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<document id="A0A8649FB7628D7CD9FE83D817C292B3" ID-DOI="10.11646/zootaxa.3974.3.8" ID-GBIF-Dataset="6f675840-f786-4135-b260-3df16a614071" ID-ISSN="1175-5326" ID-Zenodo-Dep="239449" ID-ZooBank="D93EF759-9314-4FA1-9804-FDD37B934BCA" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461220174377" checkinUser="plazi" docAuthor="Makarchenko, Eugenyi A. &amp; Makarchenko, Marina A." docDate="2015" docId="03B587C3FFB8FFC2FF2BFAB2D461C01B" docLanguage="en" docName="zt03974p423.pdf" docOrigin="Zootaxa 3974 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Parorthocladius plolabius Makarchenko et Makarchenko, sp. nov." docType="treatment" docVersion="9" lastPageNumber="418" masterDocId="FF8CFFBBFFBAFFC7FFBCFFAED53AC124" masterDocTitle="A review of the genus Parorthocladius Thienemann, 1935 (Diptera: Chironomidae: Orthocladiinae) from the Russian Far East" masterLastPageNumber="423" masterPageNumber="413" pageNumber="415" updateTime="1698603995156" updateUser="plazi">
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<mods:title id="A76E30941B55818C3C1FA16E51D1E608">A review of the genus Parorthocladius Thienemann, 1935 (Diptera: Chironomidae: Orthocladiinae) from the Russian Far East</mods:title>
</mods:titleInfo>
<mods:name id="1DE8399D3D34276DC765ACD3B51D52E0" type="personal">
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<mods:namePart id="89A7EF8F425749C519EA3BB45B09C3E0">Makarchenko, Eugenyi A.</mods:namePart>
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<mods:namePart id="B06BD1560F996FE4CDC01388FE3A76AF">Makarchenko, Marina A.</mods:namePart>
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<mods:title id="50A3DFEFBEF3887ED2597BEE26B8C2B4">Zootaxa</mods:title>
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<mods:part id="422A7490741E8497A843C2843975F675">
<mods:date id="9C8DFD2F45F75EEC340F6331DB277A49">2015</mods:date>
<mods:detail id="7902951F1C2EF8217F16415706B8131D" type="volume">
<mods:number id="5152753F1F022C821F77732EE73F98AF">3974</mods:number>
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<mods:number id="F3CF32199884B48712658B2D2004844A">3</mods:number>
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<mods:classification id="3EB81ABCCDE707FE27CEF6C932D4C10E">journal article</mods:classification>
<mods:identifier id="0E043F32D480A3068431DA05ED1D315E" type="DOI">10.11646/zootaxa.3974.3.8</mods:identifier>
<mods:identifier id="D6A0ED480BFD395C73367A8E8C8113E7" type="GBIF-Dataset">6f675840-f786-4135-b260-3df16a614071</mods:identifier>
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</mods:mods>
<treatment id="03B587C3FFB8FFC2FF2BFAB2D461C01B" ID-DOI="http://doi.org/10.5281/zenodo.5624812" ID-GBIF-Taxon="119631567" ID-Zenodo-Dep="5624812" LSID="urn:lsid:plazi:treatment:03B587C3FFB8FFC2FF2BFAB2D461C01B" httpUri="http://treatment.plazi.org/id/03B587C3FFB8FFC2FF2BFAB2D461C01B" lastPageId="5" lastPageNumber="418" pageId="2" pageNumber="415">
<subSubSection id="C306655EFFB8FFC5FF2BFAB2D41AC47C" pageId="2" pageNumber="415" type="nomenclature">
<paragraph id="8BA336D5FFB8FFC5FF2BFAB2D6FCC412" blockId="2.[151,966,1308,1368]" box="[151,966,1308,1334]" pageId="2" pageNumber="415">
<heading id="D0EB81B9FFB8FFC5FF2BFAB2D6FCC412" bold="true" box="[151,966,1308,1334]" fontSize="11" level="1" pageId="2" pageNumber="415" reason="1">
<emphasis id="B968EAC7FFB8FFC5FF2BFAB2D6FCC412" bold="true" box="[151,966,1308,1334]" pageId="2" pageNumber="415">
<taxonomicName id="4C1C4D56FFB8FFC5FF2BFAB2D666C412" ID-CoL="9HTN3" authority="Makarchenko et Makarchenko" authorityName="Makarchenko et Makarchenko" box="[151,860,1308,1334]" class="Insecta" family="Chironomidae" genus="Parorthocladius" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="415" phylum="Arthropoda" rank="species" species="plolabius" status="sp. nov.">
<emphasis id="B968EAC7FFB8FFC5FF2BFAB2D4EBC412" bold="true" box="[151,465,1308,1334]" italics="true" pageId="2" pageNumber="415">Parorthocladius plolabius</emphasis>
Makarchenko
<emphasis id="B968EAC7FFB8FFC5FD32FAB2D799C412" bold="true" box="[654,675,1308,1334]" italics="true" pageId="2" pageNumber="415">et</emphasis>
Makarchenko
</taxonomicName>
,
<taxonomicNameLabel id="A25B57BCFFB8FFC5FCD4FAB2D6FCC412" box="[872,966,1308,1334]" pageId="2" pageNumber="415" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BA336D5FFB8FFC5FF2BFA91D41AC47C" blockId="2.[151,966,1308,1368]" box="[151,288,1343,1368]" pageId="2" pageNumber="415">
(
<figureCitation id="13272A50FFB8FFC5FF23FA91D422C47C" box="[159,280,1343,1368]" captionStart-0="FIGURES 1 7" captionStart-1="FIGURES 13 17" captionStartId-0="3.[151,264,1688,1711]" captionStartId-1="6.[151,264,1709,1732]" captionTargetBox-0="[174,1417,216,1640]" captionTargetBox-1="[195,1382,183,1687]" captionTargetId-0="figure@3.[174,1426,208,1640]" captionTargetId-1="figure@6.[188,1398,177,1689]" captionTargetPageId-0="3" captionTargetPageId-1="6" captionText-0="FIGURES 1 7. Parorthocladius lazovskiensis sp. nov., male (1 3) and P. plolabius sp. nov., male (4 7). 1, 6, hypopygium in dorsal view; 2, gonocoxite and gonostylus in dorsal view; 3 5, anal point; 7, part of hypopygium without tergite IX in dorsal view. 1, male from Sakhalin Island; 2 3, male from Primorye Territory; 4 7, males from Amur River basin. Scale bars 50 µm." captionText-1="FIGURES 13 17. Parorthocladius plolabius sp. nov., pupa (13) and larva of fourth instar (14 17). 13, tergites I VIII and anal segment of male; 14 15, mentum; 16, mandible; 17, antenna. Scale bars: Fig. 13 — 200 µm; Figs. 14 17 — 20 µm." httpUri-0="https://zenodo.org/record/239450/files/figure.png" httpUri-1="https://zenodo.org/record/239451/files/figure.png" pageId="2" pageNumber="415">Figs. 417</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C306655EFFB8FFC2FF2BFA29D751C194" lastPageId="5" lastPageNumber="418" pageId="2" pageNumber="415" type="materials_examined">
<paragraph id="8BA336D5FFB8FFC5FF2BFA29D15CC7E4" blockId="2.[151,1437,1415,2019]" pageId="2" pageNumber="415">
<emphasis id="B968EAC7FFB8FFC5FF2BFA29D43CC484" bold="true" box="[151,262,1415,1440]" pageId="2" pageNumber="415">Material.</emphasis>
<typeStatus id="54A78877FFB8FFC5FEB0FA29D443C484" box="[268,377,1415,1440]" pageId="2" pageNumber="415" type="holotype">Holotype</typeStatus>
: adult male, Russian Far East, Jewish Autonomous Region, Obluchie District, Fedotkin Spring of Bidzhan River basin (Amur River basin),
<geoCoordinate id="EE285012FFB8FFC5FD1BFA02D637C4E0" box="[679,781,1452,1476]" direction="north" orientation="latitude" pageId="2" pageNumber="415" precision="925" value="48.633335">N 48º38'</geoCoordinate>
409'',
<geoCoordinate id="EE285012FFB8FFC5FCEEFA02D6F9C4E0" box="[850,963,1452,1476]" direction="east" orientation="longitude" pageId="2" pageNumber="415" precision="925" value="131.61667">E 131º37'</geoCoordinate>
217'',
<date id="FFA21015FFB8FFC5FBB4FA02D1AEC4E0" box="[1032,1172,1452,1476]" pageId="2" pageNumber="415" value="2013-03-27">27.III. 2013</date>
, leg. E. Makarchenko.
<typeStatus id="54A78877FFB8FFC5FF2BFA7ED433C4CC" box="[151,265,1488,1512]" pageId="2" pageNumber="415" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="9D1AFD5CFFB8FFC5FEAFFA7ED457C4CC" box="[275,365,1487,1512]" pageId="2" pageNumber="415" type="male">3 males</specimenCount>
, the same data as
<typeStatus id="54A78877FFB8FFC5FD86FA61D7A6C4CC" box="[570,668,1487,1512]" pageId="2" pageNumber="415" type="holotype">holotype</typeStatus>
except
<date id="FFA21015FFB8FFC5FD48FA7ED69FC4CC" box="[756,933,1488,1512]" pageId="2" pageNumber="415" value="2012-04-10" valueMax="2012-04-12" valueMin="2012-04-10">1012.IV. 2012</date>
, leg. E. Makarchenko;
<specimenCount id="9D1AFD5CFFB8FFC5FB17FA7ED03FC4CC" box="[1195,1285,1487,1512]" pageId="2" pageNumber="415" type="male">2 males</specimenCount>
,
<specimenCount id="9D1AFD5CFFB8FFC5FAADFA7ED054C4CC" box="[1297,1390,1487,1512]" pageId="2" pageNumber="415" type="larva">8 larvae</specimenCount>
, the same data as
<typeStatus id="54A78877FFB8FFC5FE85FA5DD4A1C728" box="[313,411,1523,1548]" pageId="2" pageNumber="415" type="holotype">holotype</typeStatus>
except
<date id="FFA21015FFB8FFC5FE47FA5DD7A9C728" box="[507,659,1523,1548]" pageId="2" pageNumber="415" value="2014-04-05" valueMax="2014-04-06" valueMin="2014-04-05">56.IV. 2014</date>
, leg. E. Makarchenko;
<specimenCount id="9D1AFD5CFFB8FFC5FC15FA5AD13CC728" box="[937,1030,1523,1548]" pageId="2" pageNumber="415" type="male">3 males</specimenCount>
, the same data as
<typeStatus id="54A78877FFB8FFC5FB5AFA5DD072C728" box="[1254,1352,1523,1548]" pageId="2" pageNumber="415" type="holotype">holotype</typeStatus>
except Lopatinskyi Spring,
<geoCoordinate id="EE285012FFB8FFC5FEC7F9B6D4E7C714" box="[379,477,1560,1584]" direction="north" orientation="latitude" pageId="2" pageNumber="415" precision="925" value="48.616665">N 48º37'</geoCoordinate>
810'',
<geoCoordinate id="EE285012FFB8FFC5FDA1F9B6D7B3C714" box="[541,649,1560,1584]" direction="east" orientation="longitude" pageId="2" pageNumber="415" precision="925" value="131.65">E 131º39'</geoCoordinate>
114'',
<date id="FFA21015FFB8FFC5FD74F9B6D605C714" box="[712,831,1560,1584]" pageId="2" pageNumber="415" value="2014-04-07">7.IV. 2014</date>
, leg. E. Makarchenko;
<specimenCount id="9D1AFD5CFFB8FFC5FBFCF9B6D1A2C714" box="[1088,1176,1559,1584]" pageId="2" pageNumber="415" type="male">2 males</specimenCount>
, Kamchatka Peninsula, Ozernaya River,
<date id="FFA21015FFB8FFC5FEEAF992D4E9C770" box="[342,467,1596,1620]" pageId="2" pageNumber="415" value="1998-06-09">9.VI. 1998</date>
, leg. T. Travina; 1 mature pupa,
<specimenCount id="9D1AFD5CFFB8FFC5FCF9F992D69CC770" box="[837,934,1596,1620]" pageId="2" pageNumber="415" type="pupa">7 pupae</specimenCount>
,1 pupal exuviae,
<specimenCount id="9D1AFD5CFFB8FFC5FBD4F992D1FFC770" box="[1128,1221,1595,1620]" pageId="2" pageNumber="415" type="larva">3 larvae</specimenCount>
, 1 larval skin from pupa, the same region, Plotnikova River,
<date id="FFA21015FFB8FFC5FD34F9F1D604C75C" box="[648,830,1631,1656]" pageId="2" pageNumber="415" value="2004-04-24" valueMax="2004-04-26" valueMin="2004-04-24">2426.IV. 2004</date>
, leg. T. Travina; 1 larva, Magadan Region, Olsky District, Uglikan River (Ola River Basin),
<date id="FFA21015FFB8FFC5FDCCF92AD7DDC7B8" box="[624,743,1668,1692]" pageId="2" pageNumber="415" value="2013-04-07">7.IV. 2013</date>
, E. Khamenkova; 3 mature pupae, the same region, Ola River,
<date id="FFA21015FFB8FFC5FF2BF906D407C7E4" box="[151,317,1704,1728]" pageId="2" pageNumber="415" value="2013-05-12" valueMax="2013-05-13" valueMin="2013-05-12">1213.V. 2013</date>
, E. Khamenkova; 1 larva, the same place,
<date id="FFA21015FFB8FFC5FCA5F906D6A4C7E4" box="[793,926,1704,1728]" pageId="2" pageNumber="415" value="2013-07-08">8.VII. 2013</date>
, E. Khamenkova.
</paragraph>
<paragraph id="8BA336D5FFB8FFC5FF7BF965D4B1C7C0" blockId="2.[151,1437,1415,2019]" box="[199,395,1739,1764]" pageId="2" pageNumber="415">
<emphasis id="B968EAC7FFB8FFC5FF7BF965D47CC7C0" bold="true" box="[199,326,1739,1764]" italics="true" pageId="2" pageNumber="415">Adult male</emphasis>
(n=4)
</paragraph>
<paragraph id="8BA336D5FFB8FFC5FF7BF941D186C62C" blockId="2.[151,1437,1415,2019]" box="[199,1212,1775,1800]" pageId="2" pageNumber="415">
Total length
<quantity id="4CE49B30FFB8FFC5FEE8F941D4C0C623" box="[340,506,1775,1800]" metricMagnitude="-3" metricUnit="m" metricValue="3.45" metricValueMax="4.15" metricValueMin="2.75" pageId="2" pageNumber="415" unit="mm" value="3.45" valueMax="4.15" valueMin="2.75">2.754.15 mm</quantity>
. Wing length
<quantity id="4CE49B30FFB8FFC5FD2AF95ED606C623" box="[662,828,1776,1800]" metricMagnitude="-3" metricUnit="m" metricValue="2.53" metricValueMax="2.74" metricValueMin="2.32" pageId="2" pageNumber="415" unit="mm" value="2.53" valueMax="2.74" valueMin="2.32">2.322.74 mm</quantity>
. Total length/wing length 1.21.5.
</paragraph>
<paragraph id="8BA336D5FFB8FFC5FF7BF8BDD741C650" blockId="2.[151,1437,1415,2019]" pageId="2" pageNumber="415">Head. Temporal setae 913, including 59 verticals and 4 postorbitals. Clypeus with 810 setae. Antenna with 13 flagellomeres and well developed plume; AR 1.461.85. Palp with 5 palpomeres, length of 4 distal palpomeres (in µm): 4864, 88112, 56112, 172200.</paragraph>
<paragraph id="8BA336D5FFB8FFC5FF7BF8D1D025C6B3" blockId="2.[151,1437,1415,2019]" box="[199,1311,1919,1944]" pageId="2" pageNumber="415">Thorax. Dark brown. Antepronotum without lateral setae. Dc 67, Pa 34, scutellum without setae.</paragraph>
<paragraph id="8BA336D5FFB8FFC5FF7BF80DD6EFC6C6" blockId="2.[151,1437,1415,2019]" pageId="2" pageNumber="415">Wing. R and R1 with 78 setae, R4+5 with 01 setae subapically. R4+5 ending distal of apex M3+4. Costa extension 1280 µm. Anal lobe well developed, rounded. Squama with 1419 setae.</paragraph>
<caption id="DF63665DFFB9FFC4FF2BF936D0ADC7C8" httpUri="https://zenodo.org/record/239450/files/figure.png" pageId="3" pageNumber="416" targetBox="[174,1417,216,1640]" targetPageId="3">
<paragraph id="8BA336D5FFB9FFC4FF2BF936D0ADC7C8" blockId="3.[151,1436,1688,1772]" pageId="3" pageNumber="416">
<emphasis id="B968EAC7FFB9FFC4FF2BF936D400C789" bold="true" box="[151,314,1688,1711]" pageId="3" pageNumber="416">FIGURES 17.</emphasis>
<taxonomicName id="4C1C4D56FFB9FFC4FEFDF936D74BC78A" box="[321,625,1688,1710]" class="Insecta" family="Chironomidae" genus="Parorthocladius" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="416" phylum="Arthropoda" rank="species" species="lazovskiensis" status="sp. nov.">
<emphasis id="B968EAC7FFB9FFC4FEFDF936D74BC78A" box="[321,625,1688,1710]" italics="true" pageId="3" pageNumber="416">Parorthocladius lazovskiensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25B57BCFFB9FFC4FDCBF937D7F2C78A" box="[631,712,1689,1710]" pageId="3" pageNumber="416" rank="species">
<emphasis id="B968EAC7FFB9FFC4FDCBF937D7F2C78A" bold="true" box="[631,712,1689,1710]" pageId="3" pageNumber="416">sp. nov.</emphasis>
</taxonomicNameLabel>
, male (13) and
<taxonomicName id="4C1C4D56FFB9FFC4FCCAF934D6D7C78A" box="[886,1005,1688,1711]" class="Insecta" family="Chironomidae" genus="Parorthocladius" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="416" phylum="Arthropoda" rank="species" species="plolabius" status="sp. nov.">
<emphasis id="B968EAC7FFB9FFC4FCCAF934D6D7C78A" box="[886,1005,1688,1711]" italics="true" pageId="3" pageNumber="416">P. plolabius</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25B57BCFFB9FFC4FC48F937D17FC78A" box="[1012,1093,1689,1710]" pageId="3" pageNumber="416" rank="species">
<emphasis id="B968EAC7FFB9FFC4FC48F937D17FC78A" bold="true" box="[1012,1093,1689,1710]" pageId="3" pageNumber="416">sp. nov.</emphasis>
</taxonomicNameLabel>
, male (47).
<emphasis id="B968EAC7FFB9FFC4FB72F936D1C0C789" bold="true" box="[1230,1274,1688,1709]" pageId="3" pageNumber="416">1, 6,</emphasis>
hypopygium in dorsal view;
<emphasis id="B968EAC7FFB9FFC4FEA0F919D412C7E8" bold="true" box="[284,296,1719,1740]" pageId="3" pageNumber="416">2</emphasis>
, gonocoxite and gonostylus in dorsal view;
<emphasis id="B968EAC7FFB9FFC4FD53F919D626C7E8" bold="true" box="[751,796,1719,1740]" pageId="3" pageNumber="416">35,</emphasis>
anal point;
<emphasis id="B968EAC7FFB9FFC4FC2BF919D690C7E8" bold="true" box="[919,938,1719,1740]" pageId="3" pageNumber="416">7,</emphasis>
part of hypopygium without tergite IX in dorsal view.
<emphasis id="B968EAC7FFB9FFC4FF6EF978D5DFC7CF" bold="true" box="[210,229,1750,1771]" pageId="3" pageNumber="416">1,</emphasis>
male from Sakhalin Island;
<emphasis id="B968EAC7FFB9FFC4FDBEF978D714C7CF" bold="true" box="[514,558,1750,1771]" pageId="3" pageNumber="416">23,</emphasis>
male from Primorye Territory;
<emphasis id="B968EAC7FFB9FFC4FCD0F978D6A2C7CF" bold="true" box="[876,920,1750,1771]" pageId="3" pageNumber="416">47,</emphasis>
males from Amur River basin. Scale bars 50 µm.
</paragraph>
</caption>
<paragraph id="8BA336D5FFB9FFC4FF7BF8BAD110C6B9" blockId="3.[151,1436,1811,1950]" pageId="3" pageNumber="416">
Legs. Spur of fore tibia 5260 µm long. Spurs of mid tibia 2428 µm and 20 µm long. Spurs of hind tibia 60 µm and 2024 µm long.
<collectingCountry id="F30B7645FFB9FFC4FE17F899D4D9C674" box="[427,483,1847,1872]" name="India" pageId="3" pageNumber="416">Hind</collectingCountry>
tibial comb with 1113 setae. Fore leg without pseudospurs on ta1; mid and hind legs with 2 apical pseudospurs on ta1 and ta2; sometimes hind legs with 1 apical spur on ta2. Mid leg on ta
<quantity id="4CE49B30FFB9FFC4FAB8F8C2D013C653" box="[1284,1321,1886,1914]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="3" pageNumber="416" unit="in" value="1.0">1 in</quantity>
basal part with 08 sensilla chaetica. Lengths and proportions of leg segments as in
<tableCitation id="C69E036EFFB9FFC4FC6CF82AD11CC6B9" box="[976,1062,1924,1949]" captionStart="TABLE 2" captionStartId="4.[151,239,152,175]" captionTargetBox="[151,1397,202,414]" captionTargetPageId="4" captionText="TABLE 2. Lengths (in µm) and proportions of leg segments of Parorthocladius plolabius sp. nov., male (n = 4)." httpUri="http://table.plazi.org/id/DF63665DFFBEFFC3FF2BFF36D073C18B" pageId="3" pageNumber="416" tableUuid="DF63665DFFBEFFC3FF2BFF36D073C18B">Table 2</tableCitation>
.
</paragraph>
<caption id="DF63665DFFBEFFC3FF2BFF36D073C18B" ID-Table-UUID="DF63665DFFBEFFC3FF2BFF36D073C18B" box="[151,1353,151,175]" httpUri="http://table.plazi.org/id/DF63665DFFBEFFC3FF2BFF36D073C18B" pageId="4" pageNumber="417" targetBox="[151,1397,202,414]" targetIsTable="true" targetPageId="4">
<paragraph id="8BA336D5FFBEFFC3FF2BFF36D073C18B" blockId="4.[151,1353,151,175]" box="[151,1353,151,175]" pageId="4" pageNumber="417">
<emphasis id="B968EAC7FFBEFFC3FF2BFF36D433C18B" bold="true" box="[151,265,152,175]" pageId="4" pageNumber="417">TABLE 2.</emphasis>
Lengths (in µm) and proportions of leg segments of
<taxonomicName id="4C1C4D56FFBEFFC3FC82FF39D16CC18B" box="[830,1110,151,175]" class="Insecta" family="Chironomidae" genus="Parorthocladius" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="417" phylum="Arthropoda" rank="species" species="plolabius" status="sp. nov.">
<emphasis id="B968EAC7FFBEFFC3FC82FF39D16CC18B" box="[830,1110,151,175]" italics="true" pageId="4" pageNumber="417">Parorthocladius plolabius</emphasis>
</taxonomicName>
<emphasis id="B968EAC7FFBEFFC3FBE1FF36D188C18B" bold="true" box="[1117,1202,152,175]" pageId="4" pageNumber="417">
<taxonomicNameLabel id="A25B57BCFFBEFFC3FBE1FF36D188C18B" box="[1117,1202,152,175]" pageId="4" pageNumber="417" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male (n = 4).
</paragraph>
</caption>
<paragraph id="8BA336D5FFBEFFC3FF54FF64D432C0BA" pageId="4" pageNumber="417">
<table id="F91CC475FFBE0038FF2BFF64D04FC0BA" box="[151,1397,202,414]" gridcols="6" gridrows="5" pageId="4" pageNumber="417">
<tr id="352C3497FFBE0038FF2BFF64D04FC1C0" box="[151,1397,202,228]" gridrow="0" pageId="4" pageNumber="417">
<th id="76FD5DEBFFBE0038FF2BFF64D4C8C1C0" box="[151,498,202,228]" gridcol="0" gridrow="0" pageId="4" pageNumber="417">fe ti</th>
<th id="76FD5DEBFFBE0038FDE7FF64D78EC1C0" box="[603,692,202,228]" gridcol="1" gridrow="0" pageId="4" pageNumber="417">ta1</th>
<th id="76FD5DEBFFBE0038FCB0FF64D65FC1C0" box="[780,869,202,228]" gridcol="2" gridrow="0" pageId="4" pageNumber="417">ta2</th>
<th id="76FD5DEBFFBE0038FC01FF64D12CC1C0" box="[957,1046,202,228]" gridcol="3" gridrow="0" pageId="4" pageNumber="417">ta3</th>
<th id="76FD5DEBFFBE0038FBD2FF64D1FDC1C0" box="[1134,1223,202,228]" gridcol="4" gridrow="0" pageId="4" pageNumber="417">ta4</th>
<th id="76FD5DEBFFBE0038FAA0FF64D04FC1C0" box="[1308,1397,202,228]" gridcol="5" gridrow="0" pageId="4" pageNumber="417">ta5</th>
</tr>
<tr id="352C3497FFBE0038FF2BFF58D04FC034" box="[151,1397,246,272]" gridrow="1" pageId="4" pageNumber="417">
<th id="76FD5DEBFFBE0038FF2BFF58D4C8C034" box="[151,498,246,272]" gridcol="0" gridrow="1" pageId="4" pageNumber="417">P1 7521072 8321136</th>
<td id="76FD5DEBFFBE0038FDE7FF58D78EC034" box="[603,692,246,272]" gridcol="1" gridrow="1" pageId="4" pageNumber="417">624800</td>
<td id="76FD5DEBFFBE0038FCB0FF58D65FC034" box="[780,869,246,272]" gridcol="2" gridrow="1" pageId="4" pageNumber="417">416576</td>
<td id="76FD5DEBFFBE0038FC01FF58D12CC034" box="[957,1046,246,272]" gridcol="3" gridrow="1" pageId="4" pageNumber="417">320400</td>
<td id="76FD5DEBFFBE0038FBD2FF58D1FDC034" box="[1134,1223,246,272]" gridcol="4" gridrow="1" pageId="4" pageNumber="417">224288</td>
<td id="76FD5DEBFFBE0038FAA0FF58D04FC034" box="[1308,1397,246,272]" gridcol="5" gridrow="1" pageId="4" pageNumber="417">112136</td>
</tr>
<tr id="352C3497FFBE0038FF2BFE8CD04FC018" box="[151,1397,290,316]" gridrow="2" pageId="4" pageNumber="417">
<th id="76FD5DEBFFBE0038FF2BFE8CD4C8C018" box="[151,498,290,316]" gridcol="0" gridrow="2" pageId="4" pageNumber="417">P2 752992 7841008</th>
<td id="76FD5DEBFFBE0038FDE7FE8CD78EC018" box="[603,692,290,316]" gridcol="1" gridrow="2" pageId="4" pageNumber="417">432576</td>
<td id="76FD5DEBFFBE0038FCB0FE8CD65FC018" box="[780,869,290,316]" gridcol="2" gridrow="2" pageId="4" pageNumber="417">304368</td>
<td id="76FD5DEBFFBE0038FC01FE8CD12CC018" box="[957,1046,290,316]" gridcol="3" gridrow="2" pageId="4" pageNumber="417">224288</td>
<td id="76FD5DEBFFBE0038FBD2FE8CD1FDC018" box="[1134,1223,290,316]" gridcol="4" gridrow="2" pageId="4" pageNumber="417">176224</td>
<td id="76FD5DEBFFBE0038FAA0FE8CD04FC018" box="[1308,1397,290,316]" gridcol="5" gridrow="2" pageId="4" pageNumber="417">128144</td>
</tr>
<tr id="352C3497FFBE0038FF2BFEE0D04FC043" box="[151,1397,334,359]" gridrow="3" pageId="4" pageNumber="417">
<th id="76FD5DEBFFBE0038FF2BFEE0D4C8C043" box="[151,498,334,359]" gridcol="0" gridrow="3" pageId="4" pageNumber="417">P3 8161104 9441216</th>
<td id="76FD5DEBFFBE0038FDE7FEE0D78EC043" box="[603,692,334,359]" gridcol="1" gridrow="3" pageId="4" pageNumber="417">544688</td>
<td id="76FD5DEBFFBE0038FCB0FEE0D65FC043" box="[780,869,334,359]" gridcol="2" gridrow="3" pageId="4" pageNumber="417">320432</td>
<td id="76FD5DEBFFBE0038FC01FEE0D12CC043" box="[957,1046,334,359]" gridcol="3" gridrow="3" pageId="4" pageNumber="417">256288</td>
<td id="76FD5DEBFFBE0038FBD2FEE0D1FDC043" box="[1134,1223,334,359]" gridcol="4" gridrow="3" pageId="4" pageNumber="417">176240</td>
<td id="76FD5DEBFFBE0038FAA0FEE0D04FC043" box="[1308,1397,334,359]" gridcol="5" gridrow="3" pageId="4" pageNumber="417">128144</td>
</tr>
<tr id="352C3497FFBE0038FF2BFE29D04FC0BA" box="[151,1397,391,414]" gridrow="4" pageId="4" pageNumber="417" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1">
<th id="76FD5DEBFFBE0038FF2BFE29D4C8C0BA" box="[151,498,391,414]" gridcol="0" gridrow="4" pageId="4" pageNumber="417">continued.</th>
</tr>
</table>
</paragraph>
<paragraph id="8BA336D5FFBEFFC3FEA7FE15D1C7C375" pageId="4" pageNumber="417">
<table id="F91CC475FFBE0038FF23FE14D1C4C371" box="[159,1278,442,597]" gridcols="4" gridrows="4" pageId="4" pageNumber="417">
<tr id="352C3497FFBE0038FF23FE14D1C4C0F4" box="[159,1278,442,464]" gridrow="0" pageId="4" pageNumber="417">
<th id="76FD5DEBFFBE0038FF23FE14D4BAC0F4" box="[159,384,442,464]" gridcol="0" gridrow="0" pageId="4" pageNumber="417">LR</th>
<th id="76FD5DEBFFBE0038FDE6FE14D785C0F4" box="[602,703,442,464]" gridcol="1" gridrow="0" pageId="4" pageNumber="417">BV</th>
<th id="76FD5DEBFFBE0038FC39FE14D6D1C0F4" box="[901,1003,442,464]" gridcol="2" gridrow="0" pageId="4" pageNumber="417">SV</th>
<th id="76FD5DEBFFBE0038FB0DFE14D1C4C0F4" box="[1201,1278,442,464]" gridcol="3" gridrow="0" pageId="4" pageNumber="417">BR</th>
</tr>
<tr id="352C3497FFBE0038FF23FE4DD1C4C0D9" box="[159,1278,483,509]" gridrow="1" pageId="4" pageNumber="417">
<th id="76FD5DEBFFBE0038FF23FE4DD4BAC0D9" box="[159,384,483,509]" gridcol="0" gridrow="1" pageId="4" pageNumber="417">P1 0.700.79</th>
<td id="76FD5DEBFFBE0038FDE6FE4DD785C0D9" box="[602,703,483,509]" gridcol="1" gridrow="1" pageId="4" pageNumber="417">2.012.24</td>
<td id="76FD5DEBFFBE0038FC39FE4DD6D1C0D9" box="[901,1003,483,509]" gridcol="2" gridrow="1" pageId="4" pageNumber="417">2.542.76</td>
<td id="76FD5DEBFFBE0038FB0DFE4DD1C4C0D9" box="[1201,1278,483,509]" gridcol="3" gridrow="1" pageId="4" pageNumber="417">3.0</td>
</tr>
<tr id="352C3497FFBE0038FF23FDA1D1C4C30C" box="[159,1278,527,552]" gridrow="2" pageId="4" pageNumber="417">
<th id="76FD5DEBFFBE0038FF23FDA1D4BAC30C" box="[159,384,527,552]" gridcol="0" gridrow="2" pageId="4" pageNumber="417">P2 0.540.57</th>
<td id="76FD5DEBFFBE0038FDE6FDA1D785C30C" box="[602,703,527,552]" gridcol="1" gridrow="2" pageId="4" pageNumber="417">2.362.52</td>
<td id="76FD5DEBFFBE0038FC39FDA1D6D1C30C" box="[901,1003,527,552]" gridcol="2" gridrow="2" pageId="4" pageNumber="417">3.473.69</td>
<td id="76FD5DEBFFBE0038FB0DFDA1D1C4C30C" box="[1201,1278,527,552]" gridcol="3" gridrow="2" pageId="4" pageNumber="417">3.23.8</td>
</tr>
<tr id="352C3497FFBE0038FF23FD95D1C4C371" box="[159,1278,571,597]" gridrow="3" pageId="4" pageNumber="417">
<th id="76FD5DEBFFBE0038FF23FD95D4BAC371" box="[159,384,571,597]" gridcol="0" gridrow="3" pageId="4" pageNumber="417">P3 0.570.60</th>
<td id="76FD5DEBFFBE0038FDE6FD95D785C371" box="[602,703,571,597]" gridcol="1" gridrow="3" pageId="4" pageNumber="417">2.612.72</td>
<td id="76FD5DEBFFBE0038FC39FD95D6D1C371" box="[901,1003,571,597]" gridcol="2" gridrow="3" pageId="4" pageNumber="417">3.243.37</td>
<td id="76FD5DEBFFBE0038FB0DFD95D1C4C371" box="[1201,1278,571,597]" gridcol="3" gridrow="3" pageId="4" pageNumber="417">4.45.4</td>
</tr>
</table>
</paragraph>
<paragraph id="8BA336D5FFBEFFC3FF7BFD3AD11EC278" blockId="4.[151,1437,660,2029]" pageId="4" pageNumber="417">
Hypopygium (
<figureCitation id="13272A50FFBEFFC3FED6FD3AD482C389" box="[362,440,660,685]" captionStart="FIGURES 1 7" captionStartId="3.[151,264,1688,1711]" captionTargetBox="[174,1417,216,1640]" captionTargetId="figure@3.[174,1426,208,1640]" captionTargetPageId="3" captionText="FIGURES 1 7. Parorthocladius lazovskiensis sp. nov., male (1 3) and P. plolabius sp. nov., male (4 7). 1, 6, hypopygium in dorsal view; 2, gonocoxite and gonostylus in dorsal view; 3 5, anal point; 7, part of hypopygium without tergite IX in dorsal view. 1, male from Sakhalin Island; 2 3, male from Primorye Territory; 4 7, males from Amur River basin. Scale bars 50 µm." httpUri="https://zenodo.org/record/239450/files/figure.png" pageId="4" pageNumber="417">Figs. 4</figureCitation>
12). Tergite IX and anal point with 1217 setae. Anal point nearly parallel-sided, 2836 µm long and 1012 µm wide near apex. Laterosternite IX with 58 setae on each side. Transverse sternapodeme 100112 µm long, with sharply triangular oral projections. Virga 816 µm long, like comb, consisted of 57 short setae. Gonocoxite 200240 µm long; inferior volsella with roundish naked dorsal part and roundish ventral part placed below its dorsal part, covered by microtrichia and several setae (
<figureCitation id="13272A50FFBEFFC3FC7DFC8ED12BC21C" box="[961,1041,800,825]" captionStart="FIGURES 1 7" captionStartId="3.[151,264,1688,1711]" captionTargetBox="[174,1417,216,1640]" captionTargetId="figure@3.[174,1426,208,1640]" captionTargetPageId="3" captionText="FIGURES 1 7. Parorthocladius lazovskiensis sp. nov., male (1 3) and P. plolabius sp. nov., male (4 7). 1, 6, hypopygium in dorsal view; 2, gonocoxite and gonostylus in dorsal view; 3 5, anal point; 7, part of hypopygium without tergite IX in dorsal view. 1, male from Sakhalin Island; 2 3, male from Primorye Territory; 4 7, males from Amur River basin. Scale bars 50 µm." httpUri="https://zenodo.org/record/239450/files/figure.png" pageId="4" pageNumber="417">Figs. 7</figureCitation>
, 10). Gonostylus 96100 µm long, with well developed high preapical crista dorsalis (
<figureCitation id="13272A50FFBEFFC3FD6DFCEDD62CC278" box="[721,790,835,860]" captionStart="FIGURES 1 7" captionStartId="3.[151,264,1688,1711]" captionTargetBox="[174,1417,216,1640]" captionTargetId="figure@3.[174,1426,208,1640]" captionTargetPageId="3" captionText="FIGURES 1 7. Parorthocladius lazovskiensis sp. nov., male (1 3) and P. plolabius sp. nov., male (4 7). 1, 6, hypopygium in dorsal view; 2, gonocoxite and gonostylus in dorsal view; 3 5, anal point; 7, part of hypopygium without tergite IX in dorsal view. 1, male from Sakhalin Island; 2 3, male from Primorye Territory; 4 7, males from Amur River basin. Scale bars 50 µm." httpUri="https://zenodo.org/record/239450/files/figure.png" pageId="4" pageNumber="417">Fig. 6</figureCitation>
); megaseta 12 µm long.
</paragraph>
<paragraph id="8BA336D5FFBEFFC3FF7BFCC8D47AC284" blockId="4.[151,1437,660,2029]" pageId="4" pageNumber="417">
<emphasis id="B968EAC7FFBEFFC3FF7BFCC8D43EC259" bold="true" box="[199,260,870,893]" italics="true" pageId="4" pageNumber="417">Pupa</emphasis>
(n=4). Total length
<quantity id="4CE49B30FFBEFFC3FE52FCCBD7ACC25A" box="[494,662,869,894]" metricMagnitude="-3" metricUnit="m" metricValue="3.975" metricValueMax="4.45" metricValueMin="3.5" pageId="4" pageNumber="417" unit="mm" value="3.975" valueMax="4.45" valueMin="3.5">3.504.45 mm</quantity>
. Colouration brown. Exuviae brownish, with darker lateral parts of tergites VIIX.
</paragraph>
<paragraph id="8BA336D5FFBEFFC3FF7BFC05D7EAC57C" blockId="4.[151,1437,660,2029]" pageId="4" pageNumber="417">
Cephalothorax. Frontal apotome without tubercles and frontal setae. Thoracic horn absent but present pore and 3 precorneals (length in µm): Pc1 108128, Pc2 6068, Pc
<date id="FFA21015FFBEFFC3FCF5FC73D69FC2C3" box="[841,933,975,1003]" pageId="4" pageNumber="417" value="1932-03-28">3 2832</date>
. Distance between Pc1 and Pc2 4088 µm; distance between Pc2 and Pc
<date id="FFA21015FFBEFFC3FE68FBADD713C529" box="[468,553,1013,1041]" pageId="4" pageNumber="417" value="1932-12-03">3 1232</date>
µm. Antepronotum without setae. Dorsal surface of mesonotum rugose, in the middle part with 12 rows of pointed tubercles. Three dorsocentrals weak and hair-like; distance between Dc1 and Dc2 180200 µm; between Dc2 and Dc
<date id="FFA21015FFBEFFC3FDFBFBE0D7A6C57C" box="[583,668,1087,1116]" pageId="4" pageNumber="417" value="1916-12-03">3 1216</date>
µm.
</paragraph>
<paragraph id="8BA336D5FFBEFFC3FF7BFBCBD159C4B2" blockId="4.[151,1437,660,2029]" pageId="4" pageNumber="417">
Abdomen. Tergite I with shagreen of small spinules in middle part. Shagreen of tergite II covers almost its entire surface and consists of spinules almost the same size, sometimes at the posterior edge they are slightly larger. Shagreenation of tergites IIIVIII as in
<figureCitation id="13272A50FFBEFFC3FDEEFB05D79EC5E0" box="[594,676,1195,1220]" captionStart="FIGURES 13 17" captionStartId="6.[151,264,1709,1732]" captionTargetBox="[195,1382,183,1687]" captionTargetId="figure@6.[188,1398,177,1689]" captionTargetPageId="6" captionText="FIGURES 13 17. Parorthocladius plolabius sp. nov., pupa (13) and larva of fourth instar (14 17). 13, tergites I VIII and anal segment of male; 14 15, mentum; 16, mandible; 17, antenna. Scale bars: Fig. 13 — 200 µm; Figs. 14 17 — 20 µm." httpUri="https://zenodo.org/record/239451/files/figure.png" pageId="4" pageNumber="417">Fig. 13</figureCitation>
, namely spinules of anterior and posterior parts larger than spinules deposited between these parts. Sometimes spinules in middle part very small and sparse. Sternites IIIII with shagreen composed of small spinules in middle part and with lateral longitudinal stripes of spinules. Sternites IIV, IX without shagreen. Sternites VIVIII with weak shagreen in some parts. Segment I without lateral setae or sometimes with 1 pair of setae. Segments IIV with 23 pairs of lateral setae; segments VIVIII with 3 pairs of lateral setae; all lateral setae hair-like. Anal lobe 304340 µm long. Male genital sac overreaching anal lobe. Anal macrosetae 112 µm long (
<figureCitation id="13272A50FFBEFFC3FE06FAD3D736C4B2" box="[442,524,1405,1430]" captionStart="FIGURES 13 17" captionStartId="6.[151,264,1709,1732]" captionTargetBox="[195,1382,183,1687]" captionTargetId="figure@6.[188,1398,177,1689]" captionTargetPageId="6" captionText="FIGURES 13 17. Parorthocladius plolabius sp. nov., pupa (13) and larva of fourth instar (14 17). 13, tergites I VIII and anal segment of male; 14 15, mentum; 16, mandible; 17, antenna. Scale bars: Fig. 13 — 200 µm; Figs. 14 17 — 20 µm." httpUri="https://zenodo.org/record/239451/files/figure.png" pageId="4" pageNumber="417">Fig. 13</figureCitation>
). Anal macrosetae length/anal lobe length 0.330.37.
</paragraph>
<paragraph id="8BA336D5FFBEFFC3FF7BFA31D625C49C" blockId="4.[151,1437,660,2029]" box="[199,799,1439,1465]" pageId="4" pageNumber="417">
<emphasis id="B968EAC7FFBEFFC3FF7BFA31D49FC49C" bold="true" box="[199,421,1439,1464]" italics="true" pageId="4" pageNumber="417">Fourth instar larva</emphasis>
(n
<emphasis id="B968EAC7FFBEFFC3FE75FA0FD4E6C49C" box="[457,476,1441,1464]" italics="true" pageId="4" pageNumber="417">=</emphasis>
5). Total length
<quantity id="4CE49B30FFBEFFC3FD2DFA0ED621C49C" box="[657,795,1440,1464]" metricMagnitude="-3" metricUnit="m" metricValue="3.0999999999999996" metricValueMax="3.4" metricValueMin="2.8" pageId="4" pageNumber="417" unit="mm" value="3.0999999999999996" valueMax="3.4" valueMin="2.8">2.83.4 mm</quantity>
.
</paragraph>
<paragraph id="8BA336D5FFBEFFC3FF7BFA6DD7E3C6C9" blockId="4.[151,1437,660,2029]" pageId="4" pageNumber="417">
Head. Head capsule brown or dark brown, postoccipital margin black. Head length 300325 µm, head width 225250 µm. S IIII of labrum simple and hair-like, S IV like small tubercles. Pecten epipharyngis consisting of 3 scales, middle scale narrowest. Premandible 76 µm long, in apical part expanded and rounded, with 1 tooth. Antenna with 5 segments, length of segments 15 (in µm): 5256: 2024: 1012: 610: 48; segment 3 longer than segment 4, rarely segments 3 and 4 of equal length (n=1); one large and two small ring organs in proximal 1/4 of basal segment; apex of segment 2 with stylus 12 µm long and lauterborn organs as long as 3rd segment; blade ending at base of segment 5 (
<figureCitation id="13272A50FFBEFFC3FE5FF934D70DC797" box="[483,567,1690,1715]" captionStart="FIGURES 13 17" captionStartId="6.[151,264,1709,1732]" captionTargetBox="[195,1382,183,1687]" captionTargetId="figure@6.[188,1398,177,1689]" captionTargetPageId="6" captionText="FIGURES 13 17. Parorthocladius plolabius sp. nov., pupa (13) and larva of fourth instar (14 17). 13, tergites I VIII and anal segment of male; 14 15, mentum; 16, mandible; 17, antenna. Scale bars: Fig. 13 — 200 µm; Figs. 14 17 — 20 µm." httpUri="https://zenodo.org/record/239451/files/figure.png" pageId="4" pageNumber="417">Fig. 17</figureCitation>
); AR 1.171.30. Mandible 104112 µm long, with 3 inner teeth and 1 apical tooth equal to first inner tooth or somewhat longer; other 2 inner teeth equal; seta subdentalis stout, with pointed apex (
<figureCitation id="13272A50FFBEFFC3FF62F94ED409C7DD" box="[222,307,1760,1785]" captionStart="FIGURES 13 17" captionStartId="6.[151,264,1709,1732]" captionTargetBox="[195,1382,183,1687]" captionTargetId="figure@6.[188,1398,177,1689]" captionTargetPageId="6" captionText="FIGURES 13 17. Parorthocladius plolabius sp. nov., pupa (13) and larva of fourth instar (14 17). 13, tergites I VIII and anal segment of male; 14 15, mentum; 16, mandible; 17, antenna. Scale bars: Fig. 13 — 200 µm; Figs. 14 17 — 20 µm." httpUri="https://zenodo.org/record/239451/files/figure.png" pageId="4" pageNumber="417">Fig. 16</figureCitation>
). Mentum high and subtriangular, with 3 equal median teeth and 4 pairs of lateral teeth. Beard of ventromental plates consisted of a tuft of long setae (
<figureCitation id="13272A50FFBEFFC3FD49F8ADD670C63F" box="[757,842,1795,1820]" captionStart="FIGURES 13 17" captionStartId="6.[151,264,1709,1732]" captionTargetBox="[195,1382,183,1687]" captionTargetId="figure@6.[188,1398,177,1689]" captionTargetPageId="6" captionText="FIGURES 13 17. Parorthocladius plolabius sp. nov., pupa (13) and larva of fourth instar (14 17). 13, tergites I VIII and anal segment of male; 14 15, mentum; 16, mandible; 17, antenna. Scale bars: Fig. 13 — 200 µm; Figs. 14 17 — 20 µm." httpUri="https://zenodo.org/record/239451/files/figure.png" pageId="4" pageNumber="417">Fig. 14</figureCitation>
). Very often apical part of mentum in mature larvae erased (
<figureCitation id="13272A50FFBEFFC3FF4EF88BD473C61A" box="[242,329,1829,1854]" captionStart="FIGURES 13 17" captionStartId="6.[151,264,1709,1732]" captionTargetBox="[195,1382,183,1687]" captionTargetId="figure@6.[188,1398,177,1689]" captionTargetPageId="6" captionText="FIGURES 13 17. Parorthocladius plolabius sp. nov., pupa (13) and larva of fourth instar (14 17). 13, tergites I VIII and anal segment of male; 14 15, mentum; 16, mandible; 17, antenna. Scale bars: Fig. 13 — 200 µm; Figs. 14 17 — 20 µm." httpUri="https://zenodo.org/record/239451/files/figure.png" pageId="4" pageNumber="417">Fig. 15</figureCitation>
). Posterior parapods slightly shorter than anal tubules or both structures of equal length. Anal tubules of one pair wider than anal tubules of other pair. Procercus 16 µm long and 2428 µm wide, with 2 short lateral setae and 6 apical setae 432608 µm long; rarely 5 apical setae present (n=1). Body segments IVII with 2 pairs of lateral setae: one pair of setae simple and one pair plumose (sometimes bifid); plumose setae placed at posteroventral corner of segment. Segment VIII with 1 pair of simple setae. Sometimes simple on one side and plumose seta on opposite side of the same segment.
</paragraph>
<paragraph id="8BA336D5FFBFFFC2FF7BFF39D751C194" blockId="5.[151,1436,151,320]" box="[199,619,151,176]" pageId="5" pageNumber="418">
<emphasis id="B968EAC7FFBFFFC2FF7BFF39D4EBC194" bold="true" box="[199,465,151,176]" pageId="5" pageNumber="418">Diagnostic characters.</emphasis>
See the keys.
</paragraph>
</subSubSection>
<subSubSection id="C306655EFFBFFFC2FF7BFF12D461C01B" pageId="5" pageNumber="418" type="etymology">
<paragraph id="8BA336D5FFBFFFC2FF7BFF12D461C01B" blockId="5.[151,1436,151,320]" pageId="5" pageNumber="418">
<emphasis id="B968EAC7FFBFFFC2FF7BFF12D474C1F0" bold="true" box="[199,334,188,212]" pageId="5" pageNumber="418">Etymology.</emphasis>
The specific epithet is derived from first letters of the names of rivers in the basin, where the new species was collected:
<emphasis id="B968EAC7FFBFFFC2FE29FF4ED490C1DC" box="[405,426,224,248]" italics="true" pageId="5" pageNumber="418">pl</emphasis>
—Plotnikova River,
<emphasis id="B968EAC7FFBFFFC2FD2EFF4ED78CC1DC" box="[658,694,224,248]" italics="true" pageId="5" pageNumber="418">ola</emphasis>
—Ola River,
<emphasis id="B968EAC7FFBFFFC2FCF0FF4ED658C1DC" box="[844,866,224,248]" italics="true" pageId="5" pageNumber="418">bi</emphasis>
—Bidzhan River.
<emphasis id="B968EAC7FFBFFFC2FF7BFEADD45AC038" bold="true" box="[199,352,259,284]" pageId="5" pageNumber="418">Distribution.</emphasis>
Known from the
<typeStatus id="54A78877FFBFFFC2FD8DFEABD75BC039" box="[561,609,261,285]" pageId="5" pageNumber="418">type</typeStatus>
locality in Kamchatka and Magadan Regions, Amur River basin of the Russian Far East.
</paragraph>
</subSubSection>
</treatment>
</document>