treatments-xml/data/03/B5/87/03B587F2AA354D77B6F8FAE63C11FED3.xml

<document id="94D4048A302624ABB469C98B45CA9AE7" ID-CLB-Dataset="30482" ID-DOI="10.11646/zootaxa.4398.1.1" ID-GBIF-Dataset="e5a58990-d727-440a-aab9-7638a5698954" ID-ISSN="1175-5326" ID-Zenodo-Dep="1202953" ID-ZooBank="79E3EB78-D1C3-45CF-AB13-F8E61C936252" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1521441832187" checkinUser="plazi" docAuthor="Boxshall, Geoff" docDate="2018" docId="03B587F2AA354D77B6F8FAE63C11FED3" docLanguage="en" docName="zootaxa.4398.1.1.pdf" docOrigin="Zootaxa 4398 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Caligus neoaricolus Boxshall 2018, sp. nov." docType="treatment" docVersion="9" lastPageNumber="96" masterDocId="FF8CFF8AAA6D4D28B66FFFEB3934FF93" masterDocTitle="The sea lice (Copepoda: Caligidae) of Moreton Bay (Queensland, Australia), with descriptions of thirteen new species" masterLastPageNumber="172" masterPageNumber="1" pageNumber="89" updateTime="1698532557943" updateUser="ExternalLinkService">
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<mods:title id="5FD01C0357F82F56EE81AF0C5352F426">The sea lice (Copepoda: Caligidae) of Moreton Bay (Queensland, Australia), with descriptions of thirteen new species</mods:title>
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<treatment id="03B587F2AA354D77B6F8FAE63C11FED3" ID-DOI="http://doi.org/10.5281/zenodo.5952184" ID-GBIF-Taxon="142340615" ID-Zenodo-Dep="5952184" LSID="urn:lsid:plazi:treatment:03B587F2AA354D77B6F8FAE63C11FED3" httpUri="http://treatment.plazi.org/id/03B587F2AA354D77B6F8FAE63C11FED3" lastPageId="95" lastPageNumber="96" pageId="88" pageNumber="89">
<subSubSection id="C306656FAA354D70B6F8FAE6381AFADB" pageId="88" pageNumber="89" type="nomenclature">
<paragraph id="8BA336E4AA354D70B6F8FAE638D9FAB4" blockId="88.[151,493,1293,1352]" box="[151,493,1293,1319]" pageId="88" pageNumber="89">
<heading id="D0EB8188AA354D70B6F8FAE638D9FAB4" bold="true" box="[151,493,1293,1319]" fontSize="11" level="1" pageId="88" pageNumber="89" reason="1">
<emphasis id="B968EAF6AA354D70B6F8FAE638D9FAB4" bold="true" box="[151,493,1293,1319]" pageId="88" pageNumber="89">
<taxonomicName id="4C1C4D67AA354D70B6F8FAE638BCFAB4" authority="Boxshall, 2018" authorityName="Boxshall" authorityYear="2018" box="[151,392,1293,1319]" class="Maxillopoda" family="Caligidae" genus="Caligus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siphonostomatoida" pageId="88" pageNumber="89" phylum="Arthropoda" rank="species" species="neoaricolus" status="sp. nov.">
<emphasis id="B968EAF6AA354D70B6F8FAE638BCFAB4" bold="true" box="[151,392,1293,1319]" italics="true" pageId="88" pageNumber="89">Caligus neoaricolus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25B578DAA354D70B7FFFAE638D9FAB4" box="[400,493,1293,1319]" pageId="88" pageNumber="89" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BA336E4AA354D70B6F8FAC4381AFADB" blockId="88.[151,493,1293,1352]" box="[151,302,1327,1352]" pageId="88" pageNumber="89">
(
<figureCitation id="13272A61AA354D70B6F0FAC43811FADB" box="[159,293,1327,1352]" captionStart-0="FIGURE39" captionStart-1="FIGURE40" captionStart-2="FIGURE 41" captionStartId-0="90.[151,250,1930,1952]" captionStartId-1="91.[151,250,1970,1992]" captionStartId-2="93.[151,250,1933,1955]" captionTargetBox-0="[209,1363,189,1876]" captionTargetBox-1="[222,1351,213,1933]" captionTargetBox-2="[171,1430,646,1904]" captionTargetId-0="figure@90.[185,1373,181,1895]" captionTargetId-1="figure@91.[212,1374,193,1949]" captionTargetId-2="figure@93.[151,1436,619,1926]" captionTargetPageId-0="90" captionTargetPageId-1="91" captionTargetPageId-2="93" captionText-0="FIGURE39. Caligus neoaricolus sp. nov. paratype female.A, habitus, dorsal;B, antennule, ventral; C, antenna, post-antennal process and maxillule drawn in situ; D, mandible; E, tip of maxilla; F, maxilliped; G, sternal furca; H, left corner of genital complex showing fifth leg, ventral. Scale bars:1.0 mm on A, 200 µm on B, C, G, 250 µm onF, 100 µm on D, E, H." captionText-1="FIGURE40.Caligus neoaricolus sp. nov. paratype female.A, leg 1, anterior;B,exopodof leg2, anterior; C, leg 3, ventral; D, leg 4. Scale bars:200 µm on A, B, 0.5 mm on C, 250 µm on D." captionText-2="FIGURE 41. Caligus neoaricolus sp. nov. paratype male.A, habitus, dorsal; B, left corner of genital complex showing legs 5 and 6, ventral; C, antenna; D, post-antennal process, E, maxilliped. Scale bars: 1.0 mm on A, 250 µm on E, 200 µm on C, 100 µm on B, D." httpUri-0="https://zenodo.org/record/1203031/files/figure.png" httpUri-1="https://zenodo.org/record/1203033/files/figure.png" httpUri-2="https://zenodo.org/record/1203035/files/figure.png" pageId="88" pageNumber="89">Figs. 39–41</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C306656FAA354D70B6F8FA9C38C8F923" pageId="88" pageNumber="89" type="materials_examined">
<paragraph id="8BA336E4AA354D70B6F8FA9C3DF6FA6E" blockId="88.[151,1437,1399,2037]" pageId="88" pageNumber="89">
<emphasis id="B968EAF6AA354D70B6F8FA9C3873FA03" bold="true" box="[151,327,1399,1424]" pageId="88" pageNumber="89">Type material.</emphasis>
Holotype ♀, 14 paratype ♀♀, 12 paratype ♂♂, 1 immature ♀ paratype from 
<taxonomicName id="4C1C4D67AA354D70B2BBFA9238D5FA26" authority="(Kner &amp; Steindachner, 1867)" baseAuthorityName="Kner &amp; Steindachner" baseAuthorityYear="1867" class="Actinopterygii" family="Ariidae" genus="Neoarius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="88" pageNumber="89" phylum="Chordata" rank="species" species="graeffei">
<emphasis id="B968EAF6AA354D70B2BBFA923CA8FA03" box="[1236,1436,1400,1424]" italics="true" pageId="88" pageNumber="89">Neoarius graeffei</emphasis>
(Kner &amp; Steindachner, 1867)
</taxonomicName>
(TC17881) 
<date id="FFA21024AA354D70B401FA773BC3FA26" box="[622,759,1436,1461]" pageId="88" pageNumber="89" value="2016-07-04">4 July 2016</date>
, QM Reg. Nos Holotype ♀ W53085, 1 paratype ♀♀ and 1 immature ♀ paratype W53086, 6 paratype ♂♂ W53087; 7 paratype ♀♀ and 6 paratype ♂♂ NHMUK 2017.283– 292; 1 paratype ♂ from (TC17597) 
<date id="FFA21024AA354D70B446FA0F3BF5FA6E" box="[553,705,1508,1533]" pageId="88" pageNumber="89" value="2016-06-26">26 June 2016</date>
, 1 paratype ♂ from (TC17624) 
<date id="FFA21024AA354D70B249FA0F3D8BFA6E" box="[1062,1215,1508,1533]" pageId="88" pageNumber="89" value="2016-06-27">27 June 2016</date>
.
</paragraph>
<paragraph id="8BA336E4AA354D70B6A8F9EC3DFFF9FB" blockId="88.[151,1437,1399,2037]" pageId="88" pageNumber="89">
<emphasis id="B968EAF6AA354D70B6A8F9EC3B06F9B3" bold="true" box="[199,562,1543,1568]" pageId="88" pageNumber="89">Additional non-type material</emphasis>
. 1♀ from 
<taxonomicName id="4C1C4D67AA354D70B4D1F9E23D0EF9B3" authority="(Ogilby, 1898)" baseAuthorityName="Ogilby" baseAuthorityYear="1898" box="[702,1082,1543,1568]" class="Actinopterygii" family="Ariidae" genus="Netuma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="88" pageNumber="89" phylum="Chordata" rank="species" species="proxima">
<emphasis id="B968EAF6AA354D70B4D1F9E23AB4F9B3" box="[702,896,1545,1568]" italics="true" pageId="88" pageNumber="89">Netuma proxima</emphasis>
(Ogilby, 1898)
</taxonomicName>
(TC18830) 0 
<date id="FFA21024AA354D70B28BF9E33CACF9B3" box="[1252,1432,1544,1568]" pageId="88" pageNumber="89" value="2017-08-01">1 August 2017</date>
, NHMUK 2017.294. 1♀ (possible contaminant) from 
<taxonomicName id="4C1C4D67AA354D70B49BF9C73DFEF9D7" authority="Quoy &amp; Gaimard, 1824" authorityName="Quoy &amp; Gaimard" authorityYear="1824" box="[756,1226,1580,1605]" class="Actinopterygii" family="Sillaginidae" genus="Sillago" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="88" pageNumber="89" phylum="Chordata" rank="species" species="maculata">
<emphasis id="B968EAF6AA354D70B49BF9C73A80F9D6" box="[756,948,1580,1605]" italics="true" pageId="88" pageNumber="89">Sillago maculata</emphasis>
Quoy &amp; Gaimard, 1824
</taxonomicName>
(TC17904) 
<date id="FFA21024AA354D70B338F9C739E3F9FB" pageId="88" pageNumber="89" value="2016-07-05">5 July 2016</date>
;1♂ incomplete (possible contaminant) from 
<taxonomicName id="4C1C4D67AA354D70B4A7F9BB3A03F9F4" authority="sp" authorityName="Sp" box="[712,823,1616,1639]" class="Actinopterygii" family="Gerreidae" genus="Gerres" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="88" pageNumber="64" phylum="Chordata" rank="genus">
<emphasis id="B968EAF6AA354D70B4A7F9BB3A20F9F4" box="[712,788,1616,1639]" italics="true" pageId="88" pageNumber="89">Gerres</emphasis>
sp
</taxonomicName>
, juveniles (collected 
<date id="FFA21024AA354D70B24BF9A43D8BF9FB" box="[1060,1215,1615,1640]" pageId="88" pageNumber="89" value="2016-06-26">26 June 2016</date>
).
</paragraph>
<paragraph id="8BA336E4AA354D70B6A8F99F3A52F91E" blockId="88.[151,1437,1399,2037]" box="[199,870,1652,1677]" pageId="88" pageNumber="89">
<emphasis id="B968EAF6AA354D70B6A8F99F3873F91E" bold="true" box="[199,327,1652,1677]" pageId="88" pageNumber="89">
<typeStatus id="54A78846AA354D70B6A8F99F3836F91E" box="[199,258,1652,1677]" pageId="88" pageNumber="89">Type</typeStatus>
Host.
</emphasis>
<taxonomicName id="4C1C4D67AA354D70B721F99E3A56F91E" authority="(Kner &amp; Steindachner, 1867)" baseAuthorityName="Kner &amp; Steindachner" baseAuthorityYear="1867" box="[334,866,1652,1677]" class="Actinopterygii" family="Ariidae" genus="Neoarius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="88" pageNumber="89" phylum="Chordata" rank="species" species="graeffei">
<emphasis id="B968EAF6AA354D70B721F99E3B27F91F" box="[334,531,1652,1676]" italics="true" pageId="88" pageNumber="89">Neoarius graeffei</emphasis>
(Kner &amp; Steindachner, 1867)
</taxonomicName>
.
</paragraph>
<paragraph id="8BA336E4AA354D70B6A8F97C38C8F923" blockId="88.[151,1437,1399,2037]" box="[199,508,1687,1712]" pageId="88" pageNumber="89">
<emphasis id="B968EAF6AA354D70B6A8F97C386DF923" bold="true" box="[199,345,1687,1712]" pageId="88" pageNumber="89">Site on host.</emphasis>
Body surface.
</paragraph>
</subSubSection>
<subSubSection id="C306656FAA354D70B6A8F9573D3EF946" box="[199,1034,1724,1749]" pageId="88" pageNumber="89" type="etymology">
<paragraph id="8BA336E4AA354D70B6A8F9573D3EF946" blockId="88.[151,1437,1399,2037]" box="[199,1034,1724,1749]" pageId="88" pageNumber="89">
<emphasis id="B968EAF6AA354D70B6A8F957387AF946" bold="true" box="[199,334,1724,1749]" pageId="88" pageNumber="89">Etymology.</emphasis>
The name of this species alludes to the genus of the 
<typeStatus id="54A78846AA354D70B5F3F9563AF8F946" box="[924,972,1725,1749]" pageId="88" pageNumber="89">type</typeStatus>
host.
</paragraph>
</subSubSection>
<subSubSection id="C306656FAA354D74B6A8F9343843FD5E" lastPageId="92" lastPageNumber="93" pageId="88" pageNumber="89" type="description">
<paragraph id="8BA336E4AA354D71B6A8F9343AC0FF23" blockId="88.[151,1437,1399,2037]" lastBlockId="89.[151,1437,151,1977]" lastPageId="89" lastPageNumber="90" pageId="88" pageNumber="89">
<emphasis id="B968EAF6AA354D70B6A8F934386DF96B" bold="true" box="[199,345,1759,1784]" pageId="88" pageNumber="89">Description.</emphasis>
Adult female (
<figureCitation id="13272A61AA354D70B469F9343B5AF96B" box="[518,622,1759,1784]" captionStart="FIGURE39" captionStartId="90.[151,250,1930,1952]" captionTargetBox="[209,1363,189,1876]" captionTargetId="figure@90.[185,1373,181,1895]" captionTargetPageId="90" captionText="FIGURE39. Caligus neoaricolus sp. nov. paratype female.A, habitus, dorsal;B, antennule, ventral; C, antenna, post-antennal process and maxillule drawn in situ; D, mandible; E, tip of maxilla; F, maxilliped; G, sternal furca; H, left corner of genital complex showing fifth leg, ventral. Scale bars:1.0 mm on A, 200 µm on B, C, G, 250 µm onF, 100 µm on D, E, H." httpUri="https://zenodo.org/record/1203031/files/figure.png" pageId="88" pageNumber="89">Fig. 39A</figureCitation>
) mean body length including caudal rami 
<quantity id="4CE49B01AA354D70B23BF9343D8EF964" box="[1108,1210,1759,1784]" metricMagnitude="-3" metricUnit="m" metricValue="5.91" pageId="88" pageNumber="89" unit="mm" value="5.91">5.91 mm</quantity>
(range 
<quantity id="4CE49B01AA354D70B37CF93439F0F88F" metricMagnitude="-3" metricUnit="m" metricValue="5.985" metricValueMax="6.24" metricValueMin="5.73" pageId="88" pageNumber="89" unit="mm" value="5.985" valueMax="6.24" valueMin="5.73">5.73 to 6.24 mm</quantity>
) (based on 
<specimenCount id="9D1AFD6DAA354D70B724F8EF38DDF88E" box="[331,489,1796,1821]" pageId="88" pageNumber="89" type="generic">10 specimens</specimenCount>
). Cephalothorax about 1.18 times longer than wide; comprising about 54% of total body length. Free margin of thoracic portion of dorsal cephalothoracic shield extending posteriorly just beyond rear margins of lateral portions. Lunules present ventrally on frontal plates. Genital complex 1.13 times longer than wide; narrowing slightly anteriorly and with linear lateral margins (
<figureCitation id="13272A61AA354D70B5CCF8843D3BF81B" box="[931,1039,1903,1928]" captionStart="FIGURE39" captionStartId="90.[151,250,1930,1952]" captionTargetBox="[209,1363,189,1876]" captionTargetId="figure@90.[185,1373,181,1895]" captionTargetPageId="90" captionText="FIGURE39. Caligus neoaricolus sp. nov. paratype female.A, habitus, dorsal;B, antennule, ventral; C, antenna, post-antennal process and maxillule drawn in situ; D, mandible; E, tip of maxilla; F, maxilliped; G, sternal furca; H, left corner of genital complex showing fifth leg, ventral. Scale bars:1.0 mm on A, 200 µm on B, C, G, 250 µm onF, 100 µm on D, E, H." httpUri="https://zenodo.org/record/1203031/files/figure.png" pageId="88" pageNumber="89">Fig. 39A</figureCitation>
). Genital complex about 1.4 times longer than abdomen. Abdomen 1-segmented, about 2.0 times longer than wide; carrying paired caudal rami distally; anal slit terminal. Caudal rami longer than wide, measured at midpoints of margins. Each ramus armed with short hirsute seta at inner distal angle, slightly longer hirsute seta at outer distal angle, minute hirsute seta located just ventral to outer distal seta, and 3 plumose setae on distal margin.
</paragraph>
<paragraph id="8BA336E4AA344D71B6A8FF573AB5FE3E" blockId="89.[151,1437,151,1977]" pageId="89" pageNumber="90">
Antennule (
<figureCitation id="13272A61AA344D71B724FF57389AFF47" box="[331,430,188,213]" captionStart="FIGURE39" captionStartId="90.[151,250,1930,1952]" captionTargetBox="[209,1363,189,1876]" captionTargetId="figure@90.[185,1373,181,1895]" captionTargetPageId="90" captionText="FIGURE39. Caligus neoaricolus sp. nov. paratype female.A, habitus, dorsal;B, antennule, ventral; C, antenna, post-antennal process and maxillule drawn in situ; D, mandible; E, tip of maxilla; F, maxilliped; G, sternal furca; H, left corner of genital complex showing fifth leg, ventral. Scale bars:1.0 mm on A, 200 µm on B, C, G, 250 µm onF, 100 µm on D, E, H." httpUri="https://zenodo.org/record/1203031/files/figure.png" pageId="89" pageNumber="90">Fig. 39B</figureCitation>
) 2-segmented; proximal segment with 25 plumose setae along anteroventral margin and 2 setae located dorsally; distal segment bearing 12 elements (10 setae plus 2 aesthetascs) around apex, plus isolated seta on posterior margin. Antenna (
<figureCitation id="13272A61AA344D71B439FEEF3BF1FE8F" box="[598,709,260,285]" captionStart="FIGURE39" captionStartId="90.[151,250,1930,1952]" captionTargetBox="[209,1363,189,1876]" captionTargetId="figure@90.[185,1373,181,1895]" captionTargetPageId="90" captionText="FIGURE39. Caligus neoaricolus sp. nov. paratype female.A, habitus, dorsal;B, antennule, ventral; C, antenna, post-antennal process and maxillule drawn in situ; D, mandible; E, tip of maxilla; F, maxilliped; G, sternal furca; H, left corner of genital complex showing fifth leg, ventral. Scale bars:1.0 mm on A, 200 µm on B, C, G, 250 µm onF, 100 µm on D, E, H." httpUri="https://zenodo.org/record/1203031/files/figure.png" pageId="89" pageNumber="90">Fig. 39C</figureCitation>
) comprising proximal segment with small sharply-pointed, posteriorly-directed spinous process; middle segment subrectangular, unarmed; terminal segment forming strongly curved claw with proximal sclerotized swelling bearing small seta, and armed with stout distal seta. Post-antennal process (
<figureCitation id="13272A61AA344D71B76BFE843845FE1B" box="[260,369,367,392]" captionStart="FIGURE39" captionStartId="90.[151,250,1930,1952]" captionTargetBox="[209,1363,189,1876]" captionTargetId="figure@90.[185,1373,181,1895]" captionTargetPageId="90" captionText="FIGURE39. Caligus neoaricolus sp. nov. paratype female.A, habitus, dorsal;B, antennule, ventral; C, antenna, post-antennal process and maxillule drawn in situ; D, mandible; E, tip of maxilla; F, maxilliped; G, sternal furca; H, left corner of genital complex showing fifth leg, ventral. Scale bars:1.0 mm on A, 200 µm on B, C, G, 250 µm onF, 100 µm on D, E, H." httpUri="https://zenodo.org/record/1203031/files/figure.png" pageId="89" pageNumber="90">Fig. 39C</figureCitation>
) weakly curved, ornamented with 2 multisensillate papillae on basal part; plus single multisensillate papilla on adjacent ventral cephalothoracic surface.
</paragraph>
<paragraph id="8BA336E4AA344D71B6A8FE5C3ACEFD34" blockId="89.[151,1437,151,1977]" pageId="89" pageNumber="90">
Mandible of typical stylet-like structure, with 12 marginal teeth (
<figureCitation id="13272A61AA344D71B5C4FE5C3D21FE43" box="[939,1045,439,464]" captionStart="FIGURE39" captionStartId="90.[151,250,1930,1952]" captionTargetBox="[209,1363,189,1876]" captionTargetId="figure@90.[185,1373,181,1895]" captionTargetPageId="90" captionText="FIGURE39. Caligus neoaricolus sp. nov. paratype female.A, habitus, dorsal;B, antennule, ventral; C, antenna, post-antennal process and maxillule drawn in situ; D, mandible; E, tip of maxilla; F, maxilliped; G, sternal furca; H, left corner of genital complex showing fifth leg, ventral. Scale bars:1.0 mm on A, 200 µm on B, C, G, 250 µm onF, 100 µm on D, E, H." httpUri="https://zenodo.org/record/1203031/files/figure.png" pageId="89" pageNumber="90">Fig. 39D</figureCitation>
). Maxillule (
<figureCitation id="13272A61AA344D71B2C7FE5C3C39FE43" box="[1192,1293,439,464]" captionStart="FIGURE39" captionStartId="90.[151,250,1930,1952]" captionTargetBox="[209,1363,189,1876]" captionTargetId="figure@90.[185,1373,181,1895]" captionTargetPageId="90" captionText="FIGURE39. Caligus neoaricolus sp. nov. paratype female.A, habitus, dorsal;B, antennule, ventral; C, antenna, post-antennal process and maxillule drawn in situ; D, mandible; E, tip of maxilla; F, maxilliped; G, sternal furca; H, left corner of genital complex showing fifth leg, ventral. Scale bars:1.0 mm on A, 200 µm on B, C, G, 250 µm onF, 100 µm on D, E, H." httpUri="https://zenodo.org/record/1203031/files/figure.png" pageId="89" pageNumber="90">Fig. 39C</figureCitation>
) comprising anterior papilla bearing 3 small naked setae and slender, tapering posterior process. Maxilla comprising elongate syncoxa and basis: syncoxa unarmed but bearing opening of maxillary gland; basis bearing subapical membranous flabellum on anterior margin, and terminating in 2 subequal, claw-like elements (calamus and canna) (
<figureCitation id="13272A61AA344D71B347FDCF3CA4FDAF" box="[1320,1424,548,573]" captionStart="FIGURE39" captionStartId="90.[151,250,1930,1952]" captionTargetBox="[209,1363,189,1876]" captionTargetId="figure@90.[185,1373,181,1895]" captionTargetPageId="90" captionText="FIGURE39. Caligus neoaricolus sp. nov. paratype female.A, habitus, dorsal;B, antennule, ventral; C, antenna, post-antennal process and maxillule drawn in situ; D, mandible; E, tip of maxilla; F, maxilliped; G, sternal furca; H, left corner of genital complex showing fifth leg, ventral. Scale bars:1.0 mm on A, 200 µm on B, C, G, 250 µm onF, 100 µm on D, E, H." httpUri="https://zenodo.org/record/1203031/files/figure.png" pageId="89" pageNumber="90">Fig. 39E</figureCitation>
): calamus just longer than canna, ornamented with strips of serrated membrane arranged obliquely around surface; canna with linear strips of serrated membrane. Maxilliped subchelate (
<figureCitation id="13272A61AA344D71B5D0FD873D12FD17" box="[959,1062,620,645]" captionStart="FIGURE39" captionStartId="90.[151,250,1930,1952]" captionTargetBox="[209,1363,189,1876]" captionTargetId="figure@90.[185,1373,181,1895]" captionTargetPageId="90" captionText="FIGURE39. Caligus neoaricolus sp. nov. paratype female.A, habitus, dorsal;B, antennule, ventral; C, antenna, post-antennal process and maxillule drawn in situ; D, mandible; E, tip of maxilla; F, maxilliped; G, sternal furca; H, left corner of genital complex showing fifth leg, ventral. Scale bars:1.0 mm on A, 200 µm on B, C, G, 250 µm onF, 100 µm on D, E, H." httpUri="https://zenodo.org/record/1203031/files/figure.png" pageId="89" pageNumber="90">Fig. 39F</figureCitation>
); slender proximal segment with fine transverse ridges on myxal surface; distal subchela with small inner seta.
</paragraph>
<paragraph id="8BA336E4AA344D71B6A8FD5F3AB6FD5E" blockId="89.[151,1437,151,1977]" box="[199,898,692,717]" pageId="89" pageNumber="90">
Sternal furca (
<figureCitation id="13272A61AA344D71B70AFD5F38FFFD5F" box="[357,459,692,717]" captionStart="FIGURE39" captionStartId="90.[151,250,1930,1952]" captionTargetBox="[209,1363,189,1876]" captionTargetId="figure@90.[185,1373,181,1895]" captionTargetPageId="90" captionText="FIGURE39. Caligus neoaricolus sp. nov. paratype female.A, habitus, dorsal;B, antennule, ventral; C, antenna, post-antennal process and maxillule drawn in situ; D, mandible; E, tip of maxilla; F, maxilliped; G, sternal furca; H, left corner of genital complex showing fifth leg, ventral. Scale bars:1.0 mm on A, 200 µm on B, C, G, 250 µm onF, 100 µm on D, E, H." httpUri="https://zenodo.org/record/1203031/files/figure.png" pageId="89" pageNumber="90">Fig. 39G</figureCitation>
) with weakly divergent, tapering tines.
</paragraph>
<paragraph id="8BA336E4AA344D71B6A8FD3C3C02FC5B" blockId="89.[151,1437,151,1977]" pageId="89" pageNumber="90">
First swimming leg pair (
<figureCitation id="13272A61AA344D71B78CFD3C3B7DFD63" box="[483,585,727,752]" captionStart="FIGURE40" captionStartId="91.[151,250,1970,1992]" captionTargetBox="[222,1351,213,1933]" captionTargetId="figure@91.[212,1374,193,1949]" captionTargetPageId="91" captionText="FIGURE40.Caligus neoaricolus sp. nov. paratype female.A, leg 1, anterior;B,exopodof leg2, anterior; C, leg 3, ventral; D, leg 4. Scale bars:200 µm on A, B, 0.5 mm on C, 250 µm on D." httpUri="https://zenodo.org/record/1203033/files/figure.png" pageId="89" pageNumber="90">Fig. 40A</figureCitation>
) with sympods joined by slender intercoxal sclerite; sympod with inner and outer plumose setae; endopod represented by unarmed process on posterior margin of basis, bearing minute vestiges of 2 setae apically. Exopod 2-segmented; directed laterally and forming main axis of leg; first segment robust, about 2.5 times longer than wide and armed with small outer (anterior) spine and ornamented with setule row along mid-section of posterior margin; second segment armed with 3 long plumose setae along posterior margin and 4 distal elements. Distal margin elements as follows: spine 1 (anterior-most) longest and naked; spine 2 longer than spine 3, each with accessory process; seta 4 twice as long as spine 1 and longer than segment.
</paragraph>
<paragraph id="8BA336E4AA344D71B6A8FC3F3A9BFAA3" blockId="89.[151,1437,151,1977]" pageId="89" pageNumber="90">
Second leg biramous, with flattened protopodal segments and 3-segmented rami. Coxae of leg pair joined by plate-like intercoxal sclerite bearing marginal membrane posteriorly; armed with plumose seta and surface sensilla. Basis armed with outer naked seta; ornamented with surface sensilla, marginal membrane posteriorly, and flap of membrane anteriorly, reflexed back over dorsal surface of segment. Exopodal segments 1 and 2 each with inner plumose seta and large reflexed outer spines extending obliquely across ventral surface of ramus (
<figureCitation id="13272A61AA344D71B289FB8F3C7AFBEF" box="[1254,1358,1124,1149]" captionStart="FIGURE40" captionStartId="91.[151,250,1970,1992]" captionTargetBox="[222,1351,213,1933]" captionTargetId="figure@91.[212,1374,193,1949]" captionTargetPageId="91" captionText="FIGURE40.Caligus neoaricolus sp. nov. paratype female.A, leg 1, anterior;B,exopodof leg2, anterior; C, leg 3, ventral; D, leg 4. Scale bars:200 µm on A, B, 0.5 mm on C, 250 µm on D." httpUri="https://zenodo.org/record/1203033/files/figure.png" pageId="89" pageNumber="90">Fig. 40B</figureCitation>
); outer spines ornamented with bilateral membranous strips; first exopodal segment bearing flap of membrane anteriorly, reflexed back over dorsal surface; segment 3 with 2 outer spines, proximal spine small, distal spine ornamented with membrane; apical spine with marginal membrane laterally and pinnules medially, and 5 inner plumose setae. Endopodal segments 1 and 2 armed with 1 and 2 inner plumose setae respectively; segment 3 with 6 plumose setae; outer margins of all endopodal segments ornamented with fine setules.
</paragraph>
<paragraph id="8BA336E4AA344D71B6A8FAD73D5BF973" blockId="89.[151,1437,151,1977]" pageId="89" pageNumber="90">
Third leg pair (
<figureCitation id="13272A61AA344D71B71FFAD738E0FAC7" box="[368,468,1340,1365]" captionStart="FIGURE40" captionStartId="91.[151,250,1970,1992]" captionTargetBox="[222,1351,213,1933]" captionTargetId="figure@91.[212,1374,193,1949]" captionTargetPageId="91" captionText="FIGURE40.Caligus neoaricolus sp. nov. paratype female.A, leg 1, anterior;B,exopodof leg2, anterior; C, leg 3, ventral; D, leg 4. Scale bars:200 µm on A, B, 0.5 mm on C, 250 µm on D." httpUri="https://zenodo.org/record/1203033/files/figure.png" pageId="89" pageNumber="90">Fig. 40C</figureCitation>
) forming flattened plate closing posterior part of cephalothoracic sucker, as typical for genus. Protopodal part flattened joined by plate-like, intercoxal sclerite forming apron, ornamented with marginal membrane posteriorly and along lateral margin anterior to exopod; bearing inner plumose seta at junction with intercoxal plate, and outer plumose seta dorsal to base of exopod; sensillae located adjacent to inner coxal seta and adjacent to origin of endopod. Exopod indistinctly 3-segmented; first segment armed with straight outer claw directed parallel with long axis of ramus and overlying lateral margin, lacking inner seta; second and third segments incompletely separated by partial suture; armed with slender outer spine and inner plumose seta derived from second segment and 3 outer spiniform elements and 4 inner plumose setae (derived from third segment); outer margins of segments 2 and 3 ornamented with rows of slender setules. Endopod 2-segmented; first segment expanded to form flap-like velum closing off space between rami, velum ornamented with row of fine setules along free margin; segment armed with inner plumose seta; compound distal segment with expanded and setulate lateral margin; bearing with 6 setal elements increasing in length from outermost to innermost.
</paragraph>
<paragraph id="8BA336E4AA344D71B6A8F9073A2CF8E3" blockId="89.[151,1437,151,1977]" pageId="89" pageNumber="90">
Fourth leg (
<figureCitation id="13272A61AA344D71B734F90738FEF897" box="[347,458,1772,1797]" captionStart="FIGURE40" captionStartId="91.[151,250,1970,1992]" captionTargetBox="[222,1351,213,1933]" captionTargetId="figure@91.[212,1374,193,1949]" captionTargetPageId="91" captionText="FIGURE40.Caligus neoaricolus sp. nov. paratype female.A, leg 1, anterior;B,exopodof leg2, anterior; C, leg 3, ventral; D, leg 4. Scale bars:200 µm on A, B, 0.5 mm on C, 250 µm on D." httpUri="https://zenodo.org/record/1203033/files/figure.png" pageId="89" pageNumber="90">Fig. 40D</figureCitation>
) 3-segmented, comprising slender protopodal segment and 2-segmented exopod; exopodal segments separated by oblique articulation: protopodal segment armed with outer plumose seta; first exopodal segment armed with short outer spine; second segment armed with short lateral spine and 3 unequal spines along distal margin; all spines with pecten at base.
</paragraph>
<paragraph id="8BA336E4AA344D71B6A8F8973D55F82B" blockId="89.[151,1437,151,1977]" pageId="89" pageNumber="90">
Fifth leg located posterolaterally on margin of genital complex, represented by anterior plumose seta on papilla and posterior papilla bearing 2 unequal plumose setae representing exopod (
<figureCitation id="13272A61AA344D71B583F84B3D60F82B" box="[1004,1108,1952,1977]" captionStart="FIGURE39" captionStartId="90.[151,250,1930,1952]" captionTargetBox="[209,1363,189,1876]" captionTargetId="figure@90.[185,1373,181,1895]" captionTargetPageId="90" captionText="FIGURE39. Caligus neoaricolus sp. nov. paratype female.A, habitus, dorsal;B, antennule, ventral; C, antenna, post-antennal process and maxillule drawn in situ; D, mandible; E, tip of maxilla; F, maxilliped; G, sternal furca; H, left corner of genital complex showing fifth leg, ventral. Scale bars:1.0 mm on A, 200 µm on B, C, G, 250 µm onF, 100 µm on D, E, H." httpUri="https://zenodo.org/record/1203031/files/figure.png" pageId="89" pageNumber="90">Fig. 39H</figureCitation>
).
</paragraph>
<caption id="DF63666CAA374D72B6F8F8613C1DF84E" httpUri="https://zenodo.org/record/1203031/files/figure.png" pageId="90" pageNumber="91" startId="90.[151,250,1930,1952]" targetBox="[209,1363,189,1876]" targetPageId="90">
<paragraph id="8BA336E4AA374D72B6F8F8613C1DF84E" blockId="90.[151,1436,1930,2013]" pageId="90" pageNumber="91">
<emphasis id="B968EAF6AA374D72B6F8F861382BF80C" bold="true" box="[151,287,1930,1952]" pageId="90" pageNumber="91">FIGURE 39.</emphasis>
<taxonomicName id="4C1C4D67AA374D72B74AF86138C5F833" authority="Boxshall, 2018" authorityName="Boxshall" authorityYear="2018" box="[293,497,1930,1952]" class="Maxillopoda" family="Caligidae" genus="Caligus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siphonostomatoida" pageId="90" pageNumber="91" phylum="Arthropoda" rank="species" species="neoaricolus" status="sp. nov.">
<emphasis id="B968EAF6AA374D72B74AF86138C5F833" box="[293,497,1930,1952]" italics="true" pageId="90" pageNumber="91">Caligus neoaricolus</emphasis>
</taxonomicName>
<emphasis id="B968EAF6AA374D72B798F8613B7CF80C" bold="true" box="[503,584,1930,1951]" pageId="90" pageNumber="91">
<taxonomicNameLabel id="A25B578DAA374D72B798F8613B7CF80C" box="[503,584,1930,1951]" pageId="90" pageNumber="91" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
paratype female. A, habitus, dorsal; B, antennule, ventral; C, antenna, post-antennal process and maxillule drawn 
<emphasis id="B968EAF6AA374D72B7A7F8413B3DF82C" box="[456,521,1962,1983]" italics="true" pageId="90" pageNumber="91">in situ</emphasis>
; D, mandible; E, tip of maxilla; F, maxilliped; G, sternal furca; H, left corner of genital complex showing fifth leg, ventral. Scale bars: 1.0 mm on A, 200 µm on B, C, G, 250 µm on F, 100 µm on D, E, H.
</paragraph>
</caption>
<caption id="DF63666CAA364D73B6F8F8593A2DF87A" httpUri="https://zenodo.org/record/1203033/files/figure.png" pageId="91" pageNumber="92" startId="91.[151,250,1970,1992]" targetBox="[222,1351,213,1933]" targetPageId="91">
<paragraph id="8BA336E4AA364D73B6F8F8593A2DF87A" blockId="91.[151,1435,1970,2025]" pageId="91" pageNumber="92">
<emphasis id="B968EAF6AA364D73B6F8F859382BF854" bold="true" box="[151,287,1970,1992]" pageId="91" pageNumber="92">FIGURE 40.</emphasis>
<taxonomicName id="4C1C4D67AA364D73B74BF85938C5F85B" authority="Boxshall, 2018" authorityName="Boxshall" authorityYear="2018" box="[292,497,1970,1992]" class="Maxillopoda" family="Caligidae" genus="Caligus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siphonostomatoida" pageId="91" pageNumber="92" phylum="Arthropoda" rank="species" species="neoaricolus" status="sp. nov.">
<emphasis id="B968EAF6AA364D73B74BF85938C5F85B" box="[292,497,1970,1992]" italics="true" pageId="91" pageNumber="92">Caligus neoaricolus</emphasis>
</taxonomicName>
<emphasis id="B968EAF6AA364D73B798F8593B73F854" bold="true" box="[503,583,1970,1991]" pageId="91" pageNumber="92">
<taxonomicNameLabel id="A25B578DAA364D73B798F8593B73F854" box="[503,583,1970,1991]" pageId="91" pageNumber="92" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
paratype female. A, leg 1, anterior; B, exopod of leg 2, anterior; C, leg 3, ventral; D, leg 4. Scale bars: 200 µm on A, B, 0.5 mm on C, 250 µm on D.
</paragraph>
</caption>
<paragraph id="8BA336E4AA314D74B6A8FF7C3BB8FEF6" blockId="92.[151,1437,151,2013]" pageId="92" pageNumber="93">
Male (
<figureCitation id="13272A61AA314D74B77BFF7C384AFF23" box="[276,382,151,176]" captionStart="FIGURE 41" captionStartId="93.[151,250,1933,1955]" captionTargetBox="[171,1430,646,1904]" captionTargetId="figure@93.[151,1436,619,1926]" captionTargetPageId="93" captionText="FIGURE 41. Caligus neoaricolus sp. nov. paratype male.A, habitus, dorsal; B, left corner of genital complex showing legs 5 and 6, ventral; C, antenna; D, post-antennal process, E, maxilliped. Scale bars: 1.0 mm on A, 250 µm on E, 200 µm on C, 100 µm on B, D." httpUri="https://zenodo.org/record/1203035/files/figure.png" pageId="92" pageNumber="93">Fig. 41A</figureCitation>
) mean body length including caudal rami 
<quantity id="4CE49B01AA314D74B51CFF7C3AE9FF3C" box="[883,989,151,176]" metricMagnitude="-3" metricUnit="m" metricValue="5.62" pageId="92" pageNumber="93" unit="mm" value="5.62">5.62 mm</quantity>
, range 
<quantity id="4CE49B01AA314D74B259FF7C3DCFFF3C" box="[1078,1275,151,176]" metricMagnitude="-3" metricUnit="m" metricValue="5.7" metricValueMax="5.99" metricValueMin="5.41" pageId="92" pageNumber="93" unit="mm" value="5.7" valueMax="5.99" valueMin="5.41">5.41 to 5.99 mm</quantity>
(based on 
<specimenCount id="9D1AFD6DAA314D74B3EFFF733825FF46" pageId="92" pageNumber="93" type="generic">10 specimens</specimenCount>
). Cephalothorax subcircular as in female. Fourth pedigerous somite incompletely fused to genital complex. Genital complex about 1.2 times longer than wide, measured along the mid-line; with weakly convex lateral margins. Abdomen 2-segmented, separated from genital complex dorsally but fused ventrally. Caudal rami longer than wide; armed with short plumose seta at inner distal angle, 2 short plumose setae at outer distal angle, and 3 longer plumose setae on distal margin.
</paragraph>
<paragraph id="8BA336E4AA314D74B6A8FE843D3DFE67" blockId="92.[151,1437,151,2013]" pageId="92" pageNumber="93">
Antennule, mandible, maxillule and maxilla as in female. Antenna modified (
<figureCitation id="13272A61AA314D74B24BFE843DBFFE1B" box="[1060,1163,367,392]" captionStart="FIGURE 41" captionStartId="93.[151,250,1933,1955]" captionTargetBox="[171,1430,646,1904]" captionTargetId="figure@93.[151,1436,619,1926]" captionTargetPageId="93" captionText="FIGURE 41. Caligus neoaricolus sp. nov. paratype male.A, habitus, dorsal; B, left corner of genital complex showing legs 5 and 6, ventral; C, antenna; D, post-antennal process, E, maxilliped. Scale bars: 1.0 mm on A, 250 µm on E, 200 µm on C, 100 µm on B, D." httpUri="https://zenodo.org/record/1203035/files/figure.png" pageId="92" pageNumber="93">Fig. 41C</figureCitation>
); first segment elongate; second segment reflexed, swollen proximally with surface forming corrugated adhesion pads ventrally, plus corrugated distal swelling opposing tip of claw; distal segment forming short powerful apical claw armed with 2 setae proximally. Post-antennal process more robust than in female (
<figureCitation id="13272A61AA314D74B5FBFE373AC8FE67" box="[916,1020,476,501]" captionStart="FIGURE 41" captionStartId="93.[151,250,1933,1955]" captionTargetBox="[171,1430,646,1904]" captionTargetId="figure@93.[151,1436,619,1926]" captionTargetPageId="93" captionText="FIGURE 41. Caligus neoaricolus sp. nov. paratype male.A, habitus, dorsal; B, left corner of genital complex showing legs 5 and 6, ventral; C, antenna; D, post-antennal process, E, maxilliped. Scale bars: 1.0 mm on A, 250 µm on E, 200 µm on C, 100 µm on B, D." httpUri="https://zenodo.org/record/1203035/files/figure.png" pageId="92" pageNumber="93">Fig. 41D</figureCitation>
).
</paragraph>
<paragraph id="8BA336E4AA314D74B6A8FE1438F1FDAE" blockId="92.[151,1437,151,2013]" pageId="92" pageNumber="93">
Maxilliped (
<figureCitation id="13272A61AA314D74B73EFE143887FD8B" box="[337,435,511,536]" captionStart="FIGURE 41" captionStartId="93.[151,250,1933,1955]" captionTargetBox="[171,1430,646,1904]" captionTargetId="figure@93.[151,1436,619,1926]" captionTargetPageId="93" captionText="FIGURE 41. Caligus neoaricolus sp. nov. paratype male.A, habitus, dorsal; B, left corner of genital complex showing legs 5 and 6, ventral; C, antenna; D, post-antennal process, E, maxilliped. Scale bars: 1.0 mm on A, 250 µm on E, 200 µm on C, 100 µm on B, D." httpUri="https://zenodo.org/record/1203035/files/figure.png" pageId="92" pageNumber="93">Fig. 41E</figureCitation>
) with weakly inflated myxal margin on proximal segment; distal subchela longer relative to syncoxa, than in female.
</paragraph>
<paragraph id="8BA336E4AA314D74B6A8FDA33843FD5E" blockId="92.[151,1437,151,2013]" pageId="92" pageNumber="93">
Legs 5 and 6 (
<figureCitation id="13272A61AA314D74B700FDAC38E2FDF3" box="[367,470,583,608]" captionStart="FIGURE 41" captionStartId="93.[151,250,1933,1955]" captionTargetBox="[171,1430,646,1904]" captionTargetId="figure@93.[151,1436,619,1926]" captionTargetPageId="93" captionText="FIGURE 41. Caligus neoaricolus sp. nov. paratype male.A, habitus, dorsal; B, left corner of genital complex showing legs 5 and 6, ventral; C, antenna; D, post-antennal process, E, maxilliped. Scale bars: 1.0 mm on A, 250 µm on E, 200 µm on C, 100 µm on B, D." httpUri="https://zenodo.org/record/1203035/files/figure.png" pageId="92" pageNumber="93">Fig. 41B</figureCitation>
) each forming paired flattened processes posterolaterally on genital complex. Leg 5 represented by 3 plumose setae, 1 outer seta and 2 short inner setae representing exopod. Leg 6 represented by paired opercula closing off genital apertures; each armed with 1 long and 1 short plumose seta on distal margin, plus a minute spine.
</paragraph>
</subSubSection>
<subSubSection id="C306656FAA314D77B6A8FD3C3C11FED3" lastPageId="95" lastPageNumber="96" pageId="92" pageNumber="93" type="discussion">
<paragraph id="8BA336E4AA314D74B6A8FD3C3B90FBEE" blockId="92.[151,1437,151,2013]" pageId="92" pageNumber="93">
<emphasis id="B968EAF6AA314D74B6A8FD3C380FFD63" bold="true" box="[199,315,727,752]" pageId="92" pageNumber="93">Remarks.</emphasis>
This new species possesses a 3-segmented leg 4 bearing I and IV spines on the first and second exopodal segments, respectively. This is a common configuration shared with nearly 90 other 
<taxonomicName id="4C1C4D67AA314D74B28DFD173C0EFC86" authorityName="O. F. Muller" authorityYear="1785" box="[1250,1338,764,789]" class="Maxillopoda" family="Caligidae" genus="Caligus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siphonostomatoida" pageId="92" pageNumber="93" phylum="Arthropoda" rank="genus">
<emphasis id="B968EAF6AA314D74B28DFD173C0EFC86" box="[1250,1338,764,789]" italics="true" pageId="92" pageNumber="93">Caligus</emphasis>
</taxonomicName>
species, including the 14 members of the 
<taxonomicName id="4C1C4D67AA314D74B46AFCCB3BA0FCAB" authorityName="Dana" authorityYear="1852" box="[517,660,799,824]" class="Maxillopoda" family="Caligidae" genus="Caligus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siphonostomatoida" pageId="92" pageNumber="93" phylum="Arthropoda" rank="species" species="productus">
<emphasis id="B968EAF6AA314D74B46AFCCB3BA0FCAB" box="[517,660,799,824]" italics="true" pageId="92" pageNumber="93">C. productus</emphasis>
</taxonomicName>
-group which are characterized by loss or 
<taxonomicName id="4C1C4D67AA314D74B20AFCF43D93FCAB" baseAuthorityName="non Temminck &amp; Schlegel" baseAuthorityYear="1843" box="[1125,1191,799,824]" class="Actinopterygii" family="Sparidae" genus="Pagrosomus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="92" pageNumber="93" phylum="Chordata" rank="species" species="major">major</taxonomicName>
reduction of the three plumose setae on the posterior margin of the distal exopodal segment of leg 1. These can be eliminated from further comparison since these posterior margin setae are normally developed in the new species. Another 12 species, members of the 
<emphasis id="B968EAF6AA314D74B738FC6638F8FC36" box="[343,460,908,933]" italics="true" pageId="92" pageNumber="93">C. bonito-</emphasis>
group, all of which share the presence of large denticles on the outer margin of the second endopodal segment of leg 2, can be excluded from detailed comparison as the new species has an ornamentation of fine setules along this margin. The remaining pool of similar species is reduced to 30 by restricting further comparisons to species that have a female genital complex that is as long as or longer than wide, and an abdomen that is longer than wide. Allowing for individual variation in genital complex shape in the order of 10%, generates a short list of 13 species that exhibit a genital complex with a L:W ratio in the range of 1.0 to 1.2:1 for comparison with the new species (
<tableCitation id="C69E035FAA314D74B450FB8F3BA2FBEE" box="[575,662,1124,1149]" captionStart="TABLE 6" captionStartId="94.[119,196,784,804]" captionText="TABLE 6.Selectionof species ofCaligus characterised by a3-segmented leg4bearing 4 distal spineson thesecond exopodal segment,in combination with thesetalarmature on thedistal segment ofleg 1consisting of 3plumose posterior setae, and thepresence ofan ornamentationof fine setuleson the outermargin ofthe second endopodal segmentof leg2.[Abbreviations: = abdomen length,ABW = abdomen width,GCL = genital complexlength,GCW =genital complex width]." pageId="92" pageNumber="93">Table 6</tableCitation>
).
</paragraph>
<paragraph id="8BA336E4AA314D74B6A8FB6C3DD8FAC6" blockId="92.[151,1437,151,2013]" pageId="92" pageNumber="93">
<taxonomicName id="4C1C4D67AA314D74B6A8FB6C38BDFB33" authorityName="Kroyer" authorityYear="1863" box="[199,393,1159,1184]" class="Maxillopoda" family="Caligidae" genus="Caligus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siphonostomatoida" pageId="92" pageNumber="93" phylum="Arthropoda" rank="species" species="gurnardi">
<emphasis id="B968EAF6AA314D74B6A8FB6C38BDFB33" box="[199,393,1159,1184]" italics="true" pageId="92" pageNumber="93">Caligus gurnardi</emphasis>
</taxonomicName>
differs from the new species in having a genital complex that is 2.3 times longer than the abdomen (
<bibRefCitation id="EF8D4B15AA314D74B760FB473891FB56" box="[271,421,1196,1221]" pageId="92" pageNumber="93" refId="ref86442" refString="Parker, R. R. (1965) A review and redescription of Caligus gurnardi Kroyer, 1863 (Copepoda, Caligidae). Crustaceana, 9 (1), 93 - 103. https: // doi. org / 10.1163 / 156854065 X 00226" type="journal article">Parker, 1965</bibRefCitation>
), compared to only 1.5 times longer in the new species. Four species, 
<taxonomicName id="4C1C4D67AA314D74B28FFB463C45FB56" authorityName="Gunther" authorityYear="1861" box="[1248,1393,1196,1221]" class="Actinopterygii" family="Mugilidae" genus="Myxus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="92" pageNumber="93" phylum="Chordata" rank="species" species="elongatus">
<emphasis id="B968EAF6AA314D74B28FFB463C45FB56" box="[1248,1393,1196,1221]" italics="true" pageId="92" pageNumber="93">C. elongatus</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C1C4D67AA314D74B3EDFB463836FB7B" authorityName="Heegaard" authorityYear="1972" class="Maxillopoda" family="Caligidae" genus="Caligus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siphonostomatoida" pageId="92" pageNumber="93" phylum="Arthropoda" rank="species" species="tripedalis">
<emphasis id="B968EAF6AA314D74B3EDFB463836FB7B" italics="true" pageId="92" pageNumber="93">C. tripedalis</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C1C4D67AA314D74B77EFB3B38B2FB7B" authorityName="Venmathi Maran, Ohtsuka &amp; Shang" authorityYear="2012" box="[273,390,1232,1256]" class="Maxillopoda" family="Caligidae" genus="Caligus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siphonostomatoida" pageId="92" pageNumber="93" phylum="Arthropoda" rank="species" species="ogawai">
<emphasis id="B968EAF6AA314D74B77EFB3B38B2FB7B" box="[273,390,1232,1256]" italics="true" pageId="92" pageNumber="93">C. ogawai</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C1C4D67AA314D74B7D1FB3B3B47FB7B" authorityName="Wilson" authorityYear="1937" box="[446,627,1232,1256]" class="Maxillopoda" family="Caligidae" genus="Caligus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siphonostomatoida" pageId="92" pageNumber="93" phylum="Arthropoda" rank="species" species="tenuifurcatus">
<emphasis id="B968EAF6AA314D74B7D1FB3B3B47FB7B" box="[446,627,1232,1256]" italics="true" pageId="92" pageNumber="93">C. tenuifurcatus</emphasis>
</taxonomicName>
, all have a very broad abdomen (i.e. abdomen at least 50% of width of genital complex), whereas in 
<taxonomicName id="4C1C4D67AA314D74B78FFB1E3BB0FA9E" authority="Boxshall, 2018" authorityName="Boxshall" authorityYear="2018" box="[480,644,1268,1293]" class="Maxillopoda" family="Caligidae" genus="Caligus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siphonostomatoida" pageId="92" pageNumber="93" phylum="Arthropoda" rank="species" species="neoaricolus" status="sp. nov.">
<emphasis id="B968EAF6AA314D74B78FFB1E3BB0FA9E" box="[480,644,1268,1293]" italics="true" pageId="92" pageNumber="93">C. neoaricolus</emphasis>
</taxonomicName>
<emphasis id="B968EAF6AA314D74B4E4FB1F3BD1FA9E" bold="true" box="[651,741,1268,1293]" pageId="92" pageNumber="93">
<taxonomicNameLabel id="A25B578DAA314D74B4E4FB1F3BD1FA9E" box="[651,741,1268,1293]" pageId="92" pageNumber="93" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
the abdomen is only 38% of the width of the genital complex. Three of the remaining species, 
<taxonomicName id="4C1C4D67AA314D74B46AFAF33B9DFAA3" authorityName="Heegaard" authorityYear="1962" box="[517,681,1303,1328]" class="Maxillopoda" family="Caligidae" genus="Caligus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siphonostomatoida" pageId="92" pageNumber="93" phylum="Arthropoda" rank="species" species="longirostris">
<emphasis id="B968EAF6AA314D74B46AFAF33B9DFAA3" box="[517,681,1303,1328]" italics="true" pageId="92" pageNumber="93">C. longirostris</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C1C4D67AA314D74B4D7FAF33A2FFAA3" authorityName="Milne Edwards" authorityYear="1840" box="[696,795,1304,1328]" class="Maxillopoda" family="Caligidae" genus="Caligus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siphonostomatoida" pageId="92" pageNumber="93" phylum="Arthropoda" rank="species" species="rapax">
<emphasis id="B968EAF6AA314D74B4D7FAF33A2FFAA3" box="[696,795,1304,1328]" italics="true" pageId="92" pageNumber="93">C. rapax</emphasis>
</taxonomicName>
, and 
<taxonomicName id="4C1C4D67AA314D74B533FAF33D2EFAA3" authorityName="Wilson" authorityYear="1905" box="[860,1050,1303,1328]" class="Maxillopoda" family="Caligidae" genus="Caligus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siphonostomatoida" pageId="92" pageNumber="93" phylum="Arthropoda" rank="species" species="rufimaculatus">
<emphasis id="B968EAF6AA314D74B533FAF33D2EFAA3" box="[860,1050,1303,1328]" italics="true" pageId="92" pageNumber="93">C. rufimaculatus</emphasis>
</taxonomicName>
, have a relatively short abdomen, i.e., that is only 1.4 to 1.5 times longer than wide, compared to 2.0 times longer in the new species.
</paragraph>
<paragraph id="8BA336E4AA314D74B6A8FAB438DEF9BF" blockId="92.[151,1437,151,2013]" pageId="92" pageNumber="93">
The leg 4 of 
<taxonomicName id="4C1C4D67AA314D74B73CFA8B38FBFAEB" box="[339,463,1375,1400]" class="Maxillopoda" family="Caligidae" genus="Caligus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siphonostomatoida" pageId="92" pageNumber="93" phylum="Arthropoda" rank="species" species="chiastos">
<emphasis id="B968EAF6AA314D74B73CFA8B38FBFAEB" box="[339,463,1375,1400]" italics="true" pageId="92" pageNumber="93">C. chiastos</emphasis>
</taxonomicName>
(
<figureCitation id="13272A61AA314D74B7B2FAB43B0AFAEB" box="[477,574,1375,1400]" captionStart="FIGURE 24" captionStartId="53.[151,250,1909,1931]" captionTargetBox="[229,1389,202,1868]" captionTargetId="figure@53.[181,1405,185,1887]" captionTargetPageId="53" captionText="FIGURE 24. Caligus chiastos Lin &amp; Ho, 2003, female. A, habitus, dorsal; B, antenna, post-antennal process and maxillule drawn in situ; C, sternal furca; D, distal margin of exopod of leg 1; E, exopod of leg 2, showing well developed, curved proximalouter spine of third exopodal segment;F, leg 4.Scale bars:1.0 mm on A, 200 µm on F, 100 µm on C, E, 50 µm on B, D." httpUri="https://zenodo.org/record/1203001/files/figure.png" pageId="92" pageNumber="93">Fig. 24F</figureCitation>
) has a long outer spine on exopodal segment 1 that extends beyond the origin of the lateral spine on the distal segment. In contrast, the new species has a much shorter outer spine that does not reach halfway to the base of the lateral spine on the distal segment. Other differences include the smaller posterior process on the proximal segment of the antenna in the new species, the small size of the outer spines on the distal exopodal segment of leg 2, the shape of the sternal furca, and the relative lengths of spines 1 to 3 on the distal margin of the exopod of leg 1.
</paragraph>
<paragraph id="8BA336E4AA314D74B6A8F9DC3C5DF8BB" blockId="92.[151,1437,151,2013]" pageId="92" pageNumber="93">
<taxonomicName id="4C1C4D67AA314D74B6A8F9DC38B3F9C3" authorityName="Parker &amp; Margolis" authorityYear="1964" box="[199,391,1591,1616]" class="Maxillopoda" family="Caligidae" genus="Caligus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siphonostomatoida" pageId="92" pageNumber="93" phylum="Arthropoda" rank="species" species="clemensi">
<emphasis id="B968EAF6AA314D74B6A8F9DC38B3F9C3" box="[199,391,1591,1616]" italics="true" pageId="92" pageNumber="93">Caligus clemensi</emphasis>
</taxonomicName>
exhibits a similar arrangement of outer spines on the third exopodal segment of leg 2, to that of the new species: both have a minute pointed proximal spine and a longer blunt-tipped distal spine. The configuration of spines 1 to 3 on the distal margin of the exopod of leg 1 differs: in the new species spines 1 to 3 decrease in length in order from outer to inner, whereas in 
<taxonomicName id="4C1C4D67AA314D74B533F94E3AD0F92E" authorityName="Parker &amp; Margolis" authorityYear="1964" box="[860,996,1700,1725]" class="Maxillopoda" family="Caligidae" genus="Caligus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siphonostomatoida" pageId="92" pageNumber="93" phylum="Arthropoda" rank="species" species="clemensi">
<emphasis id="B968EAF6AA314D74B533F94E3AD0F92E" box="[860,996,1700,1725]" italics="true" pageId="92" pageNumber="93">C. clemensi</emphasis>
</taxonomicName>
spines 2 and 3, with their very long accessory processes, are both longer than spine 1 (
<bibRefCitation id="EF8D4B15AA314D74B4BDF92C3AC7F973" box="[722,1011,1735,1760]" pageId="92" pageNumber="93" refId="ref86544" refString="Parker, R. R. &amp; Margolis, L. (1964) A new species of parasitic copepod, Caligus clemensi sp. nov. (Caligoida, Caligidae) from pelagic fishes in the coastal waters of British Columbia. Journal of the Fisheries Research Board of Canada, 21, 873 - 889. https: // doi. org / 10.1139 / f 64 - 085" type="journal article">Parker &amp; Margolis, 1964</bibRefCitation>
). Another noticeable difference is the outer spines on the exopodal segments of leg 4: they are much smaller in the new species, not reaching the origin of the next most distal spine, whereas in 
<taxonomicName id="4C1C4D67AA314D74B42EF8FB3BF0F8BB" authorityName="Parker &amp; Margolis" authorityYear="1964" box="[577,708,1807,1832]" class="Maxillopoda" family="Caligidae" genus="Caligus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siphonostomatoida" pageId="92" pageNumber="93" phylum="Arthropoda" rank="species" species="clemensi">
<emphasis id="B968EAF6AA314D74B42EF8FB3BF0F8BB" box="[577,708,1807,1832]" italics="true" pageId="92" pageNumber="93">C. clemensi</emphasis>
</taxonomicName>
each of the outer spines reaches the origin of the next spine.
</paragraph>
<paragraph id="8BA336E4AA314D75B6A8F8DF3C35FF47" blockId="92.[151,1437,151,2013]" lastBlockId="93.[151,1436,151,573]" lastPageId="93" lastPageNumber="94" pageId="92" pageNumber="93">
The redescription of 
<taxonomicName id="4C1C4D67AA314D74B7D8F8DE3B7BF8DF" authorityName="Bere" authorityYear="1936" box="[439,591,1845,1868]" class="Maxillopoda" family="Caligidae" genus="Caligus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siphonostomatoida" pageId="92" pageNumber="93" phylum="Arthropoda" rank="species" species="praetextus">
<emphasis id="B968EAF6AA314D74B7D8F8DE3B7BF8DF" box="[439,591,1845,1868]" italics="true" pageId="92" pageNumber="93">C. praetextus</emphasis>
</taxonomicName>
by Cressey (1991) reveals that leg 1 has an unusually squat shape in this species: the first exopodal segment is only 1.7 times longer than wide compared to 2.5 times longer in the new species. Spines 2 and 3 on the distal margin of the exopod have marginal membrane extending apically from about the origin of the accessory process, producing a spatulate appearance (Cressey, 1991: Fig. 158). These spines lack marginal membrane in 
<taxonomicName id="4C1C4D67AA314D74B7F3F82E3B76F84E" authority="Boxshall, 2018" authorityName="Boxshall" authorityYear="2018" box="[412,578,1988,2013]" class="Maxillopoda" family="Caligidae" genus="Caligus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siphonostomatoida" pageId="92" pageNumber="93" phylum="Arthropoda" rank="species" species="neoaricolus" status="sp. nov.">
<emphasis id="B968EAF6AA314D74B7F3F82E3B76F84E" box="[412,578,1988,2013]" italics="true" pageId="92" pageNumber="93">C. neoaricolus</emphasis>
</taxonomicName>
<emphasis id="B968EAF6AA314D74B426F82F3B90F84E" bold="true" box="[585,676,1988,2013]" pageId="92" pageNumber="93">
<taxonomicNameLabel id="A25B578DAA314D74B426F82F3B90F84E" box="[585,676,1988,2013]" pageId="92" pageNumber="93" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
The proximal outer spine on the third exopodal segment of leg 2 is longer than the distal spine and curves across its surface, just as highlighted for 
<taxonomicName id="4C1C4D67AA304D75B247FF733DB9FF23" box="[1064,1165,151,176]" class="Maxillopoda" family="Caligidae" genus="Caligus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siphonostomatoida" pageId="93" pageNumber="94" phylum="Arthropoda" rank="species" species="chiast">
<emphasis id="B968EAF6AA304D75B247FF733DB9FF23" box="[1064,1165,151,176]" italics="true" pageId="93" pageNumber="94">C. chiast</emphasis>
</taxonomicName>
os by 
<bibRefCitation id="EF8D4B15AA304D75B2BDFF7C3CA3FF23" box="[1234,1431,151,176]" pageId="93" pageNumber="94" refId="ref84973" refString="Lin, C. - L. &amp; &amp; Ho, J. - S. (2003) Two species of rare sea lice (Copepoda, Caligidae) parasitic on marine fishes of Taiwan. Journal of the Fisheries Society of Taiwan, 29, 313 - 332." type="journal article">Lin &amp; Ho (2003)</bibRefCitation>
, whereas in 
<taxonomicName id="4C1C4D67AA304D75B779FF56388FFF46" authority="Boxshall, 2018" authorityName="Boxshall" authorityYear="2018" box="[278,443,188,213]" class="Maxillopoda" family="Caligidae" genus="Caligus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siphonostomatoida" pageId="93" pageNumber="94" phylum="Arthropoda" rank="species" species="neoaricolus" status="sp. nov.">
<emphasis id="B968EAF6AA304D75B779FF56388FFF46" box="[278,443,188,213]" italics="true" pageId="93" pageNumber="94">C. neoaricolus</emphasis>
</taxonomicName>
<emphasis id="B968EAF6AA304D75B7ADFF573B28FF46" bold="true" box="[450,540,188,213]" pageId="93" pageNumber="94">
<taxonomicNameLabel id="A25B578DAA304D75B7ADFF573B28FF46" box="[450,540,188,213]" pageId="93" pageNumber="94" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
the proximal spine is only half as long as the distal (cf. 
<figureCitation id="13272A61AA304D75B2E0FF573DC1FF47" box="[1167,1269,188,213]" captionStart="FIGURE40" captionStartId="91.[151,250,1970,1992]" captionTargetBox="[222,1351,213,1933]" captionTargetId="figure@91.[212,1374,193,1949]" captionTargetPageId="91" captionText="FIGURE40.Caligus neoaricolus sp. nov. paratype female.A, leg 1, anterior;B,exopodof leg2, anterior; C, leg 3, ventral; D, leg 4. Scale bars:200 µm on A, B, 0.5 mm on C, 250 µm on D." httpUri="https://zenodo.org/record/1203033/files/figure.png" pageId="93" pageNumber="94">Fig. 40B</figureCitation>
).
</paragraph>
<paragraph id="8BA336E4AA304D75B6A8FF0B3B57FDAE" blockId="93.[151,1436,151,573]" pageId="93" pageNumber="94">
In terms of body proportions 
<taxonomicName id="4C1C4D67AA304D75B479FF0B3BB5FF6B" authorityName="Ho, Lin &amp; Chang" authorityYear="2007" box="[534,641,223,248]" class="Maxillopoda" family="Caligidae" genus="Caligus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siphonostomatoida" pageId="93" pageNumber="94" phylum="Arthropoda" rank="species" species="lutjani">
<emphasis id="B968EAF6AA304D75B479FF0B3BB5FF6B" box="[534,641,223,248]" italics="true" pageId="93" pageNumber="94">C. lutjani</emphasis>
</taxonomicName>
is the most similar to the new species (cf. 
<tableCitation id="C69E035FAA304D75B205FF343DF4FF6B" box="[1130,1216,223,248]" captionStart="TABLE 6" captionStartId="94.[119,196,784,804]" captionText="TABLE 6.Selectionof species ofCaligus characterised by a3-segmented leg4bearing 4 distal spineson thesecond exopodal segment,in combination with thesetalarmature on thedistal segment ofleg 1consisting of 3plumose posterior setae, and thepresence ofan ornamentationof fine setuleson the outermargin ofthe second endopodal segmentof leg2.[Abbreviations: = abdomen length,ABW = abdomen width,GCL = genital complexlength,GCW =genital complex width]." pageId="93" pageNumber="94">Table 6</tableCitation>
). Both species also share a similar configuration of spines 1 to 3 on the distal margin of the exopod of leg 1, small outer spines on the distal exopodal segment of leg 2, multisensillate papillae associated with the post-antennal process, and tapering, pointed tines on the sternal furca. The outer spines on the exopodal segments of leg 4 are much smaller in the new species, not reaching the origin of the next most distal spine, whereas in 
<taxonomicName id="4C1C4D67AA304D75B5A9FE9B3D05FE1B" authorityName="Ho, Lin &amp; Chang" authorityYear="2007" box="[966,1073,367,392]" class="Maxillopoda" family="Caligidae" genus="Caligus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siphonostomatoida" pageId="93" pageNumber="94" phylum="Arthropoda" rank="species" species="lutjani">
<emphasis id="B968EAF6AA304D75B5A9FE9B3D05FE1B" box="[966,1073,367,392]" italics="true" pageId="93" pageNumber="94">C. lutjani</emphasis>
</taxonomicName>
each of the outer spines reaches at least to the origin of the next spine. Other differences include the more strongly recurved post-antennal process in 
<taxonomicName id="4C1C4D67AA304D75B6D9FE533816FE43" authorityName="Ho, Lin &amp; Chang" authorityYear="2007" box="[182,290,439,464]" class="Maxillopoda" family="Caligidae" genus="Caligus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siphonostomatoida" pageId="93" pageNumber="94" phylum="Arthropoda" rank="species" species="lutjani">
<emphasis id="B968EAF6AA304D75B6D9FE533816FE43" box="[182,290,439,464]" italics="true" pageId="93" pageNumber="94">C. lutjani</emphasis>
</taxonomicName>
, and its slender maxilliped with its smooth myxal margin, compared to the more robust and ridged myxal area in 
<taxonomicName id="4C1C4D67AA304D75B72DFE3638DFFE66" authority="Boxshall, 2018" authorityName="Boxshall" authorityYear="2018" box="[322,491,476,501]" class="Maxillopoda" family="Caligidae" genus="Caligus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siphonostomatoida" pageId="93" pageNumber="94" phylum="Arthropoda" rank="species" species="neoaricolus" status="sp. nov.">
<emphasis id="B968EAF6AA304D75B72DFE3638DFFE66" box="[322,491,476,501]" italics="true" pageId="93" pageNumber="94">C. neoaricolus</emphasis>
</taxonomicName>
<emphasis id="B968EAF6AA304D75B799FE373B60FE66" bold="true" box="[502,596,476,501]" pageId="93" pageNumber="94">
<taxonomicNameLabel id="A25B578DAA304D75B799FE373B60FE66" box="[502,596,476,501]" pageId="93" pageNumber="94" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
The males differ in the proportional lengths of the two free abdominal somites: in 
<taxonomicName id="4C1C4D67AA304D75B776FDEB38B0FD8B" authorityName="Ho, Lin &amp; Chang" authorityYear="2007" box="[281,388,511,536]" class="Maxillopoda" family="Caligidae" genus="Caligus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siphonostomatoida" pageId="93" pageNumber="94" phylum="Arthropoda" rank="species" species="lutjani">
<emphasis id="B968EAF6AA304D75B776FDEB38B0FD8B" box="[281,388,511,536]" italics="true" pageId="93" pageNumber="94">C. lutjani</emphasis>
</taxonomicName>
the first somite is 1.24 times longer than the second, whereas in 
<taxonomicName id="4C1C4D67AA304D75B20CFDEB3C3DFD8B" authority="Boxshall, 2018" authorityName="Boxshall" authorityYear="2018" box="[1123,1289,511,536]" class="Maxillopoda" family="Caligidae" genus="Caligus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siphonostomatoida" pageId="93" pageNumber="94" phylum="Arthropoda" rank="species" species="neoaricolus" status="sp. nov.">
<emphasis id="B968EAF6AA304D75B20CFDEB3C3DFD8B" box="[1123,1289,511,536]" italics="true" pageId="93" pageNumber="94">C. neoaricolus</emphasis>
</taxonomicName>
<emphasis id="B968EAF6AA304D75B37EFE143C5FFD8B" bold="true" box="[1297,1387,511,536]" pageId="93" pageNumber="94">
<taxonomicNameLabel id="A25B578DAA304D75B37EFE143C5FFD8B" box="[1297,1387,511,536]" pageId="93" pageNumber="94" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, the first is only half the length of the second.
</paragraph>
<caption id="DF63666CAA304D75B6F8F8663821F873" httpUri="https://zenodo.org/record/1203035/files/figure.png" pageId="93" pageNumber="94" startId="93.[151,250,1933,1955]" targetBox="[171,1430,646,1904]" targetPageId="93">
<paragraph id="8BA336E4AA304D75B6F8F8663821F873" blockId="93.[151,1435,1932,2016]" pageId="93" pageNumber="94">
<emphasis id="B968EAF6AA304D75B6F8F8663814F831" bold="true" box="[151,288,1933,1955]" pageId="93" pageNumber="94">FIGURE 41.</emphasis>
<taxonomicName id="4C1C4D67AA304D75B749F86738C0F831" authority="Boxshall, 2018" authorityName="Boxshall" authorityYear="2018" box="[294,500,1932,1954]" class="Maxillopoda" family="Caligidae" genus="Caligus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siphonostomatoida" pageId="93" pageNumber="94" phylum="Arthropoda" rank="species" species="neoaricolus" status="sp. nov.">
<emphasis id="B968EAF6AA304D75B749F86738C0F831" box="[294,500,1932,1954]" italics="true" pageId="93" pageNumber="94">Caligus neoaricolus</emphasis>
</taxonomicName>
<emphasis id="B968EAF6AA304D75B795F8663B7FF831" bold="true" box="[506,587,1933,1954]" pageId="93" pageNumber="94">
<taxonomicNameLabel id="A25B578DAA304D75B795F8663B7FF831" box="[506,587,1933,1954]" pageId="93" pageNumber="94" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
paratype male. A, habitus, dorsal; B, left corner of genital complex showing legs 5 and 6, ventral; C, antenna; D, post-antennal process, E, maxilliped. Scale bars: 1.0 mm on A, 250 µm on E, 200 µm on C, 100 µm on B, D.
</paragraph>
</caption>
<caption id="DF63666CAA334D76B618FCFB3DBFFCF3" pageId="94" pageNumber="95">
<paragraph id="8BA336E4AA334D76B618FCFB3DBFFCF3" blockId="94.[119,1844,784,864]" pageId="94" pageNumber="95">
<emphasis id="B968EAF6AA334D76B618FCFB39E8FCB7" bold="true" box="[119,220,784,804]" pageId="94" pageNumber="95">TABLE 6.</emphasis>
Selection of species of 
<taxonomicName id="4C1C4D67AA334D76B7DBFCFB38CFFCB7" authorityName="O. F. Muller" authorityYear="1785" box="[436,507,784,804]" class="Maxillopoda" family="Caligidae" genus="Caligus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siphonostomatoida" pageId="94" pageNumber="95" phylum="Arthropoda" rank="genus">
<emphasis id="B968EAF6AA334D76B7DBFCFB38CFFCB7" box="[436,507,784,804]" italics="true" pageId="94" pageNumber="95">Caligus</emphasis>
</taxonomicName>
characterised by a 3-segmented leg 4 bearing 4 distal spines on the second exopodal segment, in combination with the setal armature on the distal segment of leg 1 consisting of 3 plumose posterior setae, and the presence of an ornamentation of fine setules on the outer margin of the second endopodal segment of leg 2. [Abbreviations: = abdomen length, ABW = abdomen width, GCL = genital complex length, GCW = genital complex width].
</paragraph>
</caption>
<paragraph id="8BA336E4AA334D76B6EAFC6E3FD0FA52" pageId="94" pageNumber="95">
<table id="F91CC444AA33B2D7B6EAFC6E3E75FA52" box="[133,1857,901,1473]" gridcols="6" gridrows="15" pageId="94" pageNumber="95">
<tr id="352C34A6AA33B2D7B6EAFC6E3E75FC0A" box="[133,1857,901,921]" gridrow="0" pageId="94" pageNumber="95">
<th id="76FD5DDAAA33B2D7B6EAFC6E3B5EFC0A" box="[133,618,901,921]" gridcol="0" gridrow="0" pageId="94" pageNumber="95">Species</th>
<th id="76FD5DDAAA33B2D7B4DEFC6E3A2EFC0A" box="[689,794,901,921]" gridcol="1" gridrow="0" pageId="94" pageNumber="95">GCL:GCW</th>
<th id="76FD5DDAAA33B2D7B5F4FC6E3AC8FC0A" box="[923,1020,901,921]" gridcol="2" gridrow="0" pageId="94" pageNumber="95">GCL:ABL</th>
<th id="76FD5DDAAA33B2D7B2EBFC6E3DC1FC0A" box="[1156,1269,901,921]" gridcol="3" gridrow="0" pageId="94" pageNumber="95">GCW:ABW</th>
<th id="76FD5DDAAA33B2D7B301FC6E3CE3FC0A" box="[1390,1495,901,921]" gridcol="4" gridrow="0" pageId="94" pageNumber="95">ABL:ABW</th>
<th id="76FD5DDAAA33B2D7B032FC6E3E75FC0A" box="[1629,1857,901,921]" gridcol="5" gridrow="0" pageId="94" pageNumber="95">Reference</th>
</tr>
<tr id="352C34A6AA33B2D7B6EAFC453E75FC51" box="[133,1857,942,962]" gridrow="1" pageId="94" pageNumber="95">
<th id="76FD5DDAAA33B2D7B6EAFC453B5EFC51" box="[133,618,942,962]" gridcol="0" gridrow="1" pageId="94" pageNumber="95">
<taxonomicName id="4C1C4D67AA334D76B6EAFC4538B5FC51" authority="Lin &amp; Ho, 2003" authorityName="Lin &amp; Ho" authorityYear="2003" box="[133,385,942,962]" class="Maxillopoda" family="Caligidae" genus="Caligus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siphonostomatoida" pageId="94" pageNumber="95" phylum="Arthropoda" rank="species" species="chiastos">
<emphasis id="B968EAF6AA334D76B6EAFC4539AEFC51" box="[133,154,942,962]" italics="true" pageId="94" pageNumber="95">C.</emphasis>
<emphasis id="B968EAF6AA334D76B6F0FC4539DEFC51" box="[159,234,942,962]" italics="true" pageId="94" pageNumber="95">chiastos</emphasis>
Lin &amp; Ho, 2003
</taxonomicName>
</th>
<td id="76FD5DDAAA33B2D7B4DEFC453A2EFC51" box="[689,794,942,962]" gridcol="1" gridrow="1" pageId="94" pageNumber="95">1.00:1</td>
<td id="76FD5DDAAA33B2D7B5F4FC453AC8FC51" box="[923,1020,942,962]" gridcol="2" gridrow="1" pageId="94" pageNumber="95">1.8:1</td>
<td id="76FD5DDAAA33B2D7B2EBFC453DC1FC51" box="[1156,1269,942,962]" gridcol="3" gridrow="1" pageId="94" pageNumber="95">3.1:1</td>
<td id="76FD5DDAAA33B2D7B301FC453CE3FC51" box="[1390,1495,942,962]" gridcol="4" gridrow="1" pageId="94" pageNumber="95">1.8:1</td>
<td id="76FD5DDAAA33B2D7B032FC453E75FC51" box="[1629,1857,942,962]" gridcol="5" gridrow="1" pageId="94" pageNumber="95">Ho &amp; Lin, 2004</td>
</tr>
<tr id="352C34A6AA33B2D7B6EAFC3E3E75FC7A" box="[133,1857,981,1001]" gridrow="2" pageId="94" pageNumber="95">
<th id="76FD5DDAAA33B2D7B6EAFC3E3B5EFC7A" box="[133,618,981,1001]" gridcol="0" gridrow="2" pageId="94" pageNumber="95">
<taxonomicName id="4C1C4D67AA334D76B6EAFC3E3844FC7A" authority="Kroyer, 1863" authorityName="Kroyer" authorityYear="1863" box="[133,368,981,1001]" class="Maxillopoda" family="Caligidae" genus="Caligus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siphonostomatoida" pageId="94" pageNumber="95" phylum="Arthropoda" rank="species" species="gurnardi">
<emphasis id="B968EAF6AA334D76B6EAFC3E39AFFC7A" box="[133,155,981,1001]" italics="true" pageId="94" pageNumber="95">C.</emphasis>
<emphasis id="B968EAF6AA334D76B6F0FC3E39C4FC7A" box="[159,240,981,1001]" italics="true" pageId="94" pageNumber="95">gurnardi</emphasis>
Krøyer, 1863
</taxonomicName>
</th>
<td id="76FD5DDAAA33B2D7B4DEFC3E3A2EFC7A" box="[689,794,981,1001]" gridcol="1" gridrow="2" pageId="94" pageNumber="95">1.02:1</td>
<td id="76FD5DDAAA33B2D7B5F4FC3E3AC8FC7A" box="[923,1020,981,1001]" gridcol="2" gridrow="2" pageId="94" pageNumber="95">2.3:1</td>
<td id="76FD5DDAAA33B2D7B2EBFC3E3DC1FC7A" box="[1156,1269,981,1001]" gridcol="3" gridrow="2" pageId="94" pageNumber="95">3.4:1</td>
<td id="76FD5DDAAA33B2D7B301FC3E3CE3FC7A" box="[1390,1495,981,1001]" gridcol="4" gridrow="2" pageId="94" pageNumber="95">1.6:1</td>
<td id="76FD5DDAAA33B2D7B032FC3E3E75FC7A" box="[1629,1857,981,1001]" gridcol="5" gridrow="2" pageId="94" pageNumber="95">Parker, 1965</td>
</tr>
<tr id="352C34A6AA33B2D7B6EAFC173E75FB83" box="[133,1857,1020,1040]" gridrow="3" pageId="94" pageNumber="95">
<th id="76FD5DDAAA33B2D7B6EAFC173B5EFB83" box="[133,618,1020,1040]" gridcol="0" gridrow="3" pageId="94" pageNumber="95">
<taxonomicName id="4C1C4D67AA334D76B6EAFC173894FB83" authority="Heegaard, 1962" authorityName="Heegaard" authorityYear="1962" box="[133,416,1020,1040]" class="Maxillopoda" family="Caligidae" genus="Caligus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siphonostomatoida" pageId="94" pageNumber="95" phylum="Arthropoda" rank="species" species="longirostris">
<emphasis id="B968EAF6AA334D76B6EAFC1739AEFB83" box="[133,154,1020,1040]" italics="true" pageId="94" pageNumber="95">C.</emphasis>
<emphasis id="B968EAF6AA334D76B6F0FC17383EFB83" box="[159,266,1020,1040]" italics="true" pageId="94" pageNumber="95">longirostris</emphasis>
Heegaard, 1962
</taxonomicName>
</th>
<td id="76FD5DDAAA33B2D7B4DEFC173A2EFB83" box="[689,794,1020,1040]" gridcol="1" gridrow="3" pageId="94" pageNumber="95">1.02:1</td>
<td id="76FD5DDAAA33B2D7B5F4FC173AC8FB83" box="[923,1020,1020,1040]" gridcol="2" gridrow="3" pageId="94" pageNumber="95">1.7:1</td>
<td id="76FD5DDAAA33B2D7B2EBFC173DC1FB83" box="[1156,1269,1020,1040]" gridcol="3" gridrow="3" pageId="94" pageNumber="95">2.3:1</td>
<td id="76FD5DDAAA33B2D7B301FC173CE3FB83" box="[1390,1495,1020,1040]" gridcol="4" gridrow="3" pageId="94" pageNumber="95">1.4:1</td>
<td id="76FD5DDAAA33B2D7B032FC173E75FB83" box="[1629,1857,1020,1040]" gridcol="5" gridrow="3" pageId="94" pageNumber="95">Heegaard, 1962</td>
</tr>
<tr id="352C34A6AA33B2D7B6EAFBCF3E75FBAB" box="[133,1857,1060,1080]" gridrow="4" pageId="94" pageNumber="95">
<th id="76FD5DDAAA33B2D7B6EAFBCF3B5EFBAB" box="[133,618,1060,1080]" gridcol="0" gridrow="4" pageId="94" pageNumber="95">
<taxonomicName id="4C1C4D67AA334D76B6EAFBCF38ABFBAB" authority="Milne Edwards, 1840" authorityName="Milne Edwards" authorityYear="1840" box="[133,415,1060,1080]" class="Maxillopoda" family="Caligidae" genus="Caligus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siphonostomatoida" pageId="94" pageNumber="95" phylum="Arthropoda" rank="species" species="rapax">
<emphasis id="B968EAF6AA334D76B6EAFBCF39AFFBAB" box="[133,155,1060,1080]" italics="true" pageId="94" pageNumber="95">C.</emphasis>
<emphasis id="B968EAF6AA334D76B6F0FBCF39E0FBAB" box="[159,212,1060,1080]" italics="true" pageId="94" pageNumber="95">rapax</emphasis>
Milne Edwards, 1840
</taxonomicName>
</th>
<td id="76FD5DDAAA33B2D7B4DEFBCF3A2EFBAB" box="[689,794,1060,1080]" gridcol="1" gridrow="4" pageId="94" pageNumber="95">1.02:1</td>
<td id="76FD5DDAAA33B2D7B5F4FBCF3AC8FBAB" box="[923,1020,1060,1080]" gridcol="2" gridrow="4" pageId="94" pageNumber="95">1.8:1</td>
<td id="76FD5DDAAA33B2D7B2EBFBCF3DC1FBAB" box="[1156,1269,1060,1080]" gridcol="3" gridrow="4" pageId="94" pageNumber="95">2.5:1</td>
<td id="76FD5DDAAA33B2D7B301FBCF3CE3FBAB" box="[1390,1495,1060,1080]" gridcol="4" gridrow="4" pageId="94" pageNumber="95">1.4:1</td>
<td id="76FD5DDAAA33B2D7B032FBCF3E75FBAB" box="[1629,1857,1060,1080]" gridcol="5" gridrow="4" pageId="94" pageNumber="95">Parker &amp; Margolis, 1967</td>
</tr>
<tr id="352C34A6AA33B2D7B6EAFBA03E75FBCC" box="[133,1857,1099,1119]" gridrow="5" pageId="94" pageNumber="95">
<th id="76FD5DDAAA33B2D7B6EAFBA03B5EFBCC" box="[133,618,1099,1119]" gridcol="0" gridrow="5" pageId="94" pageNumber="95">
<taxonomicName id="4C1C4D67AA334D76B6EAFBA038B0FBCC" authority="Pillai, 1985" authorityName="Pillai" authorityYear="1985" box="[133,388,1099,1119]" class="Maxillopoda" family="Caligidae" genus="Caligus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siphonostomatoida" pageId="94" pageNumber="95" phylum="Arthropoda" rank="species" species="chrysophrysi">
<emphasis id="B968EAF6AA334D76B6EAFBA039AEFBCC" box="[133,154,1099,1119]" italics="true" pageId="94" pageNumber="95">C.</emphasis>
<emphasis id="B968EAF6AA334D76B6F0FBA03821FBCC" box="[159,277,1099,1119]" italics="true" pageId="94" pageNumber="95">chrysophrysi</emphasis>
Pillai, 1985
</taxonomicName>
</th>
<td id="76FD5DDAAA33B2D7B4DEFBA03A2EFBCC" box="[689,794,1099,1119]" gridcol="1" gridrow="5" pageId="94" pageNumber="95">1.04:1</td>
<td id="76FD5DDAAA33B2D7B5F4FBA03AC8FBCC" box="[923,1020,1099,1119]" gridcol="2" gridrow="5" pageId="94" pageNumber="95">1.9:1</td>
<td id="76FD5DDAAA33B2D7B2EBFBA03DC1FBCC" box="[1156,1269,1099,1119]" gridcol="3" gridrow="5" pageId="94" pageNumber="95">3.1:1</td>
<td id="76FD5DDAAA33B2D7B301FBA03CE3FBCC" box="[1390,1495,1099,1119]" gridcol="4" gridrow="5" pageId="94" pageNumber="95">1.7:1</td>
<td id="76FD5DDAAA33B2D7B032FBA03E75FBCC" box="[1629,1857,1099,1119]" gridcol="5" gridrow="5" pageId="94" pageNumber="95">Pillai, 1985</td>
</tr>
<tr id="352C34A6AA33B2D7B6EAFB993E75FB15" box="[133,1857,1138,1158]" gridrow="6" pageId="94" pageNumber="95">
<th id="76FD5DDAAA33B2D7B6EAFB993B5EFB15" box="[133,618,1138,1158]" gridcol="0" gridrow="6" pageId="94" pageNumber="95">
<taxonomicName id="4C1C4D67AA334D76B6EAFB9938AAFB15" authority="Wilson, 1905" authorityName="Wilson" authorityYear="1905" box="[133,414,1138,1158]" class="Maxillopoda" family="Caligidae" genus="Caligus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siphonostomatoida" pageId="94" pageNumber="95" phylum="Arthropoda" rank="species" species="rufimaculatus">
<emphasis id="B968EAF6AA334D76B6EAFB9939AFFB15" box="[133,155,1138,1158]" italics="true" pageId="94" pageNumber="95">C.</emphasis>
<emphasis id="B968EAF6AA334D76B6F0FB99382AFB15" box="[159,286,1138,1158]" italics="true" pageId="94" pageNumber="95">rufimaculatus</emphasis>
Wilson, 1905
</taxonomicName>
</th>
<td id="76FD5DDAAA33B2D7B4DEFB993A2EFB15" box="[689,794,1138,1158]" gridcol="1" gridrow="6" pageId="94" pageNumber="95">1.05:1</td>
<td id="76FD5DDAAA33B2D7B5F4FB993AC8FB15" box="[923,1020,1138,1158]" gridcol="2" gridrow="6" pageId="94" pageNumber="95">1.8:1</td>
<td id="76FD5DDAAA33B2D7B2EBFB993DC1FB15" box="[1156,1269,1138,1158]" gridcol="3" gridrow="6" pageId="94" pageNumber="95">2.4:1</td>
<td id="76FD5DDAAA33B2D7B301FB993CE3FB15" box="[1390,1495,1138,1158]" gridcol="4" gridrow="6" pageId="94" pageNumber="95">1.5:1</td>
<td id="76FD5DDAAA33B2D7B032FB993E75FB15" box="[1629,1857,1138,1158]" gridcol="5" gridrow="6" pageId="94" pageNumber="95">Cressey, 1991</td>
</tr>
<tr id="352C34A6AA33B2D7B6EAFB713E75FB3D" box="[133,1857,1178,1198]" gridrow="7" pageId="94" pageNumber="95">
<th id="76FD5DDAAA33B2D7B6EAFB713B5EFB3D" box="[133,618,1178,1198]" gridcol="0" gridrow="7" pageId="94" pageNumber="95">
<taxonomicName id="4C1C4D67AA334D76B6EAFB7138F5FB3D" authority="von Nordmann, 1832" authorityName="von Nordmann" authorityYear="1832" box="[133,449,1178,1198]" class="Actinopterygii" family="Mugilidae" genus="Myxus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="94" pageNumber="95" phylum="Chordata" rank="species" species="elongatus">
<emphasis id="B968EAF6AA334D76B6EAFB7139AFFB3D" box="[133,155,1178,1198]" italics="true" pageId="94" pageNumber="95">C.</emphasis>
<emphasis id="B968EAF6AA334D76B6F0FB7139CCFB3D" box="[159,248,1178,1198]" italics="true" pageId="94" pageNumber="95">elongatus</emphasis>
von Nordmann, 1832
</taxonomicName>
</th>
<td id="76FD5DDAAA33B2D7B4DEFB713A2EFB3D" box="[689,794,1178,1198]" gridcol="1" gridrow="7" pageId="94" pageNumber="95">1.06:1</td>
<td id="76FD5DDAAA33B2D7B5F4FB713AC8FB3D" box="[923,1020,1178,1198]" gridcol="2" gridrow="7" pageId="94" pageNumber="95">1.7:1</td>
<td id="76FD5DDAAA33B2D7B2EBFB713DC1FB3D" box="[1156,1269,1178,1198]" gridcol="3" gridrow="7" pageId="94" pageNumber="95">2.0:1</td>
<td id="76FD5DDAAA33B2D7B301FB713CE3FB3D" box="[1390,1495,1178,1198]" gridcol="4" gridrow="7" pageId="94" pageNumber="95">1.4:1</td>
<td id="76FD5DDAAA33B2D7B032FB713E75FB3D" box="[1629,1857,1178,1198]" gridcol="5" gridrow="7" pageId="94" pageNumber="95">Parker, 1969</td>
</tr>
<tr id="352C34A6AA33B2D7B6EAFB2A3E75FB46" box="[133,1857,1217,1237]" gridrow="8" pageId="94" pageNumber="95">
<th id="76FD5DDAAA33B2D7B6EAFB2A3B5EFB46" box="[133,618,1217,1237]" gridcol="0" gridrow="8" pageId="94" pageNumber="95">
<taxonomicName id="4C1C4D67AA334D76B6EAFB2A38EDFB46" authority="Parker &amp; Margolis, 1964" authorityName="Parker &amp; Margolis" authorityYear="1964" box="[133,473,1217,1237]" class="Maxillopoda" family="Caligidae" genus="Caligus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siphonostomatoida" pageId="94" pageNumber="95" phylum="Arthropoda" rank="species" species="clemensi">
<emphasis id="B968EAF6AA334D76B6EAFB2A39AEFB46" box="[133,154,1217,1237]" italics="true" pageId="94" pageNumber="95">C.</emphasis>
<emphasis id="B968EAF6AA334D76B6F0FB2A39DBFB46" box="[159,239,1217,1237]" italics="true" pageId="94" pageNumber="95">clemensi</emphasis>
Parker &amp; Margolis, 1964
</taxonomicName>
</th>
<td id="76FD5DDAAA33B2D7B4DEFB2A3A2EFB46" box="[689,794,1217,1237]" gridcol="1" gridrow="8" pageId="94" pageNumber="95">1.07:1</td>
<td id="76FD5DDAAA33B2D7B5F4FB2A3AC8FB46" box="[923,1020,1217,1237]" gridcol="2" gridrow="8" pageId="94" pageNumber="95">1.8:1</td>
<td id="76FD5DDAAA33B2D7B2EBFB2A3DC1FB46" box="[1156,1269,1217,1237]" gridcol="3" gridrow="8" pageId="94" pageNumber="95">3.0:1</td>
<td id="76FD5DDAAA33B2D7B301FB2A3CE3FB46" box="[1390,1495,1217,1237]" gridcol="4" gridrow="8" pageId="94" pageNumber="95">1.8:1</td>
<td id="76FD5DDAAA33B2D7B032FB2A3E75FB46" box="[1629,1857,1217,1237]" gridcol="5" gridrow="8" pageId="94" pageNumber="95">Kabata, 1988</td>
</tr>
<tr id="352C34A6AA33B2D7B6EAFB033E75FB6F" box="[133,1857,1256,1276]" gridrow="9" pageId="94" pageNumber="95">
<th id="76FD5DDAAA33B2D7B6EAFB033B5EFB6F" box="[133,618,1256,1276]" gridcol="0" gridrow="9" pageId="94" pageNumber="95">
<taxonomicName id="4C1C4D67AA334D76B6EAFB03385DFB6F" authority="Bere, 1936" authorityName="Bere" authorityYear="1936" box="[133,361,1256,1276]" class="Maxillopoda" family="Caligidae" genus="Caligus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siphonostomatoida" pageId="94" pageNumber="95" phylum="Arthropoda" rank="species" species="praetextus">
<emphasis id="B968EAF6AA334D76B6EAFB0339AFFB6F" box="[133,155,1256,1276]" italics="true" pageId="94" pageNumber="95">C.</emphasis>
<emphasis id="B968EAF6AA334D76B6F0FB0339CAFB6F" box="[159,254,1256,1276]" italics="true" pageId="94" pageNumber="95">praetextus</emphasis>
Bere, 1936
</taxonomicName>
</th>
<td id="76FD5DDAAA33B2D7B4DEFB033A2EFB6F" box="[689,794,1256,1276]" gridcol="1" gridrow="9" pageId="94" pageNumber="95">1.10:1</td>
<td id="76FD5DDAAA33B2D7B5F4FB033AC8FB6F" box="[923,1020,1256,1276]" gridcol="2" gridrow="9" pageId="94" pageNumber="95">1.6:1</td>
<td id="76FD5DDAAA33B2D7B2EBFB033DC1FB6F" box="[1156,1269,1256,1276]" gridcol="3" gridrow="9" pageId="94" pageNumber="95">2.3:1</td>
<td id="76FD5DDAAA33B2D7B301FB033CE3FB6F" box="[1390,1495,1256,1276]" gridcol="4" gridrow="9" pageId="94" pageNumber="95">1.6:1</td>
<td id="76FD5DDAAA33B2D7B032FB033E75FB6F" box="[1629,1857,1256,1276]" gridcol="5" gridrow="9" pageId="94" pageNumber="95">Cressey, 1991</td>
</tr>
<tr id="352C34A6AA33B2D7B6EAFAFB3E75FAB7" box="[133,1857,1296,1316]" gridrow="10" pageId="94" pageNumber="95">
<th id="76FD5DDAAA33B2D7B6EAFAFB3B5EFAB7" box="[133,618,1296,1316]" gridcol="0" gridrow="10" pageId="94" pageNumber="95">
<taxonomicName id="4C1C4D67AA334D76B6EAFAFB383FFAB7" authority="Boxshall, 2018" authorityName="Boxshall" authorityYear="2018" box="[133,267,1296,1316]" class="Maxillopoda" family="Caligidae" genus="Caligus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siphonostomatoida" pageId="94" pageNumber="95" phylum="Arthropoda" rank="species" species="neoaricolus" status="sp. nov.">
<emphasis id="B968EAF6AA334D76B6EAFAFB383FFAB7" box="[133,267,1296,1316]" italics="true" pageId="94" pageNumber="95">C. neoaricolus</emphasis>
</taxonomicName>
<emphasis id="B968EAF6AA334D76B77EFAFB386FFAB7" bold="true" box="[273,347,1296,1316]" pageId="94" pageNumber="95">
<taxonomicNameLabel id="A25B578DAA334D76B77EFAFB386FFAB7" box="[273,347,1296,1316]" pageId="94" pageNumber="95" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</th>
<td id="76FD5DDAAA33B2D7B4DEFAFB3A2EFAB7" box="[689,794,1296,1316]" gridcol="1" gridrow="10" pageId="94" pageNumber="95">1.13:1</td>
<td id="76FD5DDAAA33B2D7B5F4FAFB3AC8FAB7" box="[923,1020,1296,1316]" gridcol="2" gridrow="10" pageId="94" pageNumber="95">1.5:1</td>
<td id="76FD5DDAAA33B2D7B2EBFAFB3DC1FAB7" box="[1156,1269,1296,1316]" gridcol="3" gridrow="10" pageId="94" pageNumber="95">2.6:1</td>
<td id="76FD5DDAAA33B2D7B301FAFB3CE3FAB7" box="[1390,1495,1296,1316]" gridcol="4" gridrow="10" pageId="94" pageNumber="95">2.0:1</td>
<td id="76FD5DDAAA33B2D7B032FAFB3E75FAB7" box="[1629,1857,1296,1316]" gridcol="5" gridrow="10" pageId="94" pageNumber="95">present account</td>
</tr>
<tr id="352C34A6AA33B2D7B6EAFADC3E75FAD8" box="[133,1857,1335,1355]" gridrow="11" pageId="94" pageNumber="95">
<th id="76FD5DDAAA33B2D7B6EAFADC3B5EFAD8" box="[133,618,1335,1355]" gridcol="0" gridrow="11" pageId="94" pageNumber="95">
<taxonomicName id="4C1C4D67AA334D76B6EAFADC38B9FAD8" authority="Heegaard, 1972" authorityName="Heegaard" authorityYear="1972" box="[133,397,1335,1355]" class="Maxillopoda" family="Caligidae" genus="Caligus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siphonostomatoida" pageId="94" pageNumber="95" phylum="Arthropoda" rank="species" species="tripedalis">
<emphasis id="B968EAF6AA334D76B6EAFADC39AFFAD8" box="[133,155,1335,1355]" italics="true" pageId="94" pageNumber="95">C.</emphasis>
<emphasis id="B968EAF6AA334D76B6F0FADC39C3FAD8" box="[159,247,1335,1355]" italics="true" pageId="94" pageNumber="95">tripedalis</emphasis>
Heegaard, 1972
</taxonomicName>
</th>
<td id="76FD5DDAAA33B2D7B4DEFADC3A2EFAD8" box="[689,794,1335,1355]" gridcol="1" gridrow="11" pageId="94" pageNumber="95">1.13:1</td>
<td id="76FD5DDAAA33B2D7B5F4FADC3AC8FAD8" box="[923,1020,1335,1355]" gridcol="2" gridrow="11" pageId="94" pageNumber="95">1.4:1</td>
<td id="76FD5DDAAA33B2D7B2EBFADC3DC1FAD8" box="[1156,1269,1335,1355]" gridcol="3" gridrow="11" pageId="94" pageNumber="95">1.7:1</td>
<td id="76FD5DDAAA33B2D7B301FADC3CE3FAD8" box="[1390,1495,1335,1355]" gridcol="4" gridrow="11" pageId="94" pageNumber="95">1.5:1</td>
<td id="76FD5DDAAA33B2D7B032FADC3E75FAD8" box="[1629,1857,1335,1355]" gridcol="5" gridrow="11" pageId="94" pageNumber="95">Heegaard, 1972</td>
</tr>
<tr id="352C34A6AA33B2D7B6EAFAB43E75FAE0" box="[133,1857,1375,1395]" gridrow="12" pageId="94" pageNumber="95">
<th id="76FD5DDAAA33B2D7B6EAFAB43B5EFAE0" box="[133,618,1375,1395]" gridcol="0" gridrow="12" pageId="94" pageNumber="95">
<taxonomicName id="4C1C4D67AA334D76B6EAFAB4388DFAE0" authority="Ho, Lin &amp; Chang, 2007" authorityName="Ho, Lin &amp; Chang" authorityYear="2007" box="[133,441,1375,1395]" class="Maxillopoda" family="Caligidae" genus="Caligus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siphonostomatoida" pageId="94" pageNumber="95" phylum="Arthropoda" rank="species" species="lutjani">
<emphasis id="B968EAF6AA334D76B6EAFAB439AFFAE0" box="[133,155,1375,1395]" italics="true" pageId="94" pageNumber="95">C.</emphasis>
<emphasis id="B968EAF6AA334D76B6F0FAB439EFFAE0" box="[159,219,1375,1395]" italics="true" pageId="94" pageNumber="95">lutjani</emphasis>
Ho, Lin &amp; Chang, 2007
</taxonomicName>
</th>
<td id="76FD5DDAAA33B2D7B4DEFAB43A2EFAE0" box="[689,794,1375,1395]" gridcol="1" gridrow="12" pageId="94" pageNumber="95">1.14:1</td>
<td id="76FD5DDAAA33B2D7B5F4FAB43AC8FAE0" box="[923,1020,1375,1395]" gridcol="2" gridrow="12" pageId="94" pageNumber="95">1.7:1</td>
<td id="76FD5DDAAA33B2D7B2EBFAB43DC1FAE0" box="[1156,1269,1375,1395]" gridcol="3" gridrow="12" pageId="94" pageNumber="95">2.9:1</td>
<td id="76FD5DDAAA33B2D7B301FAB43CE3FAE0" box="[1390,1495,1375,1395]" gridcol="4" gridrow="12" pageId="94" pageNumber="95">2.0:1</td>
<td id="76FD5DDAAA33B2D7B032FAB43E75FAE0" box="[1629,1857,1375,1395]" gridcol="5" gridrow="12" pageId="94" pageNumber="95">
Ho 
<emphasis id="B968EAF6AA334D76B010FAB43F99FAE0" box="[1663,1709,1375,1395]" italics="true" pageId="94" pageNumber="95">et al.</emphasis>
, 2007
</td>
</tr>
<tr id="352C34A6AA33B2D7B6EAFA6D3E75FA09" box="[133,1857,1414,1434]" gridrow="13" pageId="94" pageNumber="95">
<th id="76FD5DDAAA33B2D7B6EAFA6D3B5EFA09" box="[133,618,1414,1434]" gridcol="0" gridrow="13" pageId="94" pageNumber="95">
<taxonomicName id="4C1C4D67AA334D76B6EAFA6D3B5EFA09" authority="Venmathi Maran, Ohtsuka &amp; Shang, 2012" authorityName="Venmathi Maran, Ohtsuka &amp; Shang" authorityYear="2012" box="[133,618,1414,1434]" class="Maxillopoda" family="Caligidae" genus="Caligus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siphonostomatoida" pageId="94" pageNumber="95" phylum="Arthropoda" rank="species" species="ogawai">
<emphasis id="B968EAF6AA334D76B6EAFA6D39AFFA09" box="[133,155,1414,1434]" italics="true" pageId="94" pageNumber="95">C.</emphasis>
<emphasis id="B968EAF6AA334D76B6F0FA6D39D6FA09" box="[159,226,1414,1434]" italics="true" pageId="94" pageNumber="95">ogawai</emphasis>
Venmathi Maran, Ohtsuka &amp; Shang, 2012
</taxonomicName>
</th>
<td id="76FD5DDAAA33B2D7B4DEFA6D3A2EFA09" box="[689,794,1414,1434]" gridcol="1" gridrow="13" pageId="94" pageNumber="95">1.21:1</td>
<td id="76FD5DDAAA33B2D7B5F4FA6D3AC8FA09" box="[923,1020,1414,1434]" gridcol="2" gridrow="13" pageId="94" pageNumber="95">1.7:1</td>
<td id="76FD5DDAAA33B2D7B2EBFA6D3DC1FA09" box="[1156,1269,1414,1434]" gridcol="3" gridrow="13" pageId="94" pageNumber="95">2.0:1</td>
<td id="76FD5DDAAA33B2D7B301FA6D3CE3FA09" box="[1390,1495,1414,1434]" gridcol="4" gridrow="13" pageId="94" pageNumber="95">1.4:1</td>
<td id="76FD5DDAAA33B2D7B032FA6D3E75FA09" box="[1629,1857,1414,1434]" gridcol="5" gridrow="13" pageId="94" pageNumber="95">
V. Maran 
<emphasis id="B968EAF6AA334D76B0D5FA6D3FD6FA09" box="[1722,1762,1414,1434]" italics="true" pageId="94" pageNumber="95">et al</emphasis>
., 2012b
</td>
</tr>
<tr id="352C34A6AA33B2D7B6EAFA463E75FA52" box="[133,1857,1453,1473]" gridrow="14" pageId="94" pageNumber="95">
<th id="76FD5DDAAA33B2D7B6EAFA463B5EFA52" box="[133,618,1453,1473]" gridcol="0" gridrow="14" pageId="94" pageNumber="95">
<taxonomicName id="4C1C4D67AA334D76B6EAFA4638ACFA52" authority="Wilson, 1937" authorityName="Wilson" authorityYear="1937" box="[133,408,1453,1473]" class="Maxillopoda" family="Caligidae" genus="Caligus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siphonostomatoida" pageId="94" pageNumber="95" phylum="Arthropoda" rank="species" species="tenuifurcatus">
<emphasis id="B968EAF6AA334D76B6EAFA4639AFFA52" box="[133,155,1453,1473]" italics="true" pageId="94" pageNumber="95">C.</emphasis>
<emphasis id="B968EAF6AA334D76B6F0FA463823FA52" box="[159,279,1453,1473]" italics="true" pageId="94" pageNumber="95">tenuifurcatus</emphasis>
Wilson, 1937
</taxonomicName>
</th>
<td id="76FD5DDAAA33B2D7B4DEFA463A2EFA52" box="[689,794,1453,1473]" gridcol="1" gridrow="14" pageId="94" pageNumber="95">1.21:1</td>
<td id="76FD5DDAAA33B2D7B5F4FA463AC8FA52" box="[923,1020,1453,1473]" gridcol="2" gridrow="14" pageId="94" pageNumber="95">1.1:1</td>
<td id="76FD5DDAAA33B2D7B2EBFA463DC1FA52" box="[1156,1269,1453,1473]" gridcol="3" gridrow="14" pageId="94" pageNumber="95">1.8:1</td>
<td id="76FD5DDAAA33B2D7B301FA463CE3FA52" box="[1390,1495,1453,1473]" gridcol="4" gridrow="14" pageId="94" pageNumber="95">1.9:1</td>
<td id="76FD5DDAAA33B2D7B032FA463E75FA52" box="[1629,1857,1453,1473]" gridcol="5" gridrow="14" pageId="94" pageNumber="95">Wilson, 1937b</td>
</tr>
</table>
</paragraph>
<paragraph id="8BA336E4AA324D77B6A8FF7C3C11FED3" blockId="95.[151,1436,151,320]" pageId="95" pageNumber="96">
The material described here differs from similar congeners and these differences justify the establishment of a new species. In the 
<typeStatus id="54A78846AA324D77B71BFF563890FF46" box="[372,420,189,213]" pageId="95" pageNumber="96">type</typeStatus>
locality, Moreton 
<collectingRegion id="49D8F806AA324D77B41CFF563B95FF46" box="[627,673,189,213]" country="Somalia" name="Bay" pageId="95" pageNumber="96">Bay</collectingRegion>
, its 
<typeStatus id="54A78846AA324D77B4A0FF563BCBFF46" box="[719,767,189,213]" pageId="95" pageNumber="96">type</typeStatus>
host 
<emphasis id="B968EAF6AA324D77B553FF563D00FF47" box="[828,1076,188,212]" italics="true" pageId="95" pageNumber="96">
was 
<taxonomicName id="4C1C4D67AA324D77B500FF563D00FF47" baseAuthorityName="Kner &amp; Steindachner" baseAuthorityYear="1867" box="[879,1076,188,212]" class="Actinopterygii" family="Ariidae" genus="Neoarius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="95" pageNumber="96" phylum="Chordata" rank="species" species="graeffei">Neoarius graeffei</taxonomicName>
</emphasis>
. A single female was recovered from a wash of 
<taxonomicName id="4C1C4D67AA324D77B727FF343B33FF6B" authorityName="Quoy &amp; Gaimard" authorityYear="1824" box="[328,519,223,248]" class="Actinopterygii" family="Sillaginidae" genus="Sillago" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="95" pageNumber="96" phylum="Chordata" rank="species" species="maculata">
<emphasis id="B968EAF6AA324D77B727FF343B33FF6B" box="[328,519,223,248]" italics="true" pageId="95" pageNumber="96">Sillago maculata</emphasis>
</taxonomicName>
(TC17904) but this is thought to be a contaminant. Similarly an incomplete male was recovered from a container that had been used to hold a batch of juvenile 
<taxonomicName id="4C1C4D67AA324D77B263FEEE3DB5FE8F" authority="sp." authorityName="Sp." box="[1036,1153,261,284]" class="Actinopterygii" family="Gerreidae" genus="Gerres" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="95" pageNumber="64" phylum="Chordata" rank="genus">
<emphasis id="B968EAF6AA324D77B263FEEE3D6CFE8F" box="[1036,1112,261,284]" italics="true" pageId="95" pageNumber="96">Gerres</emphasis>
sp.
</taxonomicName>
The partly decayed state of this male suggests that it may have been left in an inadequately rinsed dish from the previous sample.
</paragraph>
</subSubSection>
</treatment>
</document>