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<document id="9A5D2A094023D4B541C328602FD87AE3" ID-DOI="10.5281/zenodo.277335" ID-GBIF-Dataset="2ce06877-be59-4249-b285-1bcbb3427746" ID-ISSN="1175-5326" ID-Zenodo-Dep="277335" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460290156032" checkinUser="plazi" docAuthor="Lisi, Oscar" docDate="2011" docId="03D7474BCF56A90C71BAFC12D618E73C" docLanguage="en" docName="zt02834p032.pdf" docOrigin="Zootaxa 2834" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Doryphoribius solidunguis Lisi, 2011, sp. nov." docType="treatment" docVersion="8" lastPageNumber="31" masterDocId="FFEE3F33CF5DA902712DFF99D36DE216" masterDocTitle="Remarks on Doryphoribius flavus (Iharos, 1966), and description of three new species (Tardigrada, Hypsibiidae)" masterLastPageNumber="32" masterPageNumber="17" pageNumber="28" updateTime="1698251089152" updateUser="plazi">
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<mods:title id="972454B9ABD1FB158AF225CC7662EF4B">Remarks on Doryphoribius flavus (Iharos, 1966), and description of three new species (Tardigrada, Hypsibiidae)</mods:title>
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<mods:namePart id="4FE481AFA228B2724A4687867D950A7D">Lisi, Oscar</mods:namePart>
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<mods:date id="AED87DA00EA0DD37D776B9C856C7C7BA">2011</mods:date>
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<treatment id="03D7474BCF56A90C71BAFC12D618E73C" ID-DOI="http://doi.org/10.5281/zenodo.5630128" ID-GBIF-Taxon="119398537" ID-Zenodo-Dep="5630128" LSID="urn:lsid:plazi:treatment:03D7474BCF56A90C71BAFC12D618E73C" httpUri="http://treatment.plazi.org/id/03D7474BCF56A90C71BAFC12D618E73C" lastPageId="14" lastPageNumber="31" pageId="11" pageNumber="28">
<subSubSection id="C364A5D6CF56A90971BAFC12D386E1D1" pageId="11" pageNumber="28" type="nomenclature">
<paragraph id="8BC1F65DCF56A90971BAFC12D141E1B2" blockId="11.[151,556,907,967]" box="[151,556,907,932]" pageId="11" pageNumber="28">
<heading id="D0894131CF56A90971BAFC12D141E1B2" bold="true" box="[151,556,907,932]" fontSize="10" level="2" pageId="11" pageNumber="28" reason="2">
<emphasis id="B90A2A4FCF56A90971BAFC12D141E1B2" bold="true" box="[151,556,907,932]" pageId="11" pageNumber="28">
<taxonomicName id="4C7E8DDECF56A90971BAFC12D2A1E1B2" box="[151,460,907,932]" class="Eutardigrada" family="Hypsibiidae" genus="Doryphoribius" kingdom="Animalia" order="Parachela" pageId="11" pageNumber="28" phylum="Tardigrada" rank="species" species="solidunguis" status="sp. nov.">
<emphasis id="B90A2A4FCF56A90971BAFC12D2A1E1B2" bold="true" box="[151,460,907,932]" italics="true" pageId="11" pageNumber="28">Doryphoribius solidunguis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2399734CF56A90970FEFC15D141E1B2" box="[467,556,908,932]" pageId="11" pageNumber="28" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BC1F65DCF56A90971BAFC36D386E1D1" blockId="11.[151,556,907,967]" box="[151,235,943,967]" pageId="11" pageNumber="28">
(
<figureCitation id="1345EAD8CF56A90971B2FC36D38FE1D1" box="[159,226,943,967]" captionStart="FIGURE 5. D" captionStartId="13.[151,250,1854,1877]" captionTargetBox="[156,1436,207,1781]" captionTargetId="figure@13.[137,1467,193,1801]" captionTargetPageId="13" captionText="FIGURE 5. D. solidunguis sp. nov. A, Cuticular sculpture of the caudal portion of a paratype; B, Bucco-pharyngeal apparatus of the holotype; C, Anterior portion of the buccal tube showing the teeth of the buccal cavity (arrow); D, Claws of the second pair of legs of the holotype; E, Claws of the fourth pair of legs of the holotype; F, Lunule of the inner claw (arrow) of the second pair of legs of the holotype (the lunule of the external claw is not in focus); G, Cuticular bar (arrow) beside the internal claw of the third pair of legs of the holotype. Scale bar = 10 µm." httpUri="https://zenodo.org/record/277340/files/figure.png" pageId="11" pageNumber="28">Fig. 5</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C364A5D6CF56A90971BAFC6ED03EE7F2" pageId="11" pageNumber="28" type="materials_examined">
<paragraph id="8BC1F65DCF56A90971BAFC6ED2F8E641" blockId="11.[151,1436,1015,1977]" pageId="11" pageNumber="28">
<emphasis id="B90A2A4FCF56A90971BAFC6ED2ECE606" bold="true" box="[151,385,1015,1040]" pageId="11" pageNumber="28">Material examined.</emphasis>
<typeStatus id="54C548FFCF56A90970A4FC6ED29CE619" box="[393,497,1015,1039]" pageId="11" pageNumber="28" type="holotype">Holotype</typeStatus>
(slide number 3796), 37
<typeStatus id="54C548FFCF56A909723DFC60D016E607" box="[784,891,1017,1041]" pageId="11" pageNumber="28" type="paratype">paratypes</typeStatus>
and 12 exuviae with eggs (slide numbers 3785- 3796, 4179, 4180) mounted in polyvinyl lactophenol; Bali: Tampaksring, from a moss sample (data available from
<bibRefCitation id="EFEF8BACCF56A90971BAFBA6D2E4E641" author="Pilato" box="[151,393,1087,1111]" pageId="11" pageNumber="28" refString="Pilato, G. &amp; Binda, M. G. (1990) Notizie sui tardigradi muscicoli di Bali (Indonesia). Animalia, 17, 209 - 218." type="journal article" year="1990">Pilato &amp; Binda, 1990</bibRefCitation>
).
</paragraph>
<paragraph id="8BC1F65DCF56A90971EAFBFAD7F6E7D6" blockId="11.[151,1436,1015,1977]" pageId="11" pageNumber="28">
<emphasis id="B90A2A4FCF56A90971EAFBFAD2CDE66A" bold="true" box="[199,416,1123,1148]" pageId="11" pageNumber="28">Specific diagnosis.</emphasis>
[Eye spots present*], very evident reticulate sculpture on dorsal and lateral cuticle forming mesh that was quite irregular in shape and size, delimited by ridges of various width (some very wide); the ridges were often rather protruding, only sometimes forming tubercles at crossings. Anterior sides of the first three pair of legs and posterior side of the fourth pair of legs, swollen; legs smooth. Bucco-pharyngeal apparatus of the
<taxonomicName id="4C7E8DDECF56A9097410FB49D38BE71A" class="Eutardigrada" family="Hypsibiidae" genus="Doryphoribius" kingdom="Animalia" order="Parachela" pageId="11" pageNumber="28" phylum="Tardigrada" rank="genus">
<emphasis id="B90A2A4FCF56A9097410FB49D38BE71A" italics="true" pageId="11" pageNumber="28">Doryphoribius</emphasis>
</taxonomicName>
<typeStatus id="54C548FFCF56A90971C3FB6CD24FE71B" box="[238,290,1269,1293]" pageId="11" pageNumber="28">type</typeStatus>
; peribuccal papulae present, some of which were divided. Two pairs of dorsal teeth in the buccal cavity, difficult to see, along the anterior margins of the stylet sheaths. Two macroplacoids and no microplacoid or septulum in the pharyngeal bulb. Claws of the
<taxonomicName id="4C7E8DDECF56A909738BFAA5D05FE742" box="[678,818,1340,1364]" class="Eutardigrada" family="Hypsibiidae" genus="Isohypsibius" kingdom="Animalia" order="Parachela" pageId="11" pageNumber="28" phylum="Tardigrada" rank="genus">
<emphasis id="B90A2A4FCF56A909738BFAA5D05FE742" box="[678,818,1340,1364]" italics="true" pageId="11" pageNumber="28">Isohypsibius</emphasis>
</taxonomicName>
<typeStatus id="54C548FFCF56A9097217FAA4D000E743" box="[826,877,1341,1365]" pageId="11" pageNumber="28">type</typeStatus>
, large and stout, with very small accessory points on the main branches; external claws with wide basal portion enlarged at the very base; internal claws with smaller basal portion; evident smooth lunules present. A cuticular bar partially divided into dots was present beside the base of the internal claws of the first three pairs of legs. Smooth eggs laid into the exuvium.
</paragraph>
<paragraph id="8BC1F65DCF56A90971E8FA54D03EE7F2" blockId="11.[151,1436,1015,1977]" box="[197,851,1484,1508]" pageId="11" pageNumber="28">[*] = Not visible after mounting in polyvinyl lactophenol.</paragraph>
</subSubSection>
<subSubSection id="C364A5D6CF56A90E71E8FA76D776E458" lastPageId="12" lastPageNumber="29" pageId="11" pageNumber="28" type="description">
<paragraph id="8BC1F65DCF56A90971E8FA76D14EE446" blockId="11.[151,1436,1015,1977]" pageId="11" pageNumber="28">
<emphasis id="B90A2A4FCF56A90971E8FA76D17FE41E" bold="true" box="[197,530,1519,1544]" pageId="11" pageNumber="28">
Description of the
<typeStatus id="54C548FFCF56A9097089FA69D160E41E" box="[420,525,1520,1544]" pageId="11" pageNumber="28" type="holotype">holotype</typeStatus>
.
</emphasis>
Unfortunately the mounting media had caused contraction to a greater of lesser extent in many of the specimens. I have therefore designated as the
<typeStatus id="54C548FFCF56A90972BAF98DD094E43A" box="[919,1017,1556,1580]" pageId="11" pageNumber="28" type="holotype">holotype</typeStatus>
a specimen that best shows the most important characters of the species.
</paragraph>
<paragraph id="8BC1F65DCF56A90971E8F9C4D10AE5AE" blockId="11.[151,1436,1015,1977]" pageId="11" pageNumber="28">
It was not possible to correctly measure body length (probably about 380 µm) but in
<tableCitation id="C6FCC3E6CF56A90975A4F9C5D7B0E462" box="[1161,1245,1628,1652]" captionStart="TABLE 4" captionStartId="12.[151,235,152,173]" captionTargetBox="[151,1436,376,883]" captionTargetPageId="12" captionText="TABLE 4. Measurements of selected morphological structures of the holotype and the smallest and largest paratype of Doryphoribius. solidunguis sp. nov.; “ Styl. supp. insert. ” indicates the stylet supports insertion point on the buccal tube; “ % bo-le ” indicates the percentage ratio with respect to the body length (calculated for the buccal tube length, the placoid row, the claws and the largest mesh)." httpUri="http://table.plazi.org/id/DF01A6D5CF51A90E71BAFF01D218E31F" pageId="11" pageNumber="28" tableUuid="DF01A6D5CF51A90E71BAFF01D218E31F">Table 4</tableCitation>
the correct measurements are provided of a
<typeStatus id="54C548FFCF56A90970FFF918D15BE48F" box="[466,566,1665,1689]" pageId="11" pageNumber="28" type="paratype">paratype</typeStatus>
; eye spots no longer visible; dorsal and lateral cuticle ornamented with ridges, often protruding and wide (especially those of the medial-dorsal cuticle), which formed a reticular design with mesh of various shape and size (
<figureCitation id="1345EAD8CF56A9097325F951D126E4F6" box="[520,587,1736,1760]" captionStart="FIGURE 5. D" captionStartId="13.[151,250,1854,1877]" captionTargetBox="[156,1436,207,1781]" captionTargetId="figure@13.[137,1467,193,1801]" captionTargetPageId="13" captionText="FIGURE 5. D. solidunguis sp. nov. A, Cuticular sculpture of the caudal portion of a paratype; B, Bucco-pharyngeal apparatus of the holotype; C, Anterior portion of the buccal tube showing the teeth of the buccal cavity (arrow); D, Claws of the second pair of legs of the holotype; E, Claws of the fourth pair of legs of the holotype; F, Lunule of the inner claw (arrow) of the second pair of legs of the holotype (the lunule of the external claw is not in focus); G, Cuticular bar (arrow) beside the internal claw of the third pair of legs of the holotype. Scale bar = 10 µm." httpUri="https://zenodo.org/record/277340/files/figure.png" pageId="11" pageNumber="28">Fig. 5</figureCitation>
A in a
<typeStatus id="54C548FFCF56A90973B7F951D192E4F6" box="[666,767,1736,1760]" pageId="11" pageNumber="28" type="paratype">paratype</typeStatus>
); the average size of the mesh increased from the lateral to the dorsal cuticle and from anterior to the posterior portion of the body, reaching the maximum between the third and the fourth pairs of legs (largest mesh about 7.3 µm;
<emphasis id="B90A2A4FCF56A9097230F888D05FE53E" box="[797,818,1809,1832]" italics="true" pageId="11" pageNumber="28">pt</emphasis>
index 16.7) and decreased more posteriorly. On some crossings a tubercle could be seen, on others it was clearly absent, while in other cases the protruding ridges made it difficult to discern whether a tubercle was present or not. Due to the orientation of the specimen in the slide, the swollen portions (anterior for the first three pairs, posterior for the fourth pair of legs) were not visible but were in the
<typeStatus id="54C548FFCF56A90971EDF838D242E5AF" box="[192,303,1953,1977]" pageId="11" pageNumber="28" type="paratype">paratypes</typeStatus>
. Cuticle of the legs smooth.
</paragraph>
<caption id="DF01A6D5CF51A90E71BAFF01D218E31F" ID-Table-UUID="DF01A6D5CF51A90E71BAFF01D218E31F" httpUri="http://table.plazi.org/id/DF01A6D5CF51A90E71BAFF01D218E31F" pageId="12" pageNumber="29" targetBox="[151,1436,376,883]" targetIsTable="true" targetPageId="12">
<paragraph id="8BC1F65DCF51A90E71BAFF01D218E31F" blockId="12.[151,1436,152,356]" pageId="12" pageNumber="29">
<emphasis id="B90A2A4FCF51A90E71BAFF01D268E2BB" bold="true" box="[151,261,152,173]" pageId="12" pageNumber="29">TABLE 4.</emphasis>
Measurements of selected morphological structures of the holotype and the smallest and largest paratype of
<emphasis id="B90A2A4FCF51A90E7472FF00D217E2DB" italics="true" pageId="12" pageNumber="29">
<taxonomicName id="4C7E8DDECF51A90E7472FF00D396E2DB" class="Eutardigrada" family="Hypsibiidae" genus="Doryphoribius" kingdom="Animalia" order="Parachela" pageId="12" pageNumber="29" phylum="Tardigrada" rank="genus">Doryphoribius</taxonomicName>
.
<taxonomicName id="4C7E8DDECF51A90E7028FF2ED217E2DB" box="[261,378,183,205]" class="Eutardigrada" family="Hypsibiidae" genus="Doryphoribius" kingdom="Animalia" order="Parachela" pageId="12" pageNumber="29" phylum="Tardigrada" rank="species" species="solidunguis" status="sp. nov.">solidunguis</taxonomicName>
</emphasis>
<emphasis id="B90A2A4FCF51A90E70ACFF2ED2BFE2DA" bold="true" box="[385,466,183,204]" pageId="12" pageNumber="29">
<taxonomicNameLabel id="A2399734CF51A90E70ACFF2ED2BFE2DA" box="[385,466,183,204]" pageId="12" pageNumber="29" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
; “Styl. supp. insert.” indicates the stylet supports insertion point on the buccal tube; “% bo-le” indicates the percentage ratio with respect to the body length (calculated for the buccal tube length, the placoid row, the claws and the largest mesh).
</paragraph>
</caption>
<paragraph id="8BC1F65DCF51A90E70B9FEBCD2F6E760" blockId="12.[151,1436,152,356]" lastBlockId="12.[151,1437,1158,2010]" pageId="12" pageNumber="29">
<typeStatus id="54C548FFCF51A90E70B9FEBCD29FE32D" box="[404,498,293,315]" pageId="12" pageNumber="29" type="holotype">Holotype</typeStatus>
Slide N. 3796 Smallest
<typeStatus id="54C548FFCF51A90E727CFEBFD0C5E32D" box="[849,936,294,315]" pageId="12" pageNumber="29" type="paratype">paratype</typeStatus>
Slide N. 3796 Largest
<typeStatus id="54C548FFCF51A90E758AFEBFD793E32D" box="[1191,1278,294,315]" pageId="12" pageNumber="29" type="paratype">paratype</typeStatus>
Slide N. 3796 Characters µm pt % bo-le µm pt % bo-le µm pt % bo-le III ex. claws 25.8 59.2???? 21.8? 52.9? 6.0? III int. claws 21.8 50.0??????? Bucco-pharyngeal apparatus of the
<taxonomicName id="4C7E8DDECF51A90E7374FB1FD193E688" box="[601,766,1158,1182]" class="Eutardigrada" family="Hypsibiidae" genus="Doryphoribius" kingdom="Animalia" order="Parachela" pageId="12" pageNumber="29" phylum="Tardigrada" rank="genus">
<emphasis id="B90A2A4FCF51A90E7374FB1FD193E688" box="[601,766,1158,1182]" italics="true" pageId="12" pageNumber="29">Doryphoribius</emphasis>
</taxonomicName>
<typeStatus id="54C548FFCF51A90E722BFB1ED05BE689" box="[774,822,1159,1183]" pageId="12" pageNumber="29">type</typeStatus>
(
<figureCitation id="1345EAD8CF51A90E726AFB1FD0E1E688" box="[839,908,1158,1182]" captionStart="FIGURE 5. D" captionStartId="13.[151,250,1854,1877]" captionTargetBox="[156,1436,207,1781]" captionTargetId="figure@13.[137,1467,193,1801]" captionTargetPageId="13" captionText="FIGURE 5. D. solidunguis sp. nov. A, Cuticular sculpture of the caudal portion of a paratype; B, Bucco-pharyngeal apparatus of the holotype; C, Anterior portion of the buccal tube showing the teeth of the buccal cavity (arrow); D, Claws of the second pair of legs of the holotype; E, Claws of the fourth pair of legs of the holotype; F, Lunule of the inner claw (arrow) of the second pair of legs of the holotype (the lunule of the external claw is not in focus); G, Cuticular bar (arrow) beside the internal claw of the third pair of legs of the holotype. Scale bar = 10 µm." httpUri="https://zenodo.org/record/277340/files/figure.png" pageId="12" pageNumber="29">Fig. 5</figureCitation>
B); peribuccal papulae present, some of which were divided; buccal armature consisted of only two pairs of dorsal teeth (
<figureCitation id="1345EAD8CF51A90E72E3FB33D77DE6D4" box="[974,1040,1194,1218]" captionStart="FIGURE 5. D" captionStartId="13.[151,250,1854,1877]" captionTargetBox="[156,1436,207,1781]" captionTargetId="figure@13.[137,1467,193,1801]" captionTargetPageId="13" captionText="FIGURE 5. D. solidunguis sp. nov. A, Cuticular sculpture of the caudal portion of a paratype; B, Bucco-pharyngeal apparatus of the holotype; C, Anterior portion of the buccal tube showing the teeth of the buccal cavity (arrow); D, Claws of the second pair of legs of the holotype; E, Claws of the fourth pair of legs of the holotype; F, Lunule of the inner claw (arrow) of the second pair of legs of the holotype (the lunule of the external claw is not in focus); G, Cuticular bar (arrow) beside the internal claw of the third pair of legs of the holotype. Scale bar = 10 µm." httpUri="https://zenodo.org/record/277340/files/figure.png" pageId="12" pageNumber="29">Fig. 5</figureCitation>
C, arrow), difficult to see, along the anterior margin of the stylet sheaths. Buccal tube 43.6 µm long and 5.9 µm wide (
<emphasis id="B90A2A4FCF51A90E7541FB49D7EFE6F1" box="[1132,1154,1232,1255]" italics="true" pageId="12" pageNumber="29">pt</emphasis>
= 13.5), stylet supports inserted on the buccal tube at 70.0 % of its length (
<emphasis id="B90A2A4FCF51A90E73F9FB6DD187E71D" box="[724,746,1268,1291]" italics="true" pageId="12" pageNumber="29">pt</emphasis>
= 70.0); ventral lamina 25.4 µm long (
<emphasis id="B90A2A4FCF51A90E758CFB6DD7DAE71D" box="[1185,1207,1268,1291]" italics="true" pageId="12" pageNumber="29">pt</emphasis>
= 58.3). Pharyngeal bulb could not be measured, with apophyses and two macroplacoids; no microplacoid or septulum present; placoid row 15.2 µm long (
<emphasis id="B90A2A4FCF51A90E7042FAA5D2E8E745" box="[367,389,1340,1363]" italics="true" pageId="12" pageNumber="29">pt</emphasis>
= 34.9), first macroplacoid slightly constricted in the middle, 8.5 µm long (
<emphasis id="B90A2A4FCF51A90E75FEFAA5D784E745" box="[1235,1257,1340,1363]" italics="true" pageId="12" pageNumber="29">pt</emphasis>
= 19.5), second one 4.5 µm (
<emphasis id="B90A2A4FCF51A90E700BFAF9D251E761" box="[294,316,1376,1399]" italics="true" pageId="12" pageNumber="29">pt</emphasis>
= 10.3).
</paragraph>
<paragraph id="8BC1F65DCF51A90E71B2FEE1D658E164" pageId="12" pageNumber="29">
<table id="F97E04FDCF5156FD71BAFEE1D6F1E165" box="[151,1436,376,883]" gridcols="8" gridrows="12" pageId="12" pageNumber="29">
<tr id="354EF41FCF5156FD71BAFEE1D6F1E3BA" box="[151,1436,376,428]" gridrow="0" pageId="12" pageNumber="29">
<th id="769F9D63CF5156FD71BAFEE1D230E3BA" box="[151,349,376,428]" gridcol="0" gridrow="0" pageId="12" pageNumber="29">Body length</th>
<th id="769F9D63CF5156FD70B9FEE1D135E3BA" box="[404,600,376,428]" gridcol="1" gridrow="0" pageId="12" pageNumber="29">&gt;325.8 - (contracted)</th>
<th id="769F9D63CF5156FD73ACFEE1D1E1E3BA" box="[641,652,376,428]" gridcol="2" gridrow="0" pageId="12" pageNumber="29">-</th>
<th id="769F9D63CF5156FD73DEFEE1D0D9E3BA" box="[755,948,376,428]" gridcol="3" gridrow="0" pageId="12" pageNumber="29">&gt;193.5 - (contracted)</th>
<th id="769F9D63CF5156FD72CBFEE1D09FE3BA" box="[998,1010,376,428]" gridcol="4" gridrow="0" pageId="12" pageNumber="29">-</th>
<th id="769F9D63CF5156FD7578FEE1D7E3E3BA" box="[1109,1166,376,428]" gridcol="5" gridrow="0" pageId="12" pageNumber="29">361.3</th>
<th id="769F9D63CF5156FD75FDFEE1D791E3BA" box="[1232,1276,376,428]" gridcol="6" gridrow="0" pageId="12" pageNumber="29">-</th>
<th id="769F9D63CF5156FD7407FEE1D6F1E3BA" box="[1322,1436,376,428]" gridcol="7" gridrow="0" pageId="12" pageNumber="29">-</th>
</tr>
<tr id="354EF41FCF5156FD71BAFE59D6F1E3C0" box="[151,1436,448,470]" gridrow="1" pageId="12" pageNumber="29">
<th id="769F9D63CF5156FD71BAFE59D230E3C0" box="[151,349,448,470]" gridcol="0" gridrow="1" pageId="12" pageNumber="29">Buccal tube length</th>
<td id="769F9D63CF5156FD70B9FE59D135E3C0" box="[404,600,448,470]" gridcol="1" gridrow="1" pageId="12" pageNumber="29">43.6 -</td>
<td id="769F9D63CF5156FD73ACFE59D1E1E3C0" box="[641,652,448,470]" gridcol="2" gridrow="1" pageId="12" pageNumber="29">?</td>
<td id="769F9D63CF5156FD73DEFE59D0D9E3C0" box="[755,948,448,470]" gridcol="3" gridrow="1" pageId="12" pageNumber="29">28.7 -</td>
<td id="769F9D63CF5156FD72CBFE59D09FE3C0" box="[998,1010,448,470]" gridcol="4" gridrow="1" pageId="12" pageNumber="29">?</td>
<td id="769F9D63CF5156FD7578FE59D7E3E3C0" box="[1109,1166,448,470]" gridcol="5" gridrow="1" pageId="12" pageNumber="29">41.2</td>
<td id="769F9D63CF5156FD75FDFE59D791E3C0" box="[1232,1276,448,470]" gridcol="6" gridrow="1" pageId="12" pageNumber="29">-</td>
<td id="769F9D63CF5156FD7407FE59D6F1E3C0" box="[1322,1436,448,470]" gridcol="7" gridrow="1" pageId="12" pageNumber="29">11.4</td>
</tr>
<tr id="354EF41FCF5156FD71BAFE70D6F1E3E9" box="[151,1436,489,511]" gridrow="2" pageId="12" pageNumber="29">
<th id="769F9D63CF5156FD71BAFE70D230E3E9" box="[151,349,489,511]" gridcol="0" gridrow="2" pageId="12" pageNumber="29">Buccal tube width</th>
<td id="769F9D63CF5156FD70B9FE70D135E3E9" box="[404,600,489,511]" gridcol="1" gridrow="2" pageId="12" pageNumber="29">5.9 13.5</td>
<td id="769F9D63CF5156FD73ACFE70D1E1E3E9" box="[641,652,489,511]" gridcol="2" gridrow="2" pageId="12" pageNumber="29">-</td>
<td id="769F9D63CF5156FD73DEFE70D0D9E3E9" box="[755,948,489,511]" gridcol="3" gridrow="2" pageId="12" pageNumber="29">3.5 12.2</td>
<td id="769F9D63CF5156FD72CBFE70D09FE3E9" box="[998,1010,489,511]" gridcol="4" gridrow="2" pageId="12" pageNumber="29">-</td>
<td id="769F9D63CF5156FD7578FE70D7E3E3E9" box="[1109,1166,489,511]" gridcol="5" gridrow="2" pageId="12" pageNumber="29">5.9</td>
<td id="769F9D63CF5156FD75FDFE70D791E3E9" box="[1232,1276,489,511]" gridcol="6" gridrow="2" pageId="12" pageNumber="29">14.3</td>
<td id="769F9D63CF5156FD7407FE70D6F1E3E9" box="[1322,1436,489,511]" gridcol="7" gridrow="2" pageId="12" pageNumber="29">-</td>
</tr>
<tr id="354EF41FCF5156FD71BAFD8BD6F1E03E" box="[151,1436,530,552]" gridrow="3" pageId="12" pageNumber="29">
<th id="769F9D63CF5156FD71BAFD8BD230E03E" box="[151,349,530,552]" gridcol="0" gridrow="3" pageId="12" pageNumber="29">Styl. supp. insert.</th>
<td id="769F9D63CF5156FD70B9FD8BD135E03E" box="[404,600,530,552]" gridcol="1" gridrow="3" pageId="12" pageNumber="29">- 70.0</td>
<td id="769F9D63CF5156FD73ACFD8BD1E1E03E" box="[641,652,530,552]" gridcol="2" gridrow="3" pageId="12" pageNumber="29">-</td>
<td id="769F9D63CF5156FD73DEFD8BD0D9E03E" box="[755,948,530,552]" gridcol="3" gridrow="3" pageId="12" pageNumber="29">- 72.3</td>
<td id="769F9D63CF5156FD72CBFD8BD09FE03E" box="[998,1010,530,552]" gridcol="4" gridrow="3" pageId="12" pageNumber="29">-</td>
<td id="769F9D63CF5156FD7578FD8BD7E3E03E" box="[1109,1166,530,552]" gridcol="5" gridrow="3" pageId="12" pageNumber="29">-</td>
<td id="769F9D63CF5156FD75FDFD8BD791E03E" box="[1232,1276,530,552]" gridcol="6" gridrow="3" pageId="12" pageNumber="29">71.8</td>
<td id="769F9D63CF5156FD7407FD8BD6F1E03E" box="[1322,1436,530,552]" gridcol="7" gridrow="3" pageId="12" pageNumber="29">-</td>
</tr>
<tr id="354EF41FCF5156FD71BAFDA2D6F1E044" box="[151,1436,571,594]" gridrow="4" pageId="12" pageNumber="29">
<th id="769F9D63CF5156FD71BAFDA2D230E044" box="[151,349,571,594]" gridcol="0" gridrow="4" pageId="12" pageNumber="29">Ventral lamina</th>
<td id="769F9D63CF5156FD70B9FDA2D135E044" box="[404,600,571,594]" gridcol="1" gridrow="4" pageId="12" pageNumber="29">25.4 58.3</td>
<td id="769F9D63CF5156FD73ACFDA2D1E1E044" box="[641,652,571,594]" gridcol="2" gridrow="4" pageId="12" pageNumber="29">-</td>
<td id="769F9D63CF5156FD73DEFDA2D0D9E044" box="[755,948,571,594]" gridcol="3" gridrow="4" pageId="12" pageNumber="29">18.1 63.1</td>
<td id="769F9D63CF5156FD72CBFDA2D09FE044" box="[998,1010,571,594]" gridcol="4" gridrow="4" pageId="12" pageNumber="29">-</td>
<td id="769F9D63CF5156FD7578FDA2D7E3E044" box="[1109,1166,571,594]" gridcol="5" gridrow="4" pageId="12" pageNumber="29">25.4</td>
<td id="769F9D63CF5156FD75FDFDA2D791E044" box="[1232,1276,571,594]" gridcol="6" gridrow="4" pageId="12" pageNumber="29">61.6</td>
<td id="769F9D63CF5156FD7407FDA2D6F1E044" box="[1322,1436,571,594]" gridcol="7" gridrow="4" pageId="12" pageNumber="29">-</td>
</tr>
<tr id="354EF41FCF5156FD71BAFDFCD6F1E06D" box="[151,1436,613,635]" gridrow="5" pageId="12" pageNumber="29">
<th id="769F9D63CF5156FD71BAFDFCD230E06D" box="[151,349,613,635]" gridcol="0" gridrow="5" pageId="12" pageNumber="29">Placoid row</th>
<td id="769F9D63CF5156FD70B9FDFCD135E06D" box="[404,600,613,635]" gridcol="1" gridrow="5" pageId="12" pageNumber="29">15.2 34.9</td>
<td id="769F9D63CF5156FD73ACFDFCD1E1E06D" box="[641,652,613,635]" gridcol="2" gridrow="5" pageId="12" pageNumber="29">?</td>
<td id="769F9D63CF5156FD73DEFDFCD0D9E06D" box="[755,948,613,635]" gridcol="3" gridrow="5" pageId="12" pageNumber="29">7.8 27.2</td>
<td id="769F9D63CF5156FD72CBFDFCD09FE06D" box="[998,1010,613,635]" gridcol="4" gridrow="5" pageId="12" pageNumber="29">?</td>
<td id="769F9D63CF5156FD7578FDFCD7E3E06D" box="[1109,1166,613,635]" gridcol="5" gridrow="5" pageId="12" pageNumber="29">14.0</td>
<td id="769F9D63CF5156FD75FDFDFCD791E06D" box="[1232,1276,613,635]" gridcol="6" gridrow="5" pageId="12" pageNumber="29">34.0</td>
<td id="769F9D63CF5156FD7407FDFCD6F1E06D" box="[1322,1436,613,635]" gridcol="7" gridrow="5" pageId="12" pageNumber="29">3.9</td>
</tr>
<tr id="354EF41FCF5156FD71BAFD17D6F1E0B2" box="[151,1436,654,676]" gridrow="6" pageId="12" pageNumber="29">
<th id="769F9D63CF5156FD71BAFD17D230E0B2" box="[151,349,654,676]" gridcol="0" gridrow="6" pageId="12" pageNumber="29">I macroplacoid</th>
<td id="769F9D63CF5156FD70B9FD17D135E0B2" box="[404,600,654,676]" gridcol="1" gridrow="6" pageId="12" pageNumber="29">8.5 19.5</td>
<td id="769F9D63CF5156FD73ACFD17D1E1E0B2" box="[641,652,654,676]" gridcol="2" gridrow="6" pageId="12" pageNumber="29">-</td>
<td id="769F9D63CF5156FD73DEFD17D0D9E0B2" box="[755,948,654,676]" gridcol="3" gridrow="6" pageId="12" pageNumber="29">4.7 16.4</td>
<td id="769F9D63CF5156FD72CBFD17D09FE0B2" box="[998,1010,654,676]" gridcol="4" gridrow="6" pageId="12" pageNumber="29">-</td>
<td id="769F9D63CF5156FD7578FD17D7E3E0B2" box="[1109,1166,654,676]" gridcol="5" gridrow="6" pageId="12" pageNumber="29">8.2</td>
<td id="769F9D63CF5156FD75FDFD17D791E0B2" box="[1232,1276,654,676]" gridcol="6" gridrow="6" pageId="12" pageNumber="29">19.9</td>
<td id="769F9D63CF5156FD7407FD17D6F1E0B2" box="[1322,1436,654,676]" gridcol="7" gridrow="6" pageId="12" pageNumber="29">-</td>
</tr>
<tr id="354EF41FCF5156FD71BAFD21D6F1E0D8" box="[151,1436,696,718]" gridrow="7" pageId="12" pageNumber="29">
<th id="769F9D63CF5156FD71BAFD21D230E0D8" box="[151,349,696,718]" gridcol="0" gridrow="7" pageId="12" pageNumber="29">II macroplacoid</th>
<td id="769F9D63CF5156FD70B9FD21D135E0D8" box="[404,600,696,718]" gridcol="1" gridrow="7" pageId="12" pageNumber="29">4.5 10.3</td>
<td id="769F9D63CF5156FD73ACFD21D1E1E0D8" box="[641,652,696,718]" gridcol="2" gridrow="7" pageId="12" pageNumber="29">-</td>
<td id="769F9D63CF5156FD73DEFD21D0D9E0D8" box="[755,948,696,718]" gridcol="3" gridrow="7" pageId="12" pageNumber="29">2.8 9.8</td>
<td id="769F9D63CF5156FD72CBFD21D09FE0D8" box="[998,1010,696,718]" gridcol="4" gridrow="7" pageId="12" pageNumber="29">-</td>
<td id="769F9D63CF5156FD7578FD21D7E3E0D8" box="[1109,1166,696,718]" gridcol="5" gridrow="7" pageId="12" pageNumber="29">5.0</td>
<td id="769F9D63CF5156FD75FDFD21D791E0D8" box="[1232,1276,696,718]" gridcol="6" gridrow="7" pageId="12" pageNumber="29">12.1</td>
<td id="769F9D63CF5156FD7407FD21D6F1E0D8" box="[1322,1436,696,718]" gridcol="7" gridrow="7" pageId="12" pageNumber="29">-</td>
</tr>
<tr id="354EF41FCF5156FD71BAFD78D6F1E0E1" box="[151,1436,737,759]" gridrow="8" pageId="12" pageNumber="29">
<th id="769F9D63CF5156FD71BAFD78D230E0E1" box="[151,349,737,759]" gridcol="0" gridrow="8" pageId="12" pageNumber="29">I ex. claws</th>
<td id="769F9D63CF5156FD70B9FD78D135E0E1" box="[404,600,737,759]" gridcol="1" gridrow="8" pageId="12" pageNumber="29">24.0 55.0</td>
<td id="769F9D63CF5156FD73ACFD78D1E1E0E1" box="[641,652,737,759]" gridcol="2" gridrow="8" pageId="12" pageNumber="29">?</td>
<td id="769F9D63CF5156FD73DEFD78D0D9E0E1" box="[755,948,737,759]" gridcol="3" gridrow="8" pageId="12" pageNumber="29">??</td>
<td id="769F9D63CF5156FD72CBFD78D09FE0E1" box="[998,1010,737,759]" gridcol="4" gridrow="8" pageId="12" pageNumber="29">?</td>
<td id="769F9D63CF5156FD7578FD78D7E3E0E1" box="[1109,1166,737,759]" gridcol="5" gridrow="8" pageId="12" pageNumber="29">?</td>
<td id="769F9D63CF5156FD75FDFD78D791E0E1" box="[1232,1276,737,759]" gridcol="6" gridrow="8" pageId="12" pageNumber="29">?</td>
<td id="769F9D63CF5156FD7407FD78D6F1E0E1" box="[1322,1436,737,759]" gridcol="7" gridrow="8" pageId="12" pageNumber="29">?</td>
</tr>
<tr id="354EF41FCF5156FD71BAFC93D6F1E136" box="[151,1436,778,800]" gridrow="9" pageId="12" pageNumber="29">
<th id="769F9D63CF5156FD71BAFC93D230E136" box="[151,349,778,800]" gridcol="0" gridrow="9" pageId="12" pageNumber="29">I int. claws</th>
<td id="769F9D63CF5156FD70B9FC93D135E136" box="[404,600,778,800]" gridcol="1" gridrow="9" pageId="12" pageNumber="29">??</td>
<td id="769F9D63CF5156FD73ACFC93D1E1E136" box="[641,652,778,800]" gridcol="2" gridrow="9" pageId="12" pageNumber="29">?</td>
<td id="769F9D63CF5156FD73DEFC93D0D9E136" box="[755,948,778,800]" gridcol="3" gridrow="9" pageId="12" pageNumber="29">10.9 38.0</td>
<td id="769F9D63CF5156FD72CBFC93D09FE136" box="[998,1010,778,800]" gridcol="4" gridrow="9" pageId="12" pageNumber="29">?</td>
<td id="769F9D63CF5156FD7578FC93D7E3E136" box="[1109,1166,778,800]" gridcol="5" gridrow="9" pageId="12" pageNumber="29">?</td>
<td id="769F9D63CF5156FD75FDFC93D791E136" box="[1232,1276,778,800]" gridcol="6" gridrow="9" pageId="12" pageNumber="29">?</td>
<td id="769F9D63CF5156FD7407FC93D6F1E136" box="[1322,1436,778,800]" gridcol="7" gridrow="9" pageId="12" pageNumber="29">?</td>
</tr>
<tr id="354EF41FCF5156FD71BAFCAAD6F1E15C" box="[151,1436,819,842]" gridrow="10" pageId="12" pageNumber="29">
<th id="769F9D63CF5156FD71BAFCAAD230E15C" box="[151,349,819,842]" gridcol="0" gridrow="10" pageId="12" pageNumber="29">II ex. claws</th>
<td id="769F9D63CF5156FD70B9FCAAD135E15C" box="[404,600,819,842]" gridcol="1" gridrow="10" pageId="12" pageNumber="29">??</td>
<td id="769F9D63CF5156FD73ACFCAAD1E1E15C" box="[641,652,819,842]" gridcol="2" gridrow="10" pageId="12" pageNumber="29">?</td>
<td id="769F9D63CF5156FD73DEFCAAD0D9E15C" box="[755,948,819,842]" gridcol="3" gridrow="10" pageId="12" pageNumber="29">??</td>
<td id="769F9D63CF5156FD72CBFCAAD09FE15C" box="[998,1010,819,842]" gridcol="4" gridrow="10" pageId="12" pageNumber="29">?</td>
<td id="769F9D63CF5156FD7578FCAAD7E3E15C" box="[1109,1166,819,842]" gridcol="5" gridrow="10" pageId="12" pageNumber="29">?</td>
<td id="769F9D63CF5156FD75FDFCAAD791E15C" box="[1232,1276,819,842]" gridcol="6" gridrow="10" pageId="12" pageNumber="29">?</td>
<td id="769F9D63CF5156FD7407FCAAD6F1E15C" box="[1322,1436,819,842]" gridcol="7" gridrow="10" pageId="12" pageNumber="29">?</td>
</tr>
<tr id="354EF41FCF5156FD71BAFCC4D6F1E165" box="[151,1436,861,883]" gridrow="11" pageId="12" pageNumber="29">
<th id="769F9D63CF5156FD71BAFCC4D230E165" box="[151,349,861,883]" gridcol="0" gridrow="11" pageId="12" pageNumber="29">II int. claws</th>
<td id="769F9D63CF5156FD70B9FCC4D135E165" box="[404,600,861,883]" gridcol="1" gridrow="11" pageId="12" pageNumber="29">??</td>
<td id="769F9D63CF5156FD73ACFCC4D1E1E165" box="[641,652,861,883]" gridcol="2" gridrow="11" pageId="12" pageNumber="29">?</td>
<td id="769F9D63CF5156FD73DEFCC4D0D9E165" box="[755,948,861,883]" gridcol="3" gridrow="11" pageId="12" pageNumber="29">??</td>
<td id="769F9D63CF5156FD72CBFCC4D09FE165" box="[998,1010,861,883]" gridcol="4" gridrow="11" pageId="12" pageNumber="29">?</td>
<td id="769F9D63CF5156FD7578FCC4D7E3E165" box="[1109,1166,861,883]" gridcol="5" gridrow="11" pageId="12" pageNumber="29">?</td>
<td id="769F9D63CF5156FD75FDFCC4D791E165" box="[1232,1276,861,883]" gridcol="6" gridrow="11" pageId="12" pageNumber="29">?</td>
<td id="769F9D63CF5156FD7407FCC4D6F1E165" box="[1322,1436,861,883]" gridcol="7" gridrow="11" pageId="12" pageNumber="29">?</td>
</tr>
</table>
</paragraph>
<paragraph id="8BC1F65DCF51A90E71B2FC43D624E657" pageId="12" pageNumber="29">
<table id="F97E04FDCF5156FD71BAFC40D6F1E654" box="[151,1436,985,1090]" gridcols="7" gridrows="3" pageId="12" pageNumber="29">
<tr id="354EF41FCF5156FD71BAFC40D6F1E1F9" box="[151,1436,985,1007]" gridrow="0" pageId="12" pageNumber="29">
<th id="769F9D63CF5156FD71BAFC40D25CE1F9" box="[151,305,985,1007]" gridcol="0" gridrow="0" pageId="12" pageNumber="29">IV ant. claws</th>
<th id="769F9D63CF5156FD70B9FC40D2ACE1F9" box="[404,449,985,1007]" gridcol="1" gridrow="0" pageId="12" pageNumber="29">22.9</th>
<th id="769F9D63CF5156FD7306FC40D1E1E1F9" box="[555,652,985,1007]" gridcol="2" gridrow="0" pageId="12" pageNumber="29">52.6?</th>
<th id="769F9D63CF5156FD73DEFC40D072E1F9" box="[755,799,985,1007]" gridcol="3" gridrow="0" pageId="12" pageNumber="29">13.0</th>
<th id="769F9D63CF5156FD72AAFC40D09CE1F9" box="[903,1009,985,1007]" gridcol="4" gridrow="0" pageId="12" pageNumber="29">45. 3?</th>
<th id="769F9D63CF5156FD7578FC40D719E1F9" box="[1109,1140,985,1007]" gridcol="5" gridrow="0" pageId="12" pageNumber="29">?</th>
<th id="769F9D63CF5156FD75FDFC40D6F1E1F9" box="[1232,1436,985,1007]" gridcol="6" gridrow="0" pageId="12" pageNumber="29">??</th>
</tr>
<tr id="354EF41FCF5156FD71BAFB9BD6F1E60E" box="[151,1436,1026,1048]" gridrow="1" pageId="12" pageNumber="29">
<th id="769F9D63CF5156FD71BAFB9BD25CE60E" box="[151,305,1026,1048]" gridcol="0" gridrow="1" pageId="12" pageNumber="29">IV post. claws</th>
<td id="769F9D63CF5156FD70B9FB9BD2ACE60E" box="[404,449,1026,1048]" gridcol="1" gridrow="1" pageId="12" pageNumber="29">25.6</td>
<td id="769F9D63CF5156FD7306FB9BD1E1E60E" box="[555,652,1026,1048]" gridcol="2" gridrow="1" pageId="12" pageNumber="29">58.7?</td>
<td id="769F9D63CF5156FD73DEFB9BD072E60E" box="[755,799,1026,1048]" gridcol="3" gridrow="1" pageId="12" pageNumber="29">?</td>
<td id="769F9D63CF5156FD72AAFB9BD09CE60E" box="[903,1009,1026,1048]" gridcol="4" gridrow="1" pageId="12" pageNumber="29">??</td>
<td id="769F9D63CF5156FD7578FB9BD719E60E" box="[1109,1140,1026,1048]" gridcol="5" gridrow="1" pageId="12" pageNumber="29">?</td>
<td id="769F9D63CF5156FD75FDFB9BD6F1E60E" box="[1232,1436,1026,1048]" gridcol="6" gridrow="1" pageId="12" pageNumber="29">??</td>
</tr>
<tr id="354EF41FCF5156FD71BAFBB2D6F1E654" box="[151,1436,1067,1090]" gridrow="2" pageId="12" pageNumber="29">
<th id="769F9D63CF5156FD71BAFBB2D25CE654" box="[151,305,1067,1090]" gridcol="0" gridrow="2" pageId="12" pageNumber="29">Largest mesh</th>
<td id="769F9D63CF5156FD70B9FBB2D2ACE654" box="[404,449,1067,1090]" gridcol="1" gridrow="2" pageId="12" pageNumber="29">7.3</td>
<td id="769F9D63CF5156FD7306FBB2D1E1E654" box="[555,652,1067,1090]" gridcol="2" gridrow="2" pageId="12" pageNumber="29">16.7?</td>
<td id="769F9D63CF5156FD73DEFBB2D072E654" box="[755,799,1067,1090]" gridcol="3" gridrow="2" pageId="12" pageNumber="29">4.0</td>
<td id="769F9D63CF5156FD72AAFBB2D09CE654" box="[903,1009,1067,1090]" gridcol="4" gridrow="2" pageId="12" pageNumber="29">13.9?</td>
<td id="769F9D63CF5156FD7578FBB2D719E654" box="[1109,1140,1067,1090]" gridcol="5" gridrow="2" pageId="12" pageNumber="29">5.9</td>
<td id="769F9D63CF5156FD75FDFBB2D6F1E654" box="[1232,1436,1067,1090]" gridcol="6" gridrow="2" pageId="12" pageNumber="29">14.4 1.6</td>
</tr>
</table>
</paragraph>
<paragraph id="8BC1F65DCF51A90E71E8FA1BD776E458" blockId="12.[151,1437,1158,2010]" pageId="12" pageNumber="29">
Claws of the
<taxonomicName id="4C7E8DDECF51A90E7073FA1AD287E78D" box="[350,490,1411,1435]" class="Eutardigrada" family="Hypsibiidae" genus="Isohypsibius" kingdom="Animalia" order="Parachela" pageId="12" pageNumber="29" phylum="Tardigrada" rank="genus">
<emphasis id="B90A2A4FCF51A90E7073FA1AD287E78D" box="[350,490,1411,1435]" italics="true" pageId="12" pageNumber="29">Isohypsibius</emphasis>
</taxonomicName>
<typeStatus id="54C548FFCF51A90E70DEFA1AD14EE78D" box="[499,547,1411,1435]" pageId="12" pageNumber="29">type</typeStatus>
(
<figureCitation id="1345EAD8CF51A90E7318FA1BD1E0E78C" box="[565,653,1410,1434]" captionStart="FIGURE 5. D" captionStartId="13.[151,250,1854,1877]" captionTargetBox="[156,1436,207,1781]" captionTargetId="figure@13.[137,1467,193,1801]" captionTargetPageId="13" captionText="FIGURE 5. D. solidunguis sp. nov. A, Cuticular sculpture of the caudal portion of a paratype; B, Bucco-pharyngeal apparatus of the holotype; C, Anterior portion of the buccal tube showing the teeth of the buccal cavity (arrow); D, Claws of the second pair of legs of the holotype; E, Claws of the fourth pair of legs of the holotype; F, Lunule of the inner claw (arrow) of the second pair of legs of the holotype (the lunule of the external claw is not in focus); G, Cuticular bar (arrow) beside the internal claw of the third pair of legs of the holotype. Scale bar = 10 µm." httpUri="https://zenodo.org/record/277340/files/figure.png" pageId="12" pageNumber="29">Fig. 5D</figureCitation>
G), slightly different in shape and size on each leg, large and stout, with poorly developed accessory points on the main branches; external claws with wide basal portion enlarged at the very base, internal with smaller basal portion; smooth lunules present, well developed under both the internal and external claws of the first three pairs of legs (
<figureCitation id="1345EAD8CF51A90E73FFFA77D078E410" box="[722,789,1518,1542]" captionStart="FIGURE 5. D" captionStartId="13.[151,250,1854,1877]" captionTargetBox="[156,1436,207,1781]" captionTargetId="figure@13.[137,1467,193,1801]" captionTargetPageId="13" captionText="FIGURE 5. D. solidunguis sp. nov. A, Cuticular sculpture of the caudal portion of a paratype; B, Bucco-pharyngeal apparatus of the holotype; C, Anterior portion of the buccal tube showing the teeth of the buccal cavity (arrow); D, Claws of the second pair of legs of the holotype; E, Claws of the fourth pair of legs of the holotype; F, Lunule of the inner claw (arrow) of the second pair of legs of the holotype (the lunule of the external claw is not in focus); G, Cuticular bar (arrow) beside the internal claw of the third pair of legs of the holotype. Scale bar = 10 µm." httpUri="https://zenodo.org/record/277340/files/figure.png" pageId="12" pageNumber="29">Fig. 5</figureCitation>
F, arrow). A cuticular bar partially divided into dots was present beside the base of the internal claws of the first three pairs of legs (
<figureCitation id="1345EAD8CF51A90E72C1F98BD743E43C" box="[1004,1070,1554,1578]" captionStart="FIGURE 5. D" captionStartId="13.[151,250,1854,1877]" captionTargetBox="[156,1436,207,1781]" captionTargetId="figure@13.[137,1467,193,1801]" captionTargetPageId="13" captionText="FIGURE 5. D. solidunguis sp. nov. A, Cuticular sculpture of the caudal portion of a paratype; B, Bucco-pharyngeal apparatus of the holotype; C, Anterior portion of the buccal tube showing the teeth of the buccal cavity (arrow); D, Claws of the second pair of legs of the holotype; E, Claws of the fourth pair of legs of the holotype; F, Lunule of the inner claw (arrow) of the second pair of legs of the holotype (the lunule of the external claw is not in focus); G, Cuticular bar (arrow) beside the internal claw of the third pair of legs of the holotype. Scale bar = 10 µm." httpUri="https://zenodo.org/record/277340/files/figure.png" pageId="12" pageNumber="29">Fig. 5</figureCitation>
G, arrow). The measurements of the claws and the other metric characters of the
<typeStatus id="54C548FFCF51A90E7383F9AFD07DE458" box="[686,784,1590,1614]" pageId="12" pageNumber="29" type="holotype">holotype</typeStatus>
are reported in
<tableCitation id="C6FCC3E6CF51A90E72EFF9AFD77AE458" box="[962,1047,1590,1614]" captionStart="TABLE 4" captionStartId="12.[151,235,152,173]" captionTargetBox="[151,1436,376,883]" captionTargetPageId="12" captionText="TABLE 4. Measurements of selected morphological structures of the holotype and the smallest and largest paratype of Doryphoribius. solidunguis sp. nov.; “ Styl. supp. insert. ” indicates the stylet supports insertion point on the buccal tube; “ % bo-le ” indicates the percentage ratio with respect to the body length (calculated for the buccal tube length, the placoid row, the claws and the largest mesh)." httpUri="http://table.plazi.org/id/DF01A6D5CF51A90E71BAFF01D218E31F" pageId="12" pageNumber="29" tableUuid="DF01A6D5CF51A90E71BAFF01D218E31F">Table 4</tableCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C364A5D6CF51A90E71E8F9C2D7ACE480" pageId="12" pageNumber="29" type="discussion">
<paragraph id="8BC1F65DCF51A90E71E8F9C2D7ACE480" blockId="12.[151,1437,1158,2010]" pageId="12" pageNumber="29">
<emphasis id="B90A2A4FCF51A90E71E8F9C2D254E465" bold="true" box="[197,313,1627,1651]" pageId="12" pageNumber="29">Remarks.</emphasis>
The
<typeStatus id="54C548FFCF51A90E7058F9C2D28CE465" box="[373,481,1627,1651]" pageId="12" pageNumber="29" type="paratype">paratypes</typeStatus>
showed the same, both qualitative and metric, characters of the
<typeStatus id="54C548FFCF51A90E7592F9C3D648E464" box="[1215,1317,1626,1650]" pageId="12" pageNumber="29" type="holotype">holotype</typeStatus>
. The measurements of some structures are reported in
<tableCitation id="C6FCC3E6CF51A90E73A3F9E7D189E480" box="[654,740,1662,1686]" captionStart="TABLE 4" captionStartId="12.[151,235,152,173]" captionTargetBox="[151,1436,376,883]" captionTargetPageId="12" captionText="TABLE 4. Measurements of selected morphological structures of the holotype and the smallest and largest paratype of Doryphoribius. solidunguis sp. nov.; “ Styl. supp. insert. ” indicates the stylet supports insertion point on the buccal tube; “ % bo-le ” indicates the percentage ratio with respect to the body length (calculated for the buccal tube length, the placoid row, the claws and the largest mesh)." httpUri="http://table.plazi.org/id/DF01A6D5CF51A90E71BAFF01D218E31F" pageId="12" pageNumber="29" tableUuid="DF01A6D5CF51A90E71BAFF01D218E31F">Table 4</tableCitation>
. Smooth eggs (47) laid into the exuvium.
</paragraph>
</subSubSection>
<subSubSection id="C364A5D6CF51A90E71E8F93AD657E4AC" box="[197,1338,1698,1723]" pageId="12" pageNumber="29" type="etymology">
<paragraph id="8BC1F65DCF51A90E71E8F93AD657E4AC" blockId="12.[151,1437,1158,2010]" box="[197,1338,1698,1723]" pageId="12" pageNumber="29">
<emphasis id="B90A2A4FCF51A90E71E8F93AD226E4AD" bold="true" box="[197,331,1699,1723]" pageId="12" pageNumber="29">Etymology.</emphasis>
The name of the species refers to the stout claws: “
<emphasis id="B90A2A4FCF51A90E72A0F93AD0B1E4AD" box="[909,988,1699,1723]" italics="true" pageId="12" pageNumber="29">solidus</emphasis>
” = “stout”, “
<emphasis id="B90A2A4FCF51A90E755DF93DD7D6E4AD" box="[1136,1211,1700,1723]" italics="true" pageId="12" pageNumber="29">unguis</emphasis>
” = “claw”.
</paragraph>
</subSubSection>
<subSubSection id="C364A5D6CF51A90C71E8F95ED618E73C" lastPageId="14" lastPageNumber="31" pageId="12" pageNumber="29" type="materials_examined">
<paragraph id="8BC1F65DCF51A90E71E8F95ED0C1E514" blockId="12.[151,1437,1158,2010]" pageId="12" pageNumber="29">
<emphasis id="B90A2A4FCF51A90E71E8F95ED2F3E4C9" bold="true" box="[197,414,1734,1759]" pageId="12" pageNumber="29">
<typeStatus id="54C548FFCF51A90E71E8F95ED26DE4C9" box="[197,256,1735,1759]" pageId="12" pageNumber="29">Type</typeStatus>
depositories.
</emphasis>
The
<typeStatus id="54C548FFCF51A90E70F7F95ED167E4C9" box="[474,522,1735,1759]" pageId="12" pageNumber="29">type</typeStatus>
material is deposited in the Binda and Pilato collection (Museum of the Department of Animal Biology “Marcello La Greca”, University of Catania).
</paragraph>
<paragraph id="8BC1F65DCF51A90E71E8F897D24AE5CC" blockId="12.[151,1437,1158,2010]" pageId="12" pageNumber="29">
<emphasis id="B90A2A4FCF51A90E71E8F897D2A1E531" bold="true" box="[197,460,1806,1831]" pageId="12" pageNumber="29">Differential diagnosis.</emphasis>
<taxonomicName id="4C7E8DDECF51A90E70F9F897D06CE530" box="[468,769,1806,1830]" class="Eutardigrada" family="Hypsibiidae" genus="Doryphoribius" kingdom="Animalia" order="Parachela" pageId="12" pageNumber="29" phylum="Tardigrada" rank="species" species="solidunguis" status="sp. nov.">
<emphasis id="B90A2A4FCF51A90E70F9F897D06CE530" box="[468,769,1806,1830]" italics="true" pageId="12" pageNumber="29">Doryphoribius solidunguis</emphasis>
</taxonomicName>
<emphasis id="B90A2A4FCF51A90E7224F896D00EE531" bold="true" box="[777,867,1807,1831]" pageId="12" pageNumber="29">
<taxonomicNameLabel id="A2399734CF51A90E7224F896D00EE531" box="[777,867,1807,1831]" pageId="12" pageNumber="29" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
differed from
<taxonomicName id="4C7E8DDECF51A90E7524F889D702E530" box="[1033,1135,1806,1831]" class="Eutardigrada" family="Hypsibiidae" genus="Doryphoribius" kingdom="Animalia" order="Parachela" pageId="12" pageNumber="29" phylum="Tardigrada" rank="species" species="flavus">
<emphasis id="B90A2A4FCF51A90E7524F889D702E530" box="[1033,1135,1806,1831]" italics="true" pageId="12" pageNumber="29">D. flavus</emphasis>
</taxonomicName>
by having cuticular sculpture with smaller mesh (
<tableCitation id="C6FCC3E6CF51A90E7089F8ABD297E55C" box="[420,506,1842,1866]" captionStart="TABLE 5" captionStartId="14.[151,235,980,1001]" captionText="TABLE 5. Measurements of selected morphological structures of a correctly measured specimen of Doryphoribius flavus (for-" pageId="12" pageNumber="29">Table 5</tableCitation>
); the reticular design appeared more irregular due to the more variable width of the ridges, which were also more protruding and formed tubercles at some crossings. The legs had no reticular ornamentation. The new species had only two pairs of dorso-lateral teeth in the buccal cavity; stouter claws (
<figureCitation id="1345EAD8CF51A90E7403F8E3D61AE584" box="[1326,1399,1914,1938]" captionStart="FIGURE 1. D" captionStartId="2.[151,250,1923,1946]" captionTargetBox="[152,1436,225,1922]" captionTargetId="figure@2.[151,1436,193,1936]" captionTargetPageId="2" captionText="FIGURE 1. D. flavus. A, Dorso-lateral teeth (arrows) of the buccal cavity of a paratype; B, Bucco-pharyngeal apparatus of another paratype; C, Claws of the second and third pairs of legs of the same paratype as Fig. 1 B; D, claws of the fourth pair of legs of the same paratype as Fig. 1 B C. Scale bar = 10 µm." httpUri="https://zenodo.org/record/277336/files/figure.png" pageId="12" pageNumber="29">Figs. 1</figureCitation>
C D, 2E and 5DG); more evident lunules of the inner claws (
<figureCitation id="1345EAD8CF51A90E7211F807D0E4E5A0" box="[828,905,1950,1974]" captionStart="FIGURE 2. D" captionStartId="4.[151,250,1888,1911]" captionTargetBox="[256,1330,187,1853]" captionTargetId="figure@4.[235,1350,171,1874]" captionTargetPageId="4" captionText="FIGURE 2. D. citrinus. A, Cuticular sculpture of the caudal portion of a paratype; B, Anterior portion of the body of another paratype; C, Bucco-pharyngeal apparatus of a third paratype in dorsal view: the dorso-lateral teeth of the buccal cavity are visible (arrows); D, Bucco-pharyngeal apparatus in ventral view of the same paratype as Fig. 2 C; E, Claws of the second and third pairs of legs of a fourth paratype; F, detail of E showing the very narrow lunule (arrow) at the base of an inner claw of the third pair of legs (the lunule of the external claw is not in focus). Scale bar = 10 µm." httpUri="https://zenodo.org/record/277337/files/figure.png" pageId="12" pageNumber="29">Figs. 2</figureCitation>
F and 5F); and a cuticular bar on the first three pairs of legs.
</paragraph>
<caption id="DF01A6D5CF50A90F71BAF8A7D1BFE5D9" httpUri="https://zenodo.org/record/277340/files/figure.png" pageId="13" pageNumber="30" targetBox="[156,1436,207,1781]" targetPageId="13">
<paragraph id="8BC1F65DCF50A90F71BAF8A7D1BFE5D9" blockId="13.[151,1437,1854,1999]" pageId="13" pageNumber="30">
<emphasis id="B90A2A4FCF50A90F71BAF8A7D27EE542" bold="true" box="[151,275,1854,1877]" pageId="13" pageNumber="30">FIGURE 5.</emphasis>
<taxonomicName id="4C7E8DDECF50A90F7034F8D9D2C0E543" box="[281,429,1855,1877]" class="Eutardigrada" family="Hypsibiidae" genus="Doryphoribius" kingdom="Animalia" order="Parachela" pageId="13" pageNumber="30" phylum="Tardigrada" rank="species" species="solidunguis" status="sp. nov.">
<emphasis id="B90A2A4FCF50A90F7034F8D9D2C0E543" box="[281,429,1855,1877]" italics="true" pageId="13" pageNumber="30">D. solidunguis</emphasis>
</taxonomicName>
<emphasis id="B90A2A4FCF50A90F7099F8A6D169E542" bold="true" box="[436,516,1855,1876]" pageId="13" pageNumber="30">
<taxonomicNameLabel id="A2399734CF50A90F7099F8A6D169E542" box="[436,516,1855,1876]" pageId="13" pageNumber="30" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
A, Cuticular sculpture of the caudal portion of a paratype; B, Bucco-pharyngeal apparatus of the holotype; C, Anterior portion of the buccal tube showing the teeth of the buccal cavity (arrow); D, Claws of the second pair of legs of the holotype; E, Claws of the fourth pair of legs of the holotype; F, Lunule of the inner claw (arrow) of the second pair of legs of the holotype (the lunule of the external claw is not in focus); G, Cuticular bar (arrow) beside the internal claw of the third pair of legs of the holotype. Scale bar = 10 µm.
</paragraph>
</caption>
<paragraph id="8BC1F65DCF53A90C71E8FF00D0C2E30A" blockId="14.[151,1437,151,932]" pageId="14" pageNumber="31">
<taxonomicName id="4C7E8DDECF53A90C71E8FF00D204E2A6" box="[197,361,152,176]" class="Eutardigrada" family="Hypsibiidae" genus="Doryphoribius" kingdom="Animalia" order="Parachela" pageId="14" pageNumber="31" phylum="Tardigrada" rank="species" species="solidunguis" status="sp. nov.">
<emphasis id="B90A2A4FCF53A90C71E8FF00D204E2A6" box="[197,361,152,176]" italics="true" pageId="14" pageNumber="31">D. solidunguis</emphasis>
</taxonomicName>
<emphasis id="B90A2A4FCF53A90C705CFF01D2A6E2A6" bold="true" box="[369,459,152,176]" pageId="14" pageNumber="31">
<taxonomicNameLabel id="A2399734CF53A90C705CFF01D2A6E2A6" box="[369,459,152,176]" pageId="14" pageNumber="31" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
differed from
<taxonomicName id="4C7E8DDECF53A90C735DFF00D065E2A6" box="[624,776,152,176]" class="Eutardigrada" family="Hypsibiidae" genus="Doryphoribius" kingdom="Animalia" order="Parachela" pageId="14" pageNumber="31" phylum="Tardigrada" rank="species" species="polynettae">
<emphasis id="B90A2A4FCF53A90C735DFF00D065E2A6" box="[624,776,152,176]" italics="true" pageId="14" pageNumber="31">D. polynettae</emphasis>
</taxonomicName>
above all in lacking cuticular sculpture on the ventral surface; by having a far more obvious dorsal cuticular sculpture with some tubercles present at the crossing of the mesh; pores nor roughness on the cuticle, absent; swollen portions of the legs; only two pairs of teeth in the oral cavity; wider buccal tube, stouter claws; and cuticular bars on the legs.
</paragraph>
<paragraph id="8BC1F65DCF53A90C71E8FEB1D14BE00E" blockId="14.[151,1437,151,932]" pageId="14" pageNumber="31">
<taxonomicName id="4C7E8DDECF53A90C71E8FEB1D299E356" box="[197,500,296,320]" class="Eutardigrada" family="Hypsibiidae" genus="Doryphoribius" kingdom="Animalia" order="Parachela" pageId="14" pageNumber="31" phylum="Tardigrada" rank="species" species="solidunguis" status="sp. nov.">
<emphasis id="B90A2A4FCF53A90C71E8FEB1D299E356" box="[197,500,296,320]" italics="true" pageId="14" pageNumber="31">Doryphoribius solidunguis</emphasis>
</taxonomicName>
<emphasis id="B90A2A4FCF53A90C70D0FEB1D135E356" bold="true" box="[509,600,296,320]" pageId="14" pageNumber="31">
<taxonomicNameLabel id="A2399734CF53A90C70D0FEB1D135E356" box="[509,600,296,320]" pageId="14" pageNumber="31" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
differed from
<taxonomicName id="4C7E8DDECF53A90C722FFEB0D0D8E356" box="[770,949,297,320]" class="Eutardigrada" family="Hypsibiidae" genus="Doryphoribius" kingdom="Animalia" order="Parachela" pageId="14" pageNumber="31" phylum="Tardigrada" rank="species" species="amazzonicus" status="sp. nov.">
<emphasis id="B90A2A4FCF53A90C722FFEB0D0D8E356" box="[770,949,297,320]" italics="true" pageId="14" pageNumber="31">D. amazzonicus</emphasis>
</taxonomicName>
<emphasis id="B90A2A4FCF53A90C7293FEB1D777E356" bold="true" box="[958,1050,296,320]" pageId="14" pageNumber="31">
<taxonomicNameLabel id="A2399734CF53A90C7293FEB1D777E356" box="[958,1050,296,320]" pageId="14" pageNumber="31" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
by having the reticulate cuticular sculpture formed by thicker, more protruding ridges forming tubercles only at some crossing of the mesh. The largest mesh had higher
<emphasis id="B90A2A4FCF53A90C70A9FEE8D2F7E39E" box="[388,410,369,392]" italics="true" pageId="14" pageNumber="31">pt</emphasis>
index values (
<tableCitation id="C6FCC3E6CF53A90C736FFEF6D1F6E391" box="[578,667,367,391]" captionStart="TABLE 5" captionStartId="14.[151,235,980,1001]" captionText="TABLE 5. Measurements of selected morphological structures of a correctly measured specimen of Doryphoribius flavus (for-" pageId="14" pageNumber="31">Table 5</tableCitation>
).
<taxonomicName id="4C7E8DDECF53A90C739FFEE8D03AE39E" box="[690,855,368,392]" class="Eutardigrada" family="Hypsibiidae" genus="Doryphoribius" kingdom="Animalia" order="Parachela" pageId="14" pageNumber="31" phylum="Tardigrada" rank="species" species="solidunguis" status="sp. nov.">
<emphasis id="B90A2A4FCF53A90C739FFEE8D03AE39E" box="[690,855,368,392]" italics="true" pageId="14" pageNumber="31">D. solidunguis</emphasis>
</taxonomicName>
<emphasis id="B90A2A4FCF53A90C724DFEE9D0D1E39E" bold="true" box="[864,956,368,392]" pageId="14" pageNumber="31">
<taxonomicNameLabel id="A2399734CF53A90C724DFEE9D0D1E39E" box="[864,956,368,392]" pageId="14" pageNumber="31" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
also had slightly shorter buccal tube with respect to the body length (
<tableCitation id="C6FCC3E6CF53A90C70E5FE0DD172E3BA" box="[456,543,404,428]" captionStart="TABLE 5" captionStartId="14.[151,235,980,1001]" captionText="TABLE 5. Measurements of selected morphological structures of a correctly measured specimen of Doryphoribius flavus (for-" pageId="14" pageNumber="31">Table 5</tableCitation>
); two pairs of teeth in the buccal cavity (not visible in
<taxonomicName id="4C7E8DDECF53A90C75ABFE0CD65AE3BA" box="[1158,1335,405,428]" class="Eutardigrada" family="Hypsibiidae" genus="Doryphoribius" kingdom="Animalia" order="Parachela" pageId="14" pageNumber="31" phylum="Tardigrada" rank="species" species="amazzonicus">
<emphasis id="B90A2A4FCF53A90C75ABFE0CD65AE3BA" box="[1158,1335,405,428]" italics="true" pageId="14" pageNumber="31">D. amazzonicus</emphasis>
</taxonomicName>
); stouter claws which were also longer with respect to the buccal tube length (compare
<tableCitation id="C6FCC3E6CF53A90C7539FE2ED705E3D9" box="[1044,1128,439,463]" captionStart="TABLE 2" captionStartId="6.[151,235,1124,1145]" captionText="TABLE 2. Measurements of selected morphological structures of the two known specimens of Doryphoribius amazzonicus sp." pageId="14" pageNumber="31">Table 2</tableCitation>
with
<tableCitation id="C6FCC3E6CF53A90C7586FE2ED66FE3D9" box="[1195,1282,439,463]" captionStart="TABLE 4" captionStartId="12.[151,235,152,173]" captionTargetBox="[151,1436,376,883]" captionTargetPageId="12" captionText="TABLE 4. Measurements of selected morphological structures of the holotype and the smallest and largest paratype of Doryphoribius. solidunguis sp. nov.; “ Styl. supp. insert. ” indicates the stylet supports insertion point on the buccal tube; “ % bo-le ” indicates the percentage ratio with respect to the body length (calculated for the buccal tube length, the placoid row, the claws and the largest mesh)." httpUri="http://table.plazi.org/id/DF01A6D5CF51A90E71BAFF01D218E31F" pageId="14" pageNumber="31" tableUuid="DF01A6D5CF51A90E71BAFF01D218E31F">Table 4</tableCitation>
). It had more evident lunules of the inner claws and a cuticular bar at the base of the inner claws on the first three pairs of legs, which is absent in
<emphasis id="B90A2A4FCF53A90C704AFD98D14BE00E" box="[359,550,513,536]" italics="true" pageId="14" pageNumber="31">
<taxonomicName id="4C7E8DDECF53A90C704AFD98D175E00E" box="[359,536,513,536]" class="Eutardigrada" family="Hypsibiidae" genus="Doryphoribius" kingdom="Animalia" order="Parachela" pageId="14" pageNumber="31" phylum="Tardigrada" rank="species" species="amazzonicus">D. amazzonicus</taxonomicName>
.
</emphasis>
</paragraph>
<paragraph id="8BC1F65DCF53A90C71E8FDBCD0D6E0DA" blockId="14.[151,1437,151,932]" pageId="14" pageNumber="31">
<taxonomicName id="4C7E8DDECF53A90C71E8FDBCD204E02A" box="[197,361,548,572]" class="Eutardigrada" family="Hypsibiidae" genus="Doryphoribius" kingdom="Animalia" order="Parachela" pageId="14" pageNumber="31" phylum="Tardigrada" rank="species" species="solidunguis" status="sp. nov.">
<emphasis id="B90A2A4FCF53A90C71E8FDBCD204E02A" box="[197,361,548,572]" italics="true" pageId="14" pageNumber="31">D. solidunguis</emphasis>
</taxonomicName>
<emphasis id="B90A2A4FCF53A90C705FFDBDD2A0E02A" bold="true" box="[370,461,548,572]" pageId="14" pageNumber="31">
<taxonomicNameLabel id="A2399734CF53A90C705FFDBDD2A0E02A" box="[370,461,548,572]" pageId="14" pageNumber="31" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
differed from
<taxonomicName id="4C7E8DDECF53A90C7359FDBCD18EE02A" box="[628,739,548,572]" class="Eutardigrada" family="Hypsibiidae" genus="Doryphoribius" kingdom="Animalia" order="Parachela" pageId="14" pageNumber="31" phylum="Tardigrada" rank="species" species="bindae" status="sp. nov.">
<emphasis id="B90A2A4FCF53A90C7359FDBCD18EE02A" box="[628,739,548,572]" italics="true" pageId="14" pageNumber="31">D. bindae</emphasis>
</taxonomicName>
<emphasis id="B90A2A4FCF53A90C73C1FDBDD02BE02A" bold="true" box="[748,838,548,572]" pageId="14" pageNumber="31">
<taxonomicNameLabel id="A2399734CF53A90C73C1FDBDD02BE02A" box="[748,838,548,572]" pageId="14" pageNumber="31" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
by having a more evident reticulate cuticular design with larger mesh (
<tableCitation id="C6FCC3E6CF53A90C7072FDDED2D9E049" box="[351,436,583,607]" captionStart="TABLE 5" captionStartId="14.[151,235,980,1001]" captionText="TABLE 5. Measurements of selected morphological structures of a correctly measured specimen of Doryphoribius flavus (for-" pageId="14" pageNumber="31">Table 5</tableCitation>
) formed by far wider and more protruding ridges; tubercles present only at some crossing of the ridges; legs unsculptured; two pairs of teeth in the buccal cavity instead of only one; longer and stouter claws (
<tableCitation id="C6FCC3E6CF53A90C71B3FD16D399E0B1" box="[158,244,655,679]" captionStart="TABLE 5" captionStartId="14.[151,235,980,1001]" captionText="TABLE 5. Measurements of selected morphological structures of a correctly measured specimen of Doryphoribius flavus (for-" pageId="14" pageNumber="31">Table 5</tableCitation>
, and compare
<tableCitation id="C6FCC3E6CF53A90C70BAFD16D284E0B1" box="[407,489,655,679]" captionStart="TABLE 3" captionStartId="9.[151,235,692,713]" captionTargetBox="[151,1436,866,1591]" captionTargetPageId="9" captionText="TABLE 3. Measurements of selected morphological structures of the holotype and the smallest and largest paratype of Doryphoribius bindae sp. nov.; “ Styl. supp. insert. ” indicates the stylet supports insertion point on the buccal tube; “ % bo-le ” indicates the percentage ratio with respect to the body length (calculated for the buccal tube length, the placoid row, the claws and the largest mesh)." httpUri="http://table.plazi.org/id/DF01A6D5CF54A90B71BAFD2DD227E130" pageId="14" pageNumber="31" tableUuid="DF01A6D5CF54A90B71BAFD2DD227E130">Table 3</tableCitation>
with
<tableCitation id="C6FCC3E6CF53A90C7304FD16D110E0B1" box="[553,637,655,679]" captionStart="TABLE 4" captionStartId="12.[151,235,152,173]" captionTargetBox="[151,1436,376,883]" captionTargetPageId="12" captionText="TABLE 4. Measurements of selected morphological structures of the holotype and the smallest and largest paratype of Doryphoribius. solidunguis sp. nov.; “ Styl. supp. insert. ” indicates the stylet supports insertion point on the buccal tube; “ % bo-le ” indicates the percentage ratio with respect to the body length (calculated for the buccal tube length, the placoid row, the claws and the largest mesh)." httpUri="http://table.plazi.org/id/DF01A6D5CF51A90E71BAFF01D218E31F" pageId="14" pageNumber="31" tableUuid="DF01A6D5CF51A90E71BAFF01D218E31F">Table 4</tableCitation>
,
<figureCitation id="1345EAD8CF53A90C73A4FD16D184E0B1" box="[649,745,655,679]" captionStart="FIGURE 4. D" captionStartId="10.[151,250,1882,1905]" captionTargetBox="[176,1406,214,1838]" captionTargetId="figure@10.[151,1436,193,1861]" captionTargetPageId="10" captionText="FIGURE 4. D. bindae sp. nov. A, Cuticular sculpture of the caudal portion of a paratype; B, Caudal portion of the holotype: some tubercles are in focus (arrows); C, Leg of the second pair of a paratype, also sculptured on the swollen portion (arrow); D, Bucco-pharyngeal apparatus and first pair of legs of the holotype; E, Dorsal teeth of the buccal cavity (arrows) of a paratype; F, Claws of the fourth pair of legs of the holotype. Scale bar = 10 µm." httpUri="https://zenodo.org/record/277339/files/figure.png" pageId="14" pageNumber="31">Figs. 4D</figureCitation>
F and 5DG); more evident lunules of the inner claws, and a cuticular bar on the first three pairs of legs, absent in
<taxonomicName id="4C7E8DDECF53A90C73C1FD2CD037E0DA" box="[748,858,692,716]" class="Eutardigrada" family="Hypsibiidae" genus="Doryphoribius" kingdom="Animalia" order="Parachela" pageId="14" pageNumber="31" phylum="Tardigrada" rank="species" species="bindae" status="sp. nov.">
<emphasis id="B90A2A4FCF53A90C73C1FD2CD037E0DA" box="[748,858,692,716]" italics="true" pageId="14" pageNumber="31">D. bindae</emphasis>
</taxonomicName>
<emphasis id="B90A2A4FCF53A90C724CFD2DD0D6E0DA" bold="true" box="[865,955,692,716]" pageId="14" pageNumber="31">
<taxonomicNameLabel id="A2399734CF53A90C724CFD2DD0D6E0DA" box="[865,955,692,716]" pageId="14" pageNumber="31" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="8BC1F65DCF53A90C71E8FD40D79BE121" blockId="14.[151,1437,151,932]" pageId="14" pageNumber="31">
<taxonomicName id="4C7E8DDECF53A90C71E8FD40D205E0E6" box="[197,360,728,752]" class="Eutardigrada" family="Hypsibiidae" genus="Doryphoribius" kingdom="Animalia" order="Parachela" pageId="14" pageNumber="31" phylum="Tardigrada" rank="species" species="solidunguis" status="sp. nov.">
<emphasis id="B90A2A4FCF53A90C71E8FD40D205E0E6" box="[197,360,728,752]" italics="true" pageId="14" pageNumber="31">D. solidunguis</emphasis>
</taxonomicName>
<emphasis id="B90A2A4FCF53A90C7043FD41D2AAE0E6" bold="true" box="[366,455,728,752]" pageId="14" pageNumber="31">
<taxonomicNameLabel id="A2399734CF53A90C7043FD41D2AAE0E6" box="[366,455,728,752]" pageId="14" pageNumber="31" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
differed from
<taxonomicName id="4C7E8DDECF53A90C7344FD40D00CE0E6" box="[617,865,728,752]" class="Eutardigrada" family="Hypsibiidae" genus="Doryphoribius" kingdom="Animalia" order="Parachela" pageId="14" pageNumber="31" phylum="Tardigrada" rank="species" species="quadrituberculatus">
<emphasis id="B90A2A4FCF53A90C7344FD40D00CE0E6" box="[617,865,728,752]" italics="true" pageId="14" pageNumber="31">D. quadrituberculatus</emphasis>
</taxonomicName>
above all by lacking dorsal gibbosities; mesh comprising ridges with tubercles at some of the crossings; swollen portions of the legs; peribuccal papulae; only two pair of dorsal teeth in the oral cavity; lunules at the base of the claws; and cuticular bars on the legs.
</paragraph>
<paragraph id="8BC1F65DCF53A90C71E8FCDCD029E1B2" blockId="14.[151,1437,151,932]" pageId="14" pageNumber="31">
<taxonomicName id="4C7E8DDECF53A90C71E8FCDCD205E14A" box="[197,360,836,860]" class="Eutardigrada" family="Hypsibiidae" genus="Doryphoribius" kingdom="Animalia" order="Parachela" pageId="14" pageNumber="31" phylum="Tardigrada" rank="species" species="solidunguis" status="sp. nov.">
<emphasis id="B90A2A4FCF53A90C71E8FCDCD205E14A" box="[197,360,836,860]" italics="true" pageId="14" pageNumber="31">D. solidunguis</emphasis>
</taxonomicName>
<emphasis id="B90A2A4FCF53A90C7042FCDDD2A5E14A" bold="true" box="[367,456,836,860]" pageId="14" pageNumber="31">
<taxonomicNameLabel id="A2399734CF53A90C7042FCDDD2A5E14A" box="[367,456,836,860]" pageId="14" pageNumber="31" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
differed from
<taxonomicName id="4C7E8DDECF53A90C7346FCDCD06BE14A" box="[619,774,836,860]" class="Eutardigrada" family="Hypsibiidae" genus="Doryphoribius" kingdom="Animalia" order="Parachela" pageId="14" pageNumber="31" phylum="Tardigrada" rank="species" species="smokiensis">
<emphasis id="B90A2A4FCF53A90C7346FCDCD06BE14A" box="[619,774,836,860]" italics="true" pageId="14" pageNumber="31">D. smokiensis</emphasis>
</taxonomicName>
by having eyes; different
<typeStatus id="54C548FFCF53A90C7507FCDCD737E14B" box="[1066,1114,837,861]" pageId="14" pageNumber="31">type</typeStatus>
of cuticular sculpture; swollen portions of the legs; only two pairs of dorsal teeth in the oral cavity; far narrower buccal tube; lunules at the claw base; cuticular bars on the legs; and other minor details.
</paragraph>
<caption id="DF01A6D5CF53A90C71BAFC4DD6F1E1FF" box="[151,1436,979,1002]" pageId="14" pageNumber="31">
<paragraph id="8BC1F65DCF53A90C71BAFC4DD6F1E1FF" blockId="14.[151,1437,979,1322]" box="[151,1436,979,1002]" pageId="14" pageNumber="31">
<emphasis id="B90A2A4FCF53A90C71BAFC4DD269E1FF" bold="true" box="[151,260,980,1001]" pageId="14" pageNumber="31">TABLE 5.</emphasis>
Measurements of selected morphological structures of a correctly measured specimen of
<taxonomicName id="4C7E8DDECF53A90C75BDFC4DD60AE1FC" box="[1168,1383,980,1002]" class="Eutardigrada" family="Hypsibiidae" genus="Doryphoribius" kingdom="Animalia" order="Parachela" pageId="14" pageNumber="31" phylum="Tardigrada" rank="species" species="flavus">
<emphasis id="B90A2A4FCF53A90C75BDFC4DD60AE1FC" box="[1168,1383,980,1002]" italics="true" pageId="14" pageNumber="31">Doryphoribius flavus</emphasis>
</taxonomicName>
(for-
</paragraph>
</caption>
<paragraph id="8BC1F65DCF53A90C71BAFC6AD6F1E61E" blockId="14.[151,1437,979,1322]" box="[151,1436,1010,1032]" pageId="14" pageNumber="31">
mer
<typeStatus id="54C548FFCF53A90C71E8FC6AD271E61E" box="[197,284,1011,1032]" pageId="14" pageNumber="31" type="paratype">paratype</typeStatus>
of
<taxonomicName id="4C7E8DDECF53A90C706DFC6BD144E61E" box="[320,553,1010,1032]" class="Eutardigrada" family="Hypsibiidae" genus="Doryphoribius" kingdom="Animalia" order="Parachela" pageId="14" pageNumber="31" phylum="Tardigrada" rank="species" species="citrinus">
<emphasis id="B90A2A4FCF53A90C706DFC6BD144E61E" box="[320,553,1010,1032]" italics="true" pageId="14" pageNumber="31">Doryphoribius citrinus</emphasis>
</taxonomicName>
; the values in brackets are from another
<typeStatus id="54C548FFCF53A90C72E1FC6AD74EE61E" box="[972,1059,1011,1032]" pageId="14" pageNumber="31" type="paratype">paratype</typeStatus>
of the same body size), the
<typeStatus id="54C548FFCF53A90C7468FC6AD6F1E61E" box="[1349,1436,1011,1032]" pageId="14" pageNumber="31" type="paratype">paratype</typeStatus>
</paragraph>
<paragraph id="8BC1F65DCF53A90C71BAFB88D6F0E631" blockId="14.[151,1437,979,1322]" box="[151,1437,1041,1063]" pageId="14" pageNumber="31">
of
<taxonomicName id="4C7E8DDECF53A90C7199FB88D2BFE631" box="[180,466,1041,1063]" class="Eutardigrada" family="Hypsibiidae" genus="Doryphoribius" kingdom="Animalia" order="Parachela" pageId="14" pageNumber="31" phylum="Tardigrada" rank="species" species="amazzonicus" status="sp. nov.">
<emphasis id="B90A2A4FCF53A90C7199FB88D2BFE631" box="[180,466,1041,1063]" italics="true" pageId="14" pageNumber="31">Doryphoribius amazzonicus</emphasis>
</taxonomicName>
<emphasis id="B90A2A4FCF53A90C70F7FB8BD141E631" bold="true" box="[474,556,1042,1063]" pageId="14" pageNumber="31">
<taxonomicNameLabel id="A2399734CF53A90C70F7FB8BD141E631" box="[474,556,1042,1063]" pageId="14" pageNumber="31" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, the
<typeStatus id="54C548FFCF53A90C734FFB88D1D6E631" box="[610,699,1041,1063]" pageId="14" pageNumber="31" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C7E8DDECF53A90C73F2FB8BD028E631" box="[735,837,1041,1063]" class="Eutardigrada" family="Hypsibiidae" genus="Doryphoribius" kingdom="Animalia" order="Parachela" pageId="14" pageNumber="31" phylum="Tardigrada" rank="species" species="bindae" status="sp. nov.">
<emphasis id="B90A2A4FCF53A90C73F2FB8BD028E631" box="[735,837,1041,1063]" italics="true" pageId="14" pageNumber="31">D. bindae</emphasis>
</taxonomicName>
<emphasis id="B90A2A4FCF53A90C7260FB8BD0F2E631" bold="true" box="[845,927,1042,1063]" pageId="14" pageNumber="31">
<taxonomicNameLabel id="A2399734CF53A90C7260FB8BD0F2E631" box="[845,927,1042,1063]" pageId="14" pageNumber="31" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and a
<typeStatus id="54C548FFCF53A90C72CAFB8BD753E631" box="[999,1086,1042,1063]" pageId="14" pageNumber="31" type="paratype">paratype</typeStatus>
of
<taxonomicName id="4C7E8DDECF53A90C754EFB8BD794E631" box="[1123,1273,1041,1063]" class="Eutardigrada" family="Hypsibiidae" genus="Doryphoribius" kingdom="Animalia" order="Parachela" pageId="14" pageNumber="31" phylum="Tardigrada" rank="species" species="solidunguis" status="sp. nov.">
<emphasis id="B90A2A4FCF53A90C754EFB8BD794E631" box="[1123,1273,1041,1063]" italics="true" pageId="14" pageNumber="31">D. solidunguis</emphasis>
</taxonomicName>
<emphasis id="B90A2A4FCF53A90C742DFB8BD63FE631" bold="true" box="[1280,1362,1042,1063]" pageId="14" pageNumber="31">
<taxonomicNameLabel id="A2399734CF53A90C742DFB8BD63FE631" box="[1280,1362,1042,1063]" pageId="14" pageNumber="31" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
; “Styl.
</paragraph>
<paragraph id="8BC1F65DCF53A90C71BAFBA8D6F1E650" blockId="14.[151,1437,979,1322]" box="[151,1436,1072,1094]" pageId="14" pageNumber="31">supp. insert.” indicates the stylet supports insertion point on the buccal tube; “% bo-le” indicates the percentage ratio with</paragraph>
<paragraph id="8BC1F65DCF53A90C71BAFBD6D679E672" blockId="14.[151,1437,979,1322]" box="[151,1300,1102,1124]" pageId="14" pageNumber="31">respect to the body length (calculated for the buccal tube length, the placoid row, the claws and the largest mesh).</paragraph>
<paragraph id="8BC1F65DCF53A90C705BFB19D647E6A4" blockId="14.[151,1437,979,1322]" pageId="14" pageNumber="31">
<emphasis id="B90A2A4FCF53A90C705BFB19D059E683" box="[374,820,1152,1173]" italics="true" pageId="14" pageNumber="31">
<taxonomicName id="4C7E8DDECF53A90C705BFB19D2A0E683" box="[374,461,1152,1173]" class="Eutardigrada" family="Hypsibiidae" genus="Doryphoribius" kingdom="Animalia" order="Parachela" pageId="14" pageNumber="31" phylum="Tardigrada" rank="species" species="flavus">D. flavus</taxonomicName>
<taxonomicName id="4C7E8DDECF53A90C73B0FB19D059E683" box="[669,820,1152,1173]" class="Eutardigrada" family="Hypsibiidae" genus="Doryphoribius" kingdom="Animalia" order="Parachela" pageId="14" pageNumber="31" phylum="Tardigrada" rank="species" species="amazzonicus" status="sp. nov.">D. amazzonicus</taxonomicName>
</emphasis>
<emphasis id="B90A2A4FCF53A90C7216FBE6D0EAE682" bold="true" box="[827,903,1151,1172]" pageId="14" pageNumber="31">
<taxonomicNameLabel id="A2399734CF53A90C7216FBE6D0EAE682" box="[827,903,1151,1172]" pageId="14" pageNumber="31" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
<taxonomicName id="4C7E8DDECF53A90C7285FB19D76AE683" box="[936,1031,1152,1173]" class="Eutardigrada" family="Hypsibiidae" genus="Doryphoribius" kingdom="Animalia" order="Parachela" pageId="14" pageNumber="31" phylum="Tardigrada" rank="species" species="bindae" status="sp. nov.">
<emphasis id="B90A2A4FCF53A90C7285FB19D76AE683" box="[936,1031,1152,1173]" italics="true" pageId="14" pageNumber="31">D. bindae</emphasis>
</taxonomicName>
<emphasis id="B90A2A4FCF53A90C7521FBE6D734E682" bold="true" box="[1036,1113,1151,1172]" pageId="14" pageNumber="31">
<taxonomicNameLabel id="A2399734CF53A90C7521FBE6D734E682" box="[1036,1113,1151,1172]" pageId="14" pageNumber="31" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
<taxonomicName id="4C7E8DDECF53A90C7587FB19D658E683" box="[1194,1333,1152,1173]" class="Eutardigrada" family="Hypsibiidae" genus="Doryphoribius" kingdom="Animalia" order="Parachela" pageId="14" pageNumber="31" phylum="Tardigrada" rank="species" species="solidunguis" status="sp. nov.">
<emphasis id="B90A2A4FCF53A90C7587FB19D658E683" box="[1194,1333,1152,1173]" italics="true" pageId="14" pageNumber="31">D. solidunguis</emphasis>
</taxonomicName>
<emphasis id="B90A2A4FCF53A90C7411FBE6D6E5E682" bold="true" box="[1340,1416,1151,1172]" pageId="14" pageNumber="31">
<taxonomicNameLabel id="A2399734CF53A90C7411FBE6D6E5E682" box="[1340,1416,1151,1172]" pageId="14" pageNumber="31" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
slide N. 1200 slide N. 4828 slide N. 3609 slide N. 3796
</paragraph>
<paragraph id="8BC1F65DCF53A90C71B2FB5CD6FAE6CC" blockId="14.[151,1437,979,1322]" box="[159,1431,1221,1242]" pageId="14" pageNumber="31">Characters µm pt % bo-le µm pt % bo-le µm pt % bo-le µm pt % bo-le</paragraph>
<paragraph id="8BC1F65DCF53A90C71B2FB74D639E714" blockId="14.[151,1437,979,1322]" box="[159,1364,1261,1282]" pageId="14" pageNumber="31">Body length 404 - - 241.2 - - 327.6 - - 361.3 - -</paragraph>
<paragraph id="8BC1F65DCF53A90C71B2FA8CD618E73C" blockId="14.[151,1437,979,1322]" box="[159,1397,1301,1322]" pageId="14" pageNumber="31">Buccal tube length 46.7 - 11.6 33.2 13.8 - 36.7 - 11.2 41.2 - 11.4</paragraph>
</subSubSection>
</treatment>
</document>