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<document id="C5AAA640B10CD6120131C17F8D8C5B43" ID-CLB-Dataset="304964" ID-DOI="10.5852/ejt.2024.967.2699" ID-GBIF-Dataset="0647b761-1496-4dcd-abb6-495b67abe85f" ID-ISSN="2118-9773" ID-Zenodo-Dep="13990819" ID-ZooBank="C08B8027-50CC-417E-BCD4-5183B9FF6738" IM.bibliography_approvedBy="guilherme" IM.illustrations_approvedBy="guilherme" IM.materialsCitations_approvedBy="guilherme" IM.metadata_approvedBy="guilherme" IM.tables_approvedBy="guilherme" IM.taxonomicNames_approvedBy="guilherme" IM.treatmentCitations_approvedBy="guilherme" IM.treatments_approvedBy="guilherme" checkinTime="1729849508920" checkinUser="plazi" docAuthor="Zonstein, Sergei L." docDate="2024" docId="03A8B7235523FFB0FDBAE03CFB3CCA11" docLanguage="en" docName="EJT.2024.967.1-185.pdf" docOrigin="European Journal of Taxonomy 967" docSource="https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2699/12459" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Raveniola Zonstein 1987" docType="treatment" docVersion="7" lastPageNumber="11" masterDocId="FF91CF5B552AFFBAFFEFE72AFFB2CE5B" masterDocTitle="A revision of the spider genus Raveniola (Araneae, Nemesiidae). II. Species from Central Asia" masterLastPageNumber="185" masterPageNumber="1" pageNumber="10" updateTime="1730721133903" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="B4472FEE9ED6FD70AD18F547BCDCAF93">A revision of the spider genus Raveniola (Araneae, Nemesiidae). II. Species from Central Asia</mods:title>
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<mods:namePart id="E43A36D5E63B963C6F7EF98AE68B01AA">Zonstein, Sergei L.</mods:namePart>
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<mods:affiliation id="253DDA7CF44256262C7C1B51240DC29C">Steinhardt Museum of Natural History, Tel-Aviv University, 69978 Tel-Aviv, Israel</mods:affiliation>
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<mods:date id="32852E65AB6C72F76AE3E670427F90A7">2024</mods:date>
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<subSubSection id="C31B55BE5523FFB3FDBAE03CFC6CC96A" box="[597,990,1814,1841]" pageId="9" pageNumber="10" type="nomenclature">
<paragraph id="8BBE06355523FFB3FDBAE03CFC6CC96A" blockId="9.[597,990,1814,1841]" box="[597,990,1814,1841]" pageId="9" pageNumber="10">
<heading id="D0F6B1595523FFB3FDBAE03CFC6CC96A" box="[597,990,1814,1841]" centered="true" fontSize="11" level="2" pageId="9" pageNumber="10" reason="2">
<emphasis id="B975DA275523FFB3FDBAE03CFC85C96A" bold="true" box="[597,823,1814,1841]" italics="true" pageId="9" pageNumber="10">
<taxonomicName id="4C017DB65523FFB3FDBAE03CFD7DC96A" ID-CoL="77BJ" authority="Zonstein, 1987" authorityName="Zonstein" authorityYear="1987" box="[597,719,1814,1841]" class="Arachnida" family="Nemesiidae" genus="Raveniola" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">Raveniola</taxonomicName>
caudata
</emphasis>
species group
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C31B55BE5523FFB0FF52E072FB02CF36" lastPageId="10" lastPageNumber="11" pageId="9" pageNumber="10" type="diagnosis">
<paragraph id="8BBE06355523FFB3FF52E072FE7EC928" blockId="9.[189,460,1880,1907]" box="[189,460,1880,1907]" pageId="9" pageNumber="10">
<heading id="D0F6B1595523FFB3FF52E072FE7EC928" bold="true" box="[189,460,1880,1907]" fontSize="11" level="3" pageId="9" pageNumber="10" reason="3">
<emphasis id="B975DA275523FFB3FF52E072FE7EC928" bold="true" box="[189,460,1880,1907]" pageId="9" pageNumber="10">Diagnostic characters</emphasis>
</heading>
</paragraph>
<paragraph id="8BBE06355523FFB0FF52E0ADFB02CF36" blockId="9.[189,1399,1927,2025]" lastBlockId="10.[189,1399,265,365]" lastPageId="10" lastPageNumber="11" pageId="9" pageNumber="10">
Maxilla with numerous cuspules arranged in a triangular area (
<figureCitation id="133A1AB05523FFB3FC41E0ADFBE5C9FA" box="[942,1111,1927,1953]" captionStart="Figs 202–210" captionStartId="129.[189,241,1851,1877]" captionTargetBox="[189,1398,265,1816]" captionTargetId="figure-88@129.[189,1398,265,1816]" captionTargetPageId="129" captionText="Figs 202–210. Raveniola spp., ♂♂, holotypes (except 204), sternum, labium and maxillae, ventral views. 202. R. caudata Zonstein, 2009 (SMNH). 203. R. inopinata sp. nov. (SMNH). 204. R. redikorzevi (Spassky, 1937), ♂ from Akar-Cheshme (SMNH). 205. R. alajensis sp. nov. (SMNH). 206. R. concolor Zonstein, 2000 (MCSN AR 18). 207. R. cucullata sp. nov. (SMNH). 208. R. dolosa sp. nov. (SMNH). 209. R. hirta sp. nov. (SMNH). 210. R. ignobilis sp. nov. (SMNH). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13990878" httpUri="https://zenodo.org/record/13990878/files/figure.png" pageId="9" pageNumber="10">Figs 202–204</figureCitation>
). PMS medium-sized to large. Apical segment of PLS elongate and considerably longer than in other
<taxonomicName id="4C017DB65523FFB3FBACE080FB43C99E" box="[1091,1265,1962,1989]" class="Arachnida" family="Nemesiidae" genus="Raveniola" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B975DA275523FFB3FBACE080FB08C99F" box="[1091,1210,1962,1988]" italics="true" pageId="9" pageNumber="10">Raveniola</emphasis>
spp.
</taxonomicName>
(
<figureCitation id="133A1AB05523FFB3FAEFE080FF5BC9B2" captionStart="Figs 555–564" captionStartId="160.[189,241,1786,1812]" captionTargetBox="[213,1375,265,1755]" captionTargetId="figure-109@160.[213,1375,265,1755]" captionText="Figs 555–564. Raveniola spp., ♂♂ (555–557, 560, 563) and ♀♀ (558–559, 561–562, 564), holotypes (except 557, 558, 561, 562, 564). Spinnerets, ventral (555–558, 560–561, 563–564) and lateral (559, 562) views. 555. R. caudata Zonstein, 2009, holotype (SMNH). 556. R. inopinata sp. nov. (SMNH). 557–558. R. redikorzevi (Spassky, 1937), ♂ from Akar-Cheshme and ♀ from Kepele, respectively (both SMNH). 559. R. afghana sp. nov. (NHMD). 560. R. alajensis sp. nov. (SMNH). 561–562. R. alajensis, paratype (SMNH). 563. R. cucullata sp. nov. (SMNH). 564. R. cucullata, paratype from Shikildara (SMNH). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.14018401" httpUri="https://zenodo.org/record/14018401/files/figure.png" pageId="9" pageNumber="10">Figs 555– 558</figureCitation>
cf.
<figureCitation id="133A1AB05523FFB3FEFCE0E4FE0BC9B2" box="[275,441,1998,2025]" captionStart-0="Figs 555–564" captionStart-1="Figs 565–574" captionStart-2="Figs 575–583" captionStart-3="Figs 584–592" captionStart-4="Figs 593–601" captionStart-5="Figs 602–610" captionStart-6="Figs 611–618" captionStartId-0="160.[189,241,1786,1812]" captionStartId-1="161.[189,241,1825,1851]" captionStartId-2="162.[189,241,1857,1883]" captionStartId-3="163.[189,241,1821,1847]" captionStartId-4="164.[189,241,1759,1785]" captionStartId-5="165.[189,241,1763,1789]" captionStartId-6="166.[189,241,1796,1822]" captionTargetBox-0="[213,1375,265,1755]" captionTargetBox-1="[202,1386,265,1765]" captionTargetBox-2="[189,1398,265,1816]" captionTargetBox-3="[189,1398,265,1789]" captionTargetBox-4="[189,1399,265,1718]" captionTargetBox-5="[189,1399,265,1718]" captionTargetBox-6="[189,1398,265,1758]" captionTargetId-0="figure-109@160.[213,1375,265,1755]" captionTargetId-1="figure-100@161.[202,1386,265,1784]" captionTargetId-2="figure-74@162.[189,1398,265,1816]" captionTargetId-3="figure-92@163.[189,1398,265,1789]" captionTargetId-4="figure-74@164.[189,1399,265,1718]" captionTargetId-5="figure-61@165.[189,1399,265,1718]" captionTargetId-6="figure-90@166.[189,1398,265,1759]" captionTargetPageId-1="161" captionTargetPageId-2="162" captionTargetPageId-3="163" captionTargetPageId-4="164" captionTargetPageId-5="165" captionTargetPageId-6="166" captionText-0="Figs 555–564. Raveniola spp., ♂♂ (555–557, 560, 563) and ♀♀ (558–559, 561–562, 564), holotypes (except 557, 558, 561, 562, 564). Spinnerets, ventral (555–558, 560–561, 563–564) and lateral (559, 562) views. 555. R. caudata Zonstein, 2009, holotype (SMNH). 556. R. inopinata sp. nov. (SMNH). 557–558. R. redikorzevi (Spassky, 1937), ♂ from Akar-Cheshme and ♀ from Kepele, respectively (both SMNH). 559. R. afghana sp. nov. (NHMD). 560. R. alajensis sp. nov. (SMNH). 561–562. R. alajensis, paratype (SMNH). 563. R. cucullata sp. nov. (SMNH). 564. R. cucullata, paratype from Shikildara (SMNH). Scale bars: 1.0 mm." captionText-1="Figs 565–574. Raveniola spp., ♂♂ (565, 568, 571, 573) and ♀♀ (566–567, 569–570, 572), paratypes (except 568, 571, 573). Spinnerets, ventral (565–566, 571–574), lateral (567–568) and ventrolateral (569–570) views. 565–566. R. cucullata sp. nov., paratypes from Shahob (SMNH). 567. R. cucullata, from Sangdara (SMNH). 568. R. dolosa sp. nov. (SMNH). 569–570. R. dolosa, paratypes (SMNH). 571. R. hirta sp. nov., holotype (SMNH). 572. R. hirta, paratype (SMNH). 573. R. ignobilis sp. nov. (SMNH). 574. R. ignobilis, paratype (SMNH). Scale bars: 1.0 mm." captionText-2="Figs 575–583. Raveniola spp., ♂♂ (575, 577, 581) and ♀♀ (576, 578–580, 582–583); holotypes (575– 577, 581) and paratypes (578–580, 582–583). Spinnerets (575–579, 581–582) and entire abdomen (580, 583), ventral (575–576, 579–580, 582–583) and lateral (577–578, 581) views. 575. R. insolita sp. nov. (ZISP). 576. R. karategensis sp. nov. (SMNH). 577–580. R. ornata sp. nov. (SMNH). 581– 583. R. ornatula sp. nov. (SMNH). Scale bars: 1.0 mm." captionText-3="Figs 584–592. Raveniola spp., ♂♂ (584, 589, 591) and ♀♀ (585–588, 590, 592), holotypes (except 585–586, 590, 592). Spinnerets, ventral (584–585, 587, 589, 591–592) and lateral (586, 588, 590) views. 584. R. pamira sp. nov. (SMNH). 585–586. R. pamira, paratype (SMNH). 587–588. R. sororcula sp. nov. (SMNH). 589. R. diluta sp. nov. (SMNH). 590. R. diluta, paratype (SMNH). 591. R. fedotovi (Charitonov, 1946), paralectotype (PSU). 592. R. fedotovi, ♀ from Hojakurgan (SMNH). Scale bars: 1.0 mm." captionText-4="Figs 593–601. Raveniola spp., ♂♂, holotypes (593, 595, 597–598, 601) and ♀♀ paratypes (594, 596, 599–600). Spinnerets, lateral (593,596–597), ventral (595,598–599, 601) and posterior (594, 600) views. 593–594. R. pallens sp. nov. (SMNH). 595–596. R. zyuzini sp. nov. (SMNH). 597–600. R. ferghanensis (Zonstein, 1984) (ZISP). 601. R. kirgizica sp. nov. (SMNH). Strictly reduced PMS indicated by white arrows. Scale bars: 1.0 mm." captionText-5="Figs 602–610. Raveniola spp., ♂♂, holotypes (602, 604–606, 609) and ♀♀ paratypes (603, 607–608, 610). Spinnerets, ventral (602–604, 606–610) and lateral (605) views. 602–603. R. kirgizica sp. nov. (SMNH). 604. R. kopetdaghensis (Fet, 1984) (ZISP). 605–608. R. mikhailovi Zonstein, 2021 (SMNH). 609–610. R. nenilini sp. nov. (SMNH). Scale bars: 1.0 mm." captionText-6="Figs 611–618. Raveniola spp., ♀♀ (611, 613, 615, 617–618) and ♂♂ (612, 614, 616). Spinnerets, ventral (611–617) and subcaudal (618) views. 611. R. nenilini sp. nov., paratype from Mazarsai (SMNH). 612. R. ovchinnikovi sp. nov., holotype (SMNH). 613. R. tarabaevi sp. nov., holotype (SMNH). 614– 615. R. virgata (Fet, 1984), from vicinity of Arslanbob (SMNH). 616. R. vulpina sp. nov., holotype (SMNH). 617–618. R. vulpina, paratype from Tumanyak (SMNH). Strictly reduced PMS indicated by white arrows. Scale bars: 1.0 mm." figureDoi-0="http://doi.org/10.5281/zenodo.14018401" figureDoi-1="http://doi.org/10.5281/zenodo.14018403" figureDoi-2="http://doi.org/10.5281/zenodo.14018406" figureDoi-3="http://doi.org/10.5281/zenodo.14018408" figureDoi-4="http://doi.org/10.5281/zenodo.14018411" figureDoi-5="http://doi.org/10.5281/zenodo.14018413" figureDoi-6="http://doi.org/10.5281/zenodo.14018415" httpUri-0="https://zenodo.org/record/14018401/files/figure.png" httpUri-1="https://zenodo.org/record/14018403/files/figure.png" httpUri-2="https://zenodo.org/record/14018406/files/figure.png" httpUri-3="https://zenodo.org/record/14018408/files/figure.png" httpUri-4="https://zenodo.org/record/14018411/files/figure.png" httpUri-5="https://zenodo.org/record/14018413/files/figure.png" httpUri-6="https://zenodo.org/record/14018415/files/figure.png" pageId="9" pageNumber="10">Figs 559–618</figureCitation>
). Males: tibiae and metatarsi I–II without modified hairs (
<figureCitation id="133A1AB05523FFB3FBBDE0E4FA9CC9B3" box="[1106,1326,1998,2025]" captionStart="Figs 256–264" captionStartId="135.[189,241,1824,1850]" captionTargetBox="[200,1388,265,1788]" captionTargetId="figure-97@135.[200,1388,265,1788]" captionTargetPageId="135" captionText="Figs 256–264. Raveniola spp., ♂♂; holotypes (except 260, 264), tibia and metatarsus I (256, 258, 260– 264) and tarsus IV (257, 259); retrolateral views. 256–257. R. caudata Zonstein, 2009 (SMNH). 258– 259. R. inopinatasp. nov. (SMNH). 260. R. redikorzevi(Spassky, 1937), ♂ fromAkar-Cheshme (SMNH). 261. R. alajensis sp. nov. (SMNH) 262. R. concolor Zonstein, 2000 (MCSN AR 18). 263. R. cucullata sp. nov. (SMNH). 264. R. cucullata, paratype from Shahob (SMNH). Modified hairs are indicated with arrows. Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13990893" httpUri="https://zenodo.org/record/13990893/files/figure.png" pageId="9" pageNumber="10">Figs 256, 258, 260</figureCitation>
,
<figureCitation id="133A1AB05523FFB3FAD6E0E4FAD8C9B3" box="[1337,1386,1998,2024]" captionStart="Figs 290–309" captionStartId="139.[189,241,1678,1704]" captionTargetBox="[189,1398,265,1651]" captionTargetId="figure-180@139.[189,1398,265,1651]" captionTargetPageId="139" captionText="Figs 290–309. Raveniola spp., ♂♂, holotypes (except 290, 294, 300, 302, 308), metatarsus I, ventral views. 290. R. redikorzevi (Spassky, 1937), ♂ from Akar-Cheshme (SMNH). 291. R. alajensis sp. nov. (SMNH). 292. R. concolor Zonstein, 2000 (MCSN AR 18). 293. R. cucullata sp. nov. (SMNH). 294. R. cucullata, paratype from Shahob (SMNH).295. R. dolosa sp. nov.(SMNH). 296. R. hirtasp. nov. (SMNH). 297. R. ignobilis sp. nov. (SMNH). 298. R. ornata sp. nov. (SMNH). 299. R. ornatula sp. nov. (SMNH). 300. R. ornatula, paratype (SMNH).301. R. pamirasp. nov. (SMNH). 302. R. pamira, paratype (SMNH). 303. R. diluta sp. nov. (SMNH). 304. R. kirgizica sp. nov. (SMNH). 305. R. kopetdaghensis (Fet, 1984) (ZISP). 306. R. mikhailovi Zonstein, 2021 (SMNH). 307. R. nenilini sp. nov. (SMNH). 308. R. virgata (Simon, 1891), ♂ from vicinity of Arslanbob (SMNH). 309. R. vulpina sp. nov. (SMNH). Modified hairs are indicated with arrows. Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13990903" httpUri="https://zenodo.org/record/13990903/files/figure.png" pageId="9" pageNumber="10">290</figureCitation>
); cymbium moderately short (as in
<figureCitation id="133A1AB05520FFB0FDA4E623FD43CF7F" box="[587,753,265,292]" captionStart="Figs 349–363" captionStartId="143.[189,241,1727,1753]" captionTargetBox="[189,1390,270,1690]" captionTargetId="figure-139@143.[189,1398,265,1690]" captionTargetPageId="143" captionText="Figs 349–363. Raveniola spp., ♂♂, holotypes (except 351, 355, 363), palpal tibia, cymbium and copulatory bulb, retrolateral (349–358, 360–361, 363) and retroventral (359, 362) views.349. R. caudata Zonstein, 2009 (SMNH). 350. R. inopinata sp. nov. (SMNH). 351. R. redikorzevi (Spassky, 1937), ♂ from Akar-Cheshme (SMNH). 352. R. alajensis sp. nov. (SMNH). 353. R. concolor Zonstein, 2000 (MCSN AR 18). 354. R. cucullata sp. nov. (SMNH). 355. R. cucullata, paratype from Shahob (SMNH). 356. R. dolosa sp. nov. (SMNH). 357. R. hirta sp. nov. (SMNH). 358–359. R. ignobilis sp. nov. (SMNH). 360. R. insolita sp. nov. (ZISP). 361. R. ornata sp. nov. (SMNH). 362. R. ornatula sp. nov. (SMNH). 363. R. pamira sp. nov., paratype (SMNH). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13990913" httpUri="https://zenodo.org/record/13990913/files/figure.png" pageId="10" pageNumber="11">Figs 349–351</figureCitation>
); embolus broadly tipped, always with well-developed subapical keel (
<figureCitation id="133A1AB05520FFB0FE95E604FD93CF13" box="[378,545,302,328]" captionStart="Figs 379–388" captionStartId="145.[189,241,1844,1870]" captionTargetBox="[189,1398,265,1806]" captionTargetId="figure-75@145.[189,1398,265,1806]" captionTargetPageId="145" captionText="Figs 379–388. Raveniola spp., ♂♂, holotypes, details of copulatoty bulb, retrolateral (379, 381, 383, 385, 388), ventral (380, 382, 384, 387) and retroventral (386) views. 379–380. R. caudata Zonstein, 2009 (SMNH). 381–382. R. inopinata sp. nov. (SMNH). 383–384. R. redikorzevi (Spassky, 1937), ♂ from Akar-Cheshme (SMNH).385–387. R. alajensis sp. nov. (SMNH). 388. R. concolor Zonstein, 2000 (MCSN AR 18). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13990917" httpUri="https://zenodo.org/record/13990917/files/figure.png" pageId="10" pageNumber="11">Figs 379–384</figureCitation>
,
<figureCitation id="133A1AB05520FFB0FDDFE604FD2ECF13" box="[560,668,302,328]" captionStart="Figs 466–474" captionStartId="154.[189,241,1854,1880]" captionTargetBox="[189,1398,265,1817]" captionTargetId="figure-57@154.[189,1398,265,1817]" captionTargetPageId="154" captionText="Figs 466–474. Raveniola spp., ♂♂, details of copulatoty bulb (466–467, 469, 471, 473) and tip of embolus (468, 470, 472, 474), ventral (466, 473–474), retrolateral (467–468) and retoventral (469–472) views. 466–468. R. redikorzevi (Spassky, 1937) (SMNH). 469–470. R. cucullata sp. nov. (SMNH). 471–474. R. diluta sp. nov. (SMNH)." figureDoi="http://doi.org/10.5281/zenodo.13990937" httpUri="https://zenodo.org/record/13990937/files/figure.png" pageId="10" pageNumber="11">466–468</figureCitation>
). Females: spermathecae (known only for one species of the group) very wide, each with two short, thin, and closely spaced stalks (
<figureCitation id="133A1AB05520FFB0FC12E679FB16CF36" box="[1021,1188,339,365]" captionStart="Figs 487–503" captionStartId="156.[189,241,1695,1721]" captionTargetBox="[189,1398,265,1653]" captionTargetId="figure-102@156.[189,1398,265,1653]" captionTargetPageId="156" captionText="Figs 487–503. Raveniola spp., ♀♀, paratypes (except 487–489), spermathecae, dorsal views. 487–488. R. redikorzevi (Spassky, 1937), Kepele (SMNH). 489. R. afghana sp. nov. (NHMD). 490. Same, holotype. 491–493. R. alajensis sp. nov., Berksu. 494. R. concolor Zonstein, 2000 (MCSN AR 19). 495–496. R. cucullata sp. nov., Shikildara (SMNH). 497. R. cucullata, Shahob (SMNH). 498– 499. R. cucullata, Sangdara (SMNH). 500–503. R. dolosa sp. nov. (SMNH). Abbreviations: isb = inner spermathecal branch; osb = outer spermathecal branch; sb = spermathecal base. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13990941" httpUri="https://zenodo.org/record/13990941/files/figure.png" pageId="10" pageNumber="11">Figs 487–488</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C31B55BE5520FFB0FF52E6BFFAB2CFBB" pageId="10" pageNumber="11" type="taxon_list">
<paragraph id="8BBE06355520FFB0FF52E6BFFE3ECFF4" blockId="10.[189,396,405,431]" box="[189,396,405,431]" pageId="10" pageNumber="11">
<heading id="D0F6B1595520FFB0FF52E6BFFE3ECFF4" bold="true" box="[189,396,405,431]" fontSize="11" level="3" pageId="10" pageNumber="11" reason="3">
<emphasis id="B975DA275520FFB0FF52E6BFFE3ECFF4" bold="true" box="[189,396,405,431]" pageId="10" pageNumber="11">Species included</emphasis>
</heading>
</paragraph>
<paragraph id="8BBE06355520FFB0FF52E6ECFAB2CFBB" blockId="10.[189,1280,454,480]" box="[189,1280,454,480]" pageId="10" pageNumber="11">
<taxonomicName id="4C017DB65520FFB0FF52E6ECFDE6CFBB" ID-CoL="4RMTM" authority="Zonstein, 2009" authorityName="Zonstein" authorityYear="2009" box="[189,596,454,480]" class="Arachnida" family="Nemesiidae" genus="Raveniola" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="caudata">
<emphasis id="B975DA275520FFB0FF52E6ECFE28CFBB" box="[189,410,454,480]" italics="true" pageId="10" pageNumber="11">Raveniola caudata</emphasis>
<bibRefCitation id="EF907BC45520FFB0FE4EE6ECFDE6CFBB" author="Zonstein" box="[417,596,454,480]" firstAuthor="Zonstein" pageId="10" pageNumber="11" pagination="37 - 41" refId="ref83362" refString="Zonstein S. 2009. A review of the mygalomorph spiders of the Raveniola caudata - species group (Araneae, Nemesiidae). Zootaxa 2217 (1): 37 - 41. https: // doi. org / 10.11646 / zootaxa. 2217.1.2" type="journal article" year="2009">Zonstein, 2009</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C017DB65520FFB0FD8FE6EDFD42CFBB" authorityName="Zonstein" authorityYear="2024" box="[608,752,455,480]" class="Arachnida" family="Nemesiidae" genus="Raveniola" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="inopinata" status="sp. nov.">
<emphasis id="B975DA275520FFB0FD8FE6EDFD42CFBB" box="[608,752,455,480]" italics="true" pageId="10" pageNumber="11">R. inopinata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A246675C5520FFB0FD17E6ECFCE7CFBB" box="[760,853,454,480]" pageId="10" pageNumber="11" rank="species">sp. nov.</taxonomicNameLabel>
, and
<taxonomicName id="4C017DB65520FFB0FC7CE6EDFB4ECFBB" ID-CoL="6X599" authority="(Spassky, 1937)" authorityName="SMNH" baseAuthorityName="Spassky" baseAuthorityYear="1937" box="[915,1276,454,480]" class="Arachnida" family="Nemesiidae" genus="Raveniola" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="redikorzevi">
<emphasis id="B975DA275520FFB0FC7CE6EDFB84CFBB" box="[915,1078,454,480]" italics="true" pageId="10" pageNumber="11">R. redikorzevi</emphasis>
(
<bibRefCitation id="EF907BC45520FFB0FBAAE6ECFB41CFBB" author="Spassky" box="[1093,1267,454,480]" firstAuthor="Spassky" pageId="10" pageNumber="11" pagination="361 - 368" refId="ref82512" refString="Spassky S. A. 1937. Araneae palaearcticae novae. Mygalomorphae. I. Festschrift Embrik Strand 3: 361 - 368." type="journal article" year="1937">Spassky, 1937</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C31B55BE5520FFB0FF52E523FEE3CA7E" pageId="10" pageNumber="11" type="key">
<paragraph id="8BBE06355520FFB0FF52E523FD10CC78" blockId="10.[189,674,520,547]" box="[189,674,520,547]" pageId="10" pageNumber="11">
<heading id="D0F6B1595520FFB0FF52E523FD10CC78" bold="true" box="[189,674,520,547]" fontSize="11" level="3" pageId="10" pageNumber="11" reason="3">
<emphasis id="B975DA275520FFB0FF52E523FD10CC78" bold="true" box="[189,674,520,547]" pageId="10" pageNumber="11">
Key to the species of the
<taxonomicName id="4C017DB65520FFB0FE1EE522FDE3CC78" ID-CoL="4RMTM" authorityName="Zonstein" authorityYear="2009" box="[497,593,520,547]" class="Arachnida" family="Nemesiidae" genus="Raveniola" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="caudata">
<emphasis id="B975DA275520FFB0FE1EE522FDE3CC78" bold="true" box="[497,593,520,547]" italics="true" pageId="10" pageNumber="11">caudata</emphasis>
</taxonomicName>
group
</emphasis>
</heading>
</paragraph>
<paragraph id="8BBE06355520FFB0FF05E513FE87CC08" blockId="10.[189,1401,569,817]" box="[234,309,569,595]" pageId="10" pageNumber="11">
<emphasis id="B975DA275520FFB0FF05E513FE87CC08" bold="true" box="[234,309,569,595]" pageId="10" pageNumber="11">Males</emphasis>
</paragraph>
<key id="FF9051045520FFB0FF52E574FAC5CDB9" pageId="10" pageNumber="11">
<keyStep id="30F51E905520FFB0FF52E574FAC5CD6A" pageId="10" pageNumber="11">
<paragraph id="8BBE06355520FFB0FF52E574FAC5CC99" blockId="10.[189,1401,569,817]" pageId="10" pageNumber="11">
<keyLead id="30F0A5005520FFB0FF52E574FAC5CC99" pageId="10" pageNumber="11">
1. Carapace and legs ginger brown or brownish orange, abdomen paler, light yellowish brown, with or without dorsal pattern (
<figureCitation id="133A1AB05520FFB0FDF3E5A9FD36CCC6" box="[540,644,643,669]" captionStart="Figs 1–9" captionStartId="107.[189,241,1848,1874]" captionTargetBox="[189,1398,268,1805]" captionTargetId="figure-84@107.[189,1398,265,1805]" captionTargetPageId="107" captionText="Figs 1–9.Raveniola spp., ♂♂, holotypes (except 3, 6), habitus, dorsal views. 1. R. caudataZonstein, 2009 (SMNH). 2. R. inopinata sp. nov. (SMNH). 3. R. redikorzevi (Spassky, 1937), ♂ from Akar-Cheshme (SMNH). 4. R. alajensis sp. nov. (SMNH). 5. R. cucullata sp. nov. (SMNH). 6. R. cucullata, paratype from Shahob (SMNH). 7. R. dolosa sp. nov. (SMNH). 8. R. hirta sp. nov. (SMNH). 9. R. ignobilis sp. nov. (SMNH). Scale bars: 5.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13990824" httpUri="https://zenodo.org/record/13990824/files/figure.png" pageId="10" pageNumber="11">Figs 1, 3</figureCitation>
); PMS larger, basal segment of PLS 1.8–2 times as long as PMS (
<figureCitation id="133A1AB05520FFB0FF1DE582FE2ACC99" box="[242,408,679,706]" captionStart="Figs 555–564" captionStartId="160.[189,241,1786,1812]" captionTargetBox="[213,1375,265,1755]" captionTargetId="figure-109@160.[213,1375,265,1755]" captionText="Figs 555–564. Raveniola spp., ♂♂ (555–557, 560, 563) and ♀♀ (558–559, 561–562, 564), holotypes (except 557, 558, 561, 562, 564). Spinnerets, ventral (555–558, 560–561, 563–564) and lateral (559, 562) views. 555. R. caudata Zonstein, 2009, holotype (SMNH). 556. R. inopinata sp. nov. (SMNH). 557–558. R. redikorzevi (Spassky, 1937), ♂ from Akar-Cheshme and ♀ from Kepele, respectively (both SMNH). 559. R. afghana sp. nov. (NHMD). 560. R. alajensis sp. nov. (SMNH). 561–562. R. alajensis, paratype (SMNH). 563. R. cucullata sp. nov. (SMNH). 564. R. cucullata, paratype from Shikildara (SMNH). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.14018401" httpUri="https://zenodo.org/record/14018401/files/figure.png" pageId="10" pageNumber="11">Figs 555, 557</figureCitation>
); embolic keel larger and higher (
<figureCitation id="133A1AB05520FFB0FCCCE582FC74CC99" box="[803,966,680,706]" captionStart="Figs 379–388" captionStartId="145.[189,241,1844,1870]" captionTargetBox="[189,1398,265,1806]" captionTargetId="figure-75@145.[189,1398,265,1806]" captionTargetPageId="145" captionText="Figs 379–388. Raveniola spp., ♂♂, holotypes, details of copulatoty bulb, retrolateral (379, 381, 383, 385, 388), ventral (380, 382, 384, 387) and retroventral (386) views. 379–380. R. caudata Zonstein, 2009 (SMNH). 381–382. R. inopinata sp. nov. (SMNH). 383–384. R. redikorzevi (Spassky, 1937), ♂ from Akar-Cheshme (SMNH).385–387. R. alajensis sp. nov. (SMNH). 388. R. concolor Zonstein, 2000 (MCSN AR 18). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13990917" httpUri="https://zenodo.org/record/13990917/files/figure.png" pageId="10" pageNumber="11">Figs 379, 383</figureCitation>
) ...................................................... 2
</keyLead>
</paragraph>
<paragraph id="8BBE06355520FFB0FF2EE5E7FAC5CD6A" blockId="10.[189,1401,569,817]" pageId="10" pageNumber="11">
<keyLead id="30F0A5005520FFB0FF2EE5E7FAC5CD6A" pageId="10" pageNumber="11">
– Carapace and legs dark sepia brown, abdomen almost uniformly greyish brown (not lighter than carapace, see
<figureCitation id="133A1AB05520FFB0FE7DE5DBFE6ECD50" box="[402,476,753,779]" captionStart="Figs 1–9" captionStartId="107.[189,241,1848,1874]" captionTargetBox="[189,1398,268,1805]" captionTargetId="figure-84@107.[189,1398,265,1805]" captionTargetPageId="107" captionText="Figs 1–9.Raveniola spp., ♂♂, holotypes (except 3, 6), habitus, dorsal views. 1. R. caudataZonstein, 2009 (SMNH). 2. R. inopinata sp. nov. (SMNH). 3. R. redikorzevi (Spassky, 1937), ♂ from Akar-Cheshme (SMNH). 4. R. alajensis sp. nov. (SMNH). 5. R. cucullata sp. nov. (SMNH). 6. R. cucullata, paratype from Shahob (SMNH). 7. R. dolosa sp. nov. (SMNH). 8. R. hirta sp. nov. (SMNH). 9. R. ignobilis sp. nov. (SMNH). Scale bars: 5.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13990824" httpUri="https://zenodo.org/record/13990824/files/figure.png" pageId="10" pageNumber="11">Fig. 2</figureCitation>
); PMS smaller, basal segment of PLS 2.5 times as long as PMS (
<figureCitation id="133A1AB05520FFB0FB10E5DBFAD8CD57" box="[1279,1386,753,780]" captionStart="Figs 555–564" captionStartId="160.[189,241,1786,1812]" captionTargetBox="[213,1375,265,1755]" captionTargetId="figure-109@160.[213,1375,265,1755]" captionText="Figs 555–564. Raveniola spp., ♂♂ (555–557, 560, 563) and ♀♀ (558–559, 561–562, 564), holotypes (except 557, 558, 561, 562, 564). Spinnerets, ventral (555–558, 560–561, 563–564) and lateral (559, 562) views. 555. R. caudata Zonstein, 2009, holotype (SMNH). 556. R. inopinata sp. nov. (SMNH). 557–558. R. redikorzevi (Spassky, 1937), ♂ from Akar-Cheshme and ♀ from Kepele, respectively (both SMNH). 559. R. afghana sp. nov. (NHMD). 560. R. alajensis sp. nov. (SMNH). 561–562. R. alajensis, paratype (SMNH). 563. R. cucullata sp. nov. (SMNH). 564. R. cucullata, paratype from Shikildara (SMNH). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.14018401" httpUri="https://zenodo.org/record/14018401/files/figure.png" pageId="10" pageNumber="11">Fig. 556</figureCitation>
); embolic keel smaller and lower (
<figureCitation id="133A1AB05520FFB0FD84E43CFD7CCD6B" box="[619,718,790,816]" captionStart="Figs 379–388" captionStartId="145.[189,241,1844,1870]" captionTargetBox="[189,1398,265,1806]" captionTargetId="figure-75@145.[189,1398,265,1806]" captionTargetPageId="145" captionText="Figs 379–388. Raveniola spp., ♂♂, holotypes, details of copulatoty bulb, retrolateral (379, 381, 383, 385, 388), ventral (380, 382, 384, 387) and retroventral (386) views. 379–380. R. caudata Zonstein, 2009 (SMNH). 381–382. R. inopinata sp. nov. (SMNH). 383–384. R. redikorzevi (Spassky, 1937), ♂ from Akar-Cheshme (SMNH).385–387. R. alajensis sp. nov. (SMNH). 388. R. concolor Zonstein, 2000 (MCSN AR 18). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13990917" httpUri="https://zenodo.org/record/13990917/files/figure.png" pageId="10" pageNumber="11">Fig. 381</figureCitation>
) .........................................................
<taxonomicName id="4C017DB65520FFB0FB90E43CFAA7CD6B" authorityName="Zonstein" authorityYear="2024" box="[1151,1301,790,816]" class="Arachnida" family="Nemesiidae" genus="Raveniola" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="inopinata" status="sp. nov.">
<emphasis id="B975DA275520FFB0FB90E43CFAA7CD6B" bold="true" box="[1151,1301,790,816]" italics="true" pageId="10" pageNumber="11">R. inopinata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A246675C5520FFB0FAF3E43DFAC5CD6A" box="[1308,1399,791,817]" pageId="10" pageNumber="11" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30F51E905520FFB0FF52E473FAC5CDB9" pageId="10" pageNumber="11">
<paragraph id="8BBE06355520FFB0FF52E473FAC5CDC3" blockId="10.[189,1399,857,994]" pageId="10" pageNumber="11">
<keyLead id="30F0A5005520FFB0FF52E473FAC5CDC3" pageId="10" pageNumber="11">
2. Abdomen with clearly developed darker dorsal pattern (
<figureCitation id="133A1AB05520FFB0FC9CE473FC0BCD28" box="[883,953,857,883]" captionStart="Figs 1–9" captionStartId="107.[189,241,1848,1874]" captionTargetBox="[189,1398,268,1805]" captionTargetId="figure-84@107.[189,1398,265,1805]" captionTargetPageId="107" captionText="Figs 1–9.Raveniola spp., ♂♂, holotypes (except 3, 6), habitus, dorsal views. 1. R. caudataZonstein, 2009 (SMNH). 2. R. inopinata sp. nov. (SMNH). 3. R. redikorzevi (Spassky, 1937), ♂ from Akar-Cheshme (SMNH). 4. R. alajensis sp. nov. (SMNH). 5. R. cucullata sp. nov. (SMNH). 6. R. cucullata, paratype from Shahob (SMNH). 7. R. dolosa sp. nov. (SMNH). 8. R. hirta sp. nov. (SMNH). 9. R. ignobilis sp. nov. (SMNH). Scale bars: 5.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13990824" httpUri="https://zenodo.org/record/13990824/files/figure.png" pageId="10" pageNumber="11">Fig. 3</figureCitation>
). Copulatory bulb 2.5 times as long as wide, with proximal part of embolus less tapering (
<figureCitation id="133A1AB05520FFB0FCAEE454FC17CDC3" box="[833,933,894,920]" captionStart="Figs 379–388" captionStartId="145.[189,241,1844,1870]" captionTargetBox="[189,1398,265,1806]" captionTargetId="figure-75@145.[189,1398,265,1806]" captionTargetPageId="145" captionText="Figs 379–388. Raveniola spp., ♂♂, holotypes, details of copulatoty bulb, retrolateral (379, 381, 383, 385, 388), ventral (380, 382, 384, 387) and retroventral (386) views. 379–380. R. caudata Zonstein, 2009 (SMNH). 381–382. R. inopinata sp. nov. (SMNH). 383–384. R. redikorzevi (Spassky, 1937), ♂ from Akar-Cheshme (SMNH).385–387. R. alajensis sp. nov. (SMNH). 388. R. concolor Zonstein, 2000 (MCSN AR 18). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13990917" httpUri="https://zenodo.org/record/13990917/files/figure.png" pageId="10" pageNumber="11">Fig. 383</figureCitation>
)...........
<taxonomicName id="4C017DB65520FFB0FBEAE454FAC1CDC3" ID-CoL="6X599" authority="(Spassky, 1937)" authorityName="SMNH" baseAuthorityName="Spassky" baseAuthorityYear="1937" box="[1029,1395,894,921]" class="Arachnida" family="Nemesiidae" genus="Raveniola" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="redikorzevi">
<emphasis id="B975DA275520FFB0FBEAE454FB1FCDC2" bold="true" box="[1029,1197,894,921]" italics="true" pageId="10" pageNumber="11">R. redikorzevi</emphasis>
(
<bibRefCitation id="EF907BC45520FFB0FB53E454FAD8CDC3" author="Spassky" box="[1212,1386,894,920]" firstAuthor="Spassky" pageId="10" pageNumber="11" pagination="361 - 368" refId="ref82512" refString="Spassky S. A. 1937. Araneae palaearcticae novae. Mygalomorphae. I. Festschrift Embrik Strand 3: 361 - 368." type="journal article" year="1937">Spassky, 1937</bibRefCitation>
)
</taxonomicName>
.
</keyLead>
</paragraph>
<paragraph id="8BBE06355520FFB0FF2EE489FAC5CDB9" blockId="10.[189,1399,857,994]" pageId="10" pageNumber="11">
<keyLead id="30F0A5005520FFB0FF2EE489FAC5CDB9" pageId="10" pageNumber="11">
– Abdomen with indistinct dorsal pattern (
<figureCitation id="133A1AB05520FFB0FD36E489FC93CDE6" box="[729,801,931,957]" captionStart="Figs 1–9" captionStartId="107.[189,241,1848,1874]" captionTargetBox="[189,1398,268,1805]" captionTargetId="figure-84@107.[189,1398,265,1805]" captionTargetPageId="107" captionText="Figs 1–9.Raveniola spp., ♂♂, holotypes (except 3, 6), habitus, dorsal views. 1. R. caudataZonstein, 2009 (SMNH). 2. R. inopinata sp. nov. (SMNH). 3. R. redikorzevi (Spassky, 1937), ♂ from Akar-Cheshme (SMNH). 4. R. alajensis sp. nov. (SMNH). 5. R. cucullata sp. nov. (SMNH). 6. R. cucullata, paratype from Shahob (SMNH). 7. R. dolosa sp. nov. (SMNH). 8. R. hirta sp. nov. (SMNH). 9. R. ignobilis sp. nov. (SMNH). Scale bars: 5.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13990824" httpUri="https://zenodo.org/record/13990824/files/figure.png" pageId="10" pageNumber="11">Fig. 1</figureCitation>
). Copulatory bulb 2 times as long as wide, with proximal part of embolus more tapering (
<figureCitation id="133A1AB05520FFB0FD20E4E2FC86CDB9" box="[719,820,968,994]" captionStart="Figs 379–388" captionStartId="145.[189,241,1844,1870]" captionTargetBox="[189,1398,265,1806]" captionTargetId="figure-75@145.[189,1398,265,1806]" captionTargetPageId="145" captionText="Figs 379–388. Raveniola spp., ♂♂, holotypes, details of copulatoty bulb, retrolateral (379, 381, 383, 385, 388), ventral (380, 382, 384, 387) and retroventral (386) views. 379–380. R. caudata Zonstein, 2009 (SMNH). 381–382. R. inopinata sp. nov. (SMNH). 383–384. R. redikorzevi (Spassky, 1937), ♂ from Akar-Cheshme (SMNH).385–387. R. alajensis sp. nov. (SMNH). 388. R. concolor Zonstein, 2000 (MCSN AR 18). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13990917" httpUri="https://zenodo.org/record/13990917/files/figure.png" pageId="10" pageNumber="11">Fig. 379</figureCitation>
) .................................
<taxonomicName id="4C017DB65520FFB0FBDAE4E2FAC0CDB9" ID-CoL="4RMTM" authority="Zonstein, 2009" authorityName="Zonstein" authorityYear="2009" box="[1077,1394,967,994]" class="Arachnida" family="Nemesiidae" genus="Raveniola" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="caudata">
<emphasis id="B975DA275520FFB0FBDAE4E2FB0ACDB9" bold="true" box="[1077,1208,967,994]" italics="true" pageId="10" pageNumber="11">R. caudata</emphasis>
<bibRefCitation id="EF907BC45520FFB0FB50E4E2FAC0CDB9" author="Zonstein" box="[1215,1394,968,994]" firstAuthor="Zonstein" pageId="10" pageNumber="11" pagination="37 - 41" refId="ref83362" refString="Zonstein S. 2009. A review of the mygalomorph spiders of the Raveniola caudata - species group (Araneae, Nemesiidae). Zootaxa 2217 (1): 37 - 41. https: // doi. org / 10.11646 / zootaxa. 2217.1.2" type="journal article" year="2009">Zonstein, 2009</bibRefCitation>
</taxonomicName>
.
</keyLead>
</paragraph>
</keyStep>
</key>
<paragraph id="8BBE06355520FFB0FF05E321FEE3CA7E" blockId="10.[189,1166,1035,1098]" box="[234,337,1035,1061]" pageId="10" pageNumber="11">
<emphasis id="B975DA275520FFB0FF05E321FEE3CA7E" bold="true" box="[234,337,1035,1061]" pageId="10" pageNumber="11">Females</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C31B55BE5520FFB0FF52E31AFB3CCA11" box="[189,1166,1072,1098]" pageId="10" pageNumber="11" type="discussion">
<paragraph id="8BBE06355520FFB0FF52E31AFB3CCA11" blockId="10.[189,1166,1035,1098]" box="[189,1166,1072,1098]" pageId="10" pageNumber="11">
Within the group, only the female of
<taxonomicName id="4C017DB65520FFB0FD9FE31AFCA6CA11" authorityName="SMNH" baseAuthorityName="Spassky" baseAuthorityYear="1937" box="[624,788,1072,1098]" class="Arachnida" family="Nemesiidae" genus="Raveniola" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="redikorzevi">
<emphasis id="B975DA275520FFB0FD9FE31AFCA6CA11" box="[624,788,1072,1098]" italics="true" pageId="10" pageNumber="11">R. redikorzevi</emphasis>
</taxonomicName>
is currently known (see below).
</paragraph>
</subSubSection>
</treatment>
</document>
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