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<mods:title id="DC41B3F54ABE8A817DBBB1A7A94BFDA6">Middle Permian (Roadian) gastropods from the Khao Khad Formation, Central Thailand: Implications for palaeogeography of the Indochina Terrane</mods:title>
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<mods:namePart id="251FEBB5161931C9A3A2B2C511647A1D">Ketwetsuriya, Chatchalerm</mods:namePart>
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<mods:namePart id="80B97D8096E38321B3CE6B9F63889B98">Karapunar, Baran</mods:namePart>
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<mods:namePart id="37C01F7294B4705EC9BF023943F0FB8E">Charoentitirat, Thasinee</mods:namePart>
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<treatment id="03B587AB4F281561FF5178E7FDB7F9AA" ID-DOI="http://doi.org/10.5281/zenodo.3803879" ID-GBIF-Taxon="163737145" ID-Zenodo-Dep="3803879" LSID="urn:lsid:plazi:treatment:03B587AB4F281561FF5178E7FDB7F9AA" httpUri="http://treatment.plazi.org/id/03B587AB4F281561FF5178E7FDB7F9AA" lastPageNumber="32" pageId="31" pageNumber="32">
<subSubSection id="C30665364F281561FF5178E7FE37FF31" box="[151,508,153,180]" pageId="31" pageNumber="32" type="nomenclature">
<paragraph id="8BA336BD4F281561FF5178E7FE37FF31" blockId="31.[151,508,153,215]" box="[151,508,153,180]" pageId="31" pageNumber="32">
<heading id="D0EB81D14F281561FF5178E7FE37FF31" bold="true" box="[151,508,153,180]" fontSize="10" level="2" pageId="31" pageNumber="32" reason="2">
<emphasis id="B968EAAF4F281561FF5178E7FE37FF31" bold="true" box="[151,508,153,180]" pageId="31" pageNumber="32">
<taxonomicName id="4C1C4D3E4F281561FF5178E7FE5CFF31" authority="Ketwetsuriya &amp; Karapunar &amp; Charoentitirat &amp; Nützel, 2020" authorityName="Ketwetsuriya &amp; Karapunar &amp; Charoentitirat &amp; Nützel" authorityYear="2020" box="[151,407,153,180]" class="Gastropoda" family="Trachyspiridae" genus="Trachyspira" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cycloneritimorpha" pageId="31" pageNumber="32" phylum="Mollusca" rank="species" species="eleganta" status="sp. nov.">
<emphasis id="B968EAAF4F281561FF5178E7FE5CFF31" bold="true" box="[151,407,153,180]" italics="true" pageId="31" pageNumber="32">Trachyspira eleganta</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25B57D44F281561FE5878E4FE37FF31" box="[414,508,154,180]" pageId="31" pageNumber="32" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C30665364F281561FF5178C3FEFBFF52" box="[151,304,189,215]" pageId="31" pageNumber="32" type="description">
<paragraph id="8BA336BD4F281561FF5178C3FEFBFF52" blockId="31.[151,508,153,215]" box="[151,304,189,215]" pageId="31" pageNumber="32">
(
<figureCitation id="13272A384F281561FF6678C3FF25FF52" box="[160,238,189,215]" captionStart="FIGURE 21" captionStartId="30.[151,250,1173,1198]" captionTargetBox="[166,1421,273,1148]" captionTargetId="figure@30.[166,1421,273,1148]" captionTargetPageId="30" captionText="FIGURE 21. AD, Trachydomia suwanneeae sp. nov., ESKU-19-LP 26, holotype. EF, Trachydomia cf. nodusum (Meek &amp; Worthen, 1866), ESKU-19-LP 92. GK, Trachyspira eleganta sp. nov., ESKU-19-LP 28, holotype. Scale bars represent: 2 mm (AD, GK); 1 mm (EF)." figureDoi="http://doi.org/10.5281/zenodo.3763949" httpUri="https://zenodo.org/record/3763949/files/figure.png" pageId="31" pageNumber="32">Fig. 21</figureCitation>
GK)
</paragraph>
</subSubSection>
<subSubSection id="C30665364F281561FF517978FD28FE9A" box="[151,739,261,288]" pageId="31" pageNumber="32" type="etymology">
<paragraph id="8BA336BD4F281561FF517978FD28FE9A" blockId="31.[151,1437,261,1584]" box="[151,739,261,288]" pageId="31" pageNumber="32">
<emphasis id="B968EAAF4F281561FF517978FED5FEA5" bold="true" box="[151,286,262,288]" pageId="31" pageNumber="32">Etymology.</emphasis>
From Latin, meaning elegant, beautiful.
</paragraph>
</subSubSection>
<subSubSection id="C30665364F281561FF017954FBBCFEED" pageId="31" pageNumber="32" type="materials_examined">
<paragraph id="8BA336BD4F281561FF017954FD28FEC6" blockId="31.[151,1437,261,1584]" box="[199,739,297,324]" pageId="31" pageNumber="32">
<materialsCitation id="3B743CE04F281561FF017954FD28FEC6" ID-GBIF-Occurrence="2608527133" box="[199,739,297,324]" collectionCode="LP" pageId="31" pageNumber="32" specimenCode="ESKU-19-LP 28" specimenCount="1" typeStatus="holotype">
<emphasis id="B968EAAF4F281561FF017954FEF0FEC1" bold="true" box="[199,315,298,324]" pageId="31" pageNumber="32">
<typeStatus id="54A7881F4F281561FF017954FEFDFEC1" box="[199,310,298,324]" pageId="31" pageNumber="32" type="holotype">Holotype</typeStatus>
.
</emphasis>
Only
<specimenCount id="9D1AFD344F281561FE477954FDD1FEC6" box="[385,538,297,324]" pageId="31" pageNumber="32">one specimen</specimenCount>
<specimenCode id="DBBA9EC64F281561FDE77957FD14FEC6" box="[545,735,297,323]" pageId="31" pageNumber="32">ESKU-19-LP 28</specimenCode>
.
</materialsCitation>
</paragraph>
<paragraph id="8BA336BD4F281561FF017933FBBCFEED" blockId="31.[151,1437,261,1584]" box="[199,1143,332,360]" pageId="31" pageNumber="32">
Dimensions (mm): ESKU-19-
<collectionCode id="ED0DAE784F281561FDD07933FDF3FEE2" box="[534,568,333,359]" country="Russia" name="Laboratory of Palaeontology" pageId="31" pageNumber="32">LP</collectionCode>
28: height = 17.6; width = 13.3; apical angle = 75
<superScript id="7C699BF54F281561FBAD7932FBBBFEDF" attach="none" box="[1131,1136,332,346]" fontSize="6" pageId="31" pageNumber="32">º</superScript>
.
</paragraph>
</subSubSection>
<subSubSection id="C30665364F281561FF01790CFE74FE56" pageId="31" pageNumber="32" type="distribution">
<paragraph id="8BA336BD4F281561FF01790CFE74FE56" blockId="31.[151,1437,261,1584]" pageId="31" pageNumber="32">
<emphasis id="B968EAAF4F281561FF01790CFD6AFE09" bold="true" box="[199,673,369,396]" pageId="31" pageNumber="32">Type locality and stratigraphical range.</emphasis>
Erawan Hill, Chong Sarika sub-district, Phatthana Nikhom district, located about
<quantity id="4CE49B584F281561FEF179EBFEB4FE2A" box="[311,383,405,431]" metricMagnitude="4" metricUnit="m" metricValue="1.3" pageId="31" pageNumber="32" unit="km" value="13.0">13 km</quantity>
east of
<collectingRegion id="49D8F85F4F281561FE1C79EBFD69FE2A" box="[474,674,405,431]" country="Thailand" name="Lop Buri" pageId="31" pageNumber="32">Lopburi Province</collectingRegion>
, Central
<collectingCountry id="F30B762D4F281561FCCF79EBFCA6FE2A" box="[777,877,405,431]" name="Thailand" pageId="31" pageNumber="32">Thailand</collectingCountry>
(
<figureCitation id="13272A384F281561FCBB79EBFC0BFE2A" box="[893,960,405,431]" captionStart="FIGURE" captionStartId="1.[151,168,1975,2001]" captionTargetBox="[243,1345,1164,1941]" captionTargetId="figure@1.[236,1350,1160,1948]" captionTargetPageId="1" captionText="FIGURE 1. Geological map of the study location at Erawan Hill, Chong Sarika sub-district, Phatthana Nikhom district, located about 13 km east of Lopburi Province, Central Thailand." figureDoi="http://doi.org/10.5281/zenodo.3763909" httpUri="https://zenodo.org/record/3763909/files/figure.png" pageId="31" pageNumber="32">Fig. 1</figureCitation>
), Khao Khad Formation,
<collectingRegion id="49D8F85F4F281561FB2179EBFA8DFE2A" box="[1255,1350,405,431]" country="Thailand" name="Sara Buri" pageId="31" pageNumber="32">Saraburi</collectingRegion>
Group, Middle Permian, Roadian.
</paragraph>
</subSubSection>
<subSubSection id="C30665364F281561FF0179A3FC22FCBE" pageId="31" pageNumber="32" type="description">
<paragraph id="8BA336BD4F281561FF0179A3FC22FCBE" blockId="31.[151,1437,261,1584]" pageId="31" pageNumber="32">
<emphasis id="B968EAAF4F281561FF0179A3FE92FE7D" bold="true" box="[199,345,477,504]" pageId="31" pageNumber="32">Description.</emphasis>
Medium-size, moderately high-spired shell, conical, subturbiniform, consisting of approximately 5 whorls; apex acute, spire angle 75°; body whorl height about 80 percent of the total height; suture adpressed; earliest two whorls without visible ornament (preservation?), convex; following whorl with fine pustules and narrow shoulder; last two preserved whorls ornamented with two categories of nodes, large nodes arranged in spiral rows and fine, densely spaced nodes arranged in opisthocline rows tending to fuse to opisthocline ribs; body whorl with three rows of nodes,1014 nodes per row; uppermost nodular row near adapical suture, forming edge of narrow ramp; middle row of nodes strongest, situated near abapical suture; third row with weakest nodes, situated on base; whorl angulated at nodular rows; whorl face concave between nodular rows; base rounded, anomphalous, ornamented with numerous spiral cords consisting of small pustules representing continuations of opisthocline rows on whorl face; aperture acute adapically, outer and anterior lip evenly rounded.
</paragraph>
</subSubSection>
<subSubSection id="C30665364F281561FF017B38FDB7F9AA" pageId="31" pageNumber="32" type="discussion">
<paragraph id="8BA336BD4F281561FF017B38FA41FC22" blockId="31.[151,1437,261,1584]" pageId="31" pageNumber="32">
<emphasis id="B968EAAF4F281561FF017B38FEF0FCE5" bold="true" box="[199,315,838,864]" pageId="31" pageNumber="32">Remarks.</emphasis>
This single specimen at hand is assigned to the genus
<taxonomicName id="4C1C4D3E4F281561FC5F7B3BFB33FCDA" authority="Gemmellaro, 1889" authorityName="Gemmellaro" authorityYear="1889" box="[921,1272,837,863]" class="Gastropoda" family="Trachyspiridae" genus="Trachyspira" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cycloneritimorpha" pageId="31" pageNumber="32" phylum="Mollusca" rank="genus">
<emphasis id="B968EAAF4F281561FC5F7B3BFBEBFCDA" box="[921,1056,837,863]" italics="true" pageId="31" pageNumber="32">Trachyspira</emphasis>
<bibRefCitation id="EF8D4B4C4F281561FBE17B3BFB33FCDA" author="Gemmellaro" box="[1063,1272,837,863]" firstAuthor="Gemmellaro" pageId="31" pageNumber="32" pagination="97 - 182" refId="ref24122" refString="Gemmellaro, G. G. (1889) La Fauna dei Calcari Con Fusulina della Valle del Fiume Sosio nella Provincia di Palermo, Fasc 2. Nautiloidea e Gastropoda. Giornale di Scienze Naturali e Economiche, 20, 97 - 182. https: // doi. org / 10.5962 / bhl. title. 10774" type="journal article" year="1889">Gemmellaro, 1889</bibRefCitation>
</taxonomicName>
which is characterized by having two different size-categories of nodes: numerous small pustules, aligned in opisthocline rows and few large nodes arranged in spiral rows (
<bibRefCitation id="EF8D4B4C4F281561FD487BF3FCABFC2D" author="Knight" box="[654,864,909,936]" etAl="et al." firstAuthor="Knight" pageId="31" pageNumber="32" pagination="310" refId="ref25325" refString="Knight, J. B., Cox, L. R., Keen, A. M., Batten, R. L., Yochelson, E. L. &amp; Robertson, R. (1960) Systematic descriptions. In: Moore, R. C. (Ed.), Treatise on invertebrate paleontology. Part I. Mollusca 1. Geological Society of America &amp; University of Kansas Press, Kansas, pp. I 169 - I 310." type="journal article" year="1960">
Knight
<emphasis id="B968EAAF4F281561FD237BF0FCDDFC22" box="[741,790,909,935]" italics="true" pageId="31" pageNumber="32">et al</emphasis>
. 1960
</bibRefCitation>
). It is the first report of this genus from
<collectingCountry id="F30B762D4F281561FAD97BF3FA4DFC22" box="[1311,1414,909,935]" name="Thailand" pageId="31" pageNumber="32">Thailand</collectingCountry>
.
</paragraph>
<paragraph id="8BA336BD4F281561FF017BCFFC13FBD9" blockId="31.[151,1437,261,1584]" pageId="31" pageNumber="32">
<taxonomicName id="4C1C4D3E4F281561FF017BCFFE7FFC4E" authority="Ketwetsuriya &amp; Karapunar &amp; Charoentitirat &amp; Nützel, 2020" authorityName="Ketwetsuriya &amp; Karapunar &amp; Charoentitirat &amp; Nützel" authorityYear="2020" box="[199,436,945,971]" class="Gastropoda" family="Trachyspiridae" genus="Trachyspira" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cycloneritimorpha" pageId="31" pageNumber="32" phylum="Mollusca" rank="species" species="eleganta" status="sp. nov.">
<emphasis id="B968EAAF4F281561FF017BCFFE7FFC4E" box="[199,436,945,971]" italics="true" pageId="31" pageNumber="32">Trachyspira eleganta</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25B57D44F281561FE7C7BCCFDC4FC49" box="[442,527,946,972]" pageId="31" pageNumber="32" rank="species">sp. nov.</taxonomicNameLabel>
from the Khao Khad Limestone resembles the specimens illustrated and described as
<taxonomicName id="4C1C4D3E4F281561FF737BA8FE31FC6A" authority="Wang, 1982" authorityName="Wang" authorityYear="1982" box="[181,506,981,1008]" class="Gastropoda" family="Trachyspiridae" genus="Trachyspira" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cycloneritimorpha" pageId="31" pageNumber="32" phylum="Mollusca" rank="species" species="obliquinodula">
<emphasis id="B968EAAF4F281561FF737BA8FF0EFC6A" box="[181,197,982,1007]" italics="true" pageId="31" pageNumber="32">T</emphasis>
.
<emphasis id="B968EAAF4F281561FF177BABFEA5FC6A" box="[209,366,981,1007]" italics="true" pageId="31" pageNumber="32">obliquinodula</emphasis>
<bibRefCitation id="EF8D4B4C4F281561FEB37BA8FE31FC6A" author="Wang" box="[373,506,981,1008]" firstAuthor="Wang" pageId="31" pageNumber="32" pagination="491 - 496" refId="ref28335" refString="Wang, H. J. (1982) Late Permian gastropods from Heshan of Laibin, Guangxi Province. Acta Palaeontologica Sinica, 21, 491 - 496." type="journal article" year="1982">Wang, 1982</bibRefCitation>
</taxonomicName>
by
<bibRefCitation id="EF8D4B4C4F281561FDE57BA8FCCFFC6A" author="Pan &amp; Erwin" box="[547,772,981,1008]" firstAuthor="Pan" pageId="31" pageNumber="32" pagination="1 - 49" refId="ref27138" refString="Pan, H. - Z. &amp; Erwin, D. H. (2002) Gastropods from the Permian of Guangxi and Yunnan provinces, South China. Journal of Paleontology, 78 (Supplement 56), 1 - 49. https: // doi. org / 10.1666 / 0022 - 3360 (2002) 76 [1: GFTPOG] 2.0. CO; 2" type="journal article" year="2002">Pan &amp; Erwin (2002)</bibRefCitation>
from the Permian of
<collectingRegion id="49D8F85F4F281561FC367BABFB9BFC6A" box="[1008,1104,981,1007]" country="China" name="Guangxi" pageId="31" pageNumber="32">Guangxi</collectingRegion>
and
<collectingRegion id="49D8F85F4F281561FB427BA8FB11FC75" box="[1156,1242,982,1008]" country="China" name="Yunnan" pageId="31" pageNumber="32">Yunnan</collectingRegion>
Provinces, South
<collectingCountry id="F30B762D4F281561FF517B87FF15FB96" box="[151,222,1017,1043]" name="China" pageId="31" pageNumber="32">China</collectingCountry>
. The
<typeStatus id="54A7881F4F281561FEDE7B84FE83FB91" box="[280,328,1018,1044]" pageId="31" pageNumber="32">type</typeStatus>
material of this species as illustrated by
<bibRefCitation id="EF8D4B4C4F281561FCCE7B84FC51FB96" author="Wang" box="[776,922,1017,1044]" firstAuthor="Wang" pageId="31" pageNumber="32" pagination="491 - 496" refId="ref28335" refString="Wang, H. J. (1982) Late Permian gastropods from Heshan of Laibin, Guangxi Province. Acta Palaeontologica Sinica, 21, 491 - 496." type="journal article" year="1982">Wang (1982)</bibRefCitation>
is poorly preserved and not well-documented. However, these specimens as well as those illustrated by
<bibRefCitation id="EF8D4B4C4F281561FCEF7C60FBD8FBB2" author="Pan &amp; Erwin" box="[809,1043,1053,1080]" firstAuthor="Pan" pageId="31" pageNumber="32" pagination="1 - 49" refId="ref27138" refString="Pan, H. - Z. &amp; Erwin, D. H. (2002) Gastropods from the Permian of Guangxi and Yunnan provinces, South China. Journal of Paleontology, 78 (Supplement 56), 1 - 49. https: // doi. org / 10.1666 / 0022 - 3360 (2002) 76 [1: GFTPOG] 2.0. CO; 2" type="journal article" year="2002">Pan &amp; Erwin (2002)</bibRefCitation>
are much broader and stouter and hence have a larger apical angle (&gt; 90°) and the nodules are much coarser.
</paragraph>
<paragraph id="8BA336BD4F281561FF017C1BFDB7F9AA" blockId="31.[151,1437,261,1584]" pageId="31" pageNumber="32">
The type species
<taxonomicName id="4C1C4D3E4F281561FE4B7C18FCE7FBFA" authority="Gemmellaro, 1889" authorityName="Gemmellaro" authorityYear="1889" box="[397,812,1125,1152]" class="Gastropoda" family="Trachyspiridae" genus="Trachyspira" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cycloneritimorpha" pageId="31" pageNumber="32" phylum="Mollusca" rank="species" species="delphinuloides">
<emphasis id="B968EAAF4F281561FE4B7C18FE56FBFA" box="[397,413,1126,1151]" italics="true" pageId="31" pageNumber="32">T</emphasis>
.
<emphasis id="B968EAAF4F281561FE6A7C1BFD9AFBFA" box="[428,593,1125,1151]" italics="true" pageId="31" pageNumber="32">delphinuloides</emphasis>
<bibRefCitation id="EF8D4B4C4F281561FD9F7C1BFCE7FBFA" author="Gemmellaro" box="[601,812,1125,1151]" firstAuthor="Gemmellaro" pageId="31" pageNumber="32" pagination="97 - 182" refId="ref24122" refString="Gemmellaro, G. G. (1889) La Fauna dei Calcari Con Fusulina della Valle del Fiume Sosio nella Provincia di Palermo, Fasc 2. Nautiloidea e Gastropoda. Giornale di Scienze Naturali e Economiche, 20, 97 - 182. https: // doi. org / 10.5962 / bhl. title. 10774" type="journal article" year="1889">Gemmellaro, 1889</bibRefCitation>
</taxonomicName>
as illustrated by
<bibRefCitation id="EF8D4B4C4F281561FC377C18FB4CFBFA" author="Batten" box="[1009,1159,1125,1152]" firstAuthor="Batten" pageId="31" pageNumber="32" pagination="1 - 26" refId="ref23086" refString="Batten, R. L. (1979) Permian gastropods from Perak, Malaysia. Part 2. The trochids, patellids, and neritids. American Museum Novitates, 2685, 1 - 26." type="journal article" year="1979">Batten (1979</bibRefCitation>
, p. 21, figs 2931) from North Africa and
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is also similar to
<taxonomicName id="4C1C4D3E4F281561FD557CF4FCD9FB26" authority="Ketwetsuriya &amp; Karapunar &amp; Charoentitirat &amp; Nützel, 2020" authorityName="Ketwetsuriya &amp; Karapunar &amp; Charoentitirat &amp; Nützel" authorityYear="2020" box="[659,786,1161,1188]" class="Gastropoda" family="Trachyspiridae" genus="Trachyspira" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cycloneritimorpha" pageId="31" pageNumber="32" phylum="Mollusca" rank="species" species="eleganta" status="sp. nov.">
<emphasis id="B968EAAF4F281561FD557CF4FD68FB26" box="[659,675,1162,1187]" italics="true" pageId="31" pageNumber="32">T</emphasis>
.
<emphasis id="B968EAAF4F281561FD747CF7FCD9FB26" box="[690,786,1161,1187]" italics="true" pageId="31" pageNumber="32">eleganta</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25B57D44F281561FCDC7CF4FCB9FB21" box="[794,882,1162,1188]" pageId="31" pageNumber="32" rank="species">sp. nov.</taxonomicNameLabel>
in having three principal rows of nodes with the second row dominant but
<bibRefCitation id="EF8D4B4C4F281561FE057CD3FDB9FB42" author="Batten's" box="[451,626,1197,1223]" firstAuthor="Batten's" pageId="31" pageNumber="32" pagination="1 - 26" refId="ref23086" refString="Batten, R. L. (1979) Permian gastropods from Perak, Malaysia. Part 2. The trochids, patellids, and neritids. American Museum Novitates, 2685, 1 - 26." type="journal article" year="1979">Battens (1979)</bibRefCitation>
specimen have a larger apical angle and the second order pustules are larger in the African specimen. However, the original illustration of
<taxonomicName id="4C1C4D3E4F281561FC487CACFB9AFB6E" authorityName="Gemmellaro" authorityYear="1889" box="[910,1105,1233,1260]" class="Gastropoda" family="Trachyspiridae" genus="Trachyspira" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cycloneritimorpha" pageId="31" pageNumber="32" phylum="Mollusca" rank="species" species="delphinuloides">
<emphasis id="B968EAAF4F281561FC487CACFC55FB6E" box="[910,926,1234,1259]" italics="true" pageId="31" pageNumber="32">T</emphasis>
.
<emphasis id="B968EAAF4F281561FC6A7CAFFB9AFB6E" box="[940,1105,1233,1259]" italics="true" pageId="31" pageNumber="32">delphinuloides</emphasis>
</taxonomicName>
given by
<bibRefCitation id="EF8D4B4C4F281561FB797CAFFA57FB6E" author="Gemmellaro" box="[1215,1436,1233,1259]" firstAuthor="Gemmellaro" pageId="31" pageNumber="32" pagination="97 - 182" refId="ref24122" refString="Gemmellaro, G. G. (1889) La Fauna dei Calcari Con Fusulina della Valle del Fiume Sosio nella Provincia di Palermo, Fasc 2. Nautiloidea e Gastropoda. Giornale di Scienze Naturali e Economiche, 20, 97 - 182. https: // doi. org / 10.5962 / bhl. title. 10774" type="journal article" year="1889">Gemmellaro (1889)</bibRefCitation>
shows a specimen with relatively small first order nodules that are spirally elongated.
<bibRefCitation id="EF8D4B4C4F281561FB9E7C8BFB3BFA8A" author="Knight" box="[1112,1264,1269,1295]" firstAuthor="Knight" pageId="31" pageNumber="32" pagination="1 - 510" refId="ref25231" refString="Knight, J. B. (1941) Paleozoic gastropod genotypes. Geological Society of America Special Paper, 32, 1 - 510. https: // doi. org / 10.1130 / SPE 32 - p 1" type="journal article" year="1941">Knight (1941</bibRefCitation>
, pl. 56) figured a specimen form the type series that differs considerably from
<bibRefCitation id="EF8D4B4C4F281561FCA47D67FB9FFAB6" author="Gemmellaro's" box="[866,1108,1305,1331]" firstAuthor="Gemmellaro's" pageId="31" pageNumber="32" pagination="97 - 182" refId="ref24122" refString="Gemmellaro, G. G. (1889) La Fauna dei Calcari Con Fusulina della Valle del Fiume Sosio nella Provincia di Palermo, Fasc 2. Nautiloidea e Gastropoda. Giornale di Scienze Naturali e Economiche, 20, 97 - 182. https: // doi. org / 10.5962 / bhl. title. 10774" type="journal article" year="1889">Gemmellaros (1889)</bibRefCitation>
figure. Nevertheless,
<bibRefCitation id="EF8D4B4C4F281561FA887D67FF29FAD2" author="Knight" firstAuthor="Knight" pageId="31" pageNumber="32" pagination="1 - 510" refId="ref25231" refString="Knight, J. B. (1941) Paleozoic gastropod genotypes. Geological Society of America Special Paper, 32, 1 - 510. https: // doi. org / 10.1130 / SPE 32 - p 1" type="journal article" year="1941">Knight (1941)</bibRefCitation>
identified this specimen as possible
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. This specimen (probably the one used for the drawing in the Treatise,
<bibRefCitation id="EF8D4B4C4F281561FF387D1FFE00FAFE" author="Knight" box="[254,459,1377,1404]" etAl="et al." firstAuthor="Knight" pageId="31" pageNumber="32" pagination="310" refId="ref25325" refString="Knight, J. B., Cox, L. R., Keen, A. M., Batten, R. L., Yochelson, E. L. &amp; Robertson, R. (1960) Systematic descriptions. In: Moore, R. C. (Ed.), Treatise on invertebrate paleontology. Part I. Mollusca 1. Geological Society of America &amp; University of Kansas Press, Kansas, pp. I 169 - I 310." type="journal article" year="1960">Knight et al. 1960</bibRefCitation>
, fig. 182/6) differs from our specimen in having a broader ramp, finer second order and stronger first order nodes, especially the adapical ones.
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<emphasis id="B968EAAF4F281561FCC57DF8FCD8FA1A" box="[771,787,1414,1439]" italics="true" pageId="31" pageNumber="32">T</emphasis>
.
<emphasis id="B968EAAF4F281561FCE67DFBFC6CFA1A" box="[800,935,1413,1439]" italics="true" pageId="31" pageNumber="32">heshanensis</emphasis>
<bibRefCitation id="EF8D4B4C4F281561FC687DF8FB4DFA1A" author="Pan &amp; Erwin" box="[942,1158,1413,1440]" firstAuthor="Pan" pageId="31" pageNumber="32" pagination="1 - 49" refId="ref27138" refString="Pan, H. - Z. &amp; Erwin, D. H. (2002) Gastropods from the Permian of Guangxi and Yunnan provinces, South China. Journal of Paleontology, 78 (Supplement 56), 1 - 49. https: // doi. org / 10.1666 / 0022 - 3360 (2002) 76 [1: GFTPOG] 2.0. CO; 2" type="journal article" year="2002">Pan &amp; Erwin, 2002</bibRefCitation>
</taxonomicName>
(p. 25, fig. 12.1112.14) from the Late Permian of
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has a larger apical angle (close to 90°), the first order nodes are weaker at comparable growth stages and it lacks a third row of first order nodes on the base.
<taxonomicName id="4C1C4D3E4F281561FC2E7DB0FABCFA62" authority="Pan &amp; Erwin, 2002" authorityName="Pan &amp; Erwin" authorityYear="2002" box="[1000,1399,1485,1512]" class="Gastropoda" family="Trachyspiridae" genus="Trachyspira" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cycloneritimorpha" pageId="31" pageNumber="32" phylum="Mollusca" rank="species" species="quangxiensis">
<emphasis id="B968EAAF4F281561FC2E7DB0FC33FA62" box="[1000,1016,1486,1511]" italics="true" pageId="31" pageNumber="32">T</emphasis>
.
<emphasis id="B968EAAF4F281561FBC07DB0FB53FA62" box="[1030,1176,1486,1511]" italics="true" pageId="31" pageNumber="32">quangxiensis</emphasis>
<bibRefCitation id="EF8D4B4C4F281561FB597DB0FABCFA62" author="Pan &amp; Erwin" box="[1183,1399,1485,1512]" firstAuthor="Pan" pageId="31" pageNumber="32" pagination="1 - 49" refId="ref27138" refString="Pan, H. - Z. &amp; Erwin, D. H. (2002) Gastropods from the Permian of Guangxi and Yunnan provinces, South China. Journal of Paleontology, 78 (Supplement 56), 1 - 49. https: // doi. org / 10.1666 / 0022 - 3360 (2002) 76 [1: GFTPOG] 2.0. CO; 2" type="journal article" year="2002">Pan &amp; Erwin, 2002</bibRefCitation>
</taxonomicName>
(p. 25, fig. 12.912.10) from the Late Permian of
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has more convex whorls, a wider ramp and three instead of two rows of first order nodes on the whorl face.
</paragraph>
</subSubSection>
</treatment>
</document>