059F6E168F3DADBFAE61B5BFAA7" box="[208,313,1352,1376]" italics="true" pageId="218" pageNumber="218">Badjcinus</emphasis>
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</taxonomicName>
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was originally described as a thylacinid by
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<bibRefCitation id="EFD711BAF6E168F3DB0EFAC71856FA46" author="Muirhead, J. & S. Wroe" box="[261,564,1385,1409]" pageId="218" pageNumber="218" pagination="612 - 626" refId="ref217610" refString="Muirhead, J., and S. Wroe. 1998. A new genus and species, Badjcinus turnbulli (Thylacinidae: Marsupialia), from the late Oligocene of Riversleigh, northern Australia, and an investigation of thylacinid phylogeny. Journal of Vertebrate Paleontology 18: 612 - 626." type="journal article" year="1998">Muirhead and Wroe (1998)</bibRefCitation>
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and is placed within
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<taxonomicName id="4C4617C8F6E168F3DB03FA241BF1FA65" authorityName="Bonaparte" authorityYear="1838" box="[264,403,1418,1442]" class="Mammalia" family="Thylacinidae" kingdom="Animalia" order="Dasyuromorphia" pageId="218" pageNumber="218" phylum="Chordata" rank="family">Thylacinidae</taxonomicName>
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in our undated totalevidence anlaysis (fig. 32) but not in our dated total-evidence analysis (
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<figureCitation id="137D70CEF6E168F3DB7CFA621BA0FA22" box="[375,450,1484,1509]" captionStart="FIG" captionStartId="181.[108,150,938,959]" captionTargetId="figure-12@181.[241,1216,224,1485]" captionTargetPageId="181" captionText="FIG. 33. (opposite page and at right) Fifty-per- cent majority rule consensus of post-burn-in trees that results from dated Bayesian analysis (using combined tip-and-node dating and separate Independent Gamma Rates [IGR] clock models for the molecular and morpho- logical partitions) of our total evidence data- set. Black dots at nodes indicate ≥0.95 Bayesian posterior probability (“strong support”); dark gray dots indicate 0.75–0.94 Bayesian poste- rior probability (“moderate support”); light gray dots indicate 0.50–0.74 Bayesian poste- rior probability (“weak support”). Nodes with- out dots were constrained a priori so that their ages could be calibrated (see tables 6, 13). Orange bars represent 95% Highest Posterior Density (HPD) intervals on the ages of nodes. For clarity, 95% HPD intervals are not shown for the ages of fossil terminals." pageId="218" pageNumber="218">fig. 33</figureCitation>
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). †
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<taxonomicName id="4C4617C8F6E168F3DBEEFA631AB5F9C1" authorityName="Murray & Megirian" authorityYear="2000" class="Mammalia" family="Thylacinidae" genus="Mutpuracinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dasyuromorphia" pageId="218" pageNumber="218" phylum="Chordata" rank="species" species="archibaldi">
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<emphasis id="B932B059F6E168F3DBEEFA631AB5F9C1" italics="true" pageId="218" pageNumber="218">Mutpuracinus archibaldi</emphasis>
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</taxonomicName>
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is another dasyuromorphian that was originally described as a thylacinid (
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<bibRefCitation id="EFD711BAF6E168F3DBF1F9A11B7AF98F" author="Murray, P. F. & D. Megirian" pageId="218" pageNumber="218" pagination="145 - 162" refId="ref218837" refString="Murray, P. F., and D. Megirian. 2000. Two new genera and three new species of Thylacinidae (Marsupialia) from the Miocene of the Northern Territory, Australia. Beagle, Records of the Museums and Art Galleries of the Northern Territory 16: 145 - 162." type="journal article" year="2000">Murray and Megirian, 2000</bibRefCitation>
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,
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<bibRefCitation id="EFD711BAF6E168F3DB2EF99E1B08F98F" author="Murray, P. F. & D. Megirian" box="[293,362,1584,1608]" pageId="218" pageNumber="218" pagination="229 - 276" refId="ref218892" refString="Murray, P. F., and D. Megirian. 2006 a. Cranial morphology of the Miocene thylacinid Mutpuracinus archibaldi (Thylacinidae, Marsupialia) and relationships within the Dasyuromorphia). Alcheringa: an Australasian Journal of Palaeontology Special Issue 1: 229 - 276." type="book chapter" year="2006">2006a</bibRefCitation>
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), but in our dated totalevidence analysis it is recovered in a clade with another fossil dasyuromorphian, †
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<taxonomicName id="4C4617C8F6E168F3DBE3F9DD1AFAF96C" authorityName="Wroe" authorityYear="1999" class="Mammalia" family="Dasyuridae" genus="Barinya" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dasyuromorphia" pageId="218" pageNumber="218" phylum="Chordata" rank="species" species="wangala">
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<emphasis id="B932B059F6E168F3DBE3F9DD1AFAF96C" italics="true" pageId="218" pageNumber="218">Barinya wangala</emphasis>
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</taxonomicName>
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, and the only extant myrmecobiid,
|
||
<taxonomicName id="4C4617C8F6E168F3D823F93A188CFF3D" baseAuthorityName="Flower" baseAuthorityYear="1867" box="[552,750,226,1707]" class="Mammalia" family="Myrmecobiidae" genus="Myrmecobius" kingdom="Animalia" order="Dasyuromorphia" pageId="218" pageNumber="218" phylum="Chordata" rank="genus">
|
||
<emphasis id="B932B059F6E168F3D823F93A188CFF3D" box="[552,750,226,1707]" italics="true" pageId="218" pageNumber="218">Myrmecobius</emphasis>
|
||
</taxonomicName>
|
||
; this clade is, in turn, sister to
|
||
<taxonomicName id="4C4617C8F6E168F3DE4BFF4C1EDEFF3D" authorityName="Goldfuss" authorityYear="1820" box="[1088,1212,226,250]" class="Mammalia" family="Dasyuridae" kingdom="Animalia" order="Dasyuromorphia" pageId="218" pageNumber="218" phylum="Chordata" rank="family">Dasyuridae</taxonomicName>
|
||
(
|
||
<figureCitation id="137D70CEF6E168F3D8BFFEAD189FFEDC" box="[692,765,259,283]" captionStart="FIG" captionStartId="181.[108,150,938,959]" captionTargetId="figure-12@181.[241,1216,224,1485]" captionTargetPageId="181" captionText="FIG. 33. (opposite page and at right) Fifty-per- cent majority rule consensus of post-burn-in trees that results from dated Bayesian analysis (using combined tip-and-node dating and separate Independent Gamma Rates [IGR] clock models for the molecular and morpho- logical partitions) of our total evidence data- set. Black dots at nodes indicate ≥0.95 Bayesian posterior probability (“strong support”); dark gray dots indicate 0.75–0.94 Bayesian poste- rior probability (“moderate support”); light gray dots indicate 0.50–0.74 Bayesian poste- rior probability (“weak support”). Nodes with- out dots were constrained a priori so that their ages could be calibrated (see tables 6, 13). Orange bars represent 95% Highest Posterior Density (HPD) intervals on the ages of nodes. For clarity, 95% HPD intervals are not shown for the ages of fossil terminals." pageId="218" pageNumber="218">fig. 33</figureCitation>
|
||
). †
|
||
<taxonomicName id="4C4617C8F6E168F3D917FEAD1913FEDD" baseAuthorityName="Wroe" baseAuthorityYear="1999" box="[796,881,259,282]" class="Mammalia" family="Dasyuridae" genus="Barinya" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dasyuromorphia" pageId="218" pageNumber="218" phylum="Chordata" rank="genus">
|
||
<emphasis id="B932B059F6E168F3D917FEAD1913FEDD" box="[796,881,259,282]" italics="true" pageId="218" pageNumber="218">Barinya</emphasis>
|
||
</taxonomicName>
|
||
itself was originally described as the oldest and most plesiomorphic known dasyurid (
|
||
<bibRefCitation id="EFD711BAF6E168F3D910FEEB19FDFE9A" author="Wroe, S." box="[795,927,325,349]" pageId="218" pageNumber="218" pagination="1 - 27" refId="ref236443" refString="Wroe, S. 1999. The geologically oldest dasyurid (Marsupialia), from the Miocene Riversleigh, northwestern Queensland. Palaeontology 42: 1 - 27." type="journal article" year="1999">Wroe, 1999</bibRefCitation>
|
||
), but this inference is not supported here. Again, our results are broadly congruent with those of
|
||
<bibRefCitation id="EFD711BAF6E168F3D9B7FE291EDAFE58" author="Kealy, S. & R. M. D. Beck" box="[956,1208,391,415]" pageId="218" pageNumber="218" pagination="240" refId="ref210569" refString="Kealy, S., and R. M. D. Beck. 2017. Total evidence phylogeny and evolutionary timescale for Australian faunivorous marsupials (Dasyuromorphia). BMC Evolutionary Biology 17 (1): 240." type="journal article" year="2017">Kealy and Beck (2017)</bibRefCitation>
|
||
, which likewise did not support thylacinid affinities for †
|
||
<taxonomicName id="4C4617C8F6E168F3D900FE6419C3FE26" authorityName="Murray & Megirian" authorityYear="2000" box="[779,929,458,481]" class="Mammalia" family="Thylacinidae" genus="Mutpuracinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dasyuromorphia" pageId="218" pageNumber="218" phylum="Chordata" rank="genus">
|
||
<emphasis id="B932B059F6E168F3D900FE6419C3FE26" box="[779,929,458,481]" italics="true" pageId="218" pageNumber="218">Mutpuracinus</emphasis>
|
||
</taxonomicName>
|
||
nor dasyurid affinities for †
|
||
<taxonomicName id="4C4617C8F6E168F3D8B2FE45196CFDC5" baseAuthorityName="Wroe" baseAuthorityYear="1999" box="[697,782,491,514]" class="Mammalia" family="Dasyuridae" genus="Barinya" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dasyuromorphia" pageId="218" pageNumber="218" phylum="Chordata" rank="genus">
|
||
<emphasis id="B932B059F6E168F3D8B2FE45196CFDC5" box="[697,782,491,514]" italics="true" pageId="218" pageNumber="218">Barinya</emphasis>
|
||
</taxonomicName>
|
||
(see also
|
||
<bibRefCitation id="EFD711BAF6E168F3D97DFE441E2FFDC5" author="Eldridge, M. D. B. & R. M. D. Beck & D. A. Croft & K. J. Travouillon & B. J. Fox" box="[886,1101,490,514]" pageId="218" pageNumber="218" pagination="802 - 837" refId="ref200965" refString="Eldridge, M. D. B., R. M. D. Beck, D. A. Croft, K. J. Travouillon, and B. J. Fox. 2019. An emerging consensus in the evolution, phylogeny, and systematics of marsupials and their fossil relatives (Metatheria). Journal of Mammalogy 100 (3): 802 - 837." type="journal article" year="2019">Eldridge et al., 2019</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFD711BAF6E168F3DE5CFE441945FDE4" author="Rovinsky" etAl="et al." firstAuthor="Rovinsky" pageId="218" pageNumber="218" pagination="7457" refId="ref225576" refString="Rovinsky, D. S., A. R. Evans, and J. W. Adams. 2019. The pre-Pleistocene fossil thylacinids (Dasyuromorphia: Thylacinidae) and the evolutionary context of the modern thylacine. PeerJ 7: e 7457." type="journal article" year="2019">Rovinsky et al., 2019</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BF96C4BF6E168F3D8C6FD821E77FA67" blockId="218.[684,1214,226,1705]" pageId="218" pageNumber="218">
|
||
The oldest putative thylacinid remains are a partial upper molar (NTM P2815-10) from the late Oligocene Pwerte Marnte Marnte Local Fauna in the
|
||
<collectingRegion id="4982A2A9F6E168F3D94DFD211E7FFD60" box="[838,1053,655,679]" country="Australia" name="Northern Territory" pageId="218" pageNumber="218">Northern Territory</collectingRegion>
|
||
, and a partial lower molar (QM
|
||
<accessionNumber id="9415F1A8F6E168F3D972FD1E19AFFD0F" box="[889,973,688,712]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/F16809" pageId="218" pageNumber="218" type="EnaNcbi">F16809</accessionNumber>
|
||
) from the late Oligocene (Faunal Zone A) D-site at Riversleigh that was originally described as a
|
||
<typeStatus id="54FDD2E9F6E168F3DE00FD5D1E0FFCCC" box="[1035,1133,755,779]" pageId="218" pageNumber="218" type="paratype">paratype</typeStatus>
|
||
of the thylacinid †
|
||
<taxonomicName id="4C4617C8F6E168F3D93DFCBA1E7CFCEB" authorityName="Muirhead & Archer" authorityYear="1990" box="[822,1054,788,812]" class="Mammalia" family="Thylacinidae" genus="Nimbacinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dasyuromorphia" pageId="218" pageNumber="218" phylum="Chordata" rank="species" species="dicksoni">
|
||
<emphasis id="B932B059F6E168F3D93DFCBA1E7CFCEB" box="[822,1054,788,812]" italics="true" pageId="218" pageNumber="218">Nimbacinus dicksoni</emphasis>
|
||
</taxonomicName>
|
||
by
|
||
<bibRefCitation id="EFD711BAF6E168F3DE46FCBA19F2FC8A" author="Muirhead, J. & M. Archer" pageId="218" pageNumber="218" pagination="203 - 221" refId="ref217483" refString="Muirhead, J., and M. Archer. 1990. Nimbacinus dicksoni, a plesiomorphic thylacine (Marsupialia, Thylacinidae) from Tertiary deposits of Queensland and the Northern Territory. Memoirs of the Queensland Museum 28: 203 - 221." type="journal article" year="1990">Muirhead and Archer (1990)</bibRefCitation>
|
||
, but which
|
||
<bibRefCitation id="EFD711BAF6E168F3DE26FC9B19FDFCA9" author="Murray, P. F. & D. Megirian" pageId="218" pageNumber="218" pagination="145 - 162" refId="ref218837" refString="Murray, P. F., and D. Megirian. 2000. Two new genera and three new species of Thylacinidae (Marsupialia) from the Miocene of the Northern Territory, Australia. Beagle, Records of the Museums and Art Galleries of the Northern Territory 16: 145 - 162." type="journal article" year="2000">Murray and Megirian (2000: 159)</bibRefCitation>
|
||
concluded “represents a different taxon, probably another genus,”
|
||
<bibRefCitation id="EFD711BAF6E168F3DE8BFCD919ABFC77" author="Wroe and Musser" firstAuthor="Wroe" page="502" pageId="218" pageNumber="218" pagination="487 - 514" refId="ref236708" refString="Wroe, S., and A. Musser. 2001. The skull of Nimbacinus dicksoni (Thylacinidae: Marsupialia). Australian Journal of Zoology 49 (5): 487 - 514." type="journal article" year="2001">Wroe and Musser (2001: 502)</bibRefCitation>
|
||
considered to be of “uncertain” taxonomic status, and
|
||
<bibRefCitation id="EFD711BAF6E168F3DE3CFC17197FFC35" author="Rovinsky" etAl="et al." firstAuthor="Rovinsky" pageId="218" pageNumber="218" pagination="7457" refId="ref225576" refString="Rovinsky, D. S., A. R. Evans, and J. W. Adams. 2019. The pre-Pleistocene fossil thylacinids (Dasyuromorphia: Thylacinidae) and the evolutionary context of the modern thylacine. PeerJ 7: e 7457." type="journal article" year="2019">Rovinsky et al. (2019)</bibRefCitation>
|
||
formally classified as
|
||
<taxonomicName id="4C4617C8F6E168F3DE27FC741EDEFC35" authorityName="Bonaparte" authorityYear="1838" box="[1068,1212,986,1010]" class="Mammalia" family="Thylacinidae" kingdom="Animalia" order="Dasyuromorphia" pageId="218" pageNumber="218" phylum="Chordata" rank="family">Thylacinidae</taxonomicName>
|
||
incertae sedis. Neither specimen has been included in published phylogenetic analyses, so even if they really are thylacinids, their relationship to the two thylacinid terminals included here, †
|
||
<taxonomicName id="4C4617C8F6E168F3D8F7FBD119E2FB50" authorityName="Muirhead & Archer" authorityYear="1990" box="[764,896,1151,1175]" class="Mammalia" family="Thylacinidae" genus="Nimbacinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dasyuromorphia" pageId="218" pageNumber="218" phylum="Chordata" rank="genus">
|
||
<emphasis id="B932B059F6E168F3D8F7FBD119E2FB50" box="[764,896,1151,1175]" italics="true" pageId="218" pageNumber="218">Nimbacinus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C4617C8F6E168F3D9B4FBD11E57FB50" authorityName="Temminck" authorityYear="1824" box="[959,1077,1151,1175]" class="Mammalia" family="Thylacinidae" genus="Thylacinus" kingdom="Animalia" order="Dasyuromorphia" pageId="218" pageNumber="218" phylum="Chordata" rank="genus">
|
||
<emphasis id="B932B059F6E168F3D9B4FBD11E57FB50" box="[959,1077,1151,1175]" italics="true" pageId="218" pageNumber="218">Thylacinus</emphasis>
|
||
</taxonomicName>
|
||
, is unclear. †
|
||
<taxonomicName id="4C4617C8F6E168F3D8B1FB0E1E62FB7F" authorityName="Muirhead" authorityYear="1997" box="[698,1024,1184,1208]" class="Mammalia" family="Thylacinidae" genus="Ngamalacinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dasyuromorphia" pageId="218" pageNumber="218" phylum="Chordata" rank="species" species="timmulvaneyi">
|
||
<emphasis id="B932B059F6E168F3D8B1FB0E1E62FB7F" box="[698,1024,1184,1208]" italics="true" pageId="218" pageNumber="218">Ngamalacinus timmulvaneyi</emphasis>
|
||
</taxonomicName>
|
||
from the early Miocene (Faunal Zone B) sites at Riversleigh (
|
||
<bibRefCitation id="EFD711BAF6E168F3D8BFFB4D191DFB3C" author="Muirhead, J." box="[692,895,1251,1275]" pageId="218" pageNumber="218" pagination="367 - 377" refId="ref217407" refString="Muirhead, J. 1997. Two new early Miocene thylacines from Riversleigh, northwestern Queensland. Memoirs of the Queensland Museum 41 (2): 367 - 377." type="journal article" year="1997">Muirhead, 1997</bibRefCitation>
|
||
) has likewise not been included here due to its incompleteness, but it was consistently recovered as a thylacinid in the total-evidence analyses of
|
||
<bibRefCitation id="EFD711BAF6E168F3D9F2FAE81895FAB8" author="Kealy, S. & R. M. D. Beck" pageId="218" pageNumber="218" pagination="240" refId="ref210569" refString="Kealy, S., and R. M. D. Beck. 2017. Total evidence phylogeny and evolutionary timescale for Australian faunivorous marsupials (Dasyuromorphia). BMC Evolutionary Biology 17 (1): 240." type="journal article" year="2017">Kealy and Beck (2017)</bibRefCitation>
|
||
, so we consider it to be the oldest definitive thylacinid currently known.
|
||
</paragraph>
|
||
<paragraph id="8BF96C4BF6E168F3D8C7FA071E17F96E" blockId="218.[684,1214,226,1705]" pageId="218" pageNumber="218">
|
||
Our estimate for the divergence between
|
||
<taxonomicName id="4C4617C8F6E168F3DE87FA07189AFA25" authorityName="Temminck" authorityYear="1824" class="Mammalia" family="Thylacinidae" genus="Thylacinus" kingdom="Animalia" order="Dasyuromorphia" pageId="218" pageNumber="218" phylum="Chordata" rank="genus">
|
||
<emphasis id="B932B059F6E168F3DE87FA07189AFA25" italics="true" pageId="218" pageNumber="218">Thylacinus</emphasis>
|
||
</taxonomicName>
|
||
and †
|
||
<taxonomicName id="4C4617C8F6E168F3D948FA6419A6FA25" authorityName="Muirhead & Archer" authorityYear="1990" box="[835,964,1482,1506]" class="Mammalia" family="Thylacinidae" genus="Nimbacinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dasyuromorphia" pageId="218" pageNumber="218" phylum="Chordata" rank="genus">
|
||
<emphasis id="B932B059F6E168F3D948FA6419A6FA25" box="[835,964,1482,1506]" italics="true" pageId="218" pageNumber="218">Nimbacinus</emphasis>
|
||
</taxonomicName>
|
||
is latest Oligocene to middle Miocene, congruent with the age of the oldest known representative of the genus
|
||
<taxonomicName id="4C4617C8F6E168F3DE6CF9A218B0F982" authorityName="Temminck" authorityYear="1824" class="Mammalia" family="Thylacinidae" genus="Thylacinus" kingdom="Animalia" order="Dasyuromorphia" pageId="218" pageNumber="218" phylum="Chordata" rank="genus">
|
||
<emphasis id="B932B059F6E168F3DE6CF9A218B0F982" italics="true" pageId="218" pageNumber="218">Thylacinus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="B932B059F6E168F3D8E9F9801895F982" box="[738,759,1582,1605]" italics="true" pageId="218" pageNumber="218">T.</emphasis>
|
||
†
|
||
<emphasis id="B932B059F6E168F3D906F983191AF982" box="[781,888,1581,1605]" italics="true" pageId="218" pageNumber="218">macknessi</emphasis>
|
||
, which is from middle Miocene (Faunal Zone C) sites at Riversleigh (
|
||
<bibRefCitation id="EFD711BAF6E168F3D8B8F9C11907F94F" author="Muirhead, J." box="[691,869,1647,1672]" pageId="218" pageNumber="218" pagination="67 - 76" refId="ref217331" refString="Muirhead, J. 1992. A specialized thylacinid, Thylacinus macknessi, (Marsupialia: Thylacinidae) from Miocene deposits of Riversleigh, northwestern Queensland. Australian Mammalogy 15: 67 - 76." type="journal article" year="1992">Muirhead, 1992</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFD711BAF6E168F3D97AF9C11EDAF940" author="Muirhead, J. & A. K. Gillespie" box="[881,1208,1647,1671]" pageId="218" pageNumber="218" pagination="55 - 60" refId="ref217565" refString="Muirhead, J., and A. K. Gillespie. 1995. Additional parts of the type specimen of Thylacinus macknessi (Marsupialia: Thylacinidae) from Miocene deposits of Riversleigh, northwestern Queensland. Australian Mammalogy 18: 55 - 60." type="journal article" year="1995">Muirhead and Gillespie, 1995</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFD711BAF6E168F3D8A5F93F194BF96E" author="Yates" box="[686,809,1681,1705]" firstAuthor="Yates" pageId="218" pageNumber="218" pagination="547" refId="ref237068" refString="Yates, A. M. 2014. New craniodental remains of Thylacinus potens (Dasyuromorphia: Thylacinidae), a carnivorous marsupial from the late Miocene Alcoota Local Fauna of central Australia. PeerJ 2: e 547." type="journal article" year="2014">Yates, 2014</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFD711BAF6E168F3D93FF93E1915F96F" author="Yates" box="[820,887,1680,1704]" firstAuthor="Yates" pageId="218" pageNumber="218" pagination="931" refId="ref237108" refString="Yates, A. M. 2015 a. Thylacinus (Marsupialia: Thylacinidae) from the Mio-Pliocene boundary and the diversity of late Neogene thylacinids in Australia. PeerJ 3: e 931." type="journal article" year="2015" yearSuffix="a">2015a</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFD711BAF6E168F3D989F93E1E0BF96E" author="Rovinsky" box="[898,1129,1680,1705]" etAl="et al." firstAuthor="Rovinsky" pageId="218" pageNumber="218" pagination="7457" refId="ref225576" refString="Rovinsky, D. S., A. R. Evans, and J. W. Adams. 2019. The pre-Pleistocene fossil thylacinids (Dasyuromorphia: Thylacinidae) and the evolutionary context of the modern thylacine. PeerJ 7: e 7457." type="journal article" year="2019">Rovinsky et al., 2019</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |