treatments-xml/data/03/BD/87/03BD87E66B11FFB7AF89F9DCFD1E3F55.xml

<document id="0D71E624370AD5D579BCCB5C6CC15EBA" ID-CLB-Dataset="8257" ID-DOI="10.11646/zootaxa.4922.1.1" ID-GBIF-Dataset="188411de-569c-4408-a016-1a04b858c142" ID-ISSN="1175-5326" ID-Zenodo-Dep="4496316" ID-ZooBank="D7F9C6A3-9C28-4F4C-8E81-BF24849FDD8C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1612331093960" checkinUser="plazi" docAuthor="Fabrizi, Silvia, Liu, Wan-Gang, Bai, Ming, Yang, Xing-Ke &amp; Ahrens, Dirk" docDate="2021" docId="03BD87E66B11FFB7AF89F9DCFD1E3F55" docLanguage="en" docName="zootaxa.4922.1.1.pdf" docOrigin="Zootaxa 4922 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Maladera laocaiensis Ahrens, Fabrizi &amp; Liu 2021, sp. n." docType="treatment" docVersion="8" lastPageNumber="115" masterDocId="FF84FF9E6B61FFC5AF1EF86AFFDE3D4C" masterDocTitle="A monograph of the genus Maladera Mulsant &amp; Rey, 1871 of China (Coleoptera: Scarabaeidae: Melolonthinae: Sericini)" masterLastPageNumber="400" masterPageNumber="1" pageNumber="113" updateTime="1698926631736" updateUser="ExternalLinkService">
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<mods:title id="60DFB795EB07C828478CFC558A503483">A monograph of the genus Maladera Mulsant &amp; Rey, 1871 of China (Coleoptera: Scarabaeidae: Melolonthinae: Sericini)</mods:title>
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<mods:namePart id="D7B40C62186CFF0AE8A742E2A349D368">Liu, Wan-Gang</mods:namePart>
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<mods:namePart id="3C42013B20430B47591C70EFEE7E64D4">Bai, Ming</mods:namePart>
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<mods:namePart id="DCF3BA927D9F46EC5D644AE496531545">Ahrens, Dirk</mods:namePart>
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<taxonomicName id="4C144D736B11FFB5AF89F9DCFD653C9C" authority="Ahrens, Fabrizi &amp; Liu" authorityName="Ahrens, Fabrizi &amp; Liu" authorityYear="2021" box="[151,699,438,465]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="112" pageNumber="113" phylum="Arthropoda" rank="species" species="laocaiensis" status="sp. nov.">
<emphasis id="B960EAE26B11FFB5AF89F9DCFE443C9D" bold="true" box="[151,410,438,465]" italics="true" pageId="112" pageNumber="113">Maladera laocaiensis</emphasis>
Ahrens, Fabrizi &amp; Liu
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, 
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<figureCitation id="132F2A756B11FFB5AF89F9B1FED03CB9" box="[151,270,475,501]" captionStart="FIGURE 24" captionStartId="113.[151,250,1897,1921]" captionTargetBox="[228,1354,244,1861]" captionTargetId="figure-41@113.[212,1375,237,1873]" captionTargetPageId="113" captionText="FIGURE 24. A–E: Maladera laocaiensis Ahrens, Fabrizi &amp; Liu, sp. n. (holotype); F–I: M. pseudoegregia Ahrens, Fabrizi &amp; Liu, sp. n. (holotype); J–M: M. gusakovi Ahrens, Fabrizi &amp; Liu, sp. n. (holotype). A, F, L: aedeagus, left side lateral view; D, H, L: aedeagus, right side lateral view; B, G, K: parameres, dorsal view; C: parameres, ventral view; E, I, M: habitus. Scale: 0.5 mm. Habitus not to scale." figureDoi="http://doi.org/10.5281/zenodo.4496368" httpUri="https://zenodo.org/record/4496368/files/figure.png" pageId="112" pageNumber="113">Figures 24</figureCitation>
A–E, 97
</paragraph>
</subSubSection>
<subSubSection id="C30E657B6B11FFB5AF89FA49FCD23FE5" pageId="112" pageNumber="113" type="materials_examined">
<paragraph id="8BAB36F06B11FFB5AF89FA49FCD23FE5" blockId="112.[151,1437,546,2013]" pageId="112" pageNumber="113">
<emphasis id="B960EAE26B11FFB5AF89FA49FE1E3F70" bold="true" box="[151,448,546,573]" pageId="112" pageNumber="113">Type material examined.</emphasis>
<materialsCitation id="3B7C3CAD6B11FFB5AED9FA49FC293F2D" ID-GBIF-Occurrence="3034555691" collectingDate="1999-05-23" collectingDateMax="1999-05-27" collectingDateMin="1999-05-23" collectionCode="ZFMK" collectorName="Ahrens" elevation="1650" location="Sa Pa" pageId="112" pageNumber="113" specimenCount="1" specimenCount-male="1" stateProvince="Lao Cai" typeStatus="holotype">
<typeStatus id="54AF88526B11FFB5AED9FA49FDEA3F71" box="[455,564,547,573]" pageId="112" pageNumber="113" type="holotype">Holotype</typeStatus>
: 
<specimenCount id="9D12FD796B11FFB5AD20FA49FD8D3F71" box="[574,595,547,573]" pageId="112" pageNumber="113" type="male">♂</specimenCount>
“N-Vietnam, Prov. 
<collectingRegion id="49D0F8126B11FFB5AC2CFA49FC513F71" box="[818,911,547,573]" country="Vietnam" name="Lao Cai" pageId="112" pageNumber="113">Lao Cai</collectingRegion>
, 
<location id="8ECB602B6B11FFB5AC84FA49FC033F71" LSID="urn:lsid:plazi:treatment:03BD87E66B11FFB7AF89F9DCFD1E3F55:8ECB602B6B11FFB5AC84FA49FC033F71" box="[922,989,547,573]" name="Sa Pa" pageId="112" pageNumber="113" stateProvince="Lao Cai">Sa Pa</location>
, 
<quantity id="4CEC9B156B11FFB5ACF6FA49FB5E3F71" box="[1000,1152,547,573]" metricMagnitude="3" metricUnit="m" metricValue="1.65" metricValueMax="1.7" metricValueMin="1.6" pageId="112" pageNumber="113" unit="m" value="1650.0" valueMax="1700.0" valueMin="1600.0">
<elevation id="0039D1C36B11FFB5ACF6FA49FB5E3F71" box="[1000,1152,547,573]" metricMagnitude="3" metricUnit="m" metricValue="1.65" metricValueMax="1.7" metricValueMin="1.6" pageId="112" pageNumber="113" unit="m" value="1650.0" valueMax="1700.0" valueMin="1600.0">1600-1700 m</elevation>
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, N22°19‘52‘‘; E103°50‘; 
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<collectingDate id="EFEEE9D86B11FFB5AF89FA2DFEE93F2D" box="[151,311,583,609]" pageId="112" pageNumber="113" value="1999-05-23" valueMax="1999-05-27" valueMin="1999-05-23">23.-27.V.1999</collectingDate>
</date>
leg. 
<collectorName id="26E153266B11FFB5AE72FA2DFE1E3F2D" box="[364,448,583,609]" pageId="112" pageNumber="113">Ahrens</collectorName>
, Jäger, Fabrizi/ 1044 
<taxonomicName id="4C144D736B11FFB5ADAEFA2DFCD93F2D" box="[688,775,583,609]" class="Insecta" family="Scarabaeidae" kingdom="Animalia" order="Coleoptera" pageId="112" pageNumber="380" phylum="Arthropoda" rank="tribe" tribe="Sericini">Sericini</taxonomicName>
Asia spec.” (
<collectionCode id="ED05AE356B11FFB5AC85FA2DFC313F2D" box="[923,1007,583,609]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="112" pageNumber="113" type="Museum">ZFMK</collectionCode>
)
</materialsCitation>
. 
<materialsCitation id="3B7C3CAD6B11FFB5AB1FFA2DFC413FC9" ID-GBIF-Occurrence="3034555881" collectionCode="CP" collectorName="P. Pacholatko" country="Vietnam" location="Vietnam" pageId="112" pageNumber="113" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<typeStatus id="54AF88526B11FFB5AB1FFA2DFBAC3F2D" box="[1025,1138,583,609]" pageId="112" pageNumber="113" type="paratype">Paratypes</typeStatus>
: 
<specimenCount id="9D12FD796B11FFB5AB63FA2DFB793F2D" box="[1149,1191,583,609]" pageId="112" pageNumber="113" type="male">1 ♂</specimenCount>
“N 
<collectingCountry id="F30376606B11FFB5ABCBFA2DFAEA3F2D" box="[1237,1332,583,609]" name="Vietnam" pageId="112" pageNumber="113">Vietnam</collectingCountry>
(Tonkin) pr. Vinh Phu 1990 Tam Dao 17.-21.v. 
<collectorName id="26E153266B11FFB5AD25FA01FD0F3FC9" box="[571,721,619,645]" pageId="112" pageNumber="113">P. Pacholatko</collectorName>
leg./ VS 79” (
<collectionCode id="ED05AE356B11FFB5AC6EFA01FC493FC9" box="[880,919,619,645]" pageId="112" pageNumber="113">CP</collectionCode>
)
</materialsCitation>
, 
<materialsCitation id="3B7C3CAD6B11FFB5ACB4FA01FCD63FE5" ID-GBIF-Occurrence="3034555953" collectingDate="2007-07-28" collectingDateMax="2007-07-29" collectingDateMin="2007-07-28" collectionCode="LSSYU" collectorName="Li Jiahui" country="Vietnam" location="Caiyanghe Nature Reserve" municipality="Pu'er" pageId="112" pageNumber="113" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="9D12FD796B11FFB5ACB4FA01FC0D3FC9" box="[938,979,619,645]" pageId="112" pageNumber="113" type="male">1 ♂</specimenCount>
“ 
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, 
<collectingMunicipality id="6BCFAC8A6B11FFB5AA00FA01FA823FC9" box="[1310,1372,619,645]" pageId="112" pageNumber="113">Pu’er</collectingMunicipality>
, Yunnan, 
<date id="FFAA10306B11FFB5AFD0FAE5FEA03FE5" box="[206,382,655,681]" pageId="112" pageNumber="113" value="2007-07-28" valueMax="2007-07-29" valueMin="2007-07-28">
<collectingDate id="EFEEE9D86B11FFB5AFD0FAE5FEA03FE5" box="[206,382,655,681]" pageId="112" pageNumber="113" value="2007-07-28" valueMax="2007-07-29" valueMin="2007-07-28">28-29.VII.2007</collectingDate>
</date>
, leg. 
<collectorName id="26E153266B11FFB5AEA4FAE5FDC23FE5" box="[442,540,655,681]" pageId="112" pageNumber="113">Li Jiahui</collectorName>
/ LW-118” (
<collectionCode id="ED05AE356B11FFB5ADBFFAE5FCDE3FE5" box="[673,768,655,681]" pageId="112" pageNumber="113">LSSYU</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C30E657B6B11FFB7AFD9FAD8FCA83DFD" lastPageId="114" lastPageNumber="115" pageId="112" pageNumber="113" type="description">
<paragraph id="8BAB36F06B11FFB5AFD9FAD8FC583FBD" blockId="112.[151,1437,546,2013]" pageId="112" pageNumber="113">
<emphasis id="B960EAE26B11FFB5AFD9FAD8FE873F80" bold="true" box="[199,345,690,716]" pageId="112" pageNumber="113">Description.</emphasis>
Length: 
<quantity id="4CEC9B156B11FFB5AEA5FAD9FDCC3F81" box="[443,530,691,717]" metricMagnitude="-3" metricUnit="m" metricValue="9.4" pageId="112" pageNumber="113" unit="mm" value="9.4">9.4 mm</quantity>
, length of elytra: 
<quantity id="4CEC9B156B11FFB5ADCCFAD9FCF73F81" box="[722,809,691,717]" metricMagnitude="-3" metricUnit="m" metricValue="6.6" pageId="112" pageNumber="113" unit="mm" value="6.6">6.6 mm</quantity>
, width: 
<quantity id="4CEC9B156B11FFB5AC9FFAD8FC063F81" box="[897,984,690,717]" metricMagnitude="-3" metricUnit="m" metricValue="5.6" pageId="112" pageNumber="113" unit="mm" value="5.6">5.6 mm</quantity>
. Body oval, dark reddish brown, antenna yellow, dorsal surface iridescent shiny, dorsal face nearly glabrous.
</paragraph>
<paragraph id="8BAB36F06B11FFB5AFD9FA91FBCF395D" blockId="112.[151,1437,546,2013]" pageId="112" pageNumber="113">
Labroclypeus wide and trapezoidal, widest at base, lateral margins straight and strongly convergent anteriorly, anterior angles strongly rounded, anterior margin distinctly emarginate, margins weakly reflexed; lateral margin and ocular canthus produce an indistinct angle; surface weakly convex, finely, densely punctate, with a few long setae anteriorly; frontoclypeal suture finely incised, weakly curved; smooth area anterior to eye convex, 1.5 times as wide as long; ocular canthus short and wide (1/4 of ocular diameter), finely densely punctate, with one terminal seta. Frons with sparse and fine punctures, with a few fine setae beside eyes. Eyes small, ratio diameter/ interocular width: 0.57. Antenna with ten antennomeres; club with three antennomeres and straight, 1.1 times as long as remaining antennomeres combined. 
<collectingRegion id="49D0F8126B11FFB5AEFFFB9DFD9E395D" box="[481,576,1015,1041]" country="Myanmar" name="Chin" pageId="112" pageNumber="113">Mentum</collectingRegion>
elevated and slightly flattened anteriorly.
</paragraph>
<paragraph id="8BAB36F06B11FFB5AFD9FC71FCF93989" blockId="112.[151,1437,546,2013]" pageId="112" pageNumber="113">Pronotum transverse, widest at base, lateral margins evenly convex and weakly convergent anteriorly, anterior angles moderately produced and right-angled, posterior angles convex; anterior margin weakly convex, with fine marginal line, base without marginal line; surface finely and densely punctate, with minute setae in punctures being as long as puncture diameter, otherwise glabrous; lateral margin densely setose; hypomeron carinate, not produced ventrally. Scutellum triangular, with fine, dense punctures.</paragraph>
<paragraph id="8BAB36F06B11FFB5AFD9FCA5FE603836" blockId="112.[151,1437,546,2013]" pageId="112" pageNumber="113">Elytra widest inposterior third, striae finely impressed, finely and densely punctate, intervals weakly convex, with fine, moderately dense punctures concentrated along striae, with minute setae in punctures, odd intervals with a few short and fine setae; epipleural margin robust, ending at apical external angle of elytra, epipleura densely setose; apical border of elytra convex before apex, membraneous, with a very fine rim of microtrichomes (visible at ca 100x magnification).</paragraph>
<paragraph id="8BAB36F06B11FFB5AFD9FDE9FEEF3B1D" blockId="112.[151,1437,546,2013]" pageId="112" pageNumber="113">Ventral surface dull, coarsely and densely punctate, metasternum including disc densely setose; metacoxa glabrous, with a few longer setae laterally. Abdominal sternites finely and densely punctate, punctures with minute setae, each sternite with a transverse row of punctures each bearing a fine seta. Mesosternum between mesocoxae as wide as mesofemur. Ratio of length of metepisternum/metacoxa: 1/1.37. Pygidium dull, convexly elevated at middle before apex, finely and densely punctate, without smooth midline, with numerous long and short, fine setae in apical half.</paragraph>
<paragraph id="8BAB36F06B11FFB4AFD9FE31FE333D99" blockId="112.[151,1437,546,2013]" lastBlockId="113.[151,1436,151,213]" lastPageId="113" lastPageNumber="114" pageId="112" pageNumber="113">Legs moderately long and narrow, shiny; femora with two longitudinal rows of setae, coarsely and moderately densely punctate. Anterior margin of metafemur acute, with adjacent continuous serrated line, anterior row of setae complete; posterior ventral margin smooth, weakly widened at ventral apex, dorsal posterior margin smooth, neither serrate, glabrous. Metatibia long and moderately wide, widest at apex, ratio of width/length: 1/3.3, moderately carinate dorsally, with two groups of spines, basal group shortly behind middle, apical group at four fifths of metatibial length, in basal half with a few short and fine single setae and a continuous serrated line beside entire dorsal margin until the basal group of spines; lateral face longitudinally convex, with sparse, fine punctures, minutely setose; ventral margin finely serrate, with four equidistant robust setae; medial face smooth and glabrous; apex finely serrate, shallowly emarginate interiorly near tarsal articulation. Tarsomeres dorsally impunctate, glabrous, neither laterally nor dorsally carinate, moderately setose ventrally; metatarsomeres with a strongly serrated ridge ventrally and a smooth subventral longitudinal carina; first metatarsomere distinctly longer than following tarsomere and slightly longer than dorsal tibial spur. Protibia moderately long, bidentate; anterior claws symmetrical, basal tooth of both claws bluntly truncate at apex.</paragraph>
<caption id="DF6B66786B10FFB4AF89FF03FE1F3AA1" ID-DOI="http://doi.org/10.5281/zenodo.4496368" ID-Zenodo-Dep="4496368" httpUri="https://zenodo.org/record/4496368/files/figure.png" pageId="113" pageNumber="114" startId="113.[151,250,1897,1921]" targetBox="[228,1354,244,1861]" targetPageId="113">
<paragraph id="8BAB36F06B10FFB4AF89FF03FE1F3AA1" blockId="113.[151,1437,1897,2029]" pageId="113" pageNumber="114">
<emphasis id="B960EAE26B10FFB4AF89FF03FE803ACD" bold="true" box="[151,350,1897,1921]" pageId="113" pageNumber="114">FIGURE 24. A–E:</emphasis>
<taxonomicName id="4C144D736B10FFB4AE78FF03FCEE3ACD" authority="Ahrens, Fabrizi &amp; Liu" authorityName="Ahrens, Fabrizi &amp; Liu" authorityYear="2021" box="[358,816,1897,1921]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="113" pageNumber="114" phylum="Arthropoda" rank="species" species="laocaiensis" status="sp. nov.">
<emphasis id="B960EAE26B10FFB4AE78FF03FD9D3ACD" box="[358,579,1897,1921]" italics="true" pageId="113" pageNumber="114">Maladera laocaiensis</emphasis>
Ahrens, Fabrizi &amp; Liu
</taxonomicName>
, 
<taxonomicNameLabel id="A25357996B10FFB4AC25FF03FCAC3ACC" box="[827,882,1897,1920]" pageId="113" pageNumber="114" rank="species">sp. n.</taxonomicNameLabel>
(holotype); 
<emphasis id="B960EAE26B10FFB4ACEFFF03FBC13ACD" bold="true" box="[1009,1055,1897,1921]" pageId="113" pageNumber="114">F–I:</emphasis>
<taxonomicName id="4C144D736B10FFB4AB38FF00FF633AE9" authority="Ahrens, Fabrizi &amp; Liu" authorityName="Ahrens, Fabrizi &amp; Liu" authorityYear="2021" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="113" pageNumber="114" phylum="Arthropoda" rank="species" species="pseudoegregia" status="sp. nov.">
<emphasis id="B960EAE26B10FFB4AB38FF00FB033ACD" box="[1062,1245,1897,1921]" italics="true" pageId="113" pageNumber="114">M. pseudoegregia</emphasis>
Ahrens, Fabrizi &amp; Liu
</taxonomicName>
, 
<taxonomicNameLabel id="A25357996B10FFB4AFD9FFE7FF203AE8" box="[199,254,1933,1956]" pageId="113" pageNumber="114" rank="species">sp. n.</taxonomicNameLabel>
(holotype); 
<emphasis id="B960EAE26B10FFB4AE62FFE7FE683AE9" bold="true" box="[380,438,1933,1957]" pageId="113" pageNumber="114">J–M:</emphasis>
<taxonomicName id="4C144D736B10FFB4AEA3FFE4FCFB3AE9" authority="Ahrens, Fabrizi &amp; Liu" authorityName="Ahrens, Fabrizi &amp; Liu" authorityYear="2021" box="[445,805,1933,1957]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="113" pageNumber="114" phylum="Arthropoda" rank="species" species="gusakovi" status="sp. nov.">
<emphasis id="B960EAE26B10FFB4AEA3FFE4FDE73AE9" box="[445,569,1933,1957]" italics="true" pageId="113" pageNumber="114">M. gusakovi</emphasis>
Ahrens, Fabrizi &amp; Liu
</taxonomicName>
, 
<taxonomicNameLabel id="A25357996B10FFB4AC31FFE7FCB83AE8" box="[815,870,1933,1956]" pageId="113" pageNumber="114" rank="species">sp. n.</taxonomicNameLabel>
(holotype). 
<emphasis id="B960EAE26B10FFB4ACFDFFE7FBEB3AE9" bold="true" box="[995,1077,1933,1957]" pageId="113" pageNumber="114">A, F, L:</emphasis>
aedeagus, left side lateral view; 
<emphasis id="B960EAE26B10FFB4AA9AFFE7FF0F3A85" bold="true" pageId="113" pageNumber="114">D, H, L:</emphasis>
aedeagus, right side lateral view; 
<emphasis id="B960EAE26B10FFB4AD30FFDBFD563A85" bold="true" box="[558,648,1969,1993]" pageId="113" pageNumber="114">B, G, K:</emphasis>
parameres, dorsal view; 
<emphasis id="B960EAE26B10FFB4AC97FFDBFC7A3A85" bold="true" box="[905,932,1969,1993]" pageId="113" pageNumber="114">C:</emphasis>
parameres, ventral view; 
<emphasis id="B960EAE26B10FFB4ABB2FFDBFADF3A85" bold="true" box="[1196,1281,1969,1993]" pageId="113" pageNumber="114">E, I, M:</emphasis>
habitus. Scale: 0.5 mm. Habitus not to scale.
</paragraph>
</caption>
<paragraph id="8BAB36F06B13FFB7AFD9F8FDFCA83DFD" blockId="114.[151,1437,151,537]" box="[199,886,151,177]" pageId="114" pageNumber="115">
Aedeagus: 
<figureCitation id="132F2A756B13FFB7AE5AF8FDFE4B3DFD" box="[324,405,151,177]" captionStart="FIGURE 24" captionStartId="113.[151,250,1897,1921]" captionTargetBox="[228,1354,244,1861]" captionTargetId="figure-41@113.[212,1375,237,1873]" captionTargetPageId="113" captionText="FIGURE 24. A–E: Maladera laocaiensis Ahrens, Fabrizi &amp; Liu, sp. n. (holotype); F–I: M. pseudoegregia Ahrens, Fabrizi &amp; Liu, sp. n. (holotype); J–M: M. gusakovi Ahrens, Fabrizi &amp; Liu, sp. n. (holotype). A, F, L: aedeagus, left side lateral view; D, H, L: aedeagus, right side lateral view; B, G, K: parameres, dorsal view; C: parameres, ventral view; E, I, M: habitus. Scale: 0.5 mm. Habitus not to scale." figureDoi="http://doi.org/10.5281/zenodo.4496368" httpUri="https://zenodo.org/record/4496368/files/figure.png" pageId="114" pageNumber="115">Fig. 24</figureCitation>
A–D. Habitus: 
<figureCitation id="132F2A756B13FFB7AD20F8FDFD7C3DFD" box="[574,674,151,177]" captionStart="FIGURE 24" captionStartId="113.[151,250,1897,1921]" captionTargetBox="[228,1354,244,1861]" captionTargetId="figure-41@113.[212,1375,237,1873]" captionTargetPageId="113" captionText="FIGURE 24. A–E: Maladera laocaiensis Ahrens, Fabrizi &amp; Liu, sp. n. (holotype); F–I: M. pseudoegregia Ahrens, Fabrizi &amp; Liu, sp. n. (holotype); J–M: M. gusakovi Ahrens, Fabrizi &amp; Liu, sp. n. (holotype). A, F, L: aedeagus, left side lateral view; D, H, L: aedeagus, right side lateral view; B, G, K: parameres, dorsal view; C: parameres, ventral view; E, I, M: habitus. Scale: 0.5 mm. Habitus not to scale." figureDoi="http://doi.org/10.5281/zenodo.4496368" httpUri="https://zenodo.org/record/4496368/files/figure.png" pageId="114" pageNumber="115">Fig. 24E</figureCitation>
. Female unknown.
</paragraph>
</subSubSection>
<subSubSection id="C30E657B6B13FFB7AFD9F8D0FEEB3CC6" pageId="114" pageNumber="115" type="diagnosis">
<paragraph id="8BAB36F06B13FFB7AFD9F8D0FEEB3CC6" blockId="114.[151,1437,151,537]" pageId="114" pageNumber="115">
<emphasis id="B960EAE26B13FFB7AFD9F8D0FE9C3D98" bold="true" box="[199,322,186,212]" pageId="114" pageNumber="115">Diagnosis.</emphasis>
<taxonomicName id="4C144D736B13FFB7AE57F8D1FC9C3D99" authority="Ahrens, Fabrizi &amp; Liu" authorityName="Ahrens, Fabrizi &amp; Liu" authorityYear="2021" box="[329,834,187,213]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="114" pageNumber="115" phylum="Arthropoda" rank="species" species="laocaiensis" status="sp. nov.">
<emphasis id="B960EAE26B13FFB7AE57F8D1FDE33D99" box="[329,573,187,213]" italics="true" pageId="114" pageNumber="115">Maladera laocaiensis</emphasis>
Ahrens, Fabrizi &amp; Liu
</taxonomicName>
, 
<taxonomicNameLabel id="A25357996B13FFB7AC50F8D1FC543D99" box="[846,906,187,213]" pageId="114" pageNumber="115" rank="species">sp. n.</taxonomicNameLabel>
differs from the similar 
<taxonomicName id="4C144D736B13FFB7AB81F8D1FE4D3DB5" authority="Ahrens, Fabrizi &amp; Liu" authorityName="Ahrens, Fabrizi &amp; Liu" authorityYear="2021" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="114" pageNumber="115" phylum="Arthropoda" rank="species" species="diaolinensis" status="sp. nov.">
<emphasis id="B960EAE26B13FFB7AB81F8D1FA423D99" box="[1183,1436,187,213]" italics="true" pageId="114" pageNumber="115">Maladera diaolinensis</emphasis>
Ahrens, Fabrizi &amp; Liu
</taxonomicName>
, 
<taxonomicNameLabel id="A25357996B13FFB7AE83F8B5FE073DB5" box="[413,473,223,249]" pageId="114" pageNumber="115" rank="species">sp. n.</taxonomicNameLabel>
and 
<taxonomicName id="4C144D736B13FFB7AD11F8B5FCB73DB5" authority="Ahrens, Fabrizi &amp; Liu" authorityName="Ahrens, Fabrizi &amp; Liu" authorityYear="2021" box="[527,873,223,249]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="114" pageNumber="115" phylum="Arthropoda" rank="species" species="tiani" status="sp. nov.">
<emphasis id="B960EAE26B13FFB7AD11F8B5FDB93DB4" box="[527,615,223,248]" italics="true" pageId="114" pageNumber="115">M. tiani</emphasis>
Ahrens, Fabrizi &amp; Liu
</taxonomicName>
, 
<taxonomicNameLabel id="A25357996B13FFB7AC6DF8B5FC713DB5" box="[883,943,223,249]" pageId="114" pageNumber="115" rank="species">sp. n.</taxonomicNameLabel>
in having the fused left and right parameres sharply carinate ventrally. From the syntopically occurring 
<taxonomicName id="4C144D736B13FFB7AC36F969FB0E3C51" authority="Ahrens, Fabrizi &amp; Liu" authorityName="Ahrens, Fabrizi &amp; Liu" authorityYear="2021" box="[808,1232,259,285]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="114" pageNumber="115" phylum="Arthropoda" rank="species" species="tiachinensis" status="sp. nov.">
<emphasis id="B960EAE26B13FFB7AC36F969FC0D3C51" box="[808,979,259,285]" italics="true" pageId="114" pageNumber="115">M. tiachinensis</emphasis>
Ahrens, Fabrizi &amp; Liu
</taxonomicName>
, 
<taxonomicNameLabel id="A25357996B13FFB7ABC4F969FAC83C51" box="[1242,1302,259,285]" pageId="114" pageNumber="115" rank="species">sp. n.</taxonomicNameLabel>
, it differs in being iridescent and shiny dorsally and the parameres being shorter and straight in distal half (lateral view), while in 
<taxonomicName id="4C144D736B13FFB7AFAAF921FE803C29" authorityName="Ahrens, Fabrizi &amp; Liu" authorityYear="2021" box="[180,350,331,357]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="114" pageNumber="115" phylum="Arthropoda" rank="species" species="tiachinensis">
<emphasis id="B960EAE26B13FFB7AFAAF921FE803C29" box="[180,350,331,357]" italics="true" pageId="114" pageNumber="115">M. tiachinensis</emphasis>
</taxonomicName>
the dorsal surface if the body is dark and dull and its parameres are curved ventrally in distal half (lateral view).
</paragraph>
</subSubSection>
<subSubSection id="C30E657B6B13FFB7AFD9F9F9FEF13C9D" pageId="114" pageNumber="115" type="etymology">
<paragraph id="8BAB36F06B13FFB7AFD9F9F9FEF13C9D" blockId="114.[151,1437,151,537]" pageId="114" pageNumber="115">
<emphasis id="B960EAE26B13FFB7AFD9F9F9FE903CE1" bold="true" box="[199,334,403,429]" pageId="114" pageNumber="115">Etymology.</emphasis>
The new species is named (adjective in the nominative singular case) after its occurrence in 
<collectingRegion id="49D0F8126B13FFB7AA6FF9F9FEF53C9D" country="Vietnam" name="Lao Cai" pageId="114" pageNumber="115">Lao Cai province</collectingRegion>
.
</paragraph>
</subSubSection>
<subSubSection id="C30E657B6B13FFB7AFD9F9B0FBBA3CB9" box="[199,1124,474,501]" pageId="114" pageNumber="115" type="description">
<paragraph id="8BAB36F06B13FFB7AFD9F9B0FBBA3CB9" blockId="114.[151,1437,151,537]" box="[199,1124,474,501]" pageId="114" pageNumber="115">
<emphasis id="B960EAE26B13FFB7AFD9F9B0FEE13CB8" bold="true" box="[199,319,474,500]" pageId="114" pageNumber="115">Variation.</emphasis>
Length: 
<quantity id="4CEC9B156B13FFB7AEBAF9B1FDF33CB9" box="[420,557,475,501]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" metricValueMax="9.4" metricValueMin="8.6" pageId="114" pageNumber="115" unit="mm" value="9.0" valueMax="9.4" valueMin="8.6">8.6–9.4 mm</quantity>
, length of elytra: 
<quantity id="4CEC9B156B13FFB7ADECF9B0FCA53CB9" box="[754,891,474,501]" metricMagnitude="-3" metricUnit="m" metricValue="6.55" metricValueMax="6.6" metricValueMin="6.5" pageId="114" pageNumber="115" unit="mm" value="6.55" valueMax="6.6" valueMin="6.5">6.5–6.6 mm</quantity>
, width: 
<quantity id="4CEC9B156B13FFB7ACCBF9B0FB813CB9" box="[981,1119,474,501]" metricMagnitude="-3" metricUnit="m" metricValue="5.35" metricValueMax="5.6" metricValueMin="5.1" pageId="114" pageNumber="115" unit="mm" value="5.35" valueMax="5.6" valueMin="5.1">5.1–5.6 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="C30E657B6B13FFB7AFD9F994FD1E3F55" box="[199,704,510,537]" pageId="114" pageNumber="115" type="distribution">
<paragraph id="8BAB36F06B13FFB7AFD9F994FD1E3F55" blockId="114.[151,1437,151,537]" box="[199,704,510,537]" pageId="114" pageNumber="115">
<emphasis id="B960EAE26B13FFB7AFD9F994FEBF3F54" bold="true" box="[199,353,510,536]" pageId="114" pageNumber="115">Distribution.</emphasis>
See map (
<figureCitation id="132F2A756B13FFB7AEC8F995FDF63F55" box="[470,552,511,537]" captionStart="FIGURE 97" captionStartId="350.[151,250,1034,1058]" captionTargetBox="[208,1378,187,1003]" captionTargetId="figure-19@350.[202,1386,181,1010]" captionTargetPageId="350" captionText="FIGURE 97. Distribution of the Maladera egregia group: Maladera diaolinensis sp. n., M. egregia (Arrow, 1946), M. gusakovi sp. n., M. hmong Ahrens, 2004, M. hubeiensis sp. n., M. laocaiensis sp. n., M. levis (Frey, 1972)." figureDoi="http://doi.org/10.5281/zenodo.4496578" httpUri="https://zenodo.org/record/4496578/files/figure.png" pageId="114" pageNumber="115">Fig. 97</figureCitation>
) and 
<tableCitation id="C696034B6B13FFB7AD78F995FD643F55" box="[614,698,511,537]" captionStart="TABLE 1" captionStartId="368.[151,239,544,570]" captionText="TABLE 1. Overview on the distribuition of the Chinese mainland species of Maladera Mulsant &amp; Rey, 1871 (x – records from material examined herein, o – records from literature, with no material of this study)." pageId="114" pageNumber="115">Table 1</tableCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>