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<document id="22473B7909F49501DCD149F7BDE7799A" ID-DOI="10.26028/cybium/2018-422-006" ID-ISSN="2101-0315" ID-Zenodo-Dep="13473358" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="guilherme" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="guilherme" checkinTime="1723040158279" checkinUser="felipe" docAuthor="Biscoito, Manuel &amp; González, José A." docDate="2018" docId="03D4260E351AB700FCC34108FE2F25CB" docLanguage="en" docName="Cybium.42.2.189-194.pdf" docOrigin="Cybium 42 (2)" docSource="http://dx.doi.org/10.26028/cybium/2018-422-006" docStyle="DocumentStyle:73DAD84611431DC6606F8B63B2C6039C.1:Cybium.2013-2018.journal_article.type1" docStyleId="73DAD84611431DC6606F8B63B2C6039C" docStyleName="Cybium.2013-2018.journal_article.type1" docStyleVersion="1" docTitle="Physiculus sudanensis Paulin 1989" docType="treatment" docVersion="3" lastPageNumber="194" masterDocId="FFED5E76351EB705FFEE4732FFD3261C" masterDocTitle="Physiculus sudanensis Paulin, 1989, a junior synonym of P. dalwigki Kaup, 1858 (Teleostei, Gadiformes, Moridae), with a redescription of P. dalwigki" masterLastPageNumber="194" masterPageNumber="189" pageNumber="193" updateTime="1724932153531" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="2C99CECF2B0E8194B375594AE51A947D">Physiculus sudanensis Paulin, 1989, a junior synonym of P. dalwigki Kaup, 1858 (Teleostei, Gadiformes, Moridae), with a redescription of P. dalwigki</mods:title>
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<paragraph id="8BC29718351AB701FCC34108FB1E204E" blockId="4.[813,1229,1593,1618]" box="[813,1229,1593,1618]" pageId="4" pageNumber="193">
<heading id="D08A2074351AB701FCC34108FB1E204E" bold="true" box="[813,1229,1593,1618]" fontSize="9" level="2" pageId="4" pageNumber="193" reason="3">
<emphasis id="B9094B0A351AB701FCC34108FB1E204E" bold="true" box="[813,1229,1593,1618]" pageId="4" pageNumber="193">
Validity of
<taxonomicName id="4C7DEC9B351AB701FC464108FB1E204E" ID-CoL="4HH7N" authority="Paulin, 1989" authorityName="Paulin" authorityYear="1989" box="[936,1229,1593,1618]" family="Moridae" genus="Physiculus" kingdom="Animalia" order="Gadiformes" pageId="4" pageNumber="193" phylum="Chordata" rank="species" species="sudanensis">
<emphasis id="B9094B0A351AB701FC464108FBEF204D" bold="true" box="[936,1084,1593,1617]" italics="true" pageId="4" pageNumber="193">P. sudanensis</emphasis>
Paulin, 1989
</taxonomicName>
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<subSubSection id="C367C493351AB700FCBC4168FE2F25CB" lastPageId="5" lastPageNumber="194" pageId="4" pageNumber="193" type="description">
<paragraph id="8BC29718351AB701FCBC4168FCA5216E" blockId="4.[813,1456,1623,1907]" pageId="4" pageNumber="193">
After a comparison of the type material of
<emphasis id="B9094B0A351AB701FAF14169FA7C206E" box="[1311,1455,1623,1650]" italics="true" pageId="4" pageNumber="193">
<taxonomicName id="4C7DEC9B351AB701FAF14169FABF206E" box="[1311,1388,1626,1650]" family="Moridae" genus="Physiculus" kingdom="Animalia" order="Gadiformes" pageId="4" pageNumber="193" phylum="Chordata" rank="species" species="suda">P. suda</taxonomicName>
<EFBFBD>e<EFBFBD>sis
</emphasis>
with the
<typeStatus id="54C629BA351AB701FC654148FC39208E" box="[907,1002,1658,1682]" pageId="4" pageNumber="193" type="holotype">holotype</typeStatus>
and other material of
<taxonomicName id="4C7DEC9B351AB701FB374149FA82208E" authorityName="Kaup" authorityYear="1858" box="[1241,1361,1658,1682]" family="Moridae" genus="Physiculus" kingdom="Animalia" order="Gadiformes" pageId="4" pageNumber="193" phylum="Chordata" rank="species" species="dalwigki">
<emphasis id="B9094B0A351AB701FB374149FA82208E" box="[1241,1361,1658,1682]" italics="true" pageId="4" pageNumber="193">P. dalwigki</emphasis>
</taxonomicName>
(
<tableCitation id="C6FFA2A3351AB701FA8F4148FA4C208E" box="[1377,1439,1658,1682]" captionStart="Table I" captionStartId="3.[132,185,217,239]" captionText="Table I. Morphometric and meristic data from the type material of Physiculus dalwigki and Physiculus suda<64>e<EFBFBD>sis." pageId="4" pageNumber="193">Tab. I</tableCitation>
), no significant differences between the two species could be observed. Most of the body proportions and meristic characters overlap or differences found are minimal. Moreover,
<bibRefCitation id="EFECEAE9351AB701FCC341C9FC1C210F" author="PAULIN C. D." box="[813,975,1787,1811]" pageId="4" pageNumber="193" pagination="93 - 133" refId="ref5191" refString="PAULIN C. D., 1989 b. - Review of the morid genera Gadella, Physiculus, and Salil <20> ta (Teleostei: Gadiformes) with descriptions of seven new species. N. Z. J. Z <20><> l., 16: 93 - 133." type="journal article" year="1989">Paulin (1989b)</bibRefCitation>
diagnosis of
<emphasis id="B9094B0A351AB701FB8C41C9FB27210E" box="[1122,1268,1783,1811]" italics="true" pageId="4" pageNumber="193">
<taxonomicName id="4C7DEC9B351AB701FB8C41C9FB7C210F" box="[1122,1199,1787,1811]" family="Moridae" genus="Physiculus" kingdom="Animalia" order="Gadiformes" pageId="4" pageNumber="193" phylum="Chordata" rank="species" species="suda">P. suda</taxonomicName>
<EFBFBD>e<EFBFBD>sis
</emphasis>
cannot be of any use, as it is the result of a mixture of three different species:
<emphasis id="B9094B0A351AB701FCC3400EFC73214F" box="[813,928,1848,1875]" italics="true" pageId="4" pageNumber="193">
<taxonomicName id="4C7DEC9B351AB701FCC3400EFCBB214F" box="[813,872,1851,1875]" family="Moridae" genus="Physiculus" kingdom="Animalia" order="Gadiformes" pageId="4" pageNumber="193" phylum="Chordata" rank="species" species="fed">P. fed</taxonomicName>
<EFBFBD><EFBFBD><EFBFBD>vi
</emphasis>
,
<emphasis id="B9094B0A351AB701FC40400EFBE1214F" box="[942,1074,1848,1875]" italics="true" pageId="4" pageNumber="193">P. be<62>telse<73>i</emphasis>
(
<bibRefCitation id="EFECEAE9351AB701FBAC4009FACE214F" author="SHCHERBACHEV Y. N." box="[1090,1309,1851,1875]" pageId="4" pageNumber="193" pagination="147 - 178" refId="ref5398" refString="SHCHERBACHEV Y. N., 1993. - Preliminary review of the genus Physiculus (Moridae, Gadiformes) in the Indian Ocean and adjacent waters of the South Atlantic. T <20> a <20> s. P. P. Shi <20> sh <20> v I <20> st. Ocea <20><> l., 128: 147 - 178." type="journal article" year="1993">Shcherbachev, 1993</bibRefCitation>
) and
<emphasis id="B9094B0A351AB701FAB7400EFCBC216F" italics="true" pageId="4" pageNumber="193">P. suda- <20>e<EFBFBD>sis</emphasis>
.
</paragraph>
<paragraph id="8BC29718351BB700FF4447EBFF1927E2" blockId="5.[132,775,214,578]" pageId="5" pageNumber="194">
The
<typeStatus id="54C629BA351BB700FF3447E8FEDB26EE" box="[218,264,218,242]" pageId="5" pageNumber="194">type</typeStatus>
locality of
<emphasis id="B9094B0A351BB700FE6D47E8FDC026ED" box="[387,531,214,241]" italics="true" pageId="5" pageNumber="194">
<taxonomicName id="4C7DEC9B351BB700FE6D47E8FE0226ED" box="[387,465,217,241]" family="Moridae" genus="Physiculus" kingdom="Animalia" order="Gadiformes" pageId="5" pageNumber="194" phylum="Chordata" rank="species" species="suda">P. suda</taxonomicName>
<EFBFBD>e<EFBFBD>sis
</emphasis>
is well within the area of distribution of
<taxonomicName id="4C7DEC9B351BB700FED147CEFE66270F" authorityName="Kaup" authorityYear="1858" box="[319,437,251,275]" family="Moridae" genus="Physiculus" kingdom="Animalia" order="Gadiformes" pageId="5" pageNumber="194" phylum="Chordata" rank="species" species="dalwigki">
<emphasis id="B9094B0A351BB700FED147CEFE66270F" box="[319,437,251,275]" italics="true" pageId="5" pageNumber="194">P. dalwigki</emphasis>
</taxonomicName>
, the eastern Atlantic Ocean and not the coast of
<collectingCountry id="F36AD788351BB700FEC7462EFEBF2728" box="[297,364,284,308]" name="Sudan" pageId="5" pageNumber="194">Sudan</collectingCountry>
in the
<collectingRegion id="49B959FA351BB700FE5F462EFDD82729" box="[433,523,284,309]" country="Sudan" name="Al Bahr al Ahmar" pageId="5" pageNumber="194">Red Sea</collectingRegion>
, as
<bibRefCitation id="EFECEAE9351BB700FDDE462EFD1D2728" author="PAULIN C. D." box="[560,718,284,308]" pageId="5" pageNumber="194" pagination="93 - 133" refId="ref5191" refString="PAULIN C. D., 1989 b. - Review of the morid genera Gadella, Physiculus, and Salil <20> ta (Teleostei: Gadiformes) with descriptions of seven new species. N. Z. J. Z <20><> l., 16: 93 - 133." type="journal article" year="1989">Paulin (1989b)</bibRefCitation>
erroneously gave. The genus
<taxonomicName id="4C7DEC9B351BB700FE79460CFDDE274A" authorityName="Kaup" authorityYear="1858" box="[407,525,318,342]" family="Moridae" genus="Physiculus" kingdom="Animalia" order="Gadiformes" pageId="5" pageNumber="194" phylum="Chordata" rank="genus">
<emphasis id="B9094B0A351BB700FE79460CFDDE274A" box="[407,525,318,342]" italics="true" pageId="5" pageNumber="194">Physiculus</emphasis>
</taxonomicName>
contains many species and is widespread in all major oceans, but individual species have fairly restricted geographical distributions (
<bibRefCitation id="EFECEAE9351BB700FD3D46B3FF2227A7" author="PAULIN C. D." pageId="5" pageNumber="194" refId="ref5139" refString="PAULIN C. D., 1989 a. - Moridae: Overview. I <20>: Papers on the Systematics of Gadiform Fishes (Cohen D. M., ed.), pp. 243 - 250. Los Angeles: Natural History Museum of Los Angeles County." type="book" year="1989">Paulin, 1989a</bibRefCitation>
) and up to the present there are no known species occurring in more than one ocean basin (
<bibRefCitation id="EFECEAE9351BB700FDAD46F7FF6D27E2" author="FROESE R. &amp; PAULY D." pageId="5" pageNumber="194" refId="ref4682" refString="FROESE R. &amp; PAULY D. (eds), 2017. - FishBase. World Wide Web electronic publication. www. fishbase. org. Accessed Feb. 2017." type="journal volume" year="2017">Froese and Pauly, 2017</bibRefCitation>
).
</paragraph>
<paragraph id="8BC29718351BB700FF44453AFD9A245D" blockId="5.[132,775,214,578]" pageId="5" pageNumber="194">
Therefore we come to the conclusion that
<emphasis id="B9094B0A351BB700FD9B453AFCD42403" box="[629,775,516,544]" italics="true" pageId="5" pageNumber="194">
<taxonomicName id="4C7DEC9B351BB700FD9B453AFD11243C" box="[629,706,520,544]" family="Moridae" genus="Physiculus" kingdom="Animalia" order="Gadiformes" pageId="5" pageNumber="194" phylum="Chordata" rank="species" species="suda">P. suda</taxonomicName>
<EFBFBD>e<EFBFBD>sis
</emphasis>
(
<figureCitation id="13468B9D351BB700FF62451BFF18245D" box="[140,203,553,577]" captionStart="Figure 4" captionStartId="2.[132,197,435,457]" captionTargetBox="[132,775,215,418]" captionTargetId="figure-381@2.[132,775,215,418]" captionTargetPageId="2" captionText="Figure 4. Physiculus suda<64>e<EFBFBD>sis, holotype, MNHN 1886-572. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.13473381" httpUri="https://zenodo.org/record/13473381/files/figure.png" pageId="5" pageNumber="194">Fig. 4</figureCitation>
) is a junior synonym of
<taxonomicName id="4C7DEC9B351BB700FE234518FD91245D" authorityName="Kaup" authorityYear="1858" box="[461,578,553,577]" family="Moridae" genus="Physiculus" kingdom="Animalia" order="Gadiformes" pageId="5" pageNumber="194" phylum="Chordata" rank="species" species="dalwigki">
<emphasis id="B9094B0A351BB700FE234518FD91245D" box="[461,578,553,577]" italics="true" pageId="5" pageNumber="194">P. dalwigki</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BC29718351BB700FF6A4546FE2F25CB" blockId="5.[132,775,628,983]" pageId="5" pageNumber="194">
<emphasis id="B9094B0A351BB700FF6A4546FE982496" bold="true" box="[132,331,628,650]" pageId="5" pageNumber="194">Acknowledgements</emphasis>
. The authors are indebted to Romain Causse and Zora Gabsi (MNHN, Paris) and James Maclaine (NHM, London) for allowing the examination of the
<typeStatus id="54C629BA351BB700FDFE459AFDEA24A1" box="[528,569,680,701]" pageId="5" pageNumber="194">type</typeStatus>
material of
<emphasis id="B9094B0A351BB700FD40459BFF6724CB" italics="true" pageId="5" pageNumber="194">
<taxonomicName id="4C7DEC9B351BB700FD40459BFD2124A1" box="[686,754,680,702]" family="Moridae" genus="Physiculus" kingdom="Animalia" order="Gadiformes" pageId="5" pageNumber="194" phylum="Chordata" rank="species" species="suda">P. suda</taxonomicName>
<EFBFBD>- e<>sis
</emphasis>
and
<emphasis id="B9094B0A351BB700FF0F45F0FE8224CA" box="[225,337,705,727]" italics="true" pageId="5" pageNumber="194">
<taxonomicName id="4C7DEC9B351BB700FF0F45F0FE9E24CA" authorityName="Kaup" authorityYear="1858" box="[225,333,705,727]" family="Moridae" genus="Physiculus" kingdom="Animalia" order="Gadiformes" pageId="5" pageNumber="194" phylum="Chordata" rank="species" species="dalwigki">P. dalwigki</taxonomicName>
,
</emphasis>
including taking radiographs. Thanks are also due to Guy Duhamel for his editorial work and to anonymous referees, which improved the original manuscript. Financial support (JAG) was received from the Government of the
<collectingRegion id="49B959FA351BB700FD9F443CFCD42538" box="[625,775,782,804]" country="Spain" name="Canarias" pageId="5" pageNumber="194">Canary Islands</collectingRegion>
and the EU ERDF in the framework of the PCT MAC 2007-2013 project MARPROF-CV (MAC//3/C124) and (MB) from the Madeira Oceanic Observatory (Project M1420-01-0145-FEDER-000001 Observatório Oceânico da Madeira
<emphasis id="B9094B0A351BB700FE174447FDEA2596" box="[505,569,885,906]" italics="true" pageId="5" pageNumber="194">-OOM</emphasis>
) and Fundação para a Ciência e Tecnologia (FCT), through the strategic project UID/ MAR/04292/2013 granted to MARE. This is contribution no. 39 of the Marine Biology Station of Funchal.
</paragraph>
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</treatment>
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