259 lines
29 KiB
XML
259 lines
29 KiB
XML
<document id="9E30F9D1CDA3962AF176BBAC8203DB6E" ID-CLB-Dataset="36820" ID-DOI="10.11646/zootaxa.3829.1.1" ID-GBIF-Dataset="a4b69269-5a4c-4b5b-9f89-6b314d3ba284" ID-ISSN="1175-5326" ID-Zenodo-Dep="286556" ID-ZooBank="75210CDC-AC6A-4624-A6F1-1BC969BC7CAA" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460452709399" checkinUser="plazi" docAuthor="Pyron, Robert Alexander & Wallach, Van" docDate="2014" docId="03B587DDC10EB178CFD7CB60FF43F9A4" docLanguage="en" docName="zt03829p081.pdf" docOrigin="Zootaxa 3829 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Madatyphlops Hedges, Marion, Lipp, Marin & Vidal 2014" docType="treatment" docVersion="7" lastPageNumber="53" masterDocId="FF8CFFA5C13AB14CCF40CA0EFFB9FFA2" masterDocTitle="Systematics of the blindsnakes (Serpentes: Scolecophidia: Typhlopoidea) based on molecular and morphological evidence" masterLastPageNumber="81" masterPageNumber="1" pageNumber="53" updateTime="1698338434491" updateUser="ExternalLinkService">
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<mods:mods id="F1DA7ECEE56D35C04F17C502FA8EAE6C" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="68FAE9CC610B91313DB38864E1E101FF">
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<mods:title id="7852D6775A9D67868A3FBA8A5CE46EFF">Systematics of the blindsnakes (Serpentes: Scolecophidia: Typhlopoidea) based on molecular and morphological evidence</mods:title>
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<mods:name id="A43FEA21C2A8C37D94B2F05949377B76" type="personal">
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<mods:namePart id="9A560527617BDC915EEE097511EA9F27">Pyron, Robert Alexander</mods:namePart>
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<mods:namePart id="348C36BE99257E092ABB24CD4AC1A9AC">Wallach, Van</mods:namePart>
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<mods:title id="5788C1558B8FCC84535A5522253FC9D3">Zootaxa</mods:title>
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<mods:part id="A29A325903197E8D3CF097AB388E0785">
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<mods:date id="473302393B45C9F182E6D6864E93B98B">2014</mods:date>
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<mods:number id="1532F9021C25C2B82807972A6B0827A3">3829</mods:number>
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<mods:identifier id="5EF5D9C2B85DE770B811BEA3495EC0B6" type="DOI">10.11646/zootaxa.3829.1.1</mods:identifier>
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<treatment id="03B587DDC10EB178CFD7CB60FF43F9A4" ID-DOI="http://doi.org/10.5281/zenodo.6127988" ID-GBIF-Taxon="127668690" ID-Zenodo-Dep="6127988" LSID="urn:lsid:plazi:treatment:03B587DDC10EB178CFD7CB60FF43F9A4" httpUri="http://treatment.plazi.org/id/03B587DDC10EB178CFD7CB60FF43F9A4" lastPageNumber="53" pageId="52" pageNumber="53">
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<subSubSection id="C3066540C10EB178CFD7CB60FCD6FE2A" box="[151,879,366,392]" pageId="52" pageNumber="53" type="nomenclature">
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<paragraph id="8BA336CBC10EB178CFD7CB60FCD6FE2A" blockId="52.[151,879,366,392]" box="[151,879,366,392]" pageId="52" pageNumber="53">
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<heading id="D0EB81A7C10EB178CFD7CB60FCD6FE2A" bold="true" box="[151,879,366,392]" fontSize="11" level="1" pageId="52" pageNumber="53" reason="1">
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<taxonomicName id="4C1C4D48C10EB178CFD7CB60FCD6FE2A" authority="Hedges, Marion, Lipp, Marin & Vidal, 2014" authorityName="Hedges, Marion, Lipp, Marin & Vidal" authorityYear="2014" box="[151,879,366,392]" class="Reptilia" family="Typhlopidae" genus="Madatyphlops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="52" pageNumber="53" phylum="Chordata" rank="genus">
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<emphasis id="B968EAD9C10EB178CFD7CB60FCD6FE2A" bold="true" box="[151,879,366,392]" pageId="52" pageNumber="53">
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<emphasis id="B968EAD9C10EB178CFD7CB60FEFBFE2A" bold="true" box="[151,322,366,392]" italics="true" pageId="52" pageNumber="53">Madatyphlops</emphasis>
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<bibRefCitation id="EF8D4B3AC10EB178CE09CB60FCD6FE2A" author="Hedges" box="[329,879,366,392]" pageId="52" pageNumber="53" refString="Hedges, S. B., Marion, A. B., Lipp, K. M., Marin, J. & Vidal, N. (2014) A taxonomic framework for typhlopid snakes from the Caribbean and other regions (Reptilia, Squamata). Caribbean Herpetology, 49, 1 - 61." type="journal article" year="2014">Hedges, Marion, Lipp, Marin & Vidal, 2014</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3066540C10EB178CFD7CBBBFD4CFD9B" pageId="52" pageNumber="53" type="materials_examined">
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<paragraph id="8BA336CBC10EB178CFD7CBBBFC95FE6C" blockId="52.[151,1437,437,1542]" box="[151,812,437,462]" pageId="52" pageNumber="53">
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<emphasis id="B968EAD9C10EB178CFD7CBBBFE8BFE6C" bold="true" box="[151,306,437,462]" pageId="52" pageNumber="53">
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<typeStatus id="54A78869C10EB178CFD7CBBBFF6BFE6C" box="[151,210,437,462]" pageId="52" pageNumber="53">Type</typeStatus>
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species.
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</emphasis>
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<taxonomicName id="4C1C4D48C10EB178CE79CBBBFC95FE6C" authority="Grandidier, 1872" authorityName="Grandidier" authorityYear="1872" box="[313,812,437,462]" class="Squamata" family="Typhlopidae" genus="Onychocephalus" higherTaxonomySource="CoL" kingdom="Animalia" pageId="52" pageNumber="53" phylum="Chordata" rank="species" species="arenarius">
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<emphasis id="B968EAD9C10EB178CE79CBBBFDDFFE6F" box="[313,614,437,462]" italics="true" pageId="52" pageNumber="53">Onychocephalus arenarius</emphasis>
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<bibRefCitation id="EF8D4B3AC10EB178CD2DCBBBFC95FE6C" author="Grandidier" box="[621,812,437,462]" pageId="52" pageNumber="53" refString="Grandidier, A. (1872) Descriptions de quelques Reptiles nouveaux decouverts a Madagascar en 1870. Annales des Sciences Naturelles, Zoologie et Paleontologie, Paris, Serie 5, 15, 6 - 11." type="journal article" year="1872">Grandidier, 1872</bibRefCitation>
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</taxonomicName>
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</paragraph>
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<paragraph id="8BA336CBC10EB178CF87CBD7FD4CFD9B" blockId="52.[151,1437,437,1542]" pageId="52" pageNumber="53">
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<emphasis id="B968EAD9C10EB178CF87CBD7FD5DFE50" bold="true" box="[199,740,473,498]" pageId="52" pageNumber="53">
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Species content.
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<taxonomicName id="4C1C4D48C10EB178CEDDCBD7FD5DFE50" authority="Mad." authorityName="Mad." box="[413,740,473,498]" class="Reptilia" family="Typhlopidae" genus="Madatyphlops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="52" pageNumber="53" phylum="Chordata" rank="species" species="andasibensis">
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<emphasis id="B968EAD9C10EB178CEDDCBD7FD5DFE50" box="[413,740,473,498]" italics="true" pageId="52" pageNumber="53">Madatyphlops andasibensis</emphasis>
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</taxonomicName>
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</emphasis>
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,
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<emphasis id="B968EAD9C10EB178CDBACBD7FC0FFE50" bold="true" box="[762,950,473,498]" italics="true" pageId="52" pageNumber="53">
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Mad.
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<taxonomicName id="4C1C4D48C10EB178CC06CBD7FC0FFE50" authority="Mad." authorityName="Mad." box="[838,950,473,498]" class="Squamata" family="Typhlopidae" genus="Onychocephalus" higherTaxonomySource="CoL" kingdom="Animalia" pageId="52" pageNumber="53" phylum="Chordata" rank="species" species="arenarius">arenarius</taxonomicName>
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</emphasis>
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, Mad.
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<taxonomicName id="4C1C4D48C10EB178CB56CBD7FB74FE50" authority="Mad." authorityName="Mad." box="[1046,1229,473,498]" class="Reptilia" family="Typhlopidae" genus="Madatyphlops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="52" pageNumber="53" phylum="Chordata" rank="species" species="boettgeri">boettgeri, Mad.</taxonomicName>
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<taxonomicName id="4C1C4D48C10EB178CB9CCBD7FA24FE50" authority="Mad." authorityName="Mad." box="[1244,1437,473,498]" class="Reptilia" family="Typhlopidae" genus="Madatyphlops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="52" pageNumber="53" phylum="Chordata" rank="species" species="calabresii">calabresii, Mad.</taxonomicName>
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<taxonomicName id="4C1C4D48C10EB178CFD7CBF0FEDBFDB4" authority="Mad." authorityName="Mad." box="[151,354,509,534]" class="Reptilia" family="Typhlopidae" genus="Madatyphlops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="52" pageNumber="53" phylum="Chordata" rank="species" species="comorensis">comorensis, Mad.</taxonomicName>
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<taxonomicName id="4C1C4D48C10EB178CE2CCBF0FD84FDB4" authority="Mad." authorityName="Mad." box="[364,573,509,534]" class="Reptilia" family="Typhlopidae" genus="Madatyphlops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="52" pageNumber="53" phylum="Chordata" rank="species" species="cuneirostris">cuneirostris, Mad.</taxonomicName>
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<taxonomicName id="4C1C4D48C10EB178CD06CBF3FD48FDB4" authority="Mad." authorityName="Mad." box="[582,753,509,534]" class="Reptilia" family="Typhlopidae" genus="Madatyphlops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="52" pageNumber="53" phylum="Chordata" rank="species" species="decorsei">decorsei, Mad.</taxonomicName>
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<taxonomicName id="4C1C4D48C10EB178CDBBCBF3FC51FDB4" authority="Mad." authorityName="Mad." box="[763,1000,509,534]" class="Reptilia" family="Typhlopidae" genus="Madatyphlops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="52" pageNumber="53" phylum="Chordata" rank="species" species="leucocephalus">leucocephalus, Mad.</taxonomicName>
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madagascariensis, Mad.
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<taxonomicName id="4C1C4D48C10EB178CA52CBF0FF6BFD98" authority="Mad." authorityName="Mad." class="Reptilia" family="Typhlopidae" genus="Madatyphlops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="52" pageNumber="53" phylum="Chordata" rank="species" species="mucronatus">mucronatus, Mad.</taxonomicName>
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<taxonomicName id="4C1C4D48C10EB178CF98C82FFEC4FD98" authority="Mad." authorityName="Mad." box="[216,381,545,570]" class="Reptilia" family="Typhlopidae" genus="Madatyphlops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="52" pageNumber="53" phylum="Chordata" rank="species" species="ocularis">ocularis, Mad.</taxonomicName>
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<taxonomicName id="4C1C4D48C10EB178CEC3C82FFD9DFD98" box="[387,548,545,570]" class="Reptilia" family="Typhlopidae" genus="Madatyphlops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="52" pageNumber="53" phylum="Chordata" rank="species" species="platyrhynchus">platyrhynchus</taxonomicName>
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, and
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<emphasis id="B968EAD9C10EB178CD1FC82FFD4CFD9B" box="[607,757,545,570]" italics="true" pageId="52" pageNumber="53">
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<emphasis id="B968EAD9C10EB178CD1FC82FFD57FD98" bold="true" box="[607,750,545,570]" italics="true" pageId="52" pageNumber="53">
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Mad.
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<taxonomicName id="4C1C4D48C10EB178CDE2C82FFD57FD98" box="[674,750,545,570]" class="Reptilia" family="Typhlopidae" genus="Madatyphlops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="52" pageNumber="53" phylum="Chordata" rank="species" species="rajeryi">rajeryi</taxonomicName>
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</emphasis>
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.
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</emphasis>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3066540C10EB178CF87C84BFE6FFB90" pageId="52" pageNumber="53" type="diagnosis">
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<paragraph id="8BA336CBC10EB178CF87C84BFD8AFC64" blockId="52.[151,1437,437,1542]" pageId="52" pageNumber="53">
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<emphasis id="B968EAD9C10EB178CF87C84BFEF8FDFC" bold="true" box="[199,321,581,606]" pageId="52" pageNumber="53">Diagnosis.</emphasis>
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<taxonomicName id="4C1C4D48C10EB178CE08C84BFE51FDFC" box="[328,488,581,606]" class="Reptilia" family="Typhlopidae" genus="Madatyphlops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="52" pageNumber="53" phylum="Chordata" rank="genus">
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<emphasis id="B968EAD9C10EB178CE08C84BFE51FDFC" box="[328,488,581,606]" italics="true" pageId="52" pageNumber="53">Madatyphlops</emphasis>
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</taxonomicName>
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can be distinguished from all other typhlopoids by the combination of the following characters: small- to large-sized (total length
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<quantity id="4CE49B2EC10EB178CDEAC867FC80FD20" box="[682,825,617,642]" metricMagnitude="-1" metricUnit="m" metricValue="3.31" metricValueMax="6.0" metricValueMin="0.62" pageId="52" pageNumber="53" unit="mm" value="331.0" valueMax="600.0" valueMin="62.0">62–600 mm</quantity>
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), stout- to slender-bodied (length/width ratio 17–85) snakes with 20–28 scale rows (with reduction, except in
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<emphasis id="B968EAD9C10EB178CC7AC883FC56FD04" box="[826,1007,653,678]" italics="true" pageId="52" pageNumber="53">
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Mad.
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<taxonomicName id="4C1C4D48C10EB178CCC0C883FC56FD04" box="[896,1007,653,678]" class="Reptilia" family="Typhlopidae" genus="Madatyphlops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="52" pageNumber="53" phylum="Chordata" rank="species" species="calabresii">calabresii</taxonomicName>
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</emphasis>
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), 196–580 total middorsals, short to moderate tail (1.2–3.0% total length) with 7–18 subcaudals (length/width ratio 0.7–2.6), and apical spine small to minute. Dorsal and lateral head profiles rounded, moderate rostral (0.29–0.54 head width), inferior nasal suture in contact with second supralabial, preocular in contact with second and third supralabials, eye small to moderate with distinct pupil, T-III or T-V
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<collectingCountry id="F30B765BC10EB178CE94C910FE45FC94" box="[468,508,798,822]" name="Cyprus" pageId="52" pageNumber="53">SIP</collectingCountry>
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, and postoculars 2–3. Lateral tongue papillae present; left lung absent, tracheal, cardiac and right lungs normally multicameral (latter unicameral in continental African species); testes segmented or unsegmented; hemipenis eversible, lacking retrocloacal sacs; and rectal caecum usually large (0.5–5.0% SVL). Coloration brown to black dorsally with lighter venter (light brown to yellow) or entirely pink (pigmentless) in a few species (yellow in preservative).
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</paragraph>
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<paragraph id="8BA336CBC10EB178CF87C9DFFE6FFB90" blockId="52.[151,1437,437,1542]" pageId="52" pageNumber="53">
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<emphasis id="B968EAD9C10EB178CF87C9DFFE5FFC48" bold="true" box="[199,486,977,1002]" pageId="52" pageNumber="53">Phylogenetic definition.</emphasis>
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Includes the MRCA of
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<taxonomicName id="4C1C4D48C10EB178CC4EC9DFFB9EFC4B" box="[782,1063,977,1002]" class="Reptilia" family="Typhlopidae" genus="Madatyphlops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="52" pageNumber="53" phylum="Chordata" rank="species" species="arenarius">
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<emphasis id="B968EAD9C10EB178CC4EC9DFFB9EFC4B" box="[782,1063,977,1002]" italics="true" pageId="52" pageNumber="53">Madatyphlops arenarius</emphasis>
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</taxonomicName>
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and
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<emphasis id="B968EAD9C10EB178CB27C9DFFA86FC48" box="[1127,1343,977,1002]" italics="true" pageId="52" pageNumber="53">
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Mad.
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<taxonomicName id="4C1C4D48C10EB178CBEEC9DFFA86FC48" box="[1198,1343,977,1002]" class="Reptilia" family="Typhlopidae" genus="Madatyphlops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="52" pageNumber="53" phylum="Chordata" rank="species" species="andasibensis">andasibensis</taxonomicName>
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</emphasis>
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and all descendants thereof, and all species more closely related to
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<emphasis id="B968EAD9C10EB178CC75C9FBFC5BFBAF" box="[821,994,1013,1038]" italics="true" pageId="52" pageNumber="53">
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Mad.
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<taxonomicName id="4C1C4D48C10EB178CC36C9F8FC5BFBAF" box="[886,994,1014,1037]" class="Reptilia" family="Typhlopidae" genus="Madatyphlops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="52" pageNumber="53" phylum="Chordata" rank="species" species="arenarius">arenarius</taxonomicName>
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</emphasis>
|
||
than to the
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<typeStatus id="54A78869C10EB178CB28C9F8FB21FBAC" box="[1128,1176,1014,1038]" pageId="52" pageNumber="53">type</typeStatus>
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species of the 15 other typhlopid genera listed here.
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</paragraph>
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</subSubSection>
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||
<subSubSection id="C3066540C10EB178CF87CE33FDE9FBD8" pageId="52" pageNumber="53" type="etymology">
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||
<paragraph id="8BA336CBC10EB178CF87CE33FDE9FBD8" blockId="52.[151,1437,437,1542]" pageId="52" pageNumber="53">
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<emphasis id="B968EAD9C10EB178CF87CE33FEF7FBF4" bold="true" box="[199,334,1085,1110]" pageId="52" pageNumber="53">Etymology.</emphasis>
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Name refers to the primarily
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<collectingCountry id="F30B765BC10EB178CDF4CE33FC98FBF4" box="[692,801,1085,1110]" name="Madagascar" pageId="52" pageNumber="53">Malagasy</collectingCountry>
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distribution of the species, similar to the skink genus
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<taxonomicName id="4C1C4D48C10EB178CFD7CE6FFE92FBD8" box="[151,299,1121,1146]" class="Reptilia" family="Scincidae" genus="Madascincus" kingdom="Animalia" order="Squamata" pageId="52" pageNumber="53" phylum="Chordata" rank="genus">
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||
<emphasis id="B968EAD9C10EB178CFD7CE6FFE92FBD8" box="[151,299,1121,1146]" italics="true" pageId="52" pageNumber="53">Madascincus</emphasis>
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</taxonomicName>
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||
(see
|
||
<bibRefCitation id="EF8D4B3AC10EB178CE25CE6FFDFDFBD8" author="Whiting" box="[357,580,1121,1146]" pageId="52" pageNumber="53" refString="Whiting, A. S., Sites, J. W. & Bauer, A. M. (2004) Molecular phylogenetics of Malagasy skinks (Squamata: Scincidae). African Journal of Herpetology, 53, 135 - 146. http: // dx. doi. org / 10.1080 / 21564574.2004.9635506" type="journal article" year="2004">
|
||
Whiting
|
||
<emphasis id="B968EAD9C10EB178CE88CE6CFDB8FBD8" box="[456,513,1121,1146]" italics="true" pageId="52" pageNumber="53">et al.</emphasis>
|
||
2004
|
||
</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3066540C10EB178CF87CE8BFD90FB44" pageId="52" pageNumber="53" type="distribution">
|
||
<paragraph id="8BA336CBC10EB178CF87CE8BFD90FB44" blockId="52.[151,1437,437,1542]" pageId="52" pageNumber="53">
|
||
<emphasis id="B968EAD9C10EB178CF87CE8BFED9FB3C" bold="true" box="[199,352,1157,1182]" pageId="52" pageNumber="53">Distribution.</emphasis>
|
||
Most species are endemic to
|
||
<collectingCountry id="F30B765BC10EB178CD84CE8BFCF2FB3C" box="[708,843,1157,1182]" name="Madagascar" pageId="52" pageNumber="53">Madagascar</collectingCountry>
|
||
(Table 1), though a few are found in northeastern Africa (
|
||
<emphasis id="B968EAD9C10EB178CFAFCEA7FC70FB60" box="[239,969,1193,1218]" italics="true" pageId="52" pageNumber="53">
|
||
<taxonomicName id="4C1C4D48C10EB178CFAFCEA7FDF7FB60" authority="Mad." authorityName="Mad." box="[239,590,1193,1218]" class="Reptilia" family="Typhlopidae" genus="Madatyphlops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="52" pageNumber="53" phylum="Chordata" rank="species" species="calabresii">Madatyphlops calabresii, Mad.</taxonomicName>
|
||
<taxonomicName id="4C1C4D48C10EB178CD15CEA4FC9BFB60" authority="Mad." authorityName="Mad." box="[597,802,1193,1218]" class="Reptilia" family="Typhlopidae" genus="Madatyphlops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="52" pageNumber="53" phylum="Chordata" rank="species" species="cuneirostris">cuneirostris, Mad.</taxonomicName>
|
||
<taxonomicName id="4C1C4D48C10EB178CC69CEA7FC70FB60" box="[809,969,1193,1218]" class="Reptilia" family="Typhlopidae" genus="Madatyphlops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="52" pageNumber="53" phylum="Chordata" rank="species" species="leucocephalus">leucocephalus</taxonomicName>
|
||
</emphasis>
|
||
, and
|
||
<emphasis id="B968EAD9C10EB178CB47CEA7FB5FFB60" box="[1031,1254,1193,1218]" italics="true" pageId="52" pageNumber="53">
|
||
Mad.
|
||
<taxonomicName id="4C1C4D48C10EB178CB08CEA7FB5FFB60" box="[1096,1254,1193,1218]" class="Reptilia" family="Typhlopidae" genus="Madatyphlops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="52" pageNumber="53" phylum="Chordata" rank="species" species="platyrhynchus">platyrhynchus</taxonomicName>
|
||
</emphasis>
|
||
), and one in the
|
||
<collectingCountry id="F30B765BC10EB178CFD7CEC0FEF0FB44" box="[151,329,1229,1254]" name="Comoros" pageId="52" pageNumber="53">Comoro Islands</collectingCountry>
|
||
(
|
||
<emphasis id="B968EAD9C10EB178CE19CEC3FDA0FB47" box="[345,537,1229,1254]" italics="true" pageId="52" pageNumber="53">
|
||
Mad.
|
||
<taxonomicName id="4C1C4D48C10EB178CEDBCEC0FDA0FB47" box="[411,537,1230,1253]" class="Reptilia" family="Typhlopidae" genus="Madatyphlops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="52" pageNumber="53" phylum="Chordata" rank="species" species="comorensis">comorensis</taxonomicName>
|
||
</emphasis>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3066540C10EB178CF87CEFFFF43F9A4" pageId="52" pageNumber="53" type="discussion">
|
||
<paragraph id="8BA336CBC10EB178CF87CEFFFF43F9A4" blockId="52.[151,1437,437,1542]" pageId="52" pageNumber="53">
|
||
<emphasis id="B968EAD9C10EB178CF87CEFFFE82FAA8" bold="true" box="[199,315,1265,1290]" pageId="52" pageNumber="53">Remarks.</emphasis>
|
||
Some species were placed in
|
||
<taxonomicName id="4C1C4D48C10EB178CDF0CEFFFCF9FAA8" box="[688,832,1265,1290]" class="Reptilia" family="Typhlopidae" genus="Afrotyphlops" kingdom="Animalia" order="Squamata" pageId="52" pageNumber="53" phylum="Chordata" rank="genus">
|
||
<emphasis id="B968EAD9C10EB178CDF0CEFFFCF9FAA8" box="[688,832,1265,1290]" italics="true" pageId="52" pageNumber="53">Afrotyphlops</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<emphasis id="B968EAD9C10EB178CC16CEFFFE9AFA8F" italics="true" pageId="52" pageNumber="53">
|
||
<taxonomicName id="4C1C4D48C10EB178CC16CEFFFB7BFAA8" authority="Mad." authorityName="Mad." box="[854,1218,1265,1290]" class="Reptilia" family="Typhlopidae" genus="Madatyphlops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="52" pageNumber="53" phylum="Chordata" rank="species" species="calabresii">Madatyphlops calabresii, Mad.</taxonomicName>
|
||
<taxonomicName id="4C1C4D48C10EB178CB8FCEFCFA24FAA8" authority="Mad." authorityName="Mad." box="[1231,1437,1265,1290]" class="Reptilia" family="Typhlopidae" genus="Madatyphlops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="52" pageNumber="53" phylum="Chordata" rank="species" species="comorensis">comorensis, Mad.</taxonomicName>
|
||
<taxonomicName id="4C1C4D48C10EB178CFD7CF18FEA6FA8F" box="[151,287,1302,1325]" class="Reptilia" family="Typhlopidae" genus="Madatyphlops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="52" pageNumber="53" phylum="Chordata" rank="species" species="cuneirostris">cuneirostris</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
and
|
||
<emphasis id="B968EAD9C10EB178CE1DCF1BFD86FA8C" box="[349,575,1301,1326]" italics="true" pageId="52" pageNumber="53">
|
||
Mad.
|
||
<taxonomicName id="4C1C4D48C10EB178CEE1CF1BFD86FA8C" box="[417,575,1301,1326]" class="Reptilia" family="Typhlopidae" genus="Madatyphlops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="52" pageNumber="53" phylum="Chordata" rank="species" species="platyrhynchus">platyrhynchus</taxonomicName>
|
||
</emphasis>
|
||
) and
|
||
<taxonomicName id="4C1C4D48C10EB178CDC3CF1BFC9CFA8C" box="[643,805,1301,1326]" class="Reptilia" family="Typhlopidae" genus="Rhinotyphlops" kingdom="Animalia" order="Squamata" pageId="52" pageNumber="53" phylum="Chordata" rank="genus">
|
||
<emphasis id="B968EAD9C10EB178CDC3CF1BFC9CFA8C" box="[643,805,1301,1326]" italics="true" pageId="52" pageNumber="53">Rhinotyphlops</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<emphasis id="B968EAD9C10EB178CC77CF1BFBA3FA8C" box="[823,1050,1301,1326]" italics="true" pageId="52" pageNumber="53">
|
||
Mad.
|
||
<taxonomicName id="4C1C4D48C10EB178CC3ACF1BFBA3FA8C" box="[890,1050,1301,1326]" class="Reptilia" family="Typhlopidae" genus="Madatyphlops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="52" pageNumber="53" phylum="Chordata" rank="species" species="leucocephalus">leucocephalus</taxonomicName>
|
||
</emphasis>
|
||
) by
|
||
<bibRefCitation id="EF8D4B3AC10EB178CB12CF1BFA86FA8C" box="[1106,1343,1301,1326]" pageId="52" pageNumber="53" refString="Hedges, S. B., Marion, A. B., Lipp, K. M., Marin, J. & Vidal, N. (2014) A taxonomic framework for typhlopid snakes from the Caribbean and other regions (Reptilia, Squamata). Caribbean Herpetology, 49, 1 - 61." type="journal article">
|
||
Hedges
|
||
<emphasis id="B968EAD9C10EB178CBEFCF18FB50FA8C" box="[1199,1257,1301,1326]" italics="true" pageId="52" pageNumber="53">et al.</emphasis>
|
||
(2014)
|
||
</bibRefCitation>
|
||
, but are likely allied with
|
||
<taxonomicName id="4C1C4D48C10EB178CE28CF37FDB1FAF0" box="[360,520,1337,1362]" class="Reptilia" family="Typhlopidae" genus="Madatyphlops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="52" pageNumber="53" phylum="Chordata" rank="genus">
|
||
<emphasis id="B968EAD9C10EB178CE28CF37FDB1FAF0" box="[360,520,1337,1362]" italics="true" pageId="52" pageNumber="53">Madatyphlops</emphasis>
|
||
</taxonomicName>
|
||
based on several characters. These include a T-V
|
||
<collectingCountry id="F30B765BC10EB178CB1ECF34FB3FFAF0" box="[1118,1158,1338,1362]" name="Cyprus" pageId="52" pageNumber="53">SIP</collectingCountry>
|
||
(
|
||
<emphasis id="B968EAD9C10EB178CBDBCF37FE9BFAD7" italics="true" pageId="52" pageNumber="53">
|
||
Mad.
|
||
<taxonomicName id="4C1C4D48C10EB178CBA0CF37FA24FAF0" authority="Mad." authorityName="Mad." box="[1248,1437,1337,1362]" class="Reptilia" family="Typhlopidae" genus="Madatyphlops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="52" pageNumber="53" phylum="Chordata" rank="species" species="calabresii">calabresii, Mad.</taxonomicName>
|
||
<taxonomicName id="4C1C4D48C10EB178CFD7CF50FEA6FAD7" box="[151,287,1374,1397]" class="Reptilia" family="Typhlopidae" genus="Madatyphlops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="52" pageNumber="53" phylum="Chordata" rank="species" species="cuneirostris">cuneirostris</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
and
|
||
<emphasis id="B968EAD9C10EB178CE1CCF53FD87FAD4" box="[348,574,1373,1398]" italics="true" pageId="52" pageNumber="53">
|
||
Mad.
|
||
<taxonomicName id="4C1C4D48C10EB178CEDECF53FD87FAD4" box="[414,574,1373,1398]" class="Reptilia" family="Typhlopidae" genus="Madatyphlops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="52" pageNumber="53" phylum="Chordata" rank="species" species="leucocephalus">leucocephalus</taxonomicName>
|
||
</emphasis>
|
||
) or a T-III
|
||
<collectingCountry id="F30B765BC10EB178CD81CF50FD50FAD4" box="[705,745,1374,1398]" name="Cyprus" pageId="52" pageNumber="53">SIP</collectingCountry>
|
||
(
|
||
<emphasis id="B968EAD9C10EB178CDBCCF53FC72FAD7" box="[764,971,1373,1398]" italics="true" pageId="52" pageNumber="53">Mad. comoroensis</emphasis>
|
||
and
|
||
<emphasis id="B968EAD9C10EB178CB44CF53FB5CFAD4" box="[1028,1253,1373,1398]" italics="true" pageId="52" pageNumber="53">
|
||
Mad.
|
||
<taxonomicName id="4C1C4D48C10EB178CB07CF53FB5CFAD4" box="[1095,1253,1373,1398]" class="Reptilia" family="Typhlopidae" genus="Madatyphlops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="52" pageNumber="53" phylum="Chordata" rank="species" species="platyrhynchus">platyrhynchus</taxonomicName>
|
||
</emphasis>
|
||
), both of which are common in Madatyphlopinae but otherwise unknown in Afrotyphlopinae (Table 2). Also, the supralabials of these species increase in size posteriorly, instead of being low and flat as in
|
||
<taxonomicName id="4C1C4D48C10EB178CB44CFABFB2DFA1C" box="[1028,1172,1445,1470]" class="Reptilia" family="Typhlopidae" genus="Afrotyphlops" kingdom="Animalia" order="Squamata" pageId="52" pageNumber="53" phylum="Chordata" rank="genus">
|
||
<emphasis id="B968EAD9C10EB178CB44CFABFB2DFA1C" box="[1028,1172,1445,1470]" italics="true" pageId="52" pageNumber="53">Afrotyphlops</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C1C4D48C10EB178CB8ECFABFAC9FA1C" box="[1230,1392,1445,1470]" class="Reptilia" family="Typhlopidae" genus="Rhinotyphlops" kingdom="Animalia" order="Squamata" pageId="52" pageNumber="53" phylum="Chordata" rank="genus">
|
||
<emphasis id="B968EAD9C10EB178CB8ECFABFAC9FA1C" box="[1230,1392,1445,1470]" italics="true" pageId="52" pageNumber="53">Rhinotyphlops</emphasis>
|
||
</taxonomicName>
|
||
. At least five new undescribed species are known from
|
||
<collectingCountry id="F30B765BC10EB178CDB7CFC7FCC7FA40" box="[759,894,1481,1506]" name="Madagascar" pageId="52" pageNumber="53">Madagascar</collectingCountry>
|
||
(Wallach unpubl. data) based on new UMMZ material.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |