194 lines
23 KiB
XML
194 lines
23 KiB
XML
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<mods:title id="82A22AD2010CA4E6D4930E1CDD7DBEFE">Middle Permian (Roadian) gastropods from the Khao Khad Formation, Central Thailand: Implications for palaeogeography of the Indochina Terrane</mods:title>
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<mods:namePart id="03EE8510C17D18ADD969D29EAE611DD2">Ketwetsuriya, Chatchalerm</mods:namePart>
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<mods:namePart id="E0E20552FBFC2F40EEAF73F35448845B">Karapunar, Baran</mods:namePart>
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<mods:namePart id="0C9B0519940F7D851F4654518B08D75E">Charoentitirat, Thasinee</mods:namePart>
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<mods:namePart id="BE23203E16ADAF0AC24808A9241595E3">Nützel, Al- Exander</mods:namePart>
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<mods:date id="7084FBCDBB4B3BB1B8535A4C8238053F">2020</mods:date>
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<treatment id="03B587AB4F221568FF517F4FFE7AFCDA" ID-DOI="http://doi.org/10.5281/zenodo.3803855" ID-GBIF-Taxon="163737157" ID-Zenodo-Dep="3803855" LSID="urn:lsid:plazi:treatment:03B587AB4F221568FF517F4FFE7AFCDA" httpUri="http://treatment.plazi.org/id/03B587AB4F221568FF517F4FFE7AFCDA" lastPageId="22" lastPageNumber="23" pageId="21" pageNumber="22">
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<subSubSection id="C30665364F22156BFF517F4FFD63F8CE" box="[151,680,1841,1868]" pageId="21" pageNumber="22" type="nomenclature">
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<paragraph id="8BA336BD4F22156BFF517F4FFD63F8CE" blockId="21.[151,680,1841,1903]" box="[151,680,1841,1868]" pageId="21" pageNumber="22">
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<heading id="D0EB81D14F22156BFF517F4FFD63F8CE" bold="true" box="[151,680,1841,1868]" fontSize="10" level="2" pageId="21" pageNumber="22" reason="2">
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<taxonomicName id="4C1C4D3E4F22156BFF517F4FFD63F8CE" ID-CoL="94LBW" authority="Yochelson, 1956" authorityName="Yochelson" authorityYear="1956" box="[151,680,1841,1868]" class="Gastropoda" family="Anomphalidae" genus="Anomphalus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Trochida" pageId="21" pageNumber="22" phylum="Mollusca" rank="species" species="vanescens">
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||
<emphasis id="B968EAAF4F22156BFF517F4FFD63F8CE" bold="true" box="[151,680,1841,1868]" pageId="21" pageNumber="22">
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||
<emphasis id="B968EAAF4F22156BFF517F4FFEF9F8C9" bold="true" box="[151,306,1841,1868]" italics="true" pageId="21" pageNumber="22">Anomphalus</emphasis>
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||
cf.
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<emphasis id="B968EAAF4F22156BFE997F4CFE12F8C9" bold="true" box="[351,473,1842,1868]" italics="true" pageId="21" pageNumber="22">vanescens</emphasis>
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||
<bibRefCitation id="EF8D4B4C4F22156BFE267F4FFD63F8CE" author="Yochelson" box="[480,680,1841,1867]" firstAuthor="Yochelson" pageId="21" pageNumber="22" pagination="179 - 275" refId="ref28700" refString="Yochelson, E. L. (1956) Permian Gastropoda of the southwestern United States. 1. Euomphalacea, Trochonematacea, Anomphalacea, Craspedostomatacea, and Platyceratacea. Bulletin of the American Museum of Natural History, 110, 179 - 275." type="journal article" year="1956">Yochelson, 1956</bibRefCitation>
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</emphasis>
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</taxonomicName>
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||
</heading>
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||
</paragraph>
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||
</subSubSection>
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<subSubSection id="C30665364F22156BFF517F2BFEE6F8EA" box="[151,301,1877,1903]" pageId="21" pageNumber="22" type="description">
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<paragraph id="8BA336BD4F22156BFF517F2BFEE6F8EA" blockId="21.[151,680,1841,1903]" box="[151,301,1877,1903]" pageId="21" pageNumber="22">
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(
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<figureCitation id="13272A384F22156BFF667F2BFF25F8EA" box="[160,238,1877,1903]" captionStart="FIGURE 17" captionStartId="22.[151,250,1941,1966]" captionTargetBox="[309,1278,885,1916]" captionTargetId="figure@22.[309,1278,885,1916]" captionTargetPageId="22" captionText="FIGURE 17. A–E, Anomphalus cf. vanescens Yochelson, 1956, A–C, ESKU-19-LP 30. D–E, ESKU-19-LP 50. F–L, Anomphalus? blancus Kues & Batten, 2001, F–I, ESKU-19-LP 67. J–L, ESKU-19-LP 96. Scale bars represent: 2 mm (A–C); 1 mm (D–L)." figureDoi="http://doi.org/10.5281/zenodo.3763941" httpUri="https://zenodo.org/record/3763941/files/figure.png" pageId="21" pageNumber="22">Fig. 17</figureCitation>
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A–E)
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</paragraph>
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</subSubSection>
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<subSubSection id="C30665364F22156BFF517FE7FB57F837" box="[151,1180,1945,1970]" pageId="21" pageNumber="22" type="reference_group">
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<paragraph id="8BA336BD4F22156BFF517FE7FB57F837" blockId="21.[151,1180,1945,1970]" box="[151,1180,1945,1970]" pageId="21" pageNumber="22">
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<treatmentCitationGroup id="AB0C11934F22156BFF517FE7FB57F837" box="[151,1180,1945,1970]" pageId="21" pageNumber="22">
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cf.
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<treatmentCitation id="0ABD10AC4F22156BFF7E7FE7FD43F837" box="[184,648,1945,1970]" pageId="21" pageNumber="22">
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<taxonomicName id="4C1C4D3E4F22156BFF7E7FE7FD43F837" ID-CoL="94LBW" authority="Yochelson, 1956: 253" authorityName="Yochelson" authorityPageNumber="253" authorityYear="1956" box="[184,648,1945,1970]" class="Gastropoda" family="Anomphalidae" genus="Anomphalus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Trochida" pageId="21" pageNumber="22" phylum="Mollusca" rank="species" species="vanescens">
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<emphasis id="B968EAAF4F22156BFF7E7FE7FE6FF834" box="[184,420,1945,1969]" italics="true" pageId="21" pageNumber="22">Anomphalus vanescens</emphasis>
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<bibRefCitation id="EF8D4B4C4F22156BFE6C7FE7FD43F837" author="Yochelson" box="[426,648,1945,1970]" firstAuthor="Yochelson" pageId="21" pageNumber="22" pagination="179 - 275" refId="ref28700" refString="Yochelson, E. L. (1956) Permian Gastropoda of the southwestern United States. 1. Euomphalacea, Trochonematacea, Anomphalacea, Craspedostomatacea, and Platyceratacea. Bulletin of the American Museum of Natural History, 110, 179 - 275." type="journal article" year="1956">Yochelson, 1956: 253</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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, pl. 22, figs 19–22;
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<treatmentCitation id="0ABD10AC4F22156BFC947FE4FBD7F837" box="[850,1052,1945,1970]" pageId="21" pageNumber="22">
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||
<bibRefCitation id="EF8D4B4C4F22156BFC947FE4FBD7F837" author="Pan & Yu" box="[850,1052,1945,1970]" firstAuthor="Pan" pageId="21" pageNumber="22" refId="ref27245" refString="Pan, Y. - T. & Yu, W. Z. (1993) Permian Gastropoda of China. Ocean Press, Beijing, 68 pp." type="book" year="1993">Pan & Yu, 1993: 46</bibRefCitation>
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</treatmentCitation>
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, pl. 9, fig. 3.
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</treatmentCitationGroup>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C30665364F221568FF517FA1FDB5FF52" lastPageId="22" lastPageNumber="23" pageId="21" pageNumber="22" type="materials_examined">
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<paragraph id="8BA336BD4F22156BFF517FA1FD24F87F" blockId="21.[151,751,2015,2042]" box="[151,751,2015,2042]" pageId="21" pageNumber="22">
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<emphasis id="B968EAAF4F22156BFF517FA1FECDF87F" bold="true" box="[151,262,2015,2042]" pageId="21" pageNumber="22">Material.</emphasis>
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Three specimens: ESKU-19-
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<collectionCode id="ED0DAE784F22156BFD887F9EFDBBF87F" box="[590,624,2016,2042]" country="Russia" name="Laboratory of Palaeontology" pageId="21" pageNumber="22">LP</collectionCode>
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30, 31, 50.
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</paragraph>
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<paragraph id="8BA336BD4F211568FF0178E7FDB5FF52" blockId="22.[151,1437,153,863]" pageId="22" pageNumber="23">
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Dimensions (mm): ESKU-19-
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<collectionCode id="ED0DAE784F211568FDDE78E7FDF1FF36" box="[536,570,153,179]" country="Russia" name="Laboratory of Palaeontology" pageId="22" pageNumber="23">LP</collectionCode>
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30: height = 4.4; width = 7.7. ESKU-19-
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<collectionCode id="ED0DAE784F211568FBCD78E7FBE6FF36" box="[1035,1069,153,179]" country="Russia" name="Laboratory of Palaeontology" pageId="22" pageNumber="23">LP</collectionCode>
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31: height =
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<emphasis id="B968EAAF4F211568FB0078E4FB19FF36" box="[1222,1234,154,179]" italics="true" pageId="22" pageNumber="23">c</emphasis>
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. 4.1; width = 6.7. ESKU-19-
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<collectionCode id="ED0DAE784F211568FECB78C3FEE4FF52" box="[269,303,189,215]" country="Russia" name="Laboratory of Palaeontology" pageId="22" pageNumber="23">LP</collectionCode>
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50: height = 3.7; width = 5.5.
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</paragraph>
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</subSubSection>
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<subSubSection id="C30665364F211568FF01789EFC16FEE2" pageId="22" pageNumber="23" type="description">
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<paragraph id="8BA336BD4F211568FF01789EFC16FEE2" blockId="22.[151,1437,153,863]" pageId="22" pageNumber="23">
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<emphasis id="B968EAAF4F211568FF01789EFE92FF7E" bold="true" box="[199,345,224,251]" pageId="22" pageNumber="23">Description.</emphasis>
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Shell small, low-spired, rotelliform comprising about three whorls; whorls smooth, convex. embracing at periphery; whorls ornamented by numerous fine and dense prosocline collabral threads; upper whorl face somewhat flattened with convex periphery at mid-whorl; suture shallowly impressed, distinct; base flatly convex, narrowly phaneromphalous; umbilicus deep; aperture not certainly known.
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</paragraph>
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</subSubSection>
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<subSubSection id="C30665364F211568FF01790FFE7AFCDA" pageId="22" pageNumber="23" type="discussion">
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<paragraph id="8BA336BD4F211568FF01790FFE7AFCDA" blockId="22.[151,1437,153,863]" pageId="22" pageNumber="23">
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<emphasis id="B968EAAF4F211568FF01790FFEF0FE0E" bold="true" box="[199,315,369,395]" pageId="22" pageNumber="23">Remarks.</emphasis>
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<taxonomicName id="4C1C4D3E4F211568FE85790FFCC6FE0E" authority="Yochelson, 1956" authorityName="Yochelson" authorityYear="1956" box="[323,781,368,395]" class="Gastropoda" family="Anomphalidae" genus="Anomphalus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Trochida" pageId="22" pageNumber="23" phylum="Mollusca" rank="species" species="vanescens">
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<emphasis id="B968EAAF4F211568FE85790FFD82FE0E" box="[323,585,369,395]" italics="true" pageId="22" pageNumber="23">Anomphalus vanescens</emphasis>
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<bibRefCitation id="EF8D4B4C4F211568FD97790FFCC6FE0E" author="Yochelson" box="[593,781,368,395]" firstAuthor="Yochelson" pageId="22" pageNumber="23" pagination="179 - 275" refId="ref28700" refString="Yochelson, E. L. (1956) Permian Gastropoda of the southwestern United States. 1. Euomphalacea, Trochonematacea, Anomphalacea, Craspedostomatacea, and Platyceratacea. Bulletin of the American Museum of Natural History, 110, 179 - 275." type="journal article" year="1956">Yochelson, 1956</bibRefCitation>
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</taxonomicName>
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from the Permian of the Southwestern
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<collectingCountry id="F30B762D4F211568FB0B790FFAC8FE0E" box="[1229,1283,369,395]" name="United States of America" pageId="22" pageNumber="23">USA</collectingCountry>
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and from the Early Permian of
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<collectingCountry id="F30B762D4F211568FE9B79EBFE6BFE2A" box="[349,416,405,431]" name="China" pageId="22" pageNumber="23">China</collectingCountry>
|
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(
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<bibRefCitation id="EF8D4B4C4F211568FE6979EBFD95FE2A" author="Pan & Yu" box="[431,606,405,431]" firstAuthor="Pan" pageId="22" pageNumber="23" refId="ref27245" refString="Pan, Y. - T. & Yu, W. Z. (1993) Permian Gastropoda of China. Ocean Press, Beijing, 68 pp." type="book" year="1993">Pan & Yu 1993</bibRefCitation>
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, p. 46, pl. 9, fig. 3) is similar but has a flatter spire, whorls embrace above periphery and whorls are lower.
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<taxonomicName id="4C1C4D3E4F211568FDCD79C7FC91FE56" authority="Yochelson, 1956" authorityName="Yochelson" authorityYear="1956" box="[523,858,440,467]" class="Gastropoda" family="Anomphalidae" genus="Anomphalus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Trochida" pageId="22" pageNumber="23" phylum="Mollusca" rank="species" species="studiosus">
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<emphasis id="B968EAAF4F211568FDCD79C7FD5FFE56" box="[523,660,441,467]" italics="true" pageId="22" pageNumber="23">A. studiosus</emphasis>
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<bibRefCitation id="EF8D4B4C4F211568FD5B79C7FC91FE56" author="Yochelson" box="[669,858,440,467]" firstAuthor="Yochelson" pageId="22" pageNumber="23" pagination="179 - 275" refId="ref28700" refString="Yochelson, E. L. (1956) Permian Gastropoda of the southwestern United States. 1. Euomphalacea, Trochonematacea, Anomphalacea, Craspedostomatacea, and Platyceratacea. Bulletin of the American Museum of Natural History, 110, 179 - 275." type="journal article" year="1956">Yochelson, 1956</bibRefCitation>
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</taxonomicName>
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(p. 254, pl. 22, figs 25–28) is also similar in shell shape as well as the rate of whorl expansion but has smaller size, a narrower umbilicus has a flatter spire and its whorls embrace above periphery.
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<taxonomicName id="4C1C4D3E4F211568FDDE7A7FFC47FD9E" authority="Pan & Erwin, 2002" authorityName="Pan & Erwin" authorityYear="2002" box="[536,908,513,539]" class="Gastropoda" family="Anomphalidae" genus="Anomphalus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Trochida" pageId="22" pageNumber="23" phylum="Mollusca" rank="species" species="fusuiensis">
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<emphasis id="B968EAAF4F211568FDDE7A7FFD6DFD9E" box="[536,678,513,539]" italics="true" pageId="22" pageNumber="23">A. fusuiensis</emphasis>
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<bibRefCitation id="EF8D4B4C4F211568FD697A7FFC47FD9E" author="Pan & Erwin" box="[687,908,513,539]" firstAuthor="Pan" pageId="22" pageNumber="23" pagination="1 - 49" refId="ref27138" refString="Pan, H. - Z. & Erwin, D. H. (2002) Gastropods from the Permian of Guangxi and Yunnan provinces, South China. Journal of Paleontology, 78 (Supplement 56), 1 - 49. https: // doi. org / 10.1666 / 0022 - 3360 (2002) 76 [1: GFTPOG] 2.0. CO; 2" type="journal article" year="2002">Pan & Erwin, 2002</bibRefCitation>
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</taxonomicName>
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from the Late Permian of South
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<collectingCountry id="F30B762D4F211568FACE7A7FFA80FD9E" box="[1288,1355,513,539]" name="China" pageId="22" pageNumber="23">China</collectingCountry>
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is also similar but has lower, more rounded whorls, a wider umbilicus and a deeper suture. The Middle Permian species
|
||
<taxonomicName id="4C1C4D3E4F211568FF517A37FD78FDE6" authority="Nutzel & Ketwetsuriya, 2016" authorityName="Nutzel & Ketwetsuriya" authorityYear="2016" box="[151,691,585,611]" class="Gastropoda" family="Anomphalidae" genus="Anomphalus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Trochida" pageId="22" pageNumber="23" phylum="Mollusca" rank="species" species="lateumbilicatus">
|
||
<emphasis id="B968EAAF4F211568FF517A37FEA8FDE6" box="[151,355,585,611]" italics="true" pageId="22" pageNumber="23">A. lateumbilicatus</emphasis>
|
||
Nützel &
|
||
<bibRefCitation id="EF8D4B4C4F211568FE1E7A37FD78FDE6" author="Ketwetsuriya" box="[472,691,585,611]" firstAuthor="Ketwetsuriya" pageId="22" pageNumber="23" pagination="481 - 513" refId="ref24896" refString="Ketwetsuriya, C., Nu ¨ tzel, A. & Kanjanapayont, P. (2016) Permian gastropods from the Tak Fa Limestone, Nakhon Sawan, Northern Thailand. Bulletin of Geosciences, 91, 481 - 513. https: // doi. org / 10.3140 / bull. geosci. 1617" type="journal article" year="2016">Ketwetsuriya, 2016</bibRefCitation>
|
||
</taxonomicName>
|
||
from the
|
||
<collectingRegion id="49D8F85F4F211568FCE67A37FC81FDE6" box="[800,842,585,611]" country="Thailand" name="Tak" pageId="22" pageNumber="23">Tak</collectingRegion>
|
||
Fa Limestone of
|
||
<collectingCountry id="F30B762D4F211568FBC87A37FBB9FDE6" box="[1038,1138,585,611]" name="Thailand" pageId="22" pageNumber="23">Thailand</collectingCountry>
|
||
has a very wide umbilicus and consists of more whorls compared to the
|
||
<taxonomicName id="4C1C4D3E4F211568FD487A13FC8AFD02" authorityName="Yochelson" authorityYear="1956" box="[654,833,621,647]" class="Gastropoda" family="Anomphalidae" genus="Anomphalus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Trochida" pageId="22" pageNumber="23" phylum="Mollusca" rank="species" species="vanescens">
|
||
<emphasis id="B968EAAF4F211568FD487A13FD6DFD02" box="[654,678,621,647]" italics="true" pageId="22" pageNumber="23">A.</emphasis>
|
||
cf.
|
||
<emphasis id="B968EAAF4F211568FD097A10FC8AFD02" box="[719,833,622,647]" italics="true" pageId="22" pageNumber="23">vanescens</emphasis>
|
||
</taxonomicName>
|
||
at comparable size.
|
||
<taxonomicName id="4C1C4D3E4F211568FBE57A13FB64FD02" authorityName="Meek & Worthen" authorityYear="1866" box="[1059,1199,621,647]" class="Gastropoda" family="Anomphalidae" genus="Anomphalus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Trochida" pageId="22" pageNumber="23" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B968EAAF4F211568FBE57A13FB64FD02" box="[1059,1199,621,647]" italics="true" pageId="22" pageNumber="23">Anomphalus</emphasis>
|
||
</taxonomicName>
|
||
sp. which is reported from the
|
||
<collectingRegion id="49D8F85F4F211568FEC67AEFFEE1FD2E" box="[256,298,657,683]" country="Thailand" name="Tak" pageId="22" pageNumber="23">Tak</collectingRegion>
|
||
Fa Limestone is also similar but has a wider umbilicus and a flatter spire.
|
||
<taxonomicName id="4C1C4D3E4F211568FBB97AEFFAC0FD2E" authorityName="Meek & Worthen" authorityYear="1866" box="[1151,1291,657,683]" class="Gastropoda" family="Anomphalidae" genus="Anomphalus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Trochida" pageId="22" pageNumber="23" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B968EAAF4F211568FBB97AEFFAC0FD2E" box="[1151,1291,657,683]" italics="true" pageId="22" pageNumber="23">Anomphalus</emphasis>
|
||
</taxonomicName>
|
||
sp. from the Permian of
|
||
<collectingCountry id="F30B762D4F211568FED87ACBFE4EFD4A" box="[286,389,693,719]" name="Malaysia" pageId="22" pageNumber="23">Malaysia</collectingCountry>
|
||
as reported by
|
||
<bibRefCitation id="EF8D4B4C4F211568FDFF7ACBFD04FD4A" author="Batten" box="[569,719,693,719]" firstAuthor="Batten" pageId="22" pageNumber="23" pagination="1 - 26" refId="ref23086" refString="Batten, R. L. (1979) Permian gastropods from Perak, Malaysia. Part 2. The trochids, patellids, and neritids. American Museum Novitates, 2685, 1 - 26." type="journal article" year="1979">Batten (1979</bibRefCitation>
|
||
, p. 8, figs 10–11) is higher spired. The present specimens also resemble
|
||
<taxonomicName id="4C1C4D3E4F211568FEC27AA7FCFEFD76" authority="Nutzel in Nutzel & Nakazawa, 2012" authorityName="Nutzel in Nutzel & Nakazawa" authorityYear="2012" box="[260,821,729,755]" class="Gastropoda" family="Anomphalidae" genus="Anomphalus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Trochida" pageId="22" pageNumber="23" phylum="Mollusca" rank="species" species="japonicus">
|
||
<emphasis id="B968EAAF4F211568FEC27AA7FE5AFD76" box="[260,401,729,755]" italics="true" pageId="22" pageNumber="23">A. japonicus</emphasis>
|
||
Nützel
|
||
<emphasis id="B968EAAF4F211568FE2A7AA4FDC9FD76" box="[492,514,730,755]" italics="true" pageId="22" pageNumber="23">in</emphasis>
|
||
Nützel &
|
||
<bibRefCitation id="EF8D4B4C4F211568FDBC7AA7FCFEFD76" author="Nakazawa" box="[634,821,729,755]" firstAuthor="Nakazawa" pageId="22" pageNumber="23" pagination="103 - 169" refId="ref27024" refString="Nu ¨ tzel, A. & Nakazawa, K. (2012) Permian (Capitanian) gastropods from the Akasaka Limestone (Gifu Prefecture, Japan). Journal of Systematic Palaeontology, 10, 103 - 169. https: // doi. org / 10.1080 / 14772019.2010.549659" type="journal article" year="2012">Nakazawa, 2012</bibRefCitation>
|
||
</taxonomicName>
|
||
from the Middle Permian of
|
||
<collectingCountry id="F30B762D4F211568FB457AA7FB08FD76" box="[1155,1219,729,755]" name="Japan" pageId="22" pageNumber="23">Japan</collectingCountry>
|
||
but the latter has a wider umbilicus and a small umbilical ridge. The present species is probably undescribed but the preservation of the material is too poor for a safe species assignment, especially the aperture and the collabral threads is hardly visible in the studied specimens.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |