225 lines
29 KiB
XML
225 lines
29 KiB
XML
<document ID-DOI="10.1206/0003-0090.457.1.1" ID-ISSN="0003-0090" ID-Zenodo-Dep="6971356" approvalRequired="59" approvalRequired_for_taxonomicNames="59" checkinTime="1659882917898" checkinUser="felipe" docAuthor="Beck, Robin M. D., Voss, Robert S. & Jansa, Sharon A." docDate="2022" docId="03EFDD5DF6D868CDD8F0FA141E72FD0F" docLanguage="en" docName="BulAmeMusNatHist.2022.457.1-350.pdf" docOrigin="Bulletin of the American Museum of Natural History 2022 (457)" docSource="https://bioone.org/journals/bulletin-of-the-american-museum-of-natural-history/volume-457/issue-1/0003-0090.457.1.1/Craniodental-Morphology-and-Phylogeny-of-Marsupials/10.1206/0003-0090.457.1.1.full" docStyle="DocumentStyle:915933466F796C9C739DF4DB6B8DCFA6.8:BulAmeMusNatHis.2011-.journal_article.1cover.type1" docStyleId="915933466F796C9C739DF4DB6B8DCFA6" docStyleName="BulAmeMusNatHis.2011-.journal_article.1cover.type1" docStyleVersion="8" docTitle="Vombatiformes Woodburne 1984" docType="treatment" docVersion="9" lastPageNumber="228" masterDocId="FFD6A525F63B6829DA0BFFAE1A62FFC7" masterDocTitle="Craniodental Morphology And Phylogeny Of Marsupials" masterLastPageNumber="353" masterPageNumber="1" pageNumber="227" updateTime="1659987007554" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:titleInfo>
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<mods:title>Craniodental Morphology And Phylogeny Of Marsupials</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Beck, Robin M. D.</mods:namePart>
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<mods:affiliation>School of Science, Engineering and Environment University of Salford, U. K. & School of Biological, Earth & Environmental Sciences University of New South Wales, Australia & Division of Vertebrate Zoology (Mammalogy) American Museum of Natural History</mods:affiliation>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Voss, Robert S.</mods:namePart>
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<mods:affiliation>Division of Vertebrate Zoology (Mammalogy) American Museum of Natural History</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Jansa, Sharon A.</mods:namePart>
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<mods:affiliation>Bell Museum and Department of Ecology, Evolution, and Behavior University of Minnesota</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:title>Bulletin of the American Museum of Natural History</mods:title>
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<mods:part>
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<mods:date>2022</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2022-06-28</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>2022</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>457</mods:number>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:url>https://bioone.org/journals/bulletin-of-the-american-museum-of-natural-history/volume-457/issue-1/0003-0090.457.1.1/Craniodental-Morphology-and-Phylogeny-of-Marsupials/10.1206/0003-0090.457.1.1.full</mods:url>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.1206/0003-0090.457.1.1</mods:identifier>
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<mods:identifier type="ISSN">0003-0090</mods:identifier>
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<mods:identifier type="Zenodo-Dep">6971356</mods:identifier>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.6974219" ID-Zenodo-Dep="6974219" LSID="urn:lsid:plazi:treatment:03EFDD5DF6D868CDD8F0FA141E72FD0F" httpUri="http://treatment.plazi.org/id/03EFDD5DF6D868CDD8F0FA141E72FD0F" lastPageId="228" lastPageNumber="228" pageId="227" pageNumber="227">
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<subSubSection box="[763,1135,1466,1490]" pageId="227" pageNumber="227" type="nomenclature">
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<paragraph blockId="227.[763,1135,1466,1490]" box="[763,1135,1466,1490]" pageId="227" pageNumber="227">
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<heading box="[763,1135,1466,1490]" centered="true" fontSize="9" level="2" pageId="227" pageNumber="227" reason="2">
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<taxonomicName authority="Woodburne, 1984" authorityName="Woodburne" authorityYear="1984" box="[763,1135,1466,1490]" genus="Vombatiformes" pageId="227" pageNumber="227" rank="genus">
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Vombatiformes
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<bibRefCitation author="Woodburne, M. O." box="[939,1135,1466,1490]" pageId="227" pageNumber="227" pagination="1062 - 1073" refId="ref234899" refString="Woodburne, M. O. 1984 a. Wakiewakie lawsoni, a new genus and species of Potoroinae (Marsupialia: Macropodidae) of medial Miocene age, South Australia. Journal of Paleontology 58 (4): 1062 - 1073." type="journal article" year="1984">Woodburne, 1984</bibRefCitation>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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||
<subSubSection pageId="227" pageNumber="227" type="synonymic_list">
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<paragraph blockId="227.[684,1212,1515,1738]" pageId="227" pageNumber="227">
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CONTENTS: †Diprotodontoidea, †
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<taxonomicName authorityName="Tedford and Woodburne" authorityYear="1987" box="[1109,1207,1515,1539]" class="Mammalia" family="Ilariidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="227" pageNumber="227" phylum="Chordata" rank="family">Ilariidae</taxonomicName>
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, †
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<taxonomicName box="[698,817,1549,1572]" class="Mammalia" family="Wynyardiidae" genus="Muramura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="227" pageNumber="227" phylum="Chordata" rank="genus">
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<emphasis box="[698,817,1549,1572]" italics="true" pageId="227" pageNumber="227">Muramura</emphasis>
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</taxonomicName>
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, †
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<taxonomicName baseAuthorityName="Pledge" baseAuthorityYear="2005" box="[850,1012,1548,1572]" class="Mammalia" family="Wynyardiidae" genus="Namilamadeta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="227" pageNumber="227" phylum="Chordata" rank="genus">
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<emphasis box="[850,1012,1548,1572]" italics="true" pageId="227" pageNumber="227">Namilamadeta</emphasis>
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</taxonomicName>
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,
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<taxonomicName authorityName="Owen" authorityYear="1839" box="[1031,1208,1548,1572]" class="Mammalia" family="Phascolarctidae" kingdom="Animalia" order="Diprotodontia" pageId="227" pageNumber="227" phylum="Chordata" rank="family">Phascolarctidae</taxonomicName>
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, and
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<taxonomicName authorityName="Burnett" authorityYear="1830" box="[732,866,1581,1605]" class="Mammalia" family="Vombatidae" kingdom="Animalia" order="Diprotodontia" pageId="227" pageNumber="227" phylum="Chordata" rank="family">Vombatidae</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="228" lastPageNumber="228" pageId="227" pageNumber="227" type="description">
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<paragraph blockId="227.[684,1212,1515,1738]" pageId="227" pageNumber="227">
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<collectionCode box="[716,775,1615,1639]" pageId="227" pageNumber="227">STEM</collectionCode>
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AGE: 39.8 Mya (95%
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<collectionCode box="[1032,1091,1615,1638]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13184" lsid="urn:lsid:biocol.org:col:13184" name="Hampstead Scientific Society" pageId="227" pageNumber="227" type="Herbarium">HPD</collectionCode>
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: 35.5–45.2 Mya).
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</paragraph>
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<paragraph blockId="227.[684,1212,1515,1738]" pageId="227" pageNumber="227">
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<collectionCode box="[716,800,1681,1705]" pageId="227" pageNumber="227">CROWN</collectionCode>
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AGE: 32.4 Mya (95%
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<collectionCode box="[1038,1097,1681,1704]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13184" lsid="urn:lsid:biocol.org:col:13184" name="Hampstead Scientific Society" pageId="227" pageNumber="227" type="Herbarium">HPD</collectionCode>
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: 29.1–36.4 Mya).
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</paragraph>
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<paragraph blockId="228.[108,637,226,1738]" pageId="228" pageNumber="228">
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UNAMBIGUOUS CRANIODENTAL
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<collectionCode box="[108,170,259,283]" pageId="228" pageNumber="228">SYNAPOMORPHIES</collectionCode>
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: Lacrimal exposure with one or more distinct tubercles (char. 8: 0→1; ci = 0.118); paroccipital process is a large erect process usually directed ventrally (char. 93: 1→2; ci = 0.100); P1 absent (char. 114: 0→1; ci = 0.200); P2 absent (char. 116: 0→1; ci = 0.333); distinct posterolingual cusp on semi- or fully sectorial P3 present (char. 125: 0→1; ci = 0.250); second lower premolar absent (char. 154: 0→1; ci = 1.000); entostylid labial to the entoconid present and cusplike (char. 174: 0→1; ci = 0.333).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="228" pageNumber="228" type="discussion">
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<paragraph blockId="228.[108,637,226,1738]" lastBlockId="228.[684,1213,225,712]" pageId="228" pageNumber="228">
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COMMENTS:
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<bibRefCitation author="Beck, R. M. D. & M. L. Taglioretti" box="[297,520,622,647]" pageId="228" pageNumber="228" pagination="385 - 417" refId="ref192802" refString="Beck, R. M. D., and M. L. Taglioretti. 2020. A nearly complete juvenile skull of the marsupial Sparassocynus derivatus from the Pliocene of Argentina, the affinities of " sparassocynids, " and the diversification of opossums (Marsupialia; Didelphimorphia; Didelphidae). Journal of Mammalian Evolution 27: 385 - 417." type="journal article" year="2020">Beck et al. (2020</bibRefCitation>
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:
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<tableCitation box="[540,625,622,647]" captionStart="TABLE 1" captionStartId="7.[611,691,226,250]" captionTargetBox="[119,1110,327,1337]" captionTargetPageId="7" captionText="TABLE 1 Higher Classification and Geographic Distribution of Recent Marsupialsa" httpUri="http://table.plazi.org/id/DF393CC3F63C682ED868FF4C1E65FED9" pageId="228" pageNumber="228" tableUuid="DF393CC3F63C682ED868FF4C1E65FED9">table 1</tableCitation>
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) defined Vombatiformes as the most inclusive clade including
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<taxonomicName baseAuthorityName="Shaw" baseAuthorityYear="1800" box="[291,490,688,712]" class="Mammalia" family="Vombatidae" genus="Vombatus" kingdom="Animalia" order="Diprotodontia" pageId="228" pageNumber="228" phylum="Chordata" rank="species">
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<emphasis box="[291,490,688,712]" italics="true" pageId="228" pageNumber="228">Vombatus ursinus</emphasis>
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</taxonomicName>
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and
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<taxonomicName baseAuthorityName="Goldfuss" baseAuthorityYear="1817" class="Mammalia" family="Phascolarctidae" genus="Phascolarctos" kingdom="Animalia" order="Diprotodontia" pageId="228" pageNumber="228" phylum="Chordata" rank="species">
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<emphasis italics="true" pageId="228" pageNumber="228">Phascolarctos cinereus</emphasis>
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</taxonomicName>
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but not
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<taxonomicName box="[398,629,722,746]" class="Mammalia" family="Phalangeridae" genus="Phalanger" kingdom="Animalia" order="Diprotodontia" pageId="228" pageNumber="228" phylum="Chordata" rank="species">
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<emphasis box="[398,629,722,746]" italics="true" pageId="228" pageNumber="228">Phalanger orientalis</emphasis>
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</taxonomicName>
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. The family †
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<taxonomicName authorityName="Gill" authorityYear="1872" box="[251,437,755,779]" class="Mammalia" family="Thylacoleonidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="228" pageNumber="228" phylum="Chordata" rank="family">Thylacoleonidae</taxonomicName>
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has usually been considered to be a member of Vombatiformes (e.g.,
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<bibRefCitation author="Archer, M." box="[174,338,821,845]" pageId="228" pageNumber="228" pagination="633 - 808" refId="ref190081" refString="Archer, M. 1984 c. The Australian marsupial radiation. In M. Archer and G. Clayton (editors), Vertebrate zoogeography and evolution in Australasia: 633 - 808. Perth: Hesperian Press." type="book chapter" year="1984">Archer, 1984c</bibRefCitation>
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;
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<bibRefCitation author="Aplin, K. P. & M. Archer" box="[353,631,821,845]" pageId="228" pageNumber="228" pagination="xv - lxxii" refId="ref189347" refString="Aplin, K. P., and M. Archer. 1987. Recent advances in marsupial systematics with a new syncretic classification. In M. Archer (editor), Possums and opossums: studies in evolution: xv - lxxii. Sydney: Surrey Beatty and Sons." type="book chapter" year="1987">Aplin and Archer, 1987</bibRefCitation>
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;
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<bibRefCitation author="Aplin, K. P." box="[108,249,854,878]" pageId="228" pageNumber="228" pagination="369 - 391" refId="ref189198" refString="Aplin, K. P. 1987. Basicranial anatomy of the early Miocene diprotodontian Wynyardia bassiana (Marsupialia: Wynyardiidae) and its implications for wynyardiid phylogeny and classification. In M. Archer (editor), Possums and opossums: studies in evolution: 369 - 391. Sydney: Surrey Beatty and Sons." type="book chapter" year="1987">Aplin, 1987</bibRefCitation>
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;
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<bibRefCitation author="Marshall, L. G. & J. A. Case & M. O. Woodburne" box="[266,517,854,878]" pageId="228" pageNumber="228" pagination="433 - 505" refId="ref215725" refString="Marshall, L. G., J. A. Case, and M. O. Woodburne. 1990. Phylogenetic relationships of the families of marsupials. Current Mammalogy 2: 433 - 505." type="journal article" year="1990">Marshall et al., 1990</bibRefCitation>
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;
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<bibRefCitation author="Munson, C. J." pageId="228" pageNumber="228" pagination="1 - 99" refId="ref218350" refString="Munson, C. J. 1992. Postcranial descriptions of Ilaria and Ngapakaldia (Vombatiformes, Marsupialia) and the phylogeny of the vombatiforms based on postcranial morphology. University of California Publications in Zoology 125: 1 - 99." type="journal article" year="1992">Munson, 1992</bibRefCitation>
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;
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<bibRefCitation author="Szalay" box="[182,328,887,911]" firstAuthor="Szalay" pageId="228" pageNumber="228" refId="ref229131" refString="Szalay, F. S. 1994. Evolutionary history of the marsupials and an analysis of osteological characters, Cambridge: Cambridge University Press." type="book" year="1994">Szalay, 1994</bibRefCitation>
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;
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<bibRefCitation author="Archer, M. & S. J. Hand" box="[345,631,887,911]" pageId="228" pageNumber="228" pagination="575 - 646" refId="ref190275" refString="Archer, M., and S. J. Hand. 2006. The Australian marsupial radiation. In J. R. Merrick, M. Archer, G. M. Hickey, and M. S. Y. Lee (editors), Evolution and biogeography of Australasian vertebrates: 575 - 646. Sydney: Auscipub Pty Ltd." type="book chapter" year="2006">Archer and Hand, 2006</bibRefCitation>
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;
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<bibRefCitation author="Gillespie, A. K." box="[108,278,920,944]" pageId="228" pageNumber="228" refId="ref205118" refString="Gillespie, A. K. 2007. Diversity and systematics of marsupial lions from the Riversleigh World Heritage Area and the evolution of the Thylacoleonidae. Ph. D. dissertation, School of Biological, Earth, and Environmental Sciences, University of New South Wales, Sydney." type="book" year="2007">Gillespie, 2007</bibRefCitation>
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;
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<bibRefCitation author="Black, K. H. & M. Archer & S. J. Hand" box="[291,502,920,944]" pageId="228" pageNumber="228" pagination="125 - 138" refId="ref194056" refString="Black, K. H., M. Archer, and S. J. Hand. 2012 a. New Tertiary koala (Marsupialia, Phascolarctidae) from Riversleigh, Australia, with a revision of phascolarctid phylogenetics, paleoecology, and paleobiodiversity. Journal of Vertebrate Paleontology 32 (1): 125 - 138." type="journal article" year="2012">Black et al., 2012a</bibRefCitation>
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,
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<bibRefCitation author="Black, K. H. & M. Archer & S. J. Hand & H. Godthelp" box="[516,586,920,944]" pageId="228" pageNumber="228" pagination="983 - 1078" refId="ref194115" refString="Black, K. H., M. Archer, S. J. Hand, and H. Godthelp. 2012 b. The rise of Australian marsupials: a synopsis of biostratigraphic, phylogenetic, palaeoecologic and palaeobiogeographic understanding. In J. A. Talent (editor), Earth and life: global biodiversity, extinction intervals and biogeographic perturbations through time: 983 - 1078. Dordrecht: Springer Verlag." type="book chapter" year="2012">2012b</bibRefCitation>
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; but see
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<bibRefCitation author="Murray, P. F. & R. Wells & M. Plane" box="[149,360,953,978]" pageId="228" pageNumber="228" pagination="433 - 466" refId="ref218994" refString="Murray, P. F., R. Wells, and M. Plane. 1987. The cranium of the Miocene thylacoleonid, Wakaleo vanderleuri: click go the shears - a fresh bite at thylacoleonid systematics. In M. Archer (editor), Possums and opossums: studies in evolution: 433 - 466. Sydney: Surrey Beatty and Sons." type="book chapter" year="1987">Murray et al., 1987</bibRefCitation>
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, for an alternative view), within which it is usually placed closer to
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<taxonomicName authorityName="E. Geoffroy Saint-Hilaire" authorityYear="1803" class="Mammalia" family="Vombatidae" genus="Vombatus" kingdom="Animalia" order="Diprotodontia" pageId="228" pageNumber="228" phylum="Chordata" rank="genus">
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<emphasis italics="true" pageId="228" pageNumber="228">Vombatus</emphasis>
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</taxonomicName>
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than to
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<taxonomicName box="[262,410,1019,1043]" class="Mammalia" family="Phascolarctidae" genus="Phascolarctos" kingdom="Animalia" order="Diprotodontia" pageId="228" pageNumber="228" phylum="Chordata" rank="genus">
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||
<emphasis box="[262,410,1019,1043]" italics="true" pageId="228" pageNumber="228">Phascolarctos</emphasis>
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||
</taxonomicName>
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—in other words, as a member of Vombatomorphia sensu
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<bibRefCitation author="Beck, R. M. D. & M. L. Taglioretti" pageId="228" pageNumber="228" pagination="385 - 417" refId="ref192802" refString="Beck, R. M. D., and M. L. Taglioretti. 2020. A nearly complete juvenile skull of the marsupial Sparassocynus derivatus from the Pliocene of Argentina, the affinities of " sparassocynids, " and the diversification of opossums (Marsupialia; Didelphimorphia; Didelphidae). Journal of Mammalian Evolution 27: 385 - 417." type="journal article" year="2020">Beck et al. (2020</bibRefCitation>
|
||
; see below). However, as noted above, †
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||
<taxonomicName authorityName="Gill" authorityYear="1872" box="[120,306,1118,1142]" class="Mammalia" family="Thylacoleonidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="228" pageNumber="228" phylum="Chordata" rank="family">Thylacoleonidae</taxonomicName>
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was placed as the sister to all other vombatiforms (i.e., outside Vombatomorphia) in our undated total-evidence analysis (fig. 32; a relationship also found by
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<bibRefCitation author="Gillespie, A. K. & M. Archer & S. J. Hand" pageId="228" pageNumber="228" pagination="1 - 26" refId="ref205243" refString="Gillespie, A. K., M. Archer, and S. J. Hand. 2016. A tiny new marsupial lion (Marsupialia, Thylacoleonidae) from the early Miocene of Australia. Palaeontologia Electronica 9.2.26 A: 1 - 26." type="book chapter" year="2016">Gillespie et al., 2016</bibRefCitation>
|
||
;
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||
<bibRefCitation author="Beck, R. M. D. & M. L. Taglioretti" box="[241,427,1251,1275]" pageId="228" pageNumber="228" pagination="385 - 417" refId="ref192802" refString="Beck, R. M. D., and M. L. Taglioretti. 2020. A nearly complete juvenile skull of the marsupial Sparassocynus derivatus from the Pliocene of Argentina, the affinities of " sparassocynids, " and the diversification of opossums (Marsupialia; Didelphimorphia; Didelphidae). Journal of Mammalian Evolution 27: 385 - 417." type="journal article" year="2020">Beck et al., 2020</bibRefCitation>
|
||
), and it was recovered in a trichotomy at the base of
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<taxonomicName authorityName="Owen" authorityYear="1866" class="Mammalia" kingdom="Animalia" order="Diprotodontia" pageId="228" pageNumber="228" phylum="Chordata" rank="order">Diprotodontia</taxonomicName>
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together with Vombatiformes and
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<taxonomicName box="[108,256,1350,1374]" pageId="228" pageNumber="234" rank="subOrder" subOrder="Phalangerida">Phalangerida</taxonomicName>
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in our dated total-evidence analysis (
|
||
<figureCitation box="[164,237,1383,1407]" captionStart="FIG" captionStartId="181.[108,150,938,959]" captionTargetId="figure-12@181.[241,1216,224,1485]" captionTargetPageId="181" captionText="FIG. 33. (opposite page and at right) Fifty-per- cent majority rule consensus of post-burn-in trees that results from dated Bayesian analysis (using combined tip-and-node dating and separate Independent Gamma Rates [IGR] clock models for the molecular and morpho- logical partitions) of our total evidence data- set. Black dots at nodes indicate ≥0.95 Bayesian posterior probability (“strong support”); dark gray dots indicate 0.75–0.94 Bayesian poste- rior probability (“moderate support”); light gray dots indicate 0.50–0.74 Bayesian poste- rior probability (“weak support”). Nodes with- out dots were constrained a priori so that their ages could be calibrated (see tables 6, 13). Orange bars represent 95% Highest Posterior Density (HPD) intervals on the ages of nodes. For clarity, 95% HPD intervals are not shown for the ages of fossil terminals." pageId="228" pageNumber="228">fig. 33</figureCitation>
|
||
). Based on these results, we suggest that †
|
||
<taxonomicName authorityName="Gill" authorityYear="1872" box="[178,368,1416,1440]" class="Mammalia" family="Thylacoleonidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="228" pageNumber="228" phylum="Chordata" rank="family">Thylacoleonidae</taxonomicName>
|
||
is best considered as
|
||
<taxonomicName authorityName="Owen" authorityYear="1866" box="[108,275,1449,1473]" class="Mammalia" kingdom="Animalia" order="Diprotodontia" pageId="228" pageNumber="228" phylum="Chordata" rank="order">Diprotodontia</taxonomicName>
|
||
incertae sedis. Thus, in our dated total-evidence analysis, Vombatiformes sensu
|
||
<bibRefCitation author="Beck, R. M. D. & M. L. Taglioretti" box="[177,372,1515,1539]" pageId="228" pageNumber="228" pagination="385 - 417" refId="ref192802" refString="Beck, R. M. D., and M. L. Taglioretti. 2020. A nearly complete juvenile skull of the marsupial Sparassocynus derivatus from the Pliocene of Argentina, the affinities of " sparassocynids, " and the diversification of opossums (Marsupialia; Didelphimorphia; Didelphidae). Journal of Mammalian Evolution 27: 385 - 417." type="journal article" year="2020">Beck et al. (2020)</bibRefCitation>
|
||
is restricted to †Diprotodontoidea, †
|
||
<taxonomicName authorityName="Tedford & Woodburne" authorityYear="1987" box="[269,330,1548,1572]" class="Mammalia" family="Ilariidae" genus="Ilaria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="228" pageNumber="228" phylum="Chordata" rank="genus">
|
||
<emphasis box="[269,330,1548,1572]" italics="true" pageId="228" pageNumber="228">Ilaria</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName authorityName="Owen" authorityYear="1839" box="[343,520,1548,1572]" class="Mammalia" family="Phascolarctidae" kingdom="Animalia" order="Diprotodontia" pageId="228" pageNumber="228" phylum="Chordata" rank="family">Phascolarctidae</taxonomicName>
|
||
,
|
||
<taxonomicName authorityName="Burnett" authorityYear="1830" class="Mammalia" family="Vombatidae" kingdom="Animalia" order="Diprotodontia" pageId="228" pageNumber="228" phylum="Chordata" rank="family">Vombatidae</taxonomicName>
|
||
, †
|
||
<taxonomicName box="[172,291,1582,1605]" class="Mammalia" family="Wynyardiidae" genus="Muramura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="228" pageNumber="228" phylum="Chordata" rank="genus">
|
||
<emphasis box="[172,291,1582,1605]" italics="true" pageId="228" pageNumber="228">Muramura</emphasis>
|
||
</taxonomicName>
|
||
, and †
|
||
<emphasis box="[364,520,1581,1605]" italics="true" pageId="228" pageNumber="228">Namilimadeta</emphasis>
|
||
. Members of the families †
|
||
<taxonomicName box="[304,431,1614,1638]" class="Mammalia" family="Maradidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="228" pageNumber="228" phylum="Chordata" rank="family">Maradidae</taxonomicName>
|
||
, †
|
||
<taxonomicName box="[460,631,1614,1638]" class="Mammalia" family="Palorchestidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="228" pageNumber="228" phylum="Chordata" rank="family">Palorchestidae</taxonomicName>
|
||
, and †
|
||
<taxonomicName box="[174,341,1647,1671]" class="Mammalia" family="Mukupirnidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="228" pageNumber="228" phylum="Chordata" rank="family">Mukupirnidae</taxonomicName>
|
||
have not been included here, but have been recovered within Vombatiformes in recent phylogenetic analyses (
|
||
<bibRefCitation author="Black, K. H. & M. Archer & S. J. Hand" box="[572,819,226,1738]" pageId="228" pageNumber="228" pagination="125 - 138" refId="ref194056" refString="Black, K. H., M. Archer, and S. J. Hand. 2012 a. New Tertiary koala (Marsupialia, Phascolarctidae) from Riversleigh, Australia, with a revision of phascolarctid phylogenetics, paleoecology, and paleobiodiversity. Journal of Vertebrate Paleontology 32 (1): 125 - 138." type="journal article" year="2012">Black et al., 2012a</bibRefCitation>
|
||
;
|
||
<bibRefCitation author="Brewer, P. & M. Archer & S. J. Hand & R. Abel" box="[829,1037,225,250]" pageId="228" pageNumber="228" pagination="1 - 40" refId="ref195250" refString="Brewer, P., M. Archer, S. J. Hand, and R. Abel. 2015. New genus of primitive wombat (Vombatidae, Marsupialia) from Miocene deposits in the Riversleigh World Heritage Area (Queensland, Australia). Palaeontologia Electronica 8.1.9 A: 1 - 40." type="book chapter" year="2015">Brewer et al., 2015</bibRefCitation>
|
||
;
|
||
<bibRefCitation author="Gillespie, A. K. & M. Archer & S. J. Hand" pageId="228" pageNumber="228" pagination="1 - 26" refId="ref205243" refString="Gillespie, A. K., M. Archer, and S. J. Hand. 2016. A tiny new marsupial lion (Marsupialia, Thylacoleonidae) from the early Miocene of Australia. Palaeontologia Electronica 9.2.26 A: 1 - 26." type="book chapter" year="2016">Gillespie et al., 2016</bibRefCitation>
|
||
;
|
||
<bibRefCitation author="Beck, R. M. D. & M. L. Taglioretti" box="[751,938,259,283]" pageId="228" pageNumber="228" pagination="385 - 417" refId="ref192802" refString="Beck, R. M. D., and M. L. Taglioretti. 2020. A nearly complete juvenile skull of the marsupial Sparassocynus derivatus from the Pliocene of Argentina, the affinities of " sparassocynids, " and the diversification of opossums (Marsupialia; Didelphimorphia; Didelphidae). Journal of Mammalian Evolution 27: 385 - 417." type="journal article" year="2020">Beck et al., 2020</bibRefCitation>
|
||
). Among the unambiguous synapomorphies of this restricted Vombatiformes are loss of P1, P2, and p2 (see above); all these teeth are retained in plesiomorphic thylacoleonids (
|
||
<bibRefCitation author="Gillespie, A. K. & M. Archer & S. J. Hand" box="[970,1208,391,415]" pageId="228" pageNumber="228" pagination="1 - 26" refId="ref205243" refString="Gillespie, A. K., M. Archer, and S. J. Hand. 2016. A tiny new marsupial lion (Marsupialia, Thylacoleonidae) from the early Miocene of Australia. Palaeontologia Electronica 9.2.26 A: 1 - 26." type="book chapter" year="2016">Gillespie et al., 2016</bibRefCitation>
|
||
;
|
||
<bibRefCitation author="Beck, R. M. D. & M. L. Taglioretti" box="[684,874,424,448]" pageId="228" pageNumber="228" pagination="385 - 417" refId="ref192802" refString="Beck, R. M. D., and M. L. Taglioretti. 2020. A nearly complete juvenile skull of the marsupial Sparassocynus derivatus from the Pliocene of Argentina, the affinities of " sparassocynids, " and the diversification of opossums (Marsupialia; Didelphimorphia; Didelphidae). Journal of Mammalian Evolution 27: 385 - 417." type="journal article" year="2020">Beck et al., 2020</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph blockId="228.[684,1213,225,712]" pageId="228" pageNumber="228">
|
||
The oldest known definitive vombatiforms are from the late Oligocene of
|
||
<collectingCountry box="[1105,1211,490,514]" name="Australia" pageId="228" pageNumber="228">Australia</collectingCountry>
|
||
(
|
||
<bibRefCitation author="Bassarova, M. & M. Archer" box="[691,910,523,547]" pageId="228" pageNumber="228" pagination="25 - 27" refId="ref192343" refString="Bassarova, M., and M. Archer. 1999. Living and extinct pseudocheirids (Marsupialia, Pseudocheiridae): Phylogenetic relationships and changes in diversity through time. Australian Mammalogy 21: 25 - 27." type="journal article" year="1999">Archer et al., 1999</bibRefCitation>
|
||
;
|
||
<bibRefCitation author="Long, J. A. & M. Archer & T. F. Flannery & S. J. Hand" box="[924,1120,523,548]" pageId="228" pageNumber="228" refId="ref213561" refString="Long, J. A., M. Archer, T. F. Flannery, and S. J. Hand. 2002. Prehistoric mammals of Australia and New Guinea: one hundred million years of evolution, Sydney: UNSW Press." type="book" year="2002">Long et al., 2002</bibRefCitation>
|
||
;
|
||
<bibRefCitation author="Archer, M. & S. J. Hand" pageId="228" pageNumber="228" pagination="575 - 646" refId="ref190275" refString="Archer, M., and S. J. Hand. 2006. The Australian marsupial radiation. In J. R. Merrick, M. Archer, G. M. Hickey, and M. S. Y. Lee (editors), Evolution and biogeography of Australasian vertebrates: 575 - 646. Sydney: Auscipub Pty Ltd." type="book chapter" year="2006">Archer and Hand, 2006</bibRefCitation>
|
||
;
|
||
<bibRefCitation author="Black, K. H. & M. Archer & S. J. Hand & H. Godthelp" box="[889,1111,556,580]" pageId="228" pageNumber="228" pagination="983 - 1078" refId="ref194115" refString="Black, K. H., M. Archer, S. J. Hand, and H. Godthelp. 2012 b. The rise of Australian marsupials: a synopsis of biostratigraphic, phylogenetic, palaeoecologic and palaeobiogeographic understanding. In J. A. Talent (editor), Earth and life: global biodiversity, extinction intervals and biogeographic perturbations through time: 983 - 1078. Dordrecht: Springer Verlag." type="book chapter" year="2012">Black et al., 2012b</bibRefCitation>
|
||
). This is congruent with our estimated divergence dates, which suggest that the initial diversification of vombatiforms took place during the late Eocene or early Oligocene.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |