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<mods:title id="DA2C06F18DBBFAA4C305085B128A367C">First Record of the Diogenid Hermit Crab Aniculus erythraeus (Crustacea: Decapoda: Anomura) from Japan</mods:title>
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<taxonomicName id="4C1C4D5EFB7EC3584C95FADE5223D505" ID-CoL="ee5bdb38-68c5-4066-a013-0ec1f6247e42" authority="Forest, 1984" authorityName="Forest" authorityYear="1984" box="[958,1295,1304,1329]" class="Malacostraca" family="Diogenidae" genus="Aniculus" kingdom="Animalia" order="Decapoda" pageId="0" pageNumber="33" phylum="Arthropoda" rank="species" species="erythraeus">
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<bibRefCitation id="EF8D4B2CFB7EC3584BBFFADF5223D505" author="Forest, J." box="[1172,1295,1305,1329]" pageId="0" pageNumber="33" pagination="1 - 91" refId="ref2911" refString="Forest, J. 1984. Revision du genre Aniculus. Crustaceana (Supplement 8): 1 - 91." type="journal article" year="1984">Forest, 1984</bibRefCitation>
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[New Japanese name: Sarasa-oni-yadokari] (
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,
<figureCitation id="13272A58FB7EC3584B4BFA9C5340D546" box="[1120,1132,1370,1394]" captionStart="Fig" captionStartId="2.[157,188,1751,1773]" captionTargetBox="[264,1323,204,1729]" captionTargetId="figure-106@2.[264,1323,204,1729]" captionTargetPageId="2" captionText="Fig. 2. Aniculus erythraeus Forest, 1984, male (sl 8.7mm), Kume Island, RUMF-ZC-02041 (AD). Aniculus ursus (Olivier, 1812), male (sl 18.6mm), KUMEJIMA 2009, Stn Dive 4, ZRC 2011.0704 (E); male (sl 21.8mm), KUMEJIMA 2009, Stn Dive 15, NTOU A01137 (F). A, left second pereopod, lateral view (most setae omitted); B, same, mesial view (most setae omitted); C, left third pereopod, lateral view (setae omitted); D, same, mesial view (setae omitted); E, left third pereopod, dactylus and propodus, lateral view (setae omitted); F, left third pereopod, propodus, lateral view (setae omitted). Scales: 2.0mm." figureDoi="http://doi.org/10.5281/zenodo.4584831" httpUri="https://zenodo.org/record/4584831/files/figure.png" pageId="0" pageNumber="33">2</figureCitation>
AD, 3)
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<taxonomicName id="4C1C4D5EFB7EC3584C1FFA5C5392D587" ID-CoL="ee5bdb38-68c5-4066-a013-0ec1f6247e42" authority="Forest, 1984: 21" authorityName="Forest" authorityPageNumber="21" authorityYear="1984" box="[820,1214,1434,1459]" class="Malacostraca" family="Diogenidae" genus="Aniculus" kingdom="Animalia" order="Decapoda" pageId="0" pageNumber="33" phylum="Arthropoda" rank="species" species="erythraeus">
<emphasis id="B968EACFFB7EC3584C1FFA5C532ED586" box="[820,1026,1434,1458]" italics="true" pageId="0" pageNumber="33">Aniculus erythraeus</emphasis>
<bibRefCitation id="EF8D4B2CFB7EC3584B3BFA5D5392D587" author="Forest, J." box="[1040,1214,1435,1459]" pageId="0" pageNumber="33" pagination="1 - 91" refId="ref2911" refString="Forest, J. 1984. Revision du genre Aniculus. Crustaceana (Supplement 8): 1 - 91." type="journal article" year="1984">Forest, 1984: 21</bibRefCitation>
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(keys), 41, figs 12, 19, 4145 (
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locality: Red Sea);
<treatmentCitation id="0ABD10CCFB7EC3584BEEFA7D5458D5C7" author="Haig, J. &amp; Ball, E. E." page="158" pageId="0" pageNumber="33" year="1988">
<bibRefCitation id="EF8D4B2CFB7EC3584BEEFA7D5458D5C7" author="Haig, J. &amp; Ball, E. E." pageId="0" pageNumber="33" pagination="151 - 196" refId="ref2932" refString="Haig, J. and Ball, E. E. 1988. Hermit crabs from northern Australian and eastern Indonesian waters (Crustacea Decapoda: Anomura: Paguroidea) collected during the 1975 Alpha Helix Expedition. Records of the Australian Museum 40: 151 - 196." type="journal article" year="1988">Haig and Ball 1988: 158</bibRefCitation>
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;
<treatmentCitation id="0ABD10CCFB7EC3584CA9FA1A53FBD5C0" author="Rahayu, D. L. &amp; Wahyundi, A. J." box="[898,1239,1500,1524]" page="23" pageId="0" pageNumber="33" year="2008">
<bibRefCitation id="EF8D4B2CFB7EC3584CA9FA1A53FBD5C0" author="Rahayu, D. L. &amp; Wahyundi, A. J." box="[898,1239,1500,1524]" pageId="0" pageNumber="33" refId="ref3457" refString="Rahayu, D. L. and Wahyundi, A. J. 2008. Common Littoral Hermit Crabs of Indonesia (Susetiono, Shirayama, Y. and Asakura, A., Eds). Kyoto University Press, Kyoto, 93 pp." type="book" year="2008">Rahayu and Wahyundi 2008: 23</bibRefCitation>
</treatmentCitation>
, unnumbered fig.;
<treatmentCitation id="0ABD10CCFB7EC3584C65FA3A534BD620" author="McLaughlin, P. A. &amp; Komai, T. &amp; Lemaitre, R. &amp; Rahayu, D. L." box="[846,1127,1532,1556]" page="18" pageId="0" pageNumber="33" year="2010">
<bibRefCitation id="EF8D4B2CFB7EC3584C65FA3A534BD620" author="McLaughlin, P. A. &amp; Komai, T. &amp; Lemaitre, R. &amp; Rahayu, D. L." box="[846,1127,1532,1556]" pageId="0" pageNumber="33" pagination="5 - 107" refId="ref3107" refString="McLaughlin, P. A., Komai, T., Lemaitre, R. and Rahayu, D. L. 2010. Annotated checklist of anomuran decapod crustaceans of the world (exclusive of the Kiwaoidea and families Chirostylidae and Galatheidae of the Galatheoidea) Part I-Lithodoidea, Lomisoidea and Paguroidea. The Raffles Bulletin of Zoology (Supplement 23): 5 - 107." type="journal article" year="2010">
McLaughlin
<emphasis id="B968EACFFB7EC3584CF9FA3B532BD620" box="[978,1031,1532,1556]" italics="true" pageId="0" pageNumber="33">et al.</emphasis>
2010: 18
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(list); Poupin 2010: 34 (list);
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<bibRefCitation id="EF8D4B2CFB7EC3584C65F9DB5352D601" author="Poupin, J. &amp; Juncker, M." box="[846,1150,1565,1589]" pageId="0" pageNumber="33" refId="ref3382" refString="Poupin, J. and Juncker, M. 2010. Guide des Crustaces Decapodes du Pacifique Sud / A Guide to Decapod Crustaceans of the South Pacific. CRISP and CPS, Noumea, 317 pp." type="book" year="2010">Poupin and Juncker 2010: 218</bibRefCitation>
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, fig. a.
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</paragraph>
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<emphasis id="B968EACFFB7EC3584C65F99B530DD641" bold="true" box="[846,1057,1629,1653]" pageId="0" pageNumber="33">Material examined</emphasis>
.
<materialsCitation id="3B743C80FB7EC3584B6AF99853A6D6A2" ID-GBIF-Occurrence="3095329302" elevation="7" latitude="26.373333" location="Aka" longLatPrecision="123" longitude="126.79667" municipality="Kume Island" pageId="0" pageNumber="33" specimenCount="1">
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,
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,
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,
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, about
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depth
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, coll
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.
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. Kawai,
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<collectingDate id="EFE6E9F5FB7EC3584BD1F9B852ADD6A2" box="[1274,1409,1662,1686]" pageId="0" pageNumber="33" value="2012-06-15">15 June 2012</collectingDate>
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,
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(sl
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), RUMF-ZC-02041
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.
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<emphasis id="B968EACFFB7EC3584C65F9795347D6E3" bold="true" box="[846,1131,1727,1751]" pageId="0" pageNumber="33">Distinguishing characters</emphasis>
. Shield (
<figureCitation id="13272A58FB7EC3584BC8F9795210D6E3" box="[1251,1340,1727,1751]" captionStart="Fig" captionStartId="1.[158,189,1669,1691]" captionTargetBox="[237,1350,204,1647]" captionTargetId="figure-155@1.[237,1350,204,1647]" captionTargetPageId="1" captionText="Fig. 1. Aniculus erythraeus Forest, 1984, male (sl 8.7mm), Kume Island, RUMF-ZC-02041. A, shield and cephalic appendages, dorsal view (setae omitted from right side); B, left cheliped, lateral view (setae omitted); C, same, mesial view (setae omitted); D, same, chela, dorsal view (setae omitted); E, coxae of chelipeds, ventral view (setae omitted); F, sixth thoracic sternite, ventral view; G, telson, dorsal view (setae omitted from right side). Scales:2.0mm." figureDoi="http://doi.org/10.5281/zenodo.4584829" httpUri="https://zenodo.org/record/4584829/files/figure.png" pageId="0" pageNumber="33">Fig. 1A</figureCitation>
) longer than broad; mesogastric region in form of irregular diamond, delimited laterally by furrows. Rostrum (
<figureCitation id="13272A58FB7EC3584A15F8C652BCD72C" box="[1342,1424,1792,1816]" captionStart="Fig" captionStartId="1.[158,189,1669,1691]" captionTargetBox="[237,1350,204,1647]" captionTargetId="figure-155@1.[237,1350,204,1647]" captionTargetPageId="1" captionText="Fig. 1. Aniculus erythraeus Forest, 1984, male (sl 8.7mm), Kume Island, RUMF-ZC-02041. A, shield and cephalic appendages, dorsal view (setae omitted from right side); B, left cheliped, lateral view (setae omitted); C, same, mesial view (setae omitted); D, same, chela, dorsal view (setae omitted); E, coxae of chelipeds, ventral view (setae omitted); F, sixth thoracic sternite, ventral view; G, telson, dorsal view (setae omitted from right side). Scales:2.0mm." figureDoi="http://doi.org/10.5281/zenodo.4584829" httpUri="https://zenodo.org/record/4584829/files/figure.png" pageId="0" pageNumber="33">Fig. 1A</figureCitation>
) triangular, overreaching lateral projections; lateral projections each with submarginal spinule directed laterally. Ocular peduncles (
<figureCitation id="13272A58FB7EC3584CF7F8A45307D74E" box="[988,1067,1890,1914]" captionStart="Fig" captionStartId="1.[158,189,1669,1691]" captionTargetBox="[237,1350,204,1647]" captionTargetId="figure-155@1.[237,1350,204,1647]" captionTargetPageId="1" captionText="Fig. 1. Aniculus erythraeus Forest, 1984, male (sl 8.7mm), Kume Island, RUMF-ZC-02041. A, shield and cephalic appendages, dorsal view (setae omitted from right side); B, left cheliped, lateral view (setae omitted); C, same, mesial view (setae omitted); D, same, chela, dorsal view (setae omitted); E, coxae of chelipeds, ventral view (setae omitted); F, sixth thoracic sternite, ventral view; G, telson, dorsal view (setae omitted from right side). Scales:2.0mm." figureDoi="http://doi.org/10.5281/zenodo.4584829" httpUri="https://zenodo.org/record/4584829/files/figure.png" pageId="0" pageNumber="33">Fig. 1A</figureCitation>
) slender, 0.70.8 as long as shield; corneas not dilated; ocular acicles each with 4 terminal spinules. Antennular peduncles (
<figureCitation id="13272A58FB7EC3584B4DF8655398D78F" box="[1126,1204,1955,1979]" captionStart="Fig" captionStartId="1.[158,189,1669,1691]" captionTargetBox="[237,1350,204,1647]" captionTargetId="figure-155@1.[237,1350,204,1647]" captionTargetPageId="1" captionText="Fig. 1. Aniculus erythraeus Forest, 1984, male (sl 8.7mm), Kume Island, RUMF-ZC-02041. A, shield and cephalic appendages, dorsal view (setae omitted from right side); B, left cheliped, lateral view (setae omitted); C, same, mesial view (setae omitted); D, same, chela, dorsal view (setae omitted); E, coxae of chelipeds, ventral view (setae omitted); F, sixth thoracic sternite, ventral view; G, telson, dorsal view (setae omitted from right side). Scales:2.0mm." figureDoi="http://doi.org/10.5281/zenodo.4584829" httpUri="https://zenodo.org/record/4584829/files/figure.png" pageId="0" pageNumber="33">Fig. 1A</figureCitation>
) when fully extended, barely reaching corneal bases. Antennal peduncles (
<figureCitation id="13272A58FB7EC3584A5BF805547FD7C8" captionStart="Fig" captionStartId="1.[158,189,1669,1691]" captionTargetBox="[237,1350,204,1647]" captionTargetId="figure-155@1.[237,1350,204,1647]" captionTargetPageId="1" captionText="Fig. 1. Aniculus erythraeus Forest, 1984, male (sl 8.7mm), Kume Island, RUMF-ZC-02041. A, shield and cephalic appendages, dorsal view (setae omitted from right side); B, left cheliped, lateral view (setae omitted); C, same, mesial view (setae omitted); D, same, chela, dorsal view (setae omitted); E, coxae of chelipeds, ventral view (setae omitted); F, sixth thoracic sternite, ventral view; G, telson, dorsal view (setae omitted from right side). Scales:2.0mm." figureDoi="http://doi.org/10.5281/zenodo.4584829" httpUri="https://zenodo.org/record/4584829/files/figure.png" pageId="0" pageNumber="33">Fig. 1A</figureCitation>
) ending far short of distal margins of corneas; antennal acicle short, reaching slightly beyond proximal margin of fifth peduncular segment. Chelipeds (
<figureCitation id="13272A58FB7FC3594D63F8E755AED70D" box="[584,642,1825,1849]" captionStart="Fig" captionStartId="1.[158,189,1669,1691]" captionTargetBox="[237,1350,204,1647]" captionTargetId="figure-155@1.[237,1350,204,1647]" captionTargetPageId="1" captionText="Fig. 1. Aniculus erythraeus Forest, 1984, male (sl 8.7mm), Kume Island, RUMF-ZC-02041. A, shield and cephalic appendages, dorsal view (setae omitted from right side); B, left cheliped, lateral view (setae omitted); C, same, mesial view (setae omitted); D, same, chela, dorsal view (setae omitted); E, coxae of chelipeds, ventral view (setae omitted); F, sixth thoracic sternite, ventral view; G, telson, dorsal view (setae omitted from right side). Scales:2.0mm." figureDoi="http://doi.org/10.5281/zenodo.4584829" httpUri="https://zenodo.org/record/4584829/files/figure.png" pageId="1" pageNumber="34">Fig. 1</figureCitation>
BD) equal and similar in armament and ornamentation. Chelae (
<figureCitation id="13272A58FB7FC3594DF3F8845788D74E" captionStart="Fig" captionStartId="1.[158,189,1669,1691]" captionTargetBox="[237,1350,204,1647]" captionTargetId="figure-155@1.[237,1350,204,1647]" captionTargetPageId="1" captionText="Fig. 1. Aniculus erythraeus Forest, 1984, male (sl 8.7mm), Kume Island, RUMF-ZC-02041. A, shield and cephalic appendages, dorsal view (setae omitted from right side); B, left cheliped, lateral view (setae omitted); C, same, mesial view (setae omitted); D, same, chela, dorsal view (setae omitted); E, coxae of chelipeds, ventral view (setae omitted); F, sixth thoracic sternite, ventral view; G, telson, dorsal view (setae omitted from right side). Scales:2.0mm." figureDoi="http://doi.org/10.5281/zenodo.4584829" httpUri="https://zenodo.org/record/4584829/files/figure.png" pageId="1" pageNumber="34">Fig. 1</figureCitation>
BD) circumscribed by transverse striae; each stria marginally with row of short setae in general, but replaced on outer upper surface by small corneous spinules accompanied by long, stiff setae. Dactyli (
<figureCitation id="13272A58FB7FC3594EC0F802550DD7E8" box="[491,545,1988,2012]" captionStart="Fig" captionStartId="1.[158,189,1669,1691]" captionTargetBox="[237,1350,204,1647]" captionTargetId="figure-155@1.[237,1350,204,1647]" captionTargetPageId="1" captionText="Fig. 1. Aniculus erythraeus Forest, 1984, male (sl 8.7mm), Kume Island, RUMF-ZC-02041. A, shield and cephalic appendages, dorsal view (setae omitted from right side); B, left cheliped, lateral view (setae omitted); C, same, mesial view (setae omitted); D, same, chela, dorsal view (setae omitted); E, coxae of chelipeds, ventral view (setae omitted); F, sixth thoracic sternite, ventral view; G, telson, dorsal view (setae omitted from right side). Scales:2.0mm." figureDoi="http://doi.org/10.5281/zenodo.4584829" httpUri="https://zenodo.org/record/4584829/files/figure.png" pageId="1" pageNumber="34">Fig. 1</figureCitation>
BD) with transverse rows of small corneous spinules partially obscured by long stiff setae. Carpi (
<figureCitation id="13272A58FB7FC3594E75F7C356FDD829" box="[350,465,2053,2077]" captionStart="Fig" captionStartId="1.[158,189,1669,1691]" captionTargetBox="[237,1350,204,1647]" captionTargetId="figure-155@1.[237,1350,204,1647]" captionTargetPageId="1" captionText="Fig. 1. Aniculus erythraeus Forest, 1984, male (sl 8.7mm), Kume Island, RUMF-ZC-02041. A, shield and cephalic appendages, dorsal view (setae omitted from right side); B, left cheliped, lateral view (setae omitted); C, same, mesial view (setae omitted); D, same, chela, dorsal view (setae omitted); E, coxae of chelipeds, ventral view (setae omitted); F, sixth thoracic sternite, ventral view; G, telson, dorsal view (setae omitted from right side). Scales:2.0mm." figureDoi="http://doi.org/10.5281/zenodo.4584829" httpUri="https://zenodo.org/record/4584829/files/figure.png" pageId="1" pageNumber="34">Fig. 1B, C</figureCitation>
) each with corneous-tipped dorsodistal spine; outer face with marginally piliferous, generally entire, transverse striae. Meri (
<figureCitation id="13272A58FB7FC3594B9DF887520ED76E" box="[1206,1314,1857,1882]" captionStart="Fig" captionStartId="1.[158,189,1669,1691]" captionTargetBox="[237,1350,204,1647]" captionTargetId="figure-155@1.[237,1350,204,1647]" captionTargetPageId="1" captionText="Fig. 1. Aniculus erythraeus Forest, 1984, male (sl 8.7mm), Kume Island, RUMF-ZC-02041. A, shield and cephalic appendages, dorsal view (setae omitted from right side); B, left cheliped, lateral view (setae omitted); C, same, mesial view (setae omitted); D, same, chela, dorsal view (setae omitted); E, coxae of chelipeds, ventral view (setae omitted); F, sixth thoracic sternite, ventral view; G, telson, dorsal view (setae omitted from right side). Scales:2.0mm." figureDoi="http://doi.org/10.5281/zenodo.4584829" httpUri="https://zenodo.org/record/4584829/files/figure.png" pageId="1" pageNumber="34">Fig. 1B, C</figureCitation>
) each with ventrolateral proximal margin greatly inflated. Coxae (
<figureCitation id="13272A58FB7FC3594A5BF8A45463D7AF" captionStart="Fig" captionStartId="1.[158,189,1669,1691]" captionTargetBox="[237,1350,204,1647]" captionTargetId="figure-155@1.[237,1350,204,1647]" captionTargetPageId="1" captionText="Fig. 1. Aniculus erythraeus Forest, 1984, male (sl 8.7mm), Kume Island, RUMF-ZC-02041. A, shield and cephalic appendages, dorsal view (setae omitted from right side); B, left cheliped, lateral view (setae omitted); C, same, mesial view (setae omitted); D, same, chela, dorsal view (setae omitted); E, coxae of chelipeds, ventral view (setae omitted); F, sixth thoracic sternite, ventral view; G, telson, dorsal view (setae omitted from right side). Scales:2.0mm." figureDoi="http://doi.org/10.5281/zenodo.4584829" httpUri="https://zenodo.org/record/4584829/files/figure.png" pageId="1" pageNumber="34">Fig. 1E</figureCitation>
) each with 2 long, transverse striae on ventral surface. Ambulatory legs (
<figureCitation id="13272A58FB7FC3594CDBF8655301D78F" box="[1008,1069,1955,1979]" captionStart="Fig" captionStartId="2.[157,188,1751,1773]" captionTargetBox="[264,1323,204,1729]" captionTargetId="figure-106@2.[264,1323,204,1729]" captionTargetPageId="2" captionText="Fig. 2. Aniculus erythraeus Forest, 1984, male (sl 8.7mm), Kume Island, RUMF-ZC-02041 (AD). Aniculus ursus (Olivier, 1812), male (sl 18.6mm), KUMEJIMA 2009, Stn Dive 4, ZRC 2011.0704 (E); male (sl 21.8mm), KUMEJIMA 2009, Stn Dive 15, NTOU A01137 (F). A, left second pereopod, lateral view (most setae omitted); B, same, mesial view (most setae omitted); C, left third pereopod, lateral view (setae omitted); D, same, mesial view (setae omitted); E, left third pereopod, dactylus and propodus, lateral view (setae omitted); F, left third pereopod, propodus, lateral view (setae omitted). Scales: 2.0mm." figureDoi="http://doi.org/10.5281/zenodo.4584831" httpUri="https://zenodo.org/record/4584831/files/figure.png" pageId="1" pageNumber="34">Fig. 2</figureCitation>
AD) with numerous tufts of long setae on dorsal and ventral margins. Dactyli (
<figureCitation id="13272A58FB7FC3594A0BF8025273D7E8" box="[1312,1375,1988,2012]" captionStart="Fig" captionStartId="2.[157,188,1751,1773]" captionTargetBox="[264,1323,204,1729]" captionTargetId="figure-106@2.[264,1323,204,1729]" captionTargetPageId="2" captionText="Fig. 2. Aniculus erythraeus Forest, 1984, male (sl 8.7mm), Kume Island, RUMF-ZC-02041 (AD). Aniculus ursus (Olivier, 1812), male (sl 18.6mm), KUMEJIMA 2009, Stn Dive 4, ZRC 2011.0704 (E); male (sl 21.8mm), KUMEJIMA 2009, Stn Dive 15, NTOU A01137 (F). A, left second pereopod, lateral view (most setae omitted); B, same, mesial view (most setae omitted); C, left third pereopod, lateral view (setae omitted); D, same, mesial view (setae omitted); E, left third pereopod, dactylus and propodus, lateral view (setae omitted); F, left third pereopod, propodus, lateral view (setae omitted). Scales: 2.0mm." figureDoi="http://doi.org/10.5281/zenodo.4584831" httpUri="https://zenodo.org/record/4584831/files/figure.png" pageId="1" pageNumber="34">Fig. 2</figureCitation>
AD) each with longitudinal row of 13 transversely arranged corneous spinules ventrally on lateral surface and with tufts of long setae, dorsal and mesial surfaces also with several corneous spinules on distal 0.7, ventral margin with row of 6 or 7 corneous spines. Propodi (
<figureCitation id="13272A58FB7CC35A4E90F80256D9D7E8" box="[443,501,1988,2012]" captionStart="Fig" captionStartId="2.[157,188,1751,1773]" captionTargetBox="[264,1323,204,1729]" captionTargetId="figure-106@2.[264,1323,204,1729]" captionTargetPageId="2" captionText="Fig. 2. Aniculus erythraeus Forest, 1984, male (sl 8.7mm), Kume Island, RUMF-ZC-02041 (AD). Aniculus ursus (Olivier, 1812), male (sl 18.6mm), KUMEJIMA 2009, Stn Dive 4, ZRC 2011.0704 (E); male (sl 21.8mm), KUMEJIMA 2009, Stn Dive 15, NTOU A01137 (F). A, left second pereopod, lateral view (most setae omitted); B, same, mesial view (most setae omitted); C, left third pereopod, lateral view (setae omitted); D, same, mesial view (setae omitted); E, left third pereopod, dactylus and propodus, lateral view (setae omitted); F, left third pereopod, propodus, lateral view (setae omitted). Scales: 2.0mm." figureDoi="http://doi.org/10.5281/zenodo.4584831" httpUri="https://zenodo.org/record/4584831/files/figure.png" pageId="2" pageNumber="35">Fig. 2</figureCitation>
AD) each with marginally piliferous, long, transverse striae on lateral surface: proximal-most and penultimate striae interrupted, but median 4 striae uninterrupted; on third pereopod, additional short stria present near ventroproximal end of lateral surface; dorsal surface with several corneous spinules on distal half (second pereopod) or only 1 subdistal mesial spinule (third pereopod); ventral surface also with several corneous spinules on distal half. Carpi (
<figureCitation id="13272A58FB7DC35B4E5FF9DB5682D601" box="[372,430,1565,1589]" captionStart="Fig" captionStartId="2.[157,188,1751,1773]" captionTargetBox="[264,1323,204,1729]" captionTargetId="figure-106@2.[264,1323,204,1729]" captionTargetPageId="2" captionText="Fig. 2. Aniculus erythraeus Forest, 1984, male (sl 8.7mm), Kume Island, RUMF-ZC-02041 (AD). Aniculus ursus (Olivier, 1812), male (sl 18.6mm), KUMEJIMA 2009, Stn Dive 4, ZRC 2011.0704 (E); male (sl 21.8mm), KUMEJIMA 2009, Stn Dive 15, NTOU A01137 (F). A, left second pereopod, lateral view (most setae omitted); B, same, mesial view (most setae omitted); C, left third pereopod, lateral view (setae omitted); D, same, mesial view (setae omitted); E, left third pereopod, dactylus and propodus, lateral view (setae omitted); F, left third pereopod, propodus, lateral view (setae omitted). Scales: 2.0mm." figureDoi="http://doi.org/10.5281/zenodo.4584831" httpUri="https://zenodo.org/record/4584831/files/figure.png" pageId="3" pageNumber="36">Fig. 2</figureCitation>
AD) each with small, corneoustipped dorsodistal spine; lateral surface with marginally piliferous, generally entire, transverse striae. Meri (
<figureCitation id="13272A58FB7DC35B4DA7F99855EAD642" box="[652,710,1630,1654]" captionStart="Fig" captionStartId="2.[157,188,1751,1773]" captionTargetBox="[264,1323,204,1729]" captionTargetId="figure-106@2.[264,1323,204,1729]" captionTargetPageId="2" captionText="Fig. 2. Aniculus erythraeus Forest, 1984, male (sl 8.7mm), Kume Island, RUMF-ZC-02041 (AD). Aniculus ursus (Olivier, 1812), male (sl 18.6mm), KUMEJIMA 2009, Stn Dive 4, ZRC 2011.0704 (E); male (sl 21.8mm), KUMEJIMA 2009, Stn Dive 15, NTOU A01137 (F). A, left second pereopod, lateral view (most setae omitted); B, same, mesial view (most setae omitted); C, left third pereopod, lateral view (setae omitted); D, same, mesial view (setae omitted); E, left third pereopod, dactylus and propodus, lateral view (setae omitted); F, left third pereopod, propodus, lateral view (setae omitted). Scales: 2.0mm." figureDoi="http://doi.org/10.5281/zenodo.4584831" httpUri="https://zenodo.org/record/4584831/files/figure.png" pageId="3" pageNumber="36">Fig. 2</figureCitation>
AD) with medially interrupted transverse striae (pronounced and much longer in third pereopod) on lateral surface, ventrolateral margin with small subdistal spine (second pereopod) or unarmed (third pereopod), ventromesial margin denticulate (second pereopod) or slightly crenulated (third pereopod). Sixth thoracic sternite (
<figureCitation id="13272A58FB7DC35B4EB9F8E756F0D70D" box="[402,476,1825,1849]" captionStart="Fig" captionStartId="1.[158,189,1669,1691]" captionTargetBox="[237,1350,204,1647]" captionTargetId="figure-155@1.[237,1350,204,1647]" captionTargetPageId="1" captionText="Fig. 1. Aniculus erythraeus Forest, 1984, male (sl 8.7mm), Kume Island, RUMF-ZC-02041. A, shield and cephalic appendages, dorsal view (setae omitted from right side); B, left cheliped, lateral view (setae omitted); C, same, mesial view (setae omitted); D, same, chela, dorsal view (setae omitted); E, coxae of chelipeds, ventral view (setae omitted); F, sixth thoracic sternite, ventral view; G, telson, dorsal view (setae omitted from right side). Scales:2.0mm." figureDoi="http://doi.org/10.5281/zenodo.4584829" httpUri="https://zenodo.org/record/4584829/files/figure.png" pageId="3" pageNumber="36">Fig. 1F</figureCitation>
) with oblong anterior lobe. Telson (
<figureCitation id="13272A58FB7DC35B4FD8F887566BD76E" box="[243,327,1857,1882]" captionStart="Fig" captionStartId="1.[158,189,1669,1691]" captionTargetBox="[237,1350,204,1647]" captionTargetId="figure-155@1.[237,1350,204,1647]" captionTargetPageId="1" captionText="Fig. 1. Aniculus erythraeus Forest, 1984, male (sl 8.7mm), Kume Island, RUMF-ZC-02041. A, shield and cephalic appendages, dorsal view (setae omitted from right side); B, left cheliped, lateral view (setae omitted); C, same, mesial view (setae omitted); D, same, chela, dorsal view (setae omitted); E, coxae of chelipeds, ventral view (setae omitted); F, sixth thoracic sternite, ventral view; G, telson, dorsal view (setae omitted from right side). Scales:2.0mm." figureDoi="http://doi.org/10.5281/zenodo.4584829" httpUri="https://zenodo.org/record/4584829/files/figure.png" pageId="3" pageNumber="36">Fig. 1G</figureCitation>
) with somewhat asymmetrical posterior lobes, left larger than right; terminal margins oblique.
</paragraph>
<caption id="DF636655FB7FC3594FB5F94356CDD6DA" ID-DOI="http://doi.org/10.5281/zenodo.4584829" ID-Zenodo-Dep="4584829" httpUri="https://zenodo.org/record/4584829/files/figure.png" pageId="1" pageNumber="34" startId="1.[158,189,1669,1691]" targetBox="[237,1350,204,1647]" targetPageId="1">
<paragraph id="8BA336DDFB7FC3594FB5F94356CDD6DA" blockId="1.[156,1432,1669,1775]" pageId="1" pageNumber="34">
Fig. 1.
<taxonomicName id="4C1C4D5EFB7FC3594FDBF943553BD6AE" authority="Forest, 1984" authorityName="Forest" authorityYear="1984" box="[240,535,1669,1691]" class="Malacostraca" family="Diogenidae" genus="Aniculus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="34" phylum="Arthropoda" rank="species" species="erythraeus">
<emphasis id="B968EACFFB7FC3594FDBF943568DD6AE" box="[240,417,1669,1690]" italics="true" pageId="1" pageNumber="34">Aniculus erythraeus</emphasis>
<bibRefCitation id="EF8D4B2CFB7FC3594E8CF940553BD6AE" author="Forest, J." box="[423,535,1669,1691]" pageId="1" pageNumber="34" pagination="1 - 91" refId="ref2911" refString="Forest, J. 1984. Revision du genre Aniculus. Crustaceana (Supplement 8): 1 - 91." type="journal article" year="1984">Forest, 1984</bibRefCitation>
</taxonomicName>
, male (sl 8.7 mm), Kume Island, RUMF-ZC-02041. A, shield and cephalic appendages, dorsal view (setae omitted from right side); B, left cheliped, lateral view (setae omitted); C, same, mesial view (setae omitted); D, same, chela, dorsal view (setae omitted); E, coxae of chelipeds, ventral view (setae omitted); F, sixth thoracic sternite, ventral view; G, telson, dorsal view (setae omitted from right side). Scales: 2.0 mm.
</paragraph>
</caption>
<caption id="DF636655FB7CC35A4FB6F9115598D768" ID-DOI="http://doi.org/10.5281/zenodo.4584831" ID-Zenodo-Dep="4584831" httpUri="https://zenodo.org/record/4584831/files/figure.png" pageId="2" pageNumber="35" startId="2.[157,188,1751,1773]" targetBox="[264,1323,204,1729]" targetPageId="2">
<paragraph id="8BA336DDFB7CC35A4FB6F9115598D768" blockId="2.[156,1432,1751,1885]" pageId="2" pageNumber="35">
Fig. 2.
<taxonomicName id="4C1C4D5EFB7CC35A4FDBF9115535D6D8" authority="Forest, 1984" authorityName="Forest" authorityYear="1984" box="[240,537,1751,1772]" class="Malacostraca" family="Diogenidae" genus="Aniculus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="35" phylum="Arthropoda" rank="species" species="erythraeus">
<emphasis id="B968EACFFB7CC35A4FDBF911568ED6D8" box="[240,418,1751,1772]" italics="true" pageId="2" pageNumber="35">Aniculus erythraeus</emphasis>
<bibRefCitation id="EF8D4B2CFB7CC35A4E83F9115535D6D8" author="Forest, J." box="[424,537,1751,1772]" pageId="2" pageNumber="35" pagination="1 - 91" refId="ref2911" refString="Forest, J. 1984. Revision du genre Aniculus. Crustaceana (Supplement 8): 1 - 91." type="journal article" year="1984">Forest, 1984</bibRefCitation>
</taxonomicName>
, male (sl 8.7mm), Kume Island, RUMF-ZC-02041 (AD).
<taxonomicName id="4C1C4D5EFB7CC35A4B1EF9115269D6D9" authority="(Olivier, 1812)" authorityName="Nomura et al." authorityYear="1996" baseAuthorityName="Olivier" baseAuthorityYear="1812" box="[1077,1349,1751,1773]" class="Malacostraca" family="Diogenidae" genus="Aniculus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="35" phylum="Arthropoda" rank="species" species="ursus">
<emphasis id="B968EACFFB7CC35A4B1EF9115395D6D9" box="[1077,1209,1751,1773]" italics="true" pageId="2" pageNumber="35">Aniculus ursus</emphasis>
(Olivier, 1812)
</taxonomicName>
, male (sl 18.6mm), KUMEJIMA 2009, Stn Dive 4, ZRC 2011.0704 (E); male (sl 21.8 mm), KUMEJIMA 2009, Stn Dive 15, NTOU
<accessionNumber id="944FAB3EFB7CC35A4BC6F9355218D73C" box="[1261,1332,1779,1800]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/A01137" pageId="2" pageNumber="35" type="EnaNcbi">A01137</accessionNumber>
(F). A, left second pereopod, lateral view (most setae omitted); B, same, mesial view (most setae omitted); C, left third pereopod, lateral view (setae omitted); D, same, mesial view (setae omitted); E, left third pereopod, dactylus and propodus, lateral view (setae omitted); F, left third pereopod, propodus, lateral view (setae omitted). Scales: 2.0mm.
</paragraph>
</caption>
<caption id="DF636655FB7DC35B4FB7FA0F523CD5CF" ID-DOI="http://doi.org/10.5281/zenodo.4584833" ID-Zenodo-Dep="4584833" httpUri="https://zenodo.org/record/4584833/files/figure.png" pageId="3" pageNumber="36" startId="3.[156,187,1481,1503]" targetBox="[239,1348,204,1458]" targetPageId="3">
<paragraph id="8BA336DDFB7DC35B4FB7FA0F523CD5CF" blockId="3.[156,1431,1481,1531]" pageId="3" pageNumber="36">
Fig. 3.
<taxonomicName id="4C1C4D5EFB7DC35B4FDBFA0F553BD5EA" authority="Forest, 1984" authorityName="Forest" authorityYear="1984" box="[240,535,1481,1502]" class="Malacostraca" family="Diogenidae" genus="Aniculus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="36" phylum="Arthropoda" rank="species" species="erythraeus">
<emphasis id="B968EACFFB7DC35B4FDBFA0F568ED5EA" box="[240,418,1481,1502]" italics="true" pageId="3" pageNumber="36">Aniculus erythraeus</emphasis>
<bibRefCitation id="EF8D4B2CFB7DC35B4E8CFA0F553BD5EA" author="Forest, J." box="[423,535,1481,1502]" pageId="3" pageNumber="36" pagination="1 - 91" refId="ref2911" refString="Forest, J. 1984. Revision du genre Aniculus. Crustaceana (Supplement 8): 1 - 91." type="journal article" year="1984">Forest, 1984</bibRefCitation>
</taxonomicName>
, male (sl 8.7 mm), Kume Island, RUMF-ZC-02041. A, entire animal, dorsal view; B, same, left, lateral view; C, anterior body, dorsal view; D, same, ventral view; E, left cheliped, lateral view; F, left ambulatory legs, lateral view.
</paragraph>
</caption>
<paragraph id="8BA336DDFB7DC35B4F9EF84454F3D828" blockId="3.[156,768,1565,2077]" lastBlockId="3.[820,1433,1565,2077]" pageId="3" pageNumber="36">
<emphasis id="B968EACFFB7DC35B4F9EF8445606D7AE" bold="true" box="[181,298,1922,1946]" pageId="3" pageNumber="36">Coloration</emphasis>
(
<figureCitation id="13272A58FB7DC35B4E13F845565FD7AF" box="[312,371,1923,1947]" captionStart="Fig" captionStartId="3.[156,187,1481,1503]" captionTargetBox="[239,1348,204,1458]" captionTargetId="figure-288@3.[239,1348,204,1458]" captionTargetPageId="3" captionText="Fig. 3. Aniculus erythraeus Forest, 1984, male (sl 8.7mm), Kume Island, RUMF-ZC-02041. A, entire animal, dorsal view; B, same, left, lateral view; C, anterior body, dorsal view; D, same, ventral view; E, left cheliped, lateral view; F, left ambulatory legs, lateral view." figureDoi="http://doi.org/10.5281/zenodo.4584833" httpUri="https://zenodo.org/record/4584833/files/figure.png" pageId="3" pageNumber="36">Fig. 3</figureCitation>
). Shield and posterior carapace (
<figureCitation id="13272A58FB7DC35B4DF3F845578AD78F" captionStart="Fig" captionStartId="3.[156,187,1481,1503]" captionTargetBox="[239,1348,204,1458]" captionTargetId="figure-288@3.[239,1348,204,1458]" captionTargetPageId="3" captionText="Fig. 3. Aniculus erythraeus Forest, 1984, male (sl 8.7mm), Kume Island, RUMF-ZC-02041. A, entire animal, dorsal view; B, same, left, lateral view; C, anterior body, dorsal view; D, same, ventral view; E, left cheliped, lateral view; F, left ambulatory legs, lateral view." figureDoi="http://doi.org/10.5281/zenodo.4584833" httpUri="https://zenodo.org/record/4584833/files/figure.png" pageId="3" pageNumber="36">Fig. 3</figureCitation>
AC) reddish purple, with irregular white streaks. Ocular peduncles (
<figureCitation id="13272A58FB7DC35B4E3BF8025657D7E8" box="[272,379,1988,2012]" captionStart="Fig" captionStartId="3.[156,187,1481,1503]" captionTargetBox="[239,1348,204,1458]" captionTargetId="figure-288@3.[239,1348,204,1458]" captionTargetPageId="3" captionText="Fig. 3. Aniculus erythraeus Forest, 1984, male (sl 8.7mm), Kume Island, RUMF-ZC-02041. A, entire animal, dorsal view; B, same, left, lateral view; C, anterior body, dorsal view; D, same, ventral view; E, left cheliped, lateral view; F, left ambulatory legs, lateral view." figureDoi="http://doi.org/10.5281/zenodo.4584833" httpUri="https://zenodo.org/record/4584833/files/figure.png" pageId="3" pageNumber="36">Fig. 3A, C</figureCitation>
) brownish purple, with white spots at bases of setae; distal margin pale yellow. Ocular acicles and antennular and antennal peduncles (
<figureCitation id="13272A58FB7DC35B4D03F7C355B2D829" box="[552,670,2053,2077]" captionStart="Fig" captionStartId="3.[156,187,1481,1503]" captionTargetBox="[239,1348,204,1458]" captionTargetId="figure-288@3.[239,1348,204,1458]" captionTargetPageId="3" captionText="Fig. 3. Aniculus erythraeus Forest, 1984, male (sl 8.7mm), Kume Island, RUMF-ZC-02041. A, entire animal, dorsal view; B, same, left, lateral view; C, anterior body, dorsal view; D, same, ventral view; E, left cheliped, lateral view; F, left ambulatory legs, lateral view." figureDoi="http://doi.org/10.5281/zenodo.4584833" httpUri="https://zenodo.org/record/4584833/files/figure.png" pageId="3" pageNumber="36">Fig. 3A, C</figureCitation>
) reddish purple, with irregular white markings. Chelipeds (
<figureCitation id="13272A58FB7DC35B4A6DF9DB544AD661" captionStart="Fig" captionStartId="3.[156,187,1481,1503]" captionTargetBox="[239,1348,204,1458]" captionTargetId="figure-288@3.[239,1348,204,1458]" captionTargetPageId="3" captionText="Fig. 3. Aniculus erythraeus Forest, 1984, male (sl 8.7mm), Kume Island, RUMF-ZC-02041. A, entire animal, dorsal view; B, same, left, lateral view; C, anterior body, dorsal view; D, same, ventral view; E, left cheliped, lateral view; F, left ambulatory legs, lateral view." figureDoi="http://doi.org/10.5281/zenodo.4584833" httpUri="https://zenodo.org/record/4584833/files/figure.png" pageId="3" pageNumber="36">Fig. 3B, D, E</figureCitation>
) reddish purple, with white streak along each transverse stria; dactyli, carpi, and meri with additional small white markings; corneous claws and spinules on chela black. Ambulatory legs (second and third pereopods;
<figureCitation id="13272A58FB7DC35B4A39F95952BCD683" box="[1298,1424,1695,1719]" captionStart="Fig" captionStartId="3.[156,187,1481,1503]" captionTargetBox="[239,1348,204,1458]" captionTargetId="figure-288@3.[239,1348,204,1458]" captionTargetPageId="3" captionText="Fig. 3. Aniculus erythraeus Forest, 1984, male (sl 8.7mm), Kume Island, RUMF-ZC-02041. A, entire animal, dorsal view; B, same, left, lateral view; C, anterior body, dorsal view; D, same, ventral view; E, left cheliped, lateral view; F, left ambulatory legs, lateral view." figureDoi="http://doi.org/10.5281/zenodo.4584833" httpUri="https://zenodo.org/record/4584833/files/figure.png" pageId="3" pageNumber="36">Fig. 3A, B, F</figureCitation>
) also reddish purple, with white streak along each transverse stria; dactyli each with black corneous claw; propodi each with broad white part proximally and narrow white part distally; carpi white on proximoventral part; meri with irregular white markings. Fourth and fifth pereopods (
<figureCitation id="13272A58FB7DC35B4A01F88752BCD76E" box="[1322,1424,1857,1882]" captionStart="Fig" captionStartId="3.[156,187,1481,1503]" captionTargetBox="[239,1348,204,1458]" captionTargetId="figure-288@3.[239,1348,204,1458]" captionTargetPageId="3" captionText="Fig. 3. Aniculus erythraeus Forest, 1984, male (sl 8.7mm), Kume Island, RUMF-ZC-02041. A, entire animal, dorsal view; B, same, left, lateral view; C, anterior body, dorsal view; D, same, ventral view; E, left cheliped, lateral view; F, left ambulatory legs, lateral view." figureDoi="http://doi.org/10.5281/zenodo.4584833" httpUri="https://zenodo.org/record/4584833/files/figure.png" pageId="3" pageNumber="36">Fig. 3B, F</figureCitation>
) with mottled pattern of reddish purple and white, propodal rasps brown. Pleon (
<figureCitation id="13272A58FB7DC35B4B22F8445358D7AF" box="[1033,1140,1922,1947]" captionStart="Fig" captionStartId="3.[156,187,1481,1503]" captionTargetBox="[239,1348,204,1458]" captionTargetId="figure-288@3.[239,1348,204,1458]" captionTargetPageId="3" captionText="Fig. 3. Aniculus erythraeus Forest, 1984, male (sl 8.7mm), Kume Island, RUMF-ZC-02041. A, entire animal, dorsal view; B, same, left, lateral view; C, anterior body, dorsal view; D, same, ventral view; E, left cheliped, lateral view; F, left ambulatory legs, lateral view." figureDoi="http://doi.org/10.5281/zenodo.4584833" httpUri="https://zenodo.org/record/4584833/files/figure.png" pageId="3" pageNumber="36">Fig. 3A, B</figureCitation>
) pale red; tergites of first to fifth somites reddish purple, with irregular white transverse lines; tergite of sixth somite and telson also reddish purple, with irregular white markings. Setae purple, with broad, white distal part.
</paragraph>
<paragraph id="8BA336DDFB7AC35C4F9EFF005672D154" blockId="4.[156,768,198,2076]" pageId="4" pageNumber="37">
<emphasis id="B968EACFFB7AC35C4F9EFF005629D0EA" bold="true" box="[181,261,198,222]" pageId="4" pageNumber="37">Habitat</emphasis>
. Under a large block of dead coral at about
<quantity id="4CE49B38FB7AC35C4DF0FF0055D3D0EA" box="[731,767,198,222]" metricMagnitude="0" metricUnit="m" metricValue="3.0" pageId="4" pageNumber="37" unit="m" value="3.0">3 m</quantity>
depth (
<bibRefCitation id="EF8D4B2CFB7AC35C4FCFFF215685D0CB" author="Haig, J. &amp; Ball, E. E." box="[228,425,231,255]" pageId="4" pageNumber="37" pagination="151 - 196" refId="ref2932" refString="Haig, J. and Ball, E. E. 1988. Hermit crabs from northern Australian and eastern Indonesian waters (Crustacea Decapoda: Anomura: Paguroidea) collected during the 1975 Alpha Helix Expedition. Records of the Australian Museum 40: 151 - 196." type="journal article" year="1988">Haig and Ball 1988</bibRefCitation>
) or on outer reef slopes at depths of
<quantity id="4CE49B38FB7AC35C4F9DFEC1562CD12B" box="[182,256,263,287]" metricMagnitude="1" metricUnit="m" metricValue="1.75" metricValueMax="3.0" metricValueMin="0.5" pageId="4" pageNumber="37" unit="m" value="17.5" valueMax="30.0" valueMin="5.0">530 m</quantity>
(
<bibRefCitation id="EF8D4B2CFB7AC35C4E27FEC1553ED114" author="Poupin, J. &amp; Juncker, M." box="[268,530,263,288]" pageId="4" pageNumber="37" refId="ref3382" refString="Poupin, J. and Juncker, M. 2010. Guide des Crustaces Decapodes du Pacifique Sud / A Guide to Decapod Crustaceans of the South Pacific. CRISP and CPS, Noumea, 317 pp." type="book" year="2010">Poupin and Juncker 2010</bibRefCitation>
). The present specimen was collected from a hard bottom on a steep coral reef slope at about
<quantity id="4CE49B38FB7AC35C4FD8FE8F563BD154" box="[243,279,329,352]" metricMagnitude="0" metricUnit="m" metricValue="7.0" pageId="4" pageNumber="37" unit="m" value="7.0">7 m</quantity>
depth.
</paragraph>
</subSubSection>
<subSubSection id="C3066556FB7AC35C4F9EFEAF5500D251" pageId="4" pageNumber="37" type="distribution">
<paragraph id="8BA336DDFB7AC35C4F9EFEAF5500D251" blockId="4.[156,768,198,2076]" pageId="4" pageNumber="37">
<emphasis id="B968EACFFB7AC35C4F9EFEAF5616D1B5" bold="true" box="[181,314,361,385]" pageId="4" pageNumber="37">Distribution</emphasis>
. Red Sea,
<collectingCountry id="F30B764DFB7AC35C4E94FEAF550BD1B5" box="[447,551,361,385]" name="Sri Lanka" pageId="4" pageNumber="37">Sri Lanka</collectingCountry>
, Banda and
<collectingCountry id="F30B764DFB7AC35C4D99FEAF55D3D1B5" box="[690,767,361,385]" name="Indonesia" pageId="4" pageNumber="37">Ambon</collectingCountry>
Islands in
<collectingCountry id="F30B764DFB7AC35C4E22FE4F5643D195" box="[265,367,393,417]" name="Indonesia" pageId="4" pageNumber="37">Indonesia</collectingCountry>
, and Tuamotu Islands in
<collectingCountry id="F30B764DFB7AC35C4D50FE4F57FCD1F6" name="French Polynesia" pageId="4" pageNumber="37">French Polynesia</collectingCountry>
(
<bibRefCitation id="EF8D4B2CFB7AC35C4FCAFE6C553FD1F6" author="Rahayu, D. L. &amp; Wahyundi, A. J." box="[225,531,426,450]" pageId="4" pageNumber="37" refId="ref3457" refString="Rahayu, D. L. and Wahyundi, A. J. 2008. Common Littoral Hermit Crabs of Indonesia (Susetiono, Shirayama, Y. and Asakura, A., Eds). Kyoto University Press, Kyoto, 93 pp." type="book" year="2008">Rahayu and Wahyundi 2008</bibRefCitation>
; Poupin 2010;
<bibRefCitation id="EF8D4B2CFB7AC35C4D9DFE6C5671D1D7" author="Poupin, J. &amp; Juncker, M." pageId="4" pageNumber="37" refId="ref3382" refString="Poupin, J. and Juncker, M. 2010. Guide des Crustaces Decapodes du Pacifique Sud / A Guide to Decapod Crustaceans of the South Pacific. CRISP and CPS, Noumea, 317 pp." type="book" year="2010">Poupin and Juncker 2010</bibRefCitation>
). In addition to these records,
<taxonomicName id="4C1C4D5EFB7AC35C4D9EFE0D57C9D237" authorityName="Forest" authorityYear="1984" class="Malacostraca" family="Diogenidae" genus="Aniculus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="37" phylum="Arthropoda" rank="species" species="erythraeus">
<emphasis id="B968EACFFB7AC35C4D9EFE0D57C9D237" italics="true" pageId="4" pageNumber="37">A. erythraeus</emphasis>
</taxonomicName>
has also been collected from
<collectingCountry id="F30B764DFB7AC35C4D33FE2D557BD237" box="[536,599,491,515]" name="Guam" pageId="4" pageNumber="37">Guam</collectingCountry>
and
<collectingCountry id="F30B764DFB7AC35C4DA0FE2A57CAD210" name="Papua New Guinea" pageId="4" pageNumber="37">Papua New Guinea</collectingCountry>
(see
<bibRefCitation id="EF8D4B2CFB7AC35C4E35FDCA553BD210" author="Legall, N. &amp; Poupin, J." box="[286,535,524,548]" pageId="4" pageNumber="37" refId="ref3013" refString="Legall, N. and Poupin, J. 2012. CRUSTA: Database of Crustacea (Decapoda and Stomatopoda), with special interest for those collected in French overseas territories. Available at http: // crustiesfroverseas. free. fr / (10 December 2012)." type="url" year="2012">Legall and Poupin 2012</bibRefCitation>
). The specimen examined from the Ryukyu Islands greatly extends the distributional range of this species to the north.
</paragraph>
</subSubSection>
<subSubSection id="C3066556FB7AC35C4F9EFDAB5243D2F1" pageId="4" pageNumber="37" type="discussion">
<paragraph id="8BA336DDFB7AC35C4F9EFDAB55BAD438" blockId="4.[156,768,198,2076]" pageId="4" pageNumber="37">
<emphasis id="B968EACFFB7AC35C4F9EFDAB563ED2B1" bold="true" box="[181,274,621,645]" pageId="4" pageNumber="37">Remarks</emphasis>
. The specimen examined is an immature male: the body size is small (sl
<quantity id="4CE49B38FB7AC35C4EC2FD48551AD292" box="[489,566,654,678]" metricMagnitude="-3" metricUnit="m" metricValue="8.7" pageId="4" pageNumber="37" unit="mm" value="8.7">8.7 mm</quantity>
) and the gonopore is not well developed and present only on the coxa of the right fifth pereopod.
<bibRefCitation id="EF8D4B2CFB7AC35C4EA9FD095525D2D3" author="Forest, J." box="[386,521,719,743]" pageId="4" pageNumber="37" pagination="1 - 91" refId="ref2911" refString="Forest, J. 1984. Revision du genre Aniculus. Crustaceana (Supplement 8): 1 - 91." type="journal article" year="1984">Forest (1984</bibRefCitation>
: fig. 12) described the sixth pleonal tergite of
<taxonomicName id="4C1C4D5EFB7AC35C4EB9FD365530D333" authorityName="Forest" authorityYear="1984" box="[402,540,751,775]" class="Malacostraca" family="Diogenidae" genus="Aniculus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="37" phylum="Arthropoda" rank="species" species="erythraeus">
<emphasis id="B968EACFFB7AC35C4EB9FD365530D333" box="[402,540,751,775]" italics="true" pageId="4" pageNumber="37">A. erythraeus</emphasis>
</taxonomicName>
as having posteriorly directed short spinules on the anterolateral part; however, in the present specimen there are no distinct spinules, only a few indistinct small protuberances. The dactylus of the left third pereopod is subequal in length to or longer than the propodus in the
<typeStatus id="54A7887FFB7AC35C4E68FC545643D39E" box="[323,367,914,938]" pageId="4" pageNumber="37">type</typeStatus>
material, but it is shorter in the present specimen. Nevertheless, the present specimen can be identified to
<taxonomicName id="4C1C4D5EFB7AC35C4FD3FC1556ADD3DF" authorityName="Forest" authorityYear="1984" box="[248,385,979,1003]" class="Malacostraca" family="Diogenidae" genus="Aniculus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="37" phylum="Arthropoda" rank="species" species="erythraeus">
<emphasis id="B968EACFFB7AC35C4FD3FC1556ADD3DF" box="[248,385,979,1003]" italics="true" pageId="4" pageNumber="37">A. erythraeus</emphasis>
</taxonomicName>
inasmuch as it has other diagnostic morphological and color characters of the species.
</paragraph>
<paragraph id="8BA336DDFB7AC35C4F9EFBD255B1D78F" blockId="4.[156,768,198,2076]" pageId="4" pageNumber="37">
<taxonomicName id="4C1C4D5EFB7AC35C4F9EFBD25651D418" authorityName="Forest" authorityYear="1984" box="[181,381,1044,1068]" class="Malacostraca" family="Diogenidae" genus="Aniculus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="37" phylum="Arthropoda" rank="species" species="erythraeus">
<emphasis id="B968EACFFB7AC35C4F9EFBD25651D418" box="[181,381,1044,1068]" italics="true" pageId="4" pageNumber="37">Aniculus erythraeus</emphasis>
</taxonomicName>
is morphologically allied to
<taxonomicName id="4C1C4D5EFB7AC35C4D8DFBD255D6D418" authorityName="Nomura et al." authorityYear="1996" baseAuthorityName="Olivier" baseAuthorityYear="1812" box="[678,762,1044,1068]" class="Malacostraca" family="Diogenidae" genus="Aniculus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="37" phylum="Arthropoda" rank="species" species="ursus">
<emphasis id="B968EACFFB7AC35C4D8DFBD255D6D418" box="[678,762,1044,1068]" italics="true" pageId="4" pageNumber="37">A. ursus</emphasis>
</taxonomicName>
. The characters shared by the two species are: mesogastric region of shield delimited laterally by furrows; ocular acicle with more than two terminal spinules; chelipeds with small corneous spinules only on outer upper surface of each palm and with two long, transverse striae on ventral surface of each coxa; and propodi of second and third pereopods with uninterrupted striae on median part of lateral surface. Preliminary comparison of the figures provided by
<bibRefCitation id="EF8D4B2CFB7AC35C4FB7FAFF5631D565" author="Forest, J." box="[156,285,1337,1361]" pageId="4" pageNumber="37" pagination="1 - 91" refId="ref2911" refString="Forest, J. 1984. Revision du genre Aniculus. Crustaceana (Supplement 8): 1 - 91." type="journal article" year="1984">Forest (1984</bibRefCitation>
: figs 33, 43, and 45) shows that the structure of the proximal-most and penultimate long, transverse striae on the lateral surface of the left third pereopod is different between the two species; however, those striae vary in the two examined specimens of
<taxonomicName id="4C1C4D5EFB7AC35C4EEFFA7D5534D5E7" authorityName="Nomura et al." authorityYear="1996" baseAuthorityName="Olivier" baseAuthorityYear="1812" box="[452,536,1467,1491]" class="Malacostraca" family="Diogenidae" genus="Aniculus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="37" phylum="Arthropoda" rank="species" species="ursus">
<emphasis id="B968EACFFB7AC35C4EEFFA7D5534D5E7" box="[452,536,1467,1491]" italics="true" pageId="4" pageNumber="37">A. ursus</emphasis>
</taxonomicName>
(
<figureCitation id="13272A58FB7AC35C4D03FA7D55BDD5E7" box="[552,657,1467,1491]" captionStart="Fig" captionStartId="2.[157,188,1751,1773]" captionTargetBox="[264,1323,204,1729]" captionTargetId="figure-106@2.[264,1323,204,1729]" captionTargetPageId="2" captionText="Fig. 2. Aniculus erythraeus Forest, 1984, male (sl 8.7mm), Kume Island, RUMF-ZC-02041 (AD). Aniculus ursus (Olivier, 1812), male (sl 18.6mm), KUMEJIMA 2009, Stn Dive 4, ZRC 2011.0704 (E); male (sl 21.8mm), KUMEJIMA 2009, Stn Dive 15, NTOU A01137 (F). A, left second pereopod, lateral view (most setae omitted); B, same, mesial view (most setae omitted); C, left third pereopod, lateral view (setae omitted); D, same, mesial view (setae omitted); E, left third pereopod, dactylus and propodus, lateral view (setae omitted); F, left third pereopod, propodus, lateral view (setae omitted). Scales: 2.0mm." figureDoi="http://doi.org/10.5281/zenodo.4584831" httpUri="https://zenodo.org/record/4584831/files/figure.png" pageId="4" pageNumber="37">Fig. 2E, F</figureCitation>
), and thus
<taxonomicName id="4C1C4D5EFB7AC35C4FB7FA1A560ED5C7" authorityName="Forest" authorityYear="1984" box="[156,290,1499,1523]" class="Malacostraca" family="Diogenidae" genus="Aniculus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="37" phylum="Arthropoda" rank="species" species="erythraeus">
<emphasis id="B968EACFFB7AC35C4FB7FA1A560ED5C7" box="[156,290,1499,1523]" italics="true" pageId="4" pageNumber="37">A. erythraeus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C1C4D5EFB7AC35C4E7EFA1A5684D5C7" authorityName="Nomura et al." authorityYear="1996" baseAuthorityName="Olivier" baseAuthorityYear="1812" box="[341,424,1500,1523]" class="Malacostraca" family="Diogenidae" genus="Aniculus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="37" phylum="Arthropoda" rank="species" species="ursus">
<emphasis id="B968EACFFB7AC35C4E7EFA1A5684D5C7" box="[341,424,1500,1523]" italics="true" pageId="4" pageNumber="37">A. ursus</emphasis>
</taxonomicName>
cannot be separated by this character. In contrast, the armature of the dactyli and propodi of the second and third pereopods does serve to distinguish the two species: the corneous spinules on the lateral surface (especially in the dorsal half) of the dactyli and dorsal surface of the propodi are much fewer in
<taxonomicName id="4C1C4D5EFB7AC35C4D15F9B855EBD6A2" authorityName="Forest" authorityYear="1984" box="[574,711,1662,1686]" class="Malacostraca" family="Diogenidae" genus="Aniculus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="37" phylum="Arthropoda" rank="species" species="erythraeus">
<emphasis id="B968EACFFB7AC35C4D15F9B855EBD6A2" box="[574,711,1662,1686]" italics="true" pageId="4" pageNumber="37">A. erythraeus</emphasis>
</taxonomicName>
than in
<taxonomicName id="4C1C4D5EFB7AC35C4F93F9595627D682" authorityName="Nomura et al." authorityYear="1996" baseAuthorityName="Olivier" baseAuthorityYear="1812" box="[184,267,1695,1718]" class="Malacostraca" family="Diogenidae" genus="Aniculus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="37" phylum="Arthropoda" rank="species" species="ursus">
<emphasis id="B968EACFFB7AC35C4F93F9595627D682" box="[184,267,1695,1718]" italics="true" pageId="4" pageNumber="37">A. ursus</emphasis>
</taxonomicName>
. Coloration of the two species is also clearly different. The shield is reddish purple and has irregular white streaks in
<taxonomicName id="4C1C4D5EFB7AC35C4E29F92656A4D6CC" authorityName="Forest" authorityYear="1984" box="[258,392,1760,1784]" class="Malacostraca" family="Diogenidae" genus="Aniculus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="37" phylum="Arthropoda" rank="species" species="erythraeus">
<emphasis id="B968EACFFB7AC35C4E29F92656A4D6CC" box="[258,392,1760,1784]" italics="true" pageId="4" pageNumber="37">A. erythraeus</emphasis>
</taxonomicName>
, whereas it is mostly green and bears reddish-brown patches anteriorly in
<taxonomicName id="4C1C4D5EFB7AC35C4D0FF8C75550D72C" authorityName="Nomura et al." authorityYear="1996" baseAuthorityName="Olivier" baseAuthorityYear="1812" box="[548,636,1793,1816]" class="Malacostraca" family="Diogenidae" genus="Aniculus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="37" phylum="Arthropoda" rank="species" species="ursus">
<emphasis id="B968EACFFB7AC35C4D0FF8C75550D72C" box="[548,636,1793,1816]" italics="true" pageId="4" pageNumber="37">A. ursus</emphasis>
</taxonomicName>
. The ocular peduncles of
<taxonomicName id="4C1C4D5EFB7AC35C4E0FF8E75680D70D" authorityName="Forest" authorityYear="1984" box="[292,428,1825,1849]" class="Malacostraca" family="Diogenidae" genus="Aniculus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="37" phylum="Arthropoda" rank="species" species="erythraeus">
<emphasis id="B968EACFFB7AC35C4E0FF8E75680D70D" box="[292,428,1825,1849]" italics="true" pageId="4" pageNumber="37">A. erythraeus</emphasis>
</taxonomicName>
are mostly reddish purple, while those of
<taxonomicName id="4C1C4D5EFB7AC35C4FDEF8845664D76D" authorityName="Nomura et al." authorityYear="1996" baseAuthorityName="Olivier" baseAuthorityYear="1812" box="[245,328,1858,1881]" class="Malacostraca" family="Diogenidae" genus="Aniculus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="37" phylum="Arthropoda" rank="species" species="ursus">
<emphasis id="B968EACFFB7AC35C4FDEF8845664D76D" box="[245,328,1858,1881]" italics="true" pageId="4" pageNumber="37">A. ursus</emphasis>
</taxonomicName>
are deep green. The chelae and the propodi of the ambulatory legs are generally reddish purple in
<taxonomicName id="4C1C4D5EFB7AC35C4DC3F8A45629D7AE" authorityName="Forest" authorityYear="1984" class="Malacostraca" family="Diogenidae" genus="Aniculus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="37" phylum="Arthropoda" rank="species" species="erythraeus">
<emphasis id="B968EACFFB7AC35C4DC3F8A45629D7AE" italics="true" pageId="4" pageNumber="37">A. erythraeus</emphasis>
</taxonomicName>
, not of olive-green as in
<taxonomicName id="4C1C4D5EFB7AC35C4ED4F845557DD7AE" authorityName="Nomura et al." authorityYear="1996" baseAuthorityName="Olivier" baseAuthorityYear="1812" box="[511,593,1923,1946]" class="Malacostraca" family="Diogenidae" genus="Aniculus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="37" phylum="Arthropoda" rank="species" species="ursus">
<emphasis id="B968EACFFB7AC35C4ED4F845557DD7AE" box="[511,593,1923,1946]" italics="true" pageId="4" pageNumber="37">A. ursus</emphasis>
</taxonomicName>
(for
<emphasis id="B968EACFFB7AC35C4DA9F84555F6D7AE" box="[642,730,1923,1946]" italics="true" pageId="4" pageNumber="37">
<taxonomicName id="4C1C4D5EFB7AC35C4DA9F84555FAD7AE" authorityName="Nomura et al." authorityYear="1996" baseAuthorityName="Olivier" baseAuthorityYear="1812" box="[642,726,1923,1946]" class="Malacostraca" family="Diogenidae" genus="Aniculus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="37" phylum="Arthropoda" rank="species" species="ursus">A. ursus</taxonomicName>
,
</emphasis>
see
<bibRefCitation id="EF8D4B2CFB7AC35C4FB7F86556AAD78F" author="McLaughlin, P. A. &amp; Rahayu, D. L. &amp; Komai, T. &amp; Chan, T. - Y." box="[156,390,1955,1979]" pageId="4" pageNumber="37" refId="ref3176" refString="McLaughlin, P. A., Rahayu, D. L., Komai, T. and Chan, T. - Y. 2007. A Catalog of the Hermit Crabs (Paguroidea) of Taiwan. National Taiwan Ocean University, Keelung, 367 pp." type="book" year="2007">
McLaughlin
<emphasis id="B968EACFFB7AC35C4E36F8625662D78F" box="[285,334,1955,1979]" italics="true" pageId="4" pageNumber="37">et al.</emphasis>
2007
</bibRefCitation>
;
<bibRefCitation id="EF8D4B2CFB7AC35C4EA4F86555BED78F" author="Poupin, J. &amp; Juncker, M." box="[399,658,1955,1979]" pageId="4" pageNumber="37" refId="ref3382" refString="Poupin, J. and Juncker, M. 2010. Guide des Crustaces Decapodes du Pacifique Sud / A Guide to Decapod Crustaceans of the South Pacific. CRISP and CPS, Noumea, 317 pp." type="book" year="2010">Poupin and Juncker 2010</bibRefCitation>
).
</paragraph>
<paragraph id="8BA336DDFB7AC35C4F9EF8055243D2F1" blockId="4.[156,768,198,2076]" lastBlockId="4.[820,1433,198,710]" pageId="4" pageNumber="37">
Besides
<taxonomicName id="4C1C4D5EFB7AC35C4E23F80256A2D7EF" authorityName="Forest" authorityYear="1984" box="[264,398,1987,2011]" class="Malacostraca" family="Diogenidae" genus="Aniculus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="37" phylum="Arthropoda" rank="species" species="erythraeus">
<emphasis id="B968EACFFB7AC35C4E23F80256A2D7EF" box="[264,398,1987,2011]" italics="true" pageId="4" pageNumber="37">A. erythraeus</emphasis>
</taxonomicName>
, four other species of
<taxonomicName id="4C1C4D5EFB7AC35C4D58F80555E6D7EF" authorityName="Dana" authorityYear="1852" box="[627,714,1987,2011]" class="Malacostraca" family="Diogenidae" genus="Aniculus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="37" phylum="Arthropoda" rank="genus">
<emphasis id="B968EACFFB7AC35C4D58F80555E6D7EF" box="[627,714,1987,2011]" italics="true" pageId="4" pageNumber="37">Aniculus</emphasis>
</taxonomicName>
have been recorded from Kume Island where the present specimen was found:
<taxonomicName id="4C1C4D5EFB7AC35C4E60F7C356EBD828" authorityName="Edmondson" authorityYear="1952" box="[331,455,2053,2076]" class="Malacostraca" family="Diogenidae" genus="Aniculus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="37" phylum="Arthropoda" rank="species" species="maximus">
<emphasis id="B968EACFFB7AC35C4E60F7C356EBD828" box="[331,455,2053,2076]" italics="true" pageId="4" pageNumber="37">A. maximus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C1C4D5EFB7AC35C4EFDF7C35519D828" authorityName="Lewinsohn" authorityYear="1982" box="[470,565,2053,2076]" class="Malacostraca" family="Diogenidae" genus="Aniculus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="37" phylum="Arthropoda" rank="species" species="retipes">
<emphasis id="B968EACFFB7AC35C4EFDF7C35519D828" box="[470,565,2053,2076]" italics="true" pageId="4" pageNumber="37">A. retipes</emphasis>
</taxonomicName>
,
<taxonomicName id="4C1C4D5EFB7AC35C4D68F7C35587D828" authorityName="Forest" authorityYear="1984" box="[579,683,2052,2076]" class="Malacostraca" family="Diogenidae" genus="Aniculus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="37" phylum="Arthropoda" rank="species" species="sibogae">
<emphasis id="B968EACFFB7AC35C4D68F7C35587D828" box="[579,683,2052,2076]" italics="true" pageId="4" pageNumber="37">A. sibogae</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C1C4D5EFB7AC35C4DC3F7C353BFD0EA" authority="(Rahayu and Osawa 2012)" baseAuthorityName="Rahayu and Osawa" baseAuthorityYear="2012" box="[744,1171,198,2076]" class="Malacostraca" family="Diogenidae" genus="Aniculus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="37" phylum="Arthropoda" rank="species" species="ursus">
<emphasis id="B968EACFFB7AC35C4DC3F7C35445D0EA" box="[744,873,199,2076]" italics="true" pageId="4" pageNumber="37">A. ursus</emphasis>
(
<bibRefCitation id="EF8D4B2CFB7AC35C4C57FF0053A7D0EA" author="Rahayu, D. L. &amp; Osawa, M." box="[892,1163,198,222]" pageId="4" pageNumber="37" pagination="176 - 190" refId="ref3420" refString="Rahayu, D. L. and Osawa, M. 2012. Hermit crabs (Crustacea: Decapoda: Diogenidae) from the KUMEJIMA 2009 Expedition, Japan. Zootaxa 3367: 176 - 190." type="journal article" year="2012">Rahayu and Osawa 2012</bibRefCitation>
)
</taxonomicName>
. There are no clear differences in habitat among these five species, all of which occur on coral and rocky reef slopes or in crevices on the reefs (
<bibRefCitation id="EF8D4B2CFB7AC35C4C5EFEEE538FD174" author="Kawamoto, T. &amp; Okuno, J." box="[885,1187,296,320]" pageId="4" pageNumber="37" refId="ref2981" refString="Kawamoto, T. and Okuno, J. 2003. Shrimps and Crabs of Kume Island, Okinawa. Hankyu Communications, Tokyo, 173 pp. [In Japanese]" type="book" year="2003">Kawamoto and Okuno 2003</bibRefCitation>
; present study).
<taxonomicName id="4C1C4D5EFB7AC35C4A7FFEEE548CD154" authorityName="Forest" authorityYear="1984" class="Malacostraca" family="Diogenidae" genus="Aniculus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="37" phylum="Arthropoda" rank="species" species="sibogae">
<emphasis id="B968EACFFB7AC35C4A7FFEEE548CD154" italics="true" pageId="4" pageNumber="37">Aniculus sibogae</emphasis>
</taxonomicName>
is a very large hermit crab with the shield length attaining
<quantity id="4CE49B38FB7AC35C4CBCFEAF54F7D1B4" box="[919,987,361,385]" metricMagnitude="-2" metricUnit="m" metricValue="5.2" pageId="4" pageNumber="37" unit="mm" value="52.0">52 mm</quantity>
(
<bibRefCitation id="EF8D4B2CFB7AC35C4CC0FEAF5344D1B5" author="Forest, J." box="[1003,1128,361,385]" pageId="4" pageNumber="37" pagination="1 - 91" refId="ref2911" refString="Forest, J. 1984. Revision du genre Aniculus. Crustaceana (Supplement 8): 1 - 91." type="journal article" year="1984">Forest 1984</bibRefCitation>
); it always uses a shell of the turbinid gastropod
<taxonomicName id="4C1C4D5EFB7AC35C4CD6FE4F5271D195" authority="Linnaeus, 1758" authorityName="Linnaeus" authorityYear="1758" box="[1021,1373,393,417]" class="Gastropoda" family="Turbinidae" genus="Turbo" higherTaxonomySource="CoL" kingdom="Animalia" order="Trochida" pageId="4" pageNumber="37" phylum="Mollusca" rank="species" species="marmoratus">
<emphasis id="B968EACFFB7AC35C4CD6FE4F5397D195" box="[1021,1211,393,417]" italics="true" pageId="4" pageNumber="37">Turbo marmoratus</emphasis>
Linnaeus, 1758
</taxonomicName>
as its domicile (
<bibRefCitation id="EF8D4B2CFB7AC35C4CB1FE6C534DD1F6" author="Nomura, K. &amp; Miyake, S. &amp; Kamezaki, N." box="[922,1121,426,450]" pageId="4" pageNumber="37" pagination="2 - 7" refId="ref3226" refString="Nomura, K., Miyake, S. and Kamezaki, N. 1996. Revision of the genus Aniculus (Anomura, Diogenidae) in Japan. I. O. P. Diving News 8: 2 - 7. [In Japanese, with English abstract]" type="journal article" year="1996">
Nomura
<emphasis id="B968EACFFB7AC35C4CDCFE6D5305D1F6" box="[1015,1065,426,450]" italics="true" pageId="4" pageNumber="37">et al.</emphasis>
1996
</bibRefCitation>
; Kawamoto and Okuno 2006). This gastropod is widely distributed in the Indo-West Pacific, but there are no records of
<taxonomicName id="4C1C4D5EFB7AC35C4BADFE2D53C0D237" authorityName="Forest" authorityYear="1984" box="[1158,1260,491,515]" class="Malacostraca" family="Diogenidae" genus="Aniculus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="37" phylum="Arthropoda" rank="species" species="sibogae">
<emphasis id="B968EACFFB7AC35C4BADFE2D53C0D237" box="[1158,1260,491,515]" italics="true" pageId="4" pageNumber="37">A. sibogae</emphasis>
</taxonomicName>
from the Indian Ocean despite recent faunal surveys there (
<emphasis id="B968EACFFB7AC35C4BCDFDCA5224D217" box="[1254,1288,524,547]" italics="true" pageId="4" pageNumber="37">e.g.</emphasis>
,
<bibRefCitation id="EF8D4B2CFB7AC35C4A38FDCD52B8D217" author="Poupin, J." box="[1299,1428,523,547]" pageId="4" pageNumber="37" refId="ref3277" refString="Poupin, J. 2009. Crustaces de la Reunion, Decapodes et Stomatopodes. IRD Editions, Marseille, 140 pp." type="book" year="2009">Poupin 2009</bibRefCitation>
; Poupin
<emphasis id="B968EACFFB7AC35C4CAFFDEB549AD270" box="[900,950,556,580]" italics="true" pageId="4" pageNumber="37">et al.</emphasis>
in press).
<taxonomicName id="4C1C4D5EFB7AC35C4B0AFDEA53EBD270" authorityName="Forest" authorityYear="1984" box="[1057,1223,556,580]" class="Malacostraca" family="Diogenidae" genus="Aniculus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="37" phylum="Arthropoda" rank="species" species="sibogae">
<emphasis id="B968EACFFB7AC35C4B0AFDEA53EBD270" box="[1057,1223,556,580]" italics="true" pageId="4" pageNumber="37">Aniculus sibogae</emphasis>
</taxonomicName>
is known only from the western Pacific, ranging from the Ryukyu Islands to
<collectingCountry id="F30B764DFB7AC35C4C1FFDAB54BCD2B1" box="[820,912,621,645]" name="Malaysia" pageId="4" pageNumber="37">Malaysia</collectingCountry>
, New
<collectingCountry id="F30B764DFB7AC35C4CE4FDAB5337D2B1" box="[975,1051,621,645]" name="Guinea-Bissau" pageId="4" pageNumber="37">Guinea</collectingCountry>
, and
<collectingCountry id="F30B764DFB7AC35C4B79FDAB5385D2B0" box="[1106,1193,621,644]" name="Vanuatu" pageId="4" pageNumber="37">Vanuatu</collectingCountry>
, whereas
<taxonomicName id="4C1C4D5EFB7AC35C4A27FDAB52BED2B1" authorityName="Forest" authorityYear="1984" box="[1292,1426,621,645]" class="Malacostraca" family="Diogenidae" genus="Aniculus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="37" phylum="Arthropoda" rank="species" species="erythraeus">
<emphasis id="B968EACFFB7AC35C4A27FDAB52BED2B1" box="[1292,1426,621,645]" italics="true" pageId="4" pageNumber="37">A. erythraeus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C1C4D5EFB7AC35C4C1FFD48549CD291" authorityName="Edmondson" authorityYear="1952" box="[820,944,654,677]" class="Malacostraca" family="Diogenidae" genus="Aniculus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="37" phylum="Arthropoda" rank="species" species="maximus">
<emphasis id="B968EACFFB7AC35C4C1FFD48549CD291" box="[820,944,654,677]" italics="true" pageId="4" pageNumber="37">A. maximus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C1C4D5EFB7AC35C4C95FD485332D291" authorityName="Lewinsohn" authorityYear="1982" box="[958,1054,654,677]" class="Malacostraca" family="Diogenidae" genus="Aniculus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="37" phylum="Arthropoda" rank="species" species="retipes">
<emphasis id="B968EACFFB7AC35C4C95FD485332D291" box="[958,1054,654,677]" italics="true" pageId="4" pageNumber="37">A. retipes</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C1C4D5EFB7AC35C4B71FD485383D291" baseAuthorityName="Rahayu and Osawa" baseAuthorityYear="2012" box="[1114,1199,654,677]" class="Malacostraca" family="Diogenidae" genus="Aniculus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="37" phylum="Arthropoda" rank="species" species="ursus">
<emphasis id="B968EACFFB7AC35C4B71FD485383D291" box="[1114,1199,654,677]" italics="true" pageId="4" pageNumber="37">A. ursus</emphasis>
</taxonomicName>
are widely distributed in the Indo-West Pacific but rather sparse in occurrence.
</paragraph>
</subSubSection>
</treatment>
</document>