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<mods:title id="E03AEC739B47100CC6F52B8054EF8F5D">A revision and one new species of Begonia L. (Begoniaceae, Cucurbitales) in Northeast India</mods:title>
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<taxonomicName id="4C3E63FA2F42FFF80049FA26E2D32BED" ID-CoL="7SG38" authority="A.DC" box="[534,729,1430,1457]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="47" pageNumber="48" phylum="Tracheophyta" rank="species" species="josephi">
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A.DC. [sect. 
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<emphasis id="B94AC46B2F42FFF80126FA26E41E2BED" box="[889,1044,1431,1457]" italics="true" pageId="47" pageNumber="48">Diploclinium</emphasis>
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<figureCitation id="130504FC2F42FFF80088FA0BE3562B88" box="[727,860,1466,1492]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="48.[189,232,1992,2018]" captionStartId-1="49.[189,232,1958,1984]" captionTargetBox-1="[403,1184,265,1921]" captionTargetId-1="figure@49.[403,1184,265,1921]" captionTargetPageId-1="49" captionText-0="Fig. 31. Map showing the location of B. josephi A.DC. specimens." captionText-1="Fig. 32. Begonia josephi A.DC. A. Plant habit. B. Male flower. Photographs courtesy of Sangeeta Rajbhandary of plants in Nepal." figureDoi-1="http://doi.org/10.5281/zenodo.3787095" httpUri-1="https://zenodo.org/record/3787095/files/figure.png" pageId="47" pageNumber="48">Figs 31–32</figureCitation>
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, 
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. 4, 
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11: 126 (de 
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)
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. – 
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<emphasis id="B94AC46B2F42FFF802E2F9D1E2332826" bold="true" box="[189,569,1632,1658]" pageId="47" pageNumber="48">Citations in other publications</emphasis>
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de 
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, 
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<bibRefCitation id="EFAF65882F42FFF803ACF93FE2D028F5" author="Clarke C. B." box="[499,730,1678,1705]" pageId="47" pageNumber="48" pagination="636 - 656" refId="ref53024" refString="Clarke C. B. 1879. Begoniaceae. In: Hooker J. D. (ed.) Flora of British India 2: 636 - 656. London." type="journal article" year="1879">Clarke (1879: 639)</bibRefCitation>
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<treatmentCitation id="0A9F3E682F42FFF800B7F93FE3C728F4" box="[744,973,1678,1704]" pageId="47" pageNumber="48">
<bibRefCitation id="EFAF65882F42FFF800B7F93FE3C728F4" author="Clarke C. B." box="[744,973,1678,1704]" pageId="47" pageNumber="48" pagination="114 - 122" refId="ref53056" refString="Clarke C. B. 1881. On Indian Begonias. Journal of the Linnean Society, Botany 18: 114 - 122, pl. 1 - 3. Available from http: // biodiversitylibrary. org / page / 230642 [accessed May 2009]." type="journal article" year="1881">Clarke (1881: 118)</bibRefCitation>
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<treatmentCitation id="0A9F3E682F42FFF80184F93FE4B928F5" box="[987,1203,1678,1705]" pageId="47" pageNumber="48">
<bibRefCitation id="EFAF65882F42FFF80184F93FE4B928F5" author="Clarke C. B." box="[987,1203,1678,1705]" pageId="47" pageNumber="48" pagination="1 - 107" refId="ref53104" refString="Clarke C. B. 1890. On the plants of Kohima and Muneypore. Journal of the Linnean Society, Botany 25: 1 - 107. Available from http: // biodiversitylibrary. org / page / 180617 [accessed May 2009]." type="journal article" year="1890">Clarke (1890: 25)</bibRefCitation>
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(1979: 181)
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Gu 
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, 
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<bibRefCitation id="EFAF65882F42FFF801FDF964E4CF28B3" author="Verma D. &amp; Roy D. K. &amp; Sinha B. K." box="[930,1221,1749,1775]" pageId="47" pageNumber="48" pagination="175 - 218" refId="ref57839" refString="Verma D., Roy D. K. &amp; Sinha B. K. 2013. Gustav Mann's contribution to ' ASSAM' herbarium, Shillong, Meghalaya, India. Pleione 7 (1): 175 - 218." type="journal article" year="2013">
Verma 
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(2013: 179)
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.
</paragraph>
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<emphasis id="B94AC46B2F42FFF802E2F8A7E173296C" bold="true" box="[189,377,1814,1840]" pageId="47" pageNumber="48">Other material</emphasis>
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: 
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<collectingRegion id="49FAD69B2F42FFF8037BF8F4E2102903" box="[292,538,1861,1887]" pageId="47" pageNumber="48">Arunachal-Pradesh</collectingRegion>
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: Bomdilla, 
<emphasis id="B94AC46B2F42FFF800F0F8F4E35F2903" box="[687,853,1861,1887]" italics="true" pageId="47" pageNumber="48">Joseph 40322</emphasis>
(
<collectionCode id="ED2F80BC2F42FFF80136F8F4E3B42903" box="[873,958,1861,1887]" country="India" httpUri="http://grbio.org/cool/1csn-k189" name="Botanical Survey of India, Arunachal Pradesh Regional Centre" pageId="47" pageNumber="48" type="Herbarium">ARUN</collectionCode>
n.v.); Senge Dzong, 
<date id="FF803EB92F42FFF80698F8F4E57B2903" box="[1223,1393,1861,1887]" pageId="47" pageNumber="48" value="1938-08-16">16 Aug. 1938</date>
, 
<emphasis id="B94AC46B2F42FFF802E2F8D9E14429DE" box="[189,334,1896,1922]" italics="true" pageId="47" pageNumber="48">Ward 14092</emphasis>
(
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, 
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<collectingRegion id="49FAD69B2F42FFF8038FF8D9E22929DE" box="[464,547,1896,1922]" country="India" name="Assam" pageId="47" pageNumber="48">Assam</collectingRegion>
</emphasis>
: Tsuga, 
<date id="FF803EB92F42FFF800D2F8D9E33E29DE" box="[653,820,1896,1923]" pageId="47" pageNumber="48" value="1928-08-25">25 Aug. 1928</date>
, 
<emphasis id="B94AC46B2F42FFF8011BF8D9E3CD29DE" box="[836,967,1896,1922]" italics="true" pageId="47" pageNumber="48">Ward 8579</emphasis>
(
<collectionCode id="ED2F80BC2F42FFF80184F8D9E3FE29DE" box="[987,1012,1896,1922]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" pageId="47" pageNumber="48" type="Herbarium">K</collectionCode>
). 
<emphasis id="B94AC46B2F42FFF80653F8D9E49D29DE" bold="true" box="[1036,1175,1896,1922]" pageId="47" pageNumber="48">
<collectingRegion id="49FAD69B2F42FFF80653F8D9E49D29DE" box="[1036,1175,1896,1922]" country="India" name="Meghalaya" pageId="47" pageNumber="48">Meghalaya</collectingRegion>
</emphasis>
: Khasi Hills, 
<date id="FF803EB92F42FFF8071CF8D9E0F029FA" pageId="47" pageNumber="48" value="1885-09">Sep. 1885</date>
, 
<emphasis id="B94AC46B2F42FFF80356F83DE14729F9" box="[265,333,1932,1957]" italics="true" pageId="47" pageNumber="48">Mann</emphasis>
s.n. (
<collectingRegion id="49FAD69B2F42FFF803CDF83AE1F329F9" box="[402,505,1931,1957]" country="India" name="Assam" pageId="47" pageNumber="48">ASSAM</collectingRegion>
n.v.); Khasi Hills, Laitlyngkot, 
<date id="FF803EB92F42FFF80124F83AE41829F9" box="[891,1042,1931,1957]" pageId="47" pageNumber="48" value="1946-07-17">17 Jul. 1946</date>
, 
<emphasis id="B94AC46B2F42FFF8067EF83AE4B829F9" box="[1057,1202,1931,1957]" italics="true" pageId="47" pageNumber="48">Ward 16047</emphasis>
(
<collectionCode id="ED2F80BC2F42FFF8069BF83AE4FC29F9" box="[1220,1270,1931,1957]" country="United Kingdom" name="Bristol Museum" pageId="47" pageNumber="48">BM</collectionCode>
, 
<collectionCode id="ED2F80BC2F42FFF8075BF83DE51329FA" box="[1284,1305,1932,1958]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15670" name="Royal Botanic Garden Edinburgh" pageId="47" pageNumber="48" type="Herbarium">E</collectionCode>
); Khasi Hills, Mairang, 
<date id="FF803EB92F42FFF80323F81FE2142994" box="[380,542,1966,1992]" pageId="47" pageNumber="48" value="1886-09-12">12 Sep. 1886</date>
, 
<emphasis id="B94AC46B2F42FFF80072F81FE2EE2994" box="[557,740,1966,1992]" italics="true" pageId="47" pageNumber="48">Clarke 45693C</emphasis>
(
<collectionCode id="ED2F80BC2F42FFF800A9F81FE3232994" box="[758,809,1966,1992]" country="United Kingdom" name="Bristol Museum" pageId="47" pageNumber="48">BM</collectionCode>
); Khasi Hills, Mawphlang, 
<date id="FF803EB92F42FFF80628F81FE5132995" box="[1143,1305,1966,1993]" pageId="47" pageNumber="48" value="1953-09-14">14 Sep. 1953</date>
, 
<emphasis id="B94AC46B2F42FFE70777F81FE0F22F7A" italics="true" lastPageId="48" lastPageNumber="49" pageId="47" pageNumber="48">Chand 7223</emphasis>
(
<collectionCode id="ED2F80BC2F5DFFE70355FEBAE1572F79" box="[266,349,267,293]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15602" lsid="urn:lsid:biocol.org:col:15602" name="University of Michigan" pageId="48" pageNumber="49" type="Herbarium">MICH</collectionCode>
); ibid., 
<date id="FF803EB92F5DFFE703E5FEBAE2502F7A" box="[442,602,267,294]" pageId="48" pageNumber="49" value="1952-10-20">20 Oct. 1952</date>
, 
<emphasis id="B94AC46B2F5DFFE70036FEBAE2F62F79" box="[617,764,267,293]" italics="true" pageId="48" pageNumber="49">Chand 6498</emphasis>
(
<collectionCode id="ED2F80BC2F5DFFE70151FEBAE36B2F79" box="[782,865,267,293]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15602" lsid="urn:lsid:biocol.org:col:15602" name="University of Michigan" pageId="48" pageNumber="49" type="Herbarium">MICH</collectionCode>
); ibid., 
<date id="FF803EB92F5DFFE701E1FEBAE4552F79" box="[958,1119,267,293]" pageId="48" pageNumber="49" value="1949-09-10">10 Sep. 1949</date>
, 
<emphasis id="B94AC46B2F5DFFE70631FEBAE50E2F7A" box="[1134,1284,267,294]" italics="true" pageId="48" pageNumber="49">Koelz 23790</emphasis>
(
<collectionCode id="ED2F80BC2F5DFFE70749FEBAE5632F79" box="[1302,1385,267,293]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15602" lsid="urn:lsid:biocol.org:col:15602" name="University of Michigan" pageId="48" pageNumber="49" type="Herbarium">MICH</collectionCode>
); ibid., 
<date id="FF803EB92F5DFFE7035EFE9EE1A92F15" box="[257,419,302,329]" pageId="48" pageNumber="49" value="1952-08-20">20 Aug. 1952</date>
, 
<emphasis id="B94AC46B2F5DFFE703F0FE9EE24B2F15" box="[431,577,303,329]" italics="true" pageId="48" pageNumber="49">Koelz 31179</emphasis>
(
<collectionCode id="ED2F80BC2F5DFFE7000EFE9EE2AE2F15" box="[593,676,303,329]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15602" lsid="urn:lsid:biocol.org:col:15602" name="University of Michigan" pageId="48" pageNumber="49" type="Herbarium">MICH</collectionCode>
); ibid., 
<date id="FF803EB92F5DFFE700A2FE9FE3922F14" box="[765,920,302,329]" pageId="48" pageNumber="49" value="1946-10-16">16 Oct. 1946</date>
, 
<emphasis id="B94AC46B2F5DFFE701FAFE9EE4392F14" box="[933,1075,302,329]" italics="true" pageId="48" pageNumber="49">Ward 16049</emphasis>
(
<collectionCode id="ED2F80BC2F5DFFE7061CFE9EE47C2F15" box="[1091,1142,303,329]" country="United Kingdom" name="Bristol Museum" pageId="48" pageNumber="49">BM</collectionCode>
); ibid., 
<date id="FF803EB92F5DFFE7068FFE9EE57B2F15" box="[1232,1393,303,329]" pageId="48" pageNumber="49" value="1949-08-10">10 Aug. 1949</date>
, 
<emphasis id="B94AC46B2F5DFFE702E2FEE3E1432F30" box="[189,329,338,364]" italics="true" pageId="48" pageNumber="49">Ward 18744</emphasis>
(
<collectionCode id="ED2F80BC2F5DFFE70306FEE3E1862F30" box="[345,396,338,364]" country="United Kingdom" name="Bristol Museum" pageId="48" pageNumber="49">BM</collectionCode>
); Khasia, 
<emphasis id="B94AC46B2F5DFFE703A1FEE3E3022F30" box="[510,776,338,364]" italics="true" pageId="48" pageNumber="49">Hooker &amp; Thomson 34</emphasis>
(iso-: 
<collectionCode id="ED2F80BC2F5DFFE7010DFEE3E3622F30" box="[850,872,338,364]" country="Germany" lsid="urn:lsid:biocol.org:col:15534" name="Botanischer Garten und Botanisches Museum Berlin-Dahlem, Zentraleinrichtung der Freien Universitaet" pageId="48" pageNumber="49" type="Herbarium">B</collectionCode>
, 
<collectionCode id="ED2F80BC2F5DFFE7012CFEE3E3A92F30" box="[883,931,338,364]" country="United Kingdom" name="Bristol Museum" pageId="48" pageNumber="49">BM</collectionCode>
, 
<collectionCode id="ED2F80BC2F5DFFE701F1FEE3E3CC2F30" box="[942,966,338,364]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" pageId="48" pageNumber="49" type="Herbarium">K</collectionCode>
, 
<collectionCode id="ED2F80BC2F5DFFE7018EFEE3E3EF2F30" box="[977,997,338,364]" country="Netherlands" lsid="urn:lsid:biocol.org:col:15678" name="Nationaal Herbarium Nederland, Leiden University branch" pageId="48" pageNumber="49" type="Herbarium">L</collectionCode>
, 
<collectionCode id="ED2F80BC2F5DFFE701AFFEE3E4152F30" box="[1008,1055,338,364]" country="USA" lsid="urn:lsid:biocol.org:col:15556" name="William and Lynda Steere Herbarium of the New York Botanical Garden" pageId="48" pageNumber="49" type="Herbarium">NY</collectionCode>
); ibid., 
<date id="FF803EB92F5DFFE7062AFEE3E4FC2F31" box="[1141,1270,338,365]" pageId="48" pageNumber="49" value="1850-07-05">5 Jul. 1850</date>
, 
<emphasis id="B94AC46B2F5DFFE7075DFEE3E15B2FD3" italics="true" pageId="48" pageNumber="49">Hooker &amp; Thomson 34</emphasis>
(iso-: 
<collectionCode id="ED2F80BC2F5DFFE703FAFEC4E1B42FD3" box="[421,446,373,399]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" pageId="48" pageNumber="49" type="Herbarium">K</collectionCode>
); ibid., 
<date id="FF803EB92F5DFFE70040FEC4E2B82FCC" box="[543,690,373,400]" pageId="48" pageNumber="49" value="1850-04-02">2 Apr. 1850</date>
, 
<emphasis id="B94AC46B2F5DFFE7009CFEC4E3D12FD3" box="[707,987,373,399]" italics="true" pageId="48" pageNumber="49">Hooker &amp; Thomson 34</emphasis>
(iso-: 
<collectionCode id="ED2F80BC2F5DFFE7066FFEC4E4432FD3" box="[1072,1097,373,399]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" pageId="48" pageNumber="49" type="Herbarium">K</collectionCode>
). 
<emphasis id="B94AC46B2F5DFFE7063EFEC4E4D02FD3" bold="true" box="[1121,1242,373,399]" pageId="48" pageNumber="49">
<collectingRegion id="49FAD69B2F5DFFE7063EFEC4E4D02FD3" box="[1121,1242,373,399]" country="India" name="Nagaland" pageId="48" pageNumber="49">Nagaland</collectingRegion>
</emphasis>
: Naga Hills, Pulebudge, 
<date id="FF803EB92F5DFFE70318FE29E1D02FEF" box="[327,474,408,435]" pageId="48" pageNumber="49" value="1935-08-04">4 Aug. 1935</date>
, 
<emphasis id="B94AC46B2F5DFFE703B9FE28E2522FEE" box="[486,600,408,434]" italics="true" pageId="48" pageNumber="49">Bon 5304</emphasis>
(
<collectionCode id="ED2F80BC2F5DFFE70037FE29E28B2FEE" box="[616,641,408,434]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" pageId="48" pageNumber="49" type="Herbarium">K</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3244BF22F5DFFE502E2FE68E17D2F15" lastPageId="50" lastPageNumber="51" pageId="48" pageNumber="49" type="description">
<paragraph id="8B8118792F5DFFE702E2FE68E1442FAF" blockId="48.[189,334,473,499]" box="[189,334,473,499]" pageId="48" pageNumber="49">
<heading id="D0C9AF152F5DFFE702E2FE68E1442FAF" bold="true" box="[189,334,473,499]" fontSize="11" level="3" pageId="48" pageNumber="49" reason="3">
<emphasis id="B94AC46B2F5DFFE702E2FE68E1442FAF" bold="true" box="[189,334,473,499]" pageId="48" pageNumber="49">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8B8118792F5DFFE502E2FDB9E17D2F15" blockId="48.[189,1399,520,1110]" lastBlockId="50.[189,1398,267,329]" lastPageId="50" lastPageNumber="51" pageId="48" pageNumber="49">
Tuberous, stemless, monoecious herb, 
<quantity id="4CC6B59C2F5DFFE700D2FDB9E3002C7F" box="[653,778,520,547]" metricMagnitude="-1" metricUnit="m" metricValue="2.0" metricValueMax="2.5" metricValueMin="1.5" pageId="48" pageNumber="49" unit="cm" value="20.0" valueMax="25.0" valueMin="15.0">15 –25 cm</quantity>
high. Stipules: lanceolate, 
<quantity id="4CC6B59C2F5DFFE7060DFDB9E4B12C7F" box="[1106,1211,520,547]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="48" pageNumber="49" unit="mm" value="2.5" valueMax="3.0" valueMin="2.0">2–3 mm</quantity>
long, glabrous, deciduous. Leaves: petiole (5–)10–20(–30) cm long, glabrous or sparsely puberulous; lamina oblongovate to broadly ovate, peltate, base rounded to truncate, 7–20(–29) × 4–15(–23) cm, symmetric, upper surface green, hispid all over, underside red-purple, sparsely hispid mostly on the veins, venation palmate-pinnate; midrib 
<quantity id="4CC6B59C2F5DFFE703B9FD24E2582CF3" box="[486,594,661,688]" metricMagnitude="-1" metricUnit="m" metricValue="1.15" metricValueMax="1.8" metricValueMin="0.5" pageId="48" pageNumber="49" unit="cm" value="11.5" valueMax="18.0" valueMin="5.0">5–18 cm</quantity>
long; margin with acute lobes, denticulate, with very sparse short hairs; apex acute to acuminate. Inflorescences: cymose, terminal, few; peduncle glabrous, branching 2–3 times, primary 9–15(–25), secondary 
<quantity id="4CC6B59C2F5DFFE700E6FD6AE32C2CAA" box="[697,806,731,758]" metricMagnitude="-2" metricUnit="m" metricValue="6.0" metricValueMax="10.0" metricValueMin="2.0" pageId="48" pageNumber="49" unit="cm" value="6.0" valueMax="10.0" valueMin="2.0">2–10 cm</quantity>
, tertiary 
<quantity id="4CC6B59C2F5DFFE701D1FD6AE3E62CAA" box="[910,1004,731,758]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="48" pageNumber="49" unit="cm" value="1.5" valueMax="2.0" valueMin="1.0">1–2 cm</quantity>
, with 2– 
<specimenCount id="9D38D3F02F5DFFE70608FD6AE4B42CA9" box="[1111,1214,731,757]" pageId="48" pageNumber="49" type="female">6 female</specimenCount>
and 4– 
<specimenCount id="9D38D3F02F5DFFE70745FD6AE57C2CA9" box="[1306,1398,731,757]" pageId="48" pageNumber="49" type="male">10 male</specimenCount>
flowers; bracts lanceolate, 
<quantity id="4CC6B59C2F5DFFE703A8FD4EE2312D45" box="[503,571,767,793]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="48" pageNumber="49" unit="mm" value="3.0">3 mm</quantity>
long, glabrous. Male flowers: pedicel 
<quantity id="4CC6B59C2F5DFFE7065FFD4FE4792D45" box="[1024,1139,766,793]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" metricValueMax="1.5" metricValueMin="0.9" pageId="48" pageNumber="49" unit="mm" value="12.0" valueMax="15.0" valueMin="9.0">9–15 mm</quantity>
long, glabrous; tepals 4; outer tepals broadly obovate, 5–10 × 
<quantity id="4CC6B59C2F5DFFE700F3FC93E3132D60" box="[684,793,802,829]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" metricValueMax="5.0" metricValueMin="3.0" pageId="48" pageNumber="49" unit="mm" value="4.0" valueMax="5.0" valueMin="3.0">3–5 mm</quantity>
, white, sparsely villose on reverse, margin entire; inner tepals spathulate, 3–5 × 
<quantity id="4CC6B59C2F5DFFE70072FCF4E29C2D03" box="[557,662,837,863]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" metricValueMax="3.0" metricValueMin="1.0" pageId="48" pageNumber="49" unit="mm" value="2.0" valueMax="3.0" valueMin="1.0">1–3 mm</quantity>
, white, glabrous; androecium with 9–15 stamens, symmetric; filaments 
<quantity id="4CC6B59C2F5DFFE7036DFCD9E1932DDF" box="[306,409,872,899]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="48" pageNumber="49" unit="mm" value="1.5" valueMax="2.0" valueMin="1.0">1–2 mm</quantity>
long, subequal, fused at base into a short column; anther ellipsoid, 
<quantity id="4CC6B59C2F5DFFE70694FCD9E5382DDF" box="[1227,1330,872,899]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="48" pageNumber="49" unit="mm" value="1.5" valueMax="2.0" valueMin="1.0">1–2 mm</quantity>
long, dehiscing through slits about half the length of the anther, not hooded, connective not extended. Female flower: pedicel 
<quantity id="4CC6B59C2F5DFFE70329FC1EE1E02D95" box="[374,490,943,969]" metricMagnitude="-2" metricUnit="m" metricValue="1.05" metricValueMax="1.7" metricValueMin="0.4" pageId="48" pageNumber="49" unit="mm" value="10.5" valueMax="17.0" valueMin="4.0">4–17 mm</quantity>
long, glabrous; bracteoles absent; tepals 4–5, unequal, 4 ovate, 1 lanceolate (not always present), outer tepals 5–10 × 
<quantity id="4CC6B59C2F5DFFE700F3FC63E31F2DB0" box="[684,789,978,1004]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" metricValueMax="7.0" metricValueMin="4.0" pageId="48" pageNumber="49" unit="mm" value="5.5" valueMax="7.0" valueMin="4.0">4–7 mm</quantity>
, white, glabrous, margin entire, inner tepals similar but smaller; ovary 3-locular, placentae bifid; capsule oblong-ellipsoid, 3–5 × 
<quantity id="4CC6B59C2F5DFFE70609FC44E4CB2A53" box="[1110,1217,1013,1039]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" metricValueMax="4.0" metricValueMin="2.0" pageId="48" pageNumber="49" unit="mm" value="3.0" valueMax="4.0" valueMin="2.0">2–4 mm</quantity>
, glabrous, with one long triangular wing and two short oblong wings; styles 3, shallowly U-shaped, persistent. Fruit: pendulous; capsule oblong-ellipsoid, 7–9 × 
<quantity id="4CC6B59C2F5DFFE7008DFB8AE3402A0A" box="[722,842,1083,1110]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" metricValueMax="1.1" metricValueMin="0.9" pageId="48" pageNumber="49" unit="mm" value="10.0" valueMax="11.0" valueMin="9.0">9–11 mm</quantity>
, glabrous; wings extending along the pedicel, unequal, rounded at the tips; longest wing rounded-oblong, 10–14 × 
<quantity id="4CC6B59C2F5FFFE501BFFEBAE4692F7A" box="[992,1123,267,294]" metricMagnitude="-2" metricUnit="m" metricValue="1.05" metricValueMax="1.1" metricValueMin="1.0" pageId="50" pageNumber="51" unit="mm" value="10.5" valueMax="11.0" valueMin="10.0">10–11 mm</quantity>
; shortest wings oblong, 1–3 × 
<quantity id="4CC6B59C2F5FFFE50354FE9EE1782F15" box="[267,370,303,329]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" metricValueMax="9.0" metricValueMin="7.0" pageId="50" pageNumber="51" unit="mm" value="8.0" valueMax="9.0" valueMin="7.0">7–9 mm</quantity>
.
</paragraph>
</subSubSection>
<caption id="DF4148F12F5DFFE702E2F879E3C029BF" box="[189,970,1992,2019]" pageId="48" pageNumber="49">
<paragraph id="8B8118792F5DFFE702E2F879E3C029BF" blockId="48.[189,970,1992,2019]" box="[189,970,1992,2019]" pageId="48" pageNumber="49">
<emphasis id="B94AC46B2F5DFFE702E2F879E11329BF" bold="true" box="[189,281,1992,2019]" pageId="48" pageNumber="49">Fig. 31.</emphasis>
Map showing the location of 
<taxonomicName id="4C3E63FA2F5DFFE70026F87BE2E529BF" box="[633,751,1993,2019]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="48" pageNumber="49" phylum="Tracheophyta" rank="species" species="josephi">
<emphasis id="B94AC46B2F5DFFE70026F87BE2E529BF" box="[633,751,1993,2019]" italics="true" pageId="48" pageNumber="49">B. josephi</emphasis>
</taxonomicName>
A.DC. specimens.
</paragraph>
</caption>
<caption id="DF4148F12F5CFFE602E2F817E23929B8" ID-DOI="http://doi.org/10.5281/zenodo.3787095" ID-Zenodo-Dep="3787095" httpUri="https://zenodo.org/record/3787095/files/figure.png" pageId="49" pageNumber="50" startId="49.[189,232,1958,1984]" targetBox="[403,1184,265,1921]" targetPageId="49">
<paragraph id="8B8118792F5CFFE602E2F817E23929B8" blockId="49.[189,1399,1958,2020]" pageId="49" pageNumber="50">
<emphasis id="B94AC46B2F5CFFE602E2F817E117299D" bold="true" box="[189,285,1958,1985]" pageId="49" pageNumber="50">Fig. 32.</emphasis>
<taxonomicName id="4C3E63FA2F5CFFE60375F819E1E7299D" box="[298,493,1959,1985]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="49" pageNumber="50" phylum="Tracheophyta" rank="species" species="josephi">
<emphasis id="B94AC46B2F5CFFE60375F819E1E7299D" box="[298,493,1959,1985]" italics="true" pageId="49" pageNumber="50">Begonia josephi</emphasis>
</taxonomicName>
A.DC. 
<emphasis id="B94AC46B2F5CFFE60010F817E26E299C" bold="true" box="[591,612,1958,1984]" pageId="49" pageNumber="50">A</emphasis>
. Plant habit. 
<emphasis id="B94AC46B2F5CFFE60151F816E328299D" bold="true" box="[782,802,1959,1985]" pageId="49" pageNumber="50">B</emphasis>
. Male flower. Photographs courtesy of Sangeeta Rajbhandary of plants in Nepal.
</paragraph>
</caption>
<subSubSection id="C3244BF22F5FFFE502E2FEC1E49C2CCE" pageId="50" pageNumber="51" type="distribution">
<paragraph id="8B8118792F5FFFE502E2FEC1E21F2FD6" blockId="50.[189,533,368,394]" box="[189,533,368,394]" pageId="50" pageNumber="51">
<heading id="D0C9AF152F5FFFE502E2FEC1E21F2FD6" bold="true" box="[189,533,368,394]" fontSize="11" level="3" pageId="50" pageNumber="51" reason="3">
<emphasis id="B94AC46B2F5FFFE502E2FEC1E21F2FD6" bold="true" box="[189,533,368,394]" pageId="50" pageNumber="51">Distribution and phenology</emphasis>
</heading>
</paragraph>
<paragraph id="8B8118792F5FFFE502E2FE2FE3C42F80" blockId="50.[189,1398,414,476]" pageId="50" pageNumber="51">
Arunachal-Pradesh, Assam, Meghalaya and Nagaland; also in 
<collectingCountry id="F32958E92F5FFFE50197FE2FE41B2FE4" box="[968,1041,414,440]" name="China" pageId="50" pageNumber="51">China</collectingCountry>
, 
<collectingCountry id="F32958E92F5FFFE5067BFE2FE4672FE4" box="[1060,1133,414,440]" name="Nepal" pageId="50" pageNumber="51">Nepal</collectingCountry>
, Sikkim, 
<collectingCountry id="F32958E92F5FFFE506B6FE2FE5342FE4" box="[1257,1342,414,440]" name="Bhutan" pageId="50" pageNumber="51">Bhutan</collectingCountry>
and 
<collectingCountry id="F32958E92F5FFFE502E2FE73E1392F80" box="[189,307,450,476]" name="Myanmar" pageId="50" pageNumber="51">Myanmar</collectingCountry>
; 
<quantity id="4CC6B59C2F5FFFE50360FE70E1E02F80" box="[319,490,449,476]" metricMagnitude="3" metricUnit="m" metricValue="1.975" metricValueMax="2.45" metricValueMin="1.5" pageId="50" pageNumber="51" unit="m" value="1975.0" valueMax="2450.0" valueMin="1500.0">1500–2450 m</quantity>
. Flowering and fruiting: June to October.
</paragraph>
<paragraph id="8B8118792F5FFFE502E2FDB2E1BF2C41" blockId="50.[189,437,515,541]" box="[189,437,515,541]" pageId="50" pageNumber="51">
<heading id="D0C9AF152F5FFFE502E2FDB2E1BF2C41" bold="true" box="[189,437,515,541]" fontSize="11" level="3" pageId="50" pageNumber="51" reason="3">
<emphasis id="B94AC46B2F5FFFE502E2FDB2E1BF2C41" bold="true" box="[189,437,515,541]" pageId="50" pageNumber="51">Conservation status</emphasis>
</heading>
</paragraph>
<paragraph id="8B8118792F5FFFE502E2FD83E49C2CCE" blockId="50.[189,1398,560,658]" pageId="50" pageNumber="51">
Least Concern. 
<taxonomicName id="4C3E63FA2F5FFFE50325FD82E2302C10" box="[378,570,562,588]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="50" pageNumber="51" phylum="Tracheophyta" rank="species" species="josephi">
<emphasis id="B94AC46B2F5FFFE50325FD82E2302C10" box="[378,570,562,588]" italics="true" pageId="50" pageNumber="51">Begonia josephi</emphasis>
</taxonomicName>
has an AOO of 
<quantity id="4CC6B59C2F5FFFE5015EFD83E3562C10" box="[769,860,562,588]" metricMagnitude="5" metricUnit="m" metricValue="1.72" pageId="50" pageNumber="51" unit="km" value="172.0">172 km</quantity>
<superScript id="7C4BB5312F5FFFE50103FD81E36F2C62" attach="left" box="[860,869,560,574]" fontSize="6" pageId="50" pageNumber="51">2</superScript>
and an EOO of 
<quantity id="4CC6B59C2F5FFFE50671FD83E4B62C10" box="[1070,1212,562,588]" metricMagnitude="8" metricUnit="m" metricValue="2.7" pageId="50" pageNumber="51" unit="km" value="270000.0">270,000 km</quantity>
<superScript id="7C4BB5312F5FFFE506E3FD81E4CF2C62" attach="left" box="[1212,1221,560,574]" fontSize="6" pageId="50" pageNumber="51">2</superScript>
with plenty of suitable habitat from which it is commonly collected in the Eastern Himalayas. The species has been found in several national parks in 
<collectingCountry id="F32958E92F5FFFE50013FDC9E2982CCE" box="[588,658,632,658]" name="Nepal" pageId="50" pageNumber="51">Nepal</collectingCountry>
and from Sakteng National Park in 
<collectingCountry id="F32958E92F5FFFE50666FDC9E4982CCE" box="[1081,1170,632,658]" name="Bhutan" pageId="50" pageNumber="51">Bhutan</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C3244BF22F5FFFE502E2FD0BE39B2D98" pageId="50" pageNumber="51" type="discussion">
<paragraph id="8B8118792F5FFFE502E2FD0BE1252C88" blockId="50.[189,303,698,724]" box="[189,303,698,724]" pageId="50" pageNumber="51">
<heading id="D0C9AF152F5FFFE502E2FD0BE1252C88" bold="true" box="[189,303,698,724]" fontSize="11" level="3" pageId="50" pageNumber="51" reason="3">
<emphasis id="B94AC46B2F5FFFE502E2FD0BE1252C88" bold="true" box="[189,303,698,724]" pageId="50" pageNumber="51">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8B8118792F5FFFE502E2FD59E57D2D79" blockId="50.[189,1399,744,806]" pageId="50" pageNumber="51">
This is the only species known in the area with a peltate leaf. 
<emphasis id="B94AC46B2F5FFFE501F0FD59E4492D5E" box="[943,1091,744,770]" italics="true" pageId="50" pageNumber="51">Koelz 31179</emphasis>
has a larger and a barely peltate leaf with a deeply cordate leaf base; however, the flowers compare very well to other specimens.
</paragraph>
<paragraph id="8B8118792F5FFFE50395FCD2E39B2D98" blockId="50.[458,1129,867,965]" pageId="50" pageNumber="51">
<heading id="D0C9AF152F5FFFE50395FCD2E4632D22" box="[458,1129,867,894]" centered="true" fontSize="11" level="2" pageId="50" pageNumber="51" reason="2">
<taxonomicName id="4C3E63FA2F5FFFE50395FCD2E3192D21" authority="R.Camfield" authorityName="R. Camfield" authorityYear="2018" box="[458,787,867,894]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="50" pageNumber="51" phylum="Tracheophyta" rank="species" species="koelzii" status="sp. nov.">
<emphasis id="B94AC46B2F5FFFE50395FCD2E28F2D22" bold="true" box="[458,645,867,894]" italics="true" pageId="50" pageNumber="51">Begonia koelzii</emphasis>
R.Camfield
</taxonomicName>
<taxonomicNameLabel id="A27979102F5FFFE50145FCD5E37F2D22" box="[794,885,868,894]" pageId="50" pageNumber="51" rank="species">sp. nov.</taxonomicNameLabel>
[sect. 
<taxonomicName id="4C3E63FA2F5FFFE5019EFCD2E4552D21" box="[961,1119,867,893]" class="Magnoliopsida" family="Begoniaceae" genus="Platycentrum" higherTaxonomySource="CoL" kingdom="Plantae" order="Cucurbitales" pageId="50" pageNumber="51" phylum="Tracheophyta" rank="genus">
<emphasis id="B94AC46B2F5FFFE5019EFCD2E4552D21" box="[961,1119,867,893]" italics="true" pageId="50" pageNumber="51">Platycentrum</emphasis>
</taxonomicName>
]
</heading>
urn:lsid:ipni.org:names:77175032-1 
<figureCitation id="130504FC2F5FFFE500FDFC1BE3242D99" box="[674,814,938,965]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStartId-0="50.[189,232,1992,2018]" captionStartId-1="51.[189,232,1718,1744]" captionStartId-2="52.[189,232,1889,1915]" captionTargetBox-0="[428,1159,1224,1951]" captionTargetBox-1="[189,1399,265,1681]" captionTargetBox-2="[238,1350,274,1842]" captionTargetId-0="figure@50.[416,1172,1213,1966]" captionTargetId-1="figure@51.[189,1399,265,1681]" captionTargetId-2="figure@52.[234,1354,266,1846]" captionTargetPageId-0="50" captionTargetPageId-1="51" captionTargetPageId-2="52" captionText-0="Fig. 33. Map showing the location of B. koelzii R.Camfield sp. nov. specimens." captionText-1="Fig. 34. Begonia koelzii R.Camfield sp. nov. A. Plant habit. B. Leaf. C. Bulbil. D. Young female flowers. E. Fruit. Photographs courtesy of Nick Macer, of a plant in Manipur." captionText-2="Fig. 35. Illustration of B. koelzii R.Camfield sp. nov. A. Plant habit. B. Adaxial leaf surface. C. Abaxial leaf surface. D. Inflorescence. E. Style. F. Male flower. G. Stamens. H. Female buds. I. Female flower. J. Mature fruit. Drawn by Rebecca Camfield. A, H from photographs by Nick Macer. B–D, J from Koelz 267739; E–G, I from Koelz 26053; all MICH." figureDoi-0="http://doi.org/10.5281/zenodo.3787097" figureDoi-1="http://doi.org/10.5281/zenodo.3787099" figureDoi-2="http://doi.org/10.5281/zenodo.3787101" httpUri-0="https://zenodo.org/record/3787097/files/figure.png" httpUri-1="https://zenodo.org/record/3787099/files/figure.png" httpUri-2="https://zenodo.org/record/3787101/files/figure.png" pageId="50" pageNumber="51">Figs 33–35</figureCitation>
; 
<tableCitation id="C6BC2DC22F5FFFE50165FC1BE39B2D98" box="[826,913,938,964]" captionStart="Table 1" captionStartId="53.[183,252,558,586]" captionText="Table 1. Comparison of the key vegetative and floral characters of the new species B. koelzii R.Camfield sp. nov. and several similar species. Data collected from herbarium specimens at E, HAST and K, species protologues and the Flora of China (Gu et al. 2007)." pageId="50" pageNumber="51">Table 1</tableCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3244BF22F5FFFE502E2FC5AE49F2A0B" pageId="50" pageNumber="51" type="diagnosis">
<paragraph id="8B8118792F5FFFE502E2FC5AE13C2A59" blockId="50.[189,310,1003,1029]" box="[189,310,1003,1029]" pageId="50" pageNumber="51">
<heading id="D0C9AF152F5FFFE502E2FC5AE13C2A59" bold="true" box="[189,310,1003,1029]" fontSize="11" level="3" pageId="50" pageNumber="51" reason="3">
<emphasis id="B94AC46B2F5FFFE502E2FC5AE13C2A59" bold="true" box="[189,310,1003,1029]" pageId="50" pageNumber="51">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8B8118792F5FFFE502E2FBABE49F2A0B" blockId="50.[189,1399,1050,1112]" pageId="50" pageNumber="51">
Similar to 
<taxonomicName id="4C3E63FA2F5FFFE50360FBAAE2D92A68" authority="Irmsch. (1951: 41)" authorityName="Irmsch. (Irmscher" authorityPageNumber="41" authorityYear="1951" box="[319,723,1050,1077]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="50" pageNumber="51" phylum="Tracheophyta" rank="species" species="macrotoma">
<emphasis id="B94AC46B2F5FFFE50360FBAAE1E22A68" box="[319,488,1051,1076]" italics="true" pageId="50" pageNumber="51">B. macrotoma</emphasis>
Irmsch. (1951: 41)
</taxonomicName>
in having lacerate leaves, but differs in having a larger lamina (
<quantity id="4CC6B59C2F5FFFE50342FB8CE1922A0B" box="[285,408,1085,1111]" metricMagnitude="-1" metricUnit="m" metricValue="3.0" metricValueMax="4.0" metricValueMin="2.0" pageId="50" pageNumber="51" unit="cm" value="30.0" valueMax="40.0" valueMin="20.0">20–40 cm</quantity>
long, not 
<quantity id="4CC6B59C2F5FFFE50050FB8CE2832A0B" box="[527,649,1085,1112]" metricMagnitude="-1" metricUnit="m" metricValue="1.35" metricValueMax="1.5" metricValueMin="1.2" pageId="50" pageNumber="51" unit="cm" value="13.5" valueMax="15.0" valueMin="12.0">12–15 cm</quantity>
) and female flowers with 4–6 (not 3) tepals.
</paragraph>
</subSubSection>
<caption id="DF4148F12F5FFFE502E2F879E45529BF" ID-DOI="http://doi.org/10.5281/zenodo.3787097" ID-Zenodo-Dep="3787097" box="[189,1119,1992,2019]" httpUri="https://zenodo.org/record/3787097/files/figure.png" pageId="50" pageNumber="51" startId="50.[189,232,1992,2018]" targetBox="[428,1159,1224,1951]" targetPageId="50">
<paragraph id="8B8118792F5FFFE502E2F879E45529BF" blockId="50.[189,1119,1992,2019]" box="[189,1119,1992,2019]" pageId="50" pageNumber="51">
<emphasis id="B94AC46B2F5FFFE502E2F879E11029BF" bold="true" box="[189,282,1992,2019]" pageId="50" pageNumber="51">Fig. 33.</emphasis>
Map showing the location of 
<taxonomicName id="4C3E63FA2F5FFFE50026F87BE37E29BF" authority="R.Camfield" authorityName="R. Camfield" authorityYear="2018" box="[633,884,1993,2019]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="50" pageNumber="51" phylum="Tracheophyta" rank="species" species="koelzii" status="sp. nov.">
<emphasis id="B94AC46B2F5FFFE50026F87BE2ED29BF" box="[633,743,1993,2019]" italics="true" pageId="50" pageNumber="51">B. koelzii</emphasis>
R.Camfield
</taxonomicName>
<taxonomicNameLabel id="A27979102F5FFFE50124F878E3DC29BF" box="[891,982,1993,2019]" pageId="50" pageNumber="51" rank="species">sp. nov.</taxonomicNameLabel>
specimens.
</paragraph>
</caption>
<caption id="DF4148F12F5EFFE402E2F907E3EE28A8" ID-DOI="http://doi.org/10.5281/zenodo.3787099" ID-Zenodo-Dep="3787099" httpUri="https://zenodo.org/record/3787099/files/figure.png" pageId="51" pageNumber="52" startId="51.[189,232,1718,1744]" targetBox="[189,1399,265,1681]" targetPageId="51">
<paragraph id="8B8118792F5EFFE402E2F907E3EE28A8" blockId="51.[189,1399,1718,1780]" pageId="51" pageNumber="52">
<emphasis id="B94AC46B2F5EFFE402E2F907E11D288D" bold="true" box="[189,279,1718,1745]" pageId="51" pageNumber="52">Fig. 34.</emphasis>
<taxonomicName id="4C3E63FA2F5EFFE40341F909E255288D" authority="R.Camfield" authorityName="R. Camfield" authorityYear="2018" box="[286,607,1719,1745]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="51" pageNumber="52" phylum="Tracheophyta" rank="species" species="koelzii" status="sp. nov.">
<emphasis id="B94AC46B2F5EFFE40341F909E1D9288D" box="[286,467,1719,1745]" italics="true" pageId="51" pageNumber="52">Begonia koelzii</emphasis>
R.Camfield
</taxonomicName>
<taxonomicNameLabel id="A27979102F5EFFE4003AF906E2B4288D" box="[613,702,1719,1745]" pageId="51" pageNumber="52" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="B94AC46B2F5EFFE4009BF907E2D3288C" bold="true" box="[708,729,1718,1744]" pageId="51" pageNumber="52">A</emphasis>
. Plant habit. 
<emphasis id="B94AC46B2F5EFFE4012EF906E38F288D" bold="true" box="[881,901,1719,1745]" pageId="51" pageNumber="52">B</emphasis>
. Leaf. 
<emphasis id="B94AC46B2F5EFFE4018AF907E3E0288C" bold="true" box="[981,1002,1718,1744]" pageId="51" pageNumber="52">C</emphasis>
. Bulbil. 
<emphasis id="B94AC46B2F5EFFE40611F906E469288D" bold="true" box="[1102,1123,1719,1745]" pageId="51" pageNumber="52">D</emphasis>
. Young female flowers. 
<emphasis id="B94AC46B2F5EFFE402E2F96BE0DB28A8" bold="true" box="[189,209,1754,1780]" pageId="51" pageNumber="52">E</emphasis>
. Fruit. Photographs courtesy of Nick Macer, of a plant in Manipur.
</paragraph>
</caption>
<caption id="DF4148F12F59FFE302E2F8D0E2D629B8" ID-DOI="http://doi.org/10.5281/zenodo.3787101" ID-Zenodo-Dep="3787101" httpUri="https://zenodo.org/record/3787101/files/figure.png" pageId="52" pageNumber="53" startId="52.[189,232,1889,1915]" targetBox="[238,1350,274,1842]" targetPageId="52">
<paragraph id="8B8118792F59FFE302E2F8D0E2D629B8" blockId="52.[189,1399,1889,2021]" pageId="52" pageNumber="53">
<emphasis id="B94AC46B2F59FFE302E2F8D0E1132927" bold="true" box="[189,281,1889,1915]" pageId="52" pageNumber="53">Fig. 35.</emphasis>
Illustration of 
<taxonomicName id="4C3E63FA2F59FFE30397F8D3E2C92927" authority="R.Camfield" authorityName="R. Camfield" authorityYear="2018" box="[456,707,1889,1915]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="52" pageNumber="53" phylum="Tracheophyta" rank="species" species="koelzii" status="sp. nov.">
<emphasis id="B94AC46B2F59FFE30397F8D3E23C2927" box="[456,566,1889,1915]" italics="true" pageId="52" pageNumber="53">B. koelzii</emphasis>
R.Camfield
</taxonomicName>
<taxonomicNameLabel id="A27979102F59FFE30095F8D0E32F2927" box="[714,805,1889,1915]" pageId="52" pageNumber="53" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="B94AC46B2F59FFE30173F8D0E34B2927" bold="true" box="[812,833,1889,1915]" pageId="52" pageNumber="53">A</emphasis>
. Plant habit. 
<emphasis id="B94AC46B2F59FFE30183F8D0E3FA2927" bold="true" box="[988,1008,1889,1915]" pageId="52" pageNumber="53">B</emphasis>
. Adaxial leaf surface. 
<emphasis id="B94AC46B2F59FFE306A8F8D0E5062927" bold="true" box="[1271,1292,1889,1915]" pageId="52" pageNumber="53">C</emphasis>
. Abaxial leaf surface. 
<emphasis id="B94AC46B2F59FFE3030BF835E16329C2" bold="true" box="[340,361,1924,1950]" pageId="52" pageNumber="53">D</emphasis>
. Inflorescence. 
<emphasis id="B94AC46B2F59FFE3007CF835E23D29C2" bold="true" box="[547,567,1924,1950]" pageId="52" pageNumber="53">E</emphasis>
. Style. 
<emphasis id="B94AC46B2F59FFE300CFF835E2A829C2" bold="true" box="[656,674,1924,1950]" pageId="52" pageNumber="53">F</emphasis>
. Male flower. 
<emphasis id="B94AC46B2F59FFE30112F835E36E29C2" bold="true" box="[845,868,1924,1950]" pageId="52" pageNumber="53">G</emphasis>
. Stamens. 
<emphasis id="B94AC46B2F59FFE301BBF835E3F129C2" bold="true" box="[996,1019,1924,1950]" pageId="52" pageNumber="53">H</emphasis>
. Female buds. 
<emphasis id="B94AC46B2F59FFE306F1F835E4B329C2" bold="true" box="[1198,1209,1924,1950]" pageId="52" pageNumber="53">I</emphasis>
. Female flower. 
<emphasis id="B94AC46B2F59FFE302E2F816E0C6299D" bold="true" box="[189,204,1959,1985]" pageId="52" pageNumber="53">J</emphasis>
. Mature fruit. Drawn by Rebecca Camfield. A, H from photographs by Nick Macer. B–D, J from 
<emphasis id="B94AC46B2F59FFE3076BF816E11F29B8" italics="true" pageId="52" pageNumber="53">Koelz 267739</emphasis>
; E–G, I from 
<emphasis id="B94AC46B2F59FFE3039FF87BE25929B8" box="[448,595,1994,2020]" italics="true" pageId="52" pageNumber="53">Koelz 26053</emphasis>
; all MICH.
</paragraph>
</caption>
<caption id="DF4148F12F58FFE202E8FD9FE61B2C31" pageId="53" pageNumber="54">
<paragraph id="8B8118792F58FFE202E8FD9FE61B2C31" blockId="53.[183,1937,558,621]" pageId="53" pageNumber="54">
<emphasis id="B94AC46B2F58FFE202E8FD9FE1162C16" bold="true" box="[183,284,558,586]" pageId="53" pageNumber="54">Table 1.</emphasis>
Comparison of the key vegetative and floral characters of the new species 
<taxonomicName id="4C3E63FA2F58FFE206C0FD9FE5942C16" authority="R.Camfield" authorityName="R. Camfield" authorityYear="2018" box="[1183,1438,558,586]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="53" pageNumber="54" phylum="Tracheophyta" rank="species" species="koelzii" status="sp. nov.">
<emphasis id="B94AC46B2F58FFE206C0FD9FE5042C16" box="[1183,1294,558,586]" italics="true" pageId="53" pageNumber="54">B. koelzii</emphasis>
R.Camfield
</taxonomicName>
<taxonomicNameLabel id="A27979102F58FFE207F9FD9FE6092C16" box="[1446,1539,558,586]" pageId="53" pageNumber="54" rank="species">sp. nov.</taxonomicNameLabel>
and several similar species. Data collected from herbarium specimens at E, HAST and K, species protologues and the Flora of China (
<bibRefCitation id="EFAF65882F58FFE20709FDE0E60E2C31" author="Gu C. &amp; Peng C. &amp; Turland N. J." box="[1366,1540,593,621]" pageId="53" pageNumber="54" pagination="153 - 207" refId="ref54157" refString="Gu C., Peng C. - I &amp; Turland N. J. 2007. Begoniaceae. In: Wu Z. Y., Raven P. H. &amp; Hong D. Y. (eds) Flora of China 13: 153 - 207. Missouri Botanical Garden Press, Missouri." type="journal article" year="2007">
Gu 
<emphasis id="B94AC46B2F58FFE207DDFDE0E5B72C31" box="[1410,1469,593,621]" italics="true" pageId="53" pageNumber="54">et al.</emphasis>
2007
</bibRefCitation>
).
</paragraph>
</caption>
<paragraph id="8B8118792F58FFE202E4FD2EE72228E1" pageId="53" pageNumber="54">
<table id="F93EEAD92F58002802E4FD2EE78E28E1" box="[187,1924,671,1725]" gridcols="8" gridrows="18" pageId="53" pageNumber="54">
<tr id="350E1A3B2F58002802E4FD2EE78E2C97" box="[187,1924,671,715]" gridrow="0" pageId="53" pageNumber="54">
<th id="76DF73472F58002802E4FD2EE1BA2C97" box="[187,432,671,715]" gridcol="0" gridrow="0" pageId="53" pageNumber="54">
<emphasis id="B94AC46B2F58FFE202E4FD2EE12C2CE9" bold="true" box="[187,294,671,693]" pageId="53" pageNumber="54">Characters</emphasis>
</th>
<th id="76DF73472F5800280396FD2EE2542C97" box="[457,606,671,715]" gridcol="1" gridrow="0" pageId="53" pageNumber="54">
<taxonomicName id="4C3E63FA2F58FFE20396FD2EE2142C97" authority="R.Camfield" authorityName="R. Camfield" authorityYear="2018" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="53" pageNumber="54" phylum="Tracheophyta" rank="species" species="koelzii" status="sp. nov.">
<emphasis id="B94AC46B2F58FFE20396FD2EE2152CE9" bold="true" box="[457,543,671,693]" italics="true" pageId="53" pageNumber="54">B. koelzii</emphasis>
R.Camfield
</taxonomicName>
<taxonomicNameLabel id="A27979102F58FFE2007CFD08E2542C97" box="[547,606,697,715]" pageId="53" pageNumber="54" rank="species">sp. nov.</taxonomicNameLabel>
</th>
<th id="76DF73472F58002800CCFD2EE3312C97" box="[659,827,671,715]" gridcol="2" gridrow="0" pageId="53" pageNumber="54">
<taxonomicName id="4C3E63FA2F58FFE200CCFD2EE3312C97" authority="K. Y. Guan &amp; D. K. Tian" authorityName="K. Y. Guan &amp; D. K. Tian (Guan &amp; Tian" authorityYear="2000" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="53" pageNumber="54" phylum="Tracheophyta" rank="species" species="longialata">
<emphasis id="B94AC46B2F58FFE200CCFD2EE3002CE9" bold="true" box="[659,778,671,693]" italics="true" pageId="53" pageNumber="54">B. longialata</emphasis>
K.Y.Guan &amp; D.K.Tian
</taxonomicName>
</th>
<th id="76DF73472F580028012EFD2EE4062C97" box="[881,1036,671,715]" gridcol="3" gridrow="0" pageId="53" pageNumber="54">
<taxonomicName id="4C3E63FA2F58FFE2012EFD2EE3A12C97" authority="Irmsch." authorityName="Irmsch. (Irmscher" authorityYear="1951" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="53" pageNumber="54" phylum="Tracheophyta" rank="species" species="macrotoma">
<emphasis id="B94AC46B2F58FFE2012EFD2EE3FE2CE9" bold="true" box="[881,1012,671,693]" italics="true" pageId="53" pageNumber="54">B. macrotoma</emphasis>
Irmsch.
</taxonomicName>
</th>
<th id="76DF73472F5800280671FD2EE4DC2C97" box="[1070,1238,671,715]" gridcol="4" gridrow="0" pageId="53" pageNumber="54">
<taxonomicName id="4C3E63FA2F58FFE20671FD2EE47A2C97" authority="S. Rajbh." class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="53" pageNumber="54" phylum="Tracheophyta" rank="species" species="panchtharensis">
<emphasis id="B94AC46B2F58FFE20671FD2EE4DC2CE9" bold="true" box="[1070,1238,671,693]" italics="true" pageId="53" pageNumber="54">B. panchtharensis</emphasis>
S.Rajbh.
</taxonomicName>
</th>
<th id="76DF73472F58002806A9FD2EE5812C97" box="[1270,1419,671,715]" gridcol="5" gridrow="0" pageId="53" pageNumber="54">
<taxonomicName id="4C3E63FA2F58FFE206A9FD2EE52F2C97" authority="H. Lev" authorityName="H. Lev. (Leveille" authorityYear="1909" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="53" pageNumber="54" phylum="Tracheophyta" rank="species" species="pedatifida">
<emphasis id="B94AC46B2F58FFE206A9FD2EE5612CE9" bold="true" box="[1270,1387,671,693]" italics="true" pageId="53" pageNumber="54">B. pedatifida</emphasis>
H.Lév
</taxonomicName>
</th>
<th id="76DF73472F58002807EFFD2EE66B2C97" box="[1456,1633,671,715]" gridcol="6" gridrow="0" pageId="53" pageNumber="54">
<taxonomicName id="4C3E63FA2F58FFE207EEFD2EE66B2C97" authority="S. H. Huang &amp; Y. M. Shui" authorityName="S. H. Huang &amp; Y. M. Shui (Huang &amp; Shui" authorityYear="1994" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="53" pageNumber="54" phylum="Tracheophyta" rank="species" species="rubropunctata">
<emphasis id="B94AC46B2F58FFE207EEFD2EE65A2CE9" bold="true" box="[1457,1616,671,693]" italics="true" pageId="53" pageNumber="54">B. rubropunctata</emphasis>
S.H.Huang &amp; Y.M.Shui
</taxonomicName>
</th>
<th id="76DF73472F58002804C4FD2EE78E2C97" box="[1691,1924,671,715]" gridcol="7" gridrow="0" pageId="53" pageNumber="54">
<taxonomicName id="4C3E63FA2F58FFE204C4FD2EE78E2C97" authority="Rekha Morris &amp; P. D. McMillan" authorityName="Rekha Morris &amp; P. D. McMillan (Morris &amp; McMillan" authorityYear="2012" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="53" pageNumber="54" phylum="Tracheophyta" rank="species" species="shilendrae">
<emphasis id="B94AC46B2F58FFE204C4FD2EE71C2CE9" bold="true" box="[1691,1814,671,693]" italics="true" pageId="53" pageNumber="54">B. shilendrae</emphasis>
Rekha Morris &amp; P.D.McMillan
</taxonomicName>
</th>
</tr>
<tr id="350E1A3B2F58002802E4FD50E78E2CAB" box="[187,1924,737,759]" gridrow="1" pageId="53" pageNumber="54">
<th id="76DF73472F58002802E4FD50E1BA2CAB" box="[187,432,737,759]" gridcol="0" gridrow="1" pageId="53" pageNumber="54">cauline leaves</th>
<td id="76DF73472F5800280396FD50E2542CAB" box="[457,606,737,759]" gridcol="1" gridrow="1" pageId="53" pageNumber="54">no</td>
<td id="76DF73472F58002800CCFD50E3312CAB" box="[659,827,737,759]" gridcol="2" gridrow="1" pageId="53" pageNumber="54">yes</td>
<td id="76DF73472F580028012EFD50E4062CAB" box="[881,1036,737,759]" gridcol="3" gridrow="1" pageId="53" pageNumber="54">yes</td>
<td id="76DF73472F5800280671FD50E4DC2CAB" box="[1070,1238,737,759]" gridcol="4" gridrow="1" pageId="53" pageNumber="54">no</td>
<td id="76DF73472F58002806A9FD50E5812CAB" box="[1270,1419,737,759]" gridcol="5" gridrow="1" pageId="53" pageNumber="54">no</td>
<td id="76DF73472F58002807EFFD50E66B2CAB" box="[1456,1633,737,759]" gridcol="6" gridrow="1" pageId="53" pageNumber="54">no</td>
<td id="76DF73472F58002804C4FD50E78E2CAB" box="[1691,1924,737,759]" gridcol="7" gridrow="1" pageId="53" pageNumber="54">no</td>
</tr>
<tr id="350E1A3B2F58002802E4FCADE78E2D6E" box="[187,1924,796,818]" gridrow="2" pageId="53" pageNumber="54">
<th id="76DF73472F58002802E4FCADE1BA2D6E" box="[187,432,796,818]" gridcol="0" gridrow="2" pageId="53" pageNumber="54">bulbils</th>
<td id="76DF73472F5800280396FCADE2542D6E" box="[457,606,796,818]" gridcol="1" gridrow="2" pageId="53" pageNumber="54">yes, frequently</td>
<td id="76DF73472F58002800CCFCADE3312D6E" box="[659,827,796,818]" gridcol="2" gridrow="2" pageId="53" pageNumber="54">no</td>
<td id="76DF73472F580028012EFCADE4062D6E" box="[881,1036,796,818]" gridcol="3" gridrow="2" pageId="53" pageNumber="54">no</td>
<td id="76DF73472F5800280671FCADE4DC2D6E" box="[1070,1238,796,818]" gridcol="4" gridrow="2" pageId="53" pageNumber="54">very occasionally?</td>
<td id="76DF73472F58002806A9FCADE66B2D6E" box="[1270,1633,796,818]" colspan="2" colspanRight="1" gridcol="5" gridrow="2" pageId="53" pageNumber="54">very occasionally? very occasionally?</td>
<td id="76DF73472F58002804C4FCADE78E2D6E" box="[1691,1924,796,818]" gridcol="7" gridrow="2" pageId="53" pageNumber="54">no</td>
</tr>
<tr id="350E1A3B2F58002802E4FCE6E78E2D31" box="[187,1924,855,877]" gridrow="3" pageId="53" pageNumber="54">
<th id="76DF73472F58002802E4FCE6E1BA2D31" box="[187,432,855,877]" gridcol="0" gridrow="3" pageId="53" pageNumber="54">lamina shape</th>
<td id="76DF73472F5800280396FCE6E2542D31" box="[457,606,855,877]" gridcol="1" gridrow="3" pageId="53" pageNumber="54">ovate</td>
<td id="76DF73472F58002800CCFCE6E3312D31" box="[659,827,855,877]" gridcol="2" gridrow="3" pageId="53" pageNumber="54">suborbicular</td>
<td id="76DF73472F580028012EFCE6E4062D31" box="[881,1036,855,877]" gridcol="3" gridrow="3" pageId="53" pageNumber="54">broad ovate</td>
<td id="76DF73472F5800280671FCE6E4DC2D31" box="[1070,1238,855,877]" gridcol="4" gridrow="3" pageId="53" pageNumber="54">suborbicular</td>
<td id="76DF73472F58002806A9FCE6E5812D31" box="[1270,1419,855,877]" gridcol="5" gridrow="3" pageId="53" pageNumber="54">suborbicular</td>
<td id="76DF73472F58002807EFFCE6E66B2D31" box="[1456,1633,855,877]" gridcol="6" gridrow="3" pageId="53" pageNumber="54">suborbicular</td>
<td id="76DF73472F58002804C4FCE6E78E2D31" box="[1691,1924,855,877]" gridcol="7" gridrow="3" pageId="53" pageNumber="54">broad ovate</td>
</tr>
<tr id="350E1A3B2F58002802E4FC20E78E2DFB" box="[187,1924,913,935]" gridrow="4" pageId="53" pageNumber="54">
<th id="76DF73472F58002802E4FC20E1BA2DFB" box="[187,432,913,935]" gridcol="0" gridrow="4" pageId="53" pageNumber="54">lamina lobes</th>
<td id="76DF73472F5800280396FC20E2542DFB" box="[457,606,913,935]" gridcol="1" gridrow="4" pageId="53" pageNumber="54">deeply incised</td>
<td id="76DF73472F58002800CCFC20E3312DFB" box="[659,827,913,935]" gridcol="2" gridrow="4" pageId="53" pageNumber="54">incised</td>
<td id="76DF73472F580028012EFC20E4062DFB" box="[881,1036,913,935]" gridcol="3" gridrow="4" pageId="53" pageNumber="54">shallowly incised</td>
<td id="76DF73472F5800280671FC20E4DC2DFB" box="[1070,1238,913,935]" gridcol="4" gridrow="4" pageId="53" pageNumber="54">deeply incised</td>
<td id="76DF73472F58002806A9FC20E5812DFB" box="[1270,1419,913,935]" gridcol="5" gridrow="4" pageId="53" pageNumber="54">deeply incised</td>
<td id="76DF73472F58002807EFFC20E66B2DFB" box="[1456,1633,913,935]" gridcol="6" gridrow="4" pageId="53" pageNumber="54">deeply incised</td>
<td id="76DF73472F58002804C4FC20E78E2DFB" box="[1691,1924,913,935]" gridcol="7" gridrow="4" pageId="53" pageNumber="54">linear</td>
</tr>
<tr id="350E1A3B2F58002802E4FC7DE78E2DBE" box="[187,1924,972,994]" gridrow="5" pageId="53" pageNumber="54">
<th id="76DF73472F58002802E4FC7DE1BA2DBE" box="[187,432,972,994]" gridcol="0" gridrow="5" pageId="53" pageNumber="54">lamina upper surface</th>
<td id="76DF73472F5800280396FC7DE2542DBE" box="[457,606,972,994]" gridcol="1" gridrow="5" pageId="53" pageNumber="54">sparsely strigose</td>
<td id="76DF73472F58002800CCFC7DE3312DBE" box="[659,827,972,994]" gridcol="2" gridrow="5" pageId="53" pageNumber="54">glabrous</td>
<td id="76DF73472F580028012EFC7DE4062DBE" box="[881,1036,972,994]" gridcol="3" gridrow="5" pageId="53" pageNumber="54">sparsely hirsute</td>
<td id="76DF73472F5800280671FC7DE4DC2DBE" box="[1070,1238,972,994]" gridcol="4" gridrow="5" pageId="53" pageNumber="54">sparsely hirsute</td>
<td id="76DF73472F58002806A9FC7DE5812DBE" box="[1270,1419,972,994]" gridcol="5" gridrow="5" pageId="53" pageNumber="54">sparsely strigose</td>
<td id="76DF73472F58002807EFFC7DE66B2DBE" box="[1456,1633,972,994]" gridcol="6" gridrow="5" pageId="53" pageNumber="54">sparsely strigose</td>
<td id="76DF73472F58002804C4FC7DE78E2DBE" box="[1691,1924,972,994]" gridcol="7" gridrow="5" pageId="53" pageNumber="54">glabrous</td>
</tr>
<tr id="350E1A3B2F58002802E4FBB6E78E2A41" box="[187,1924,1031,1053]" gridrow="6" pageId="53" pageNumber="54">
<th id="76DF73472F58002802E4FBB6E1BA2A41" box="[187,432,1031,1053]" gridcol="0" gridrow="6" pageId="53" pageNumber="54">lamina length (cm)</th>
<td id="76DF73472F5800280396FBB6E2542A41" box="[457,606,1031,1053]" gridcol="1" gridrow="6" pageId="53" pageNumber="54">20– 50</td>
<td id="76DF73472F58002800CCFBB6E3312A41" box="[659,827,1031,1053]" gridcol="2" gridrow="6" pageId="53" pageNumber="54">25 –45</td>
<td id="76DF73472F580028012EFBB6E4062A41" box="[881,1036,1031,1053]" gridcol="3" gridrow="6" pageId="53" pageNumber="54">12–15</td>
<td id="76DF73472F5800280671FBB6E4DC2A41" box="[1070,1238,1031,1053]" gridcol="4" gridrow="6" pageId="53" pageNumber="54">25– 40</td>
<td id="76DF73472F58002806A9FBB6E5812A41" box="[1270,1419,1031,1053]" gridcol="5" gridrow="6" pageId="53" pageNumber="54">10– 40</td>
<td id="76DF73472F58002807EFFBB6E66B2A41" box="[1456,1633,1031,1053]" gridcol="6" gridrow="6" pageId="53" pageNumber="54">15– 20</td>
<td id="76DF73472F58002804C4FBB6E78E2A41" box="[1691,1924,1031,1053]" gridcol="7" gridrow="6" pageId="53" pageNumber="54">18 –35</td>
</tr>
<tr id="350E1A3B2F58002802E4FBF0E78E2A0B" box="[187,1924,1089,1111]" gridrow="7" pageId="53" pageNumber="54">
<th id="76DF73472F58002802E4FBF0E1BA2A0B" box="[187,432,1089,1111]" gridcol="0" gridrow="7" pageId="53" pageNumber="54">petiole length (cm)</th>
<td id="76DF73472F5800280396FBF0E2542A0B" box="[457,606,1089,1111]" gridcol="1" gridrow="7" pageId="53" pageNumber="54">30– 50</td>
<td id="76DF73472F58002800CCFBF0E3312A0B" box="[659,827,1089,1111]" gridcol="2" gridrow="7" pageId="53" pageNumber="54">10 –30</td>
<td id="76DF73472F580028012EFBF0E4062A0B" box="[881,1036,1089,1111]" gridcol="3" gridrow="7" pageId="53" pageNumber="54">5–15</td>
<td id="76DF73472F5800280671FBF0E4DC2A0B" box="[1070,1238,1089,1111]" gridcol="4" gridrow="7" pageId="53" pageNumber="54">30– 45</td>
<td id="76DF73472F58002806A9FBF0E5812A0B" box="[1270,1419,1089,1111]" gridcol="5" gridrow="7" pageId="53" pageNumber="54">10– 55</td>
<td id="76DF73472F58002807EFFBF0E66B2A0B" box="[1456,1633,1089,1111]" gridcol="6" gridrow="7" pageId="53" pageNumber="54">15– 25</td>
<td id="76DF73472F58002804C4FBF0E78E2A0B" box="[1691,1924,1089,1111]" gridcol="7" gridrow="7" pageId="53" pageNumber="54">up to 40</td>
</tr>
<tr id="350E1A3B2F58002802E4FBCDE78E2AF1" box="[187,1924,1148,1197]" gridrow="8" pageId="53" pageNumber="54">
<th id="76DF73472F58002802E4FBCDE1BA2AF1" box="[187,432,1148,1197]" gridcol="0" gridrow="8" pageId="53" pageNumber="54">inflorescence structure</th>
<td id="76DF73472F5800280396FBCDE2542AF1" box="[457,606,1148,1197]" gridcol="1" gridrow="8" pageId="53" pageNumber="54">cymose</td>
<td id="76DF73472F58002800CCFBCDE3312AF1" box="[659,827,1148,1197]" gridcol="2" gridrow="8" pageId="53" pageNumber="54">cymose</td>
<td id="76DF73472F580028012EFBCDE4062AF1" box="[881,1036,1148,1197]" gridcol="3" gridrow="8" pageId="53" pageNumber="54">cymose</td>
<td id="76DF73472F5800280671FBCDE4DC2AF1" box="[1070,1238,1148,1197]" gridcol="4" gridrow="8" pageId="53" pageNumber="54">cymose</td>
<td id="76DF73472F58002806A9FBCDE5812AF1" box="[1270,1419,1148,1197]" gridcol="5" gridrow="8" pageId="53" pageNumber="54">cymose</td>
<td id="76DF73472F58002807EFFBCDE66B2AF1" box="[1456,1633,1148,1197]" gridcol="6" gridrow="8" pageId="53" pageNumber="54">cymose</td>
<td id="76DF73472F58002804C4FBCDE78E2AF1" box="[1691,1924,1148,1197]" gridcol="7" gridrow="8" pageId="53" pageNumber="54">paniculate or raceme of cymes</td>
</tr>
<tr id="350E1A3B2F58002802E4FB60E78E2ABB" box="[187,1924,1233,1255]" gridrow="9" pageId="53" pageNumber="54">
<th id="76DF73472F58002802E4FB60E1BA2ABB" box="[187,432,1233,1255]" gridcol="0" gridrow="9" pageId="53" pageNumber="54">peduncle length (cm)</th>
<td id="76DF73472F5800280396FB60E2542ABB" box="[457,606,1233,1255]" gridcol="1" gridrow="9" pageId="53" pageNumber="54">30– 60</td>
<td id="76DF73472F58002800CCFB60E3312ABB" box="[659,827,1233,1255]" gridcol="2" gridrow="9" pageId="53" pageNumber="54">5 –10</td>
<td id="76DF73472F580028012EFB60E4062ABB" box="[881,1036,1233,1255]" gridcol="3" gridrow="9" pageId="53" pageNumber="54">5–15</td>
<td id="76DF73472F5800280671FB60E4DC2ABB" box="[1070,1238,1233,1255]" gridcol="4" gridrow="9" pageId="53" pageNumber="54">40– 50</td>
<td id="76DF73472F58002806A9FB60E5812ABB" box="[1270,1419,1233,1255]" gridcol="5" gridrow="9" pageId="53" pageNumber="54">15– 20</td>
<td id="76DF73472F58002807EFFB60E66B2ABB" box="[1456,1633,1233,1255]" gridcol="6" gridrow="9" pageId="53" pageNumber="54">50– 60</td>
<td id="76DF73472F58002804C4FB60E78E2ABB" box="[1691,1924,1233,1255]" gridcol="7" gridrow="9" pageId="53" pageNumber="54">10 –30</td>
</tr>
<tr id="350E1A3B2F58002802E4FABDE78E2B7E" box="[187,1924,1292,1314]" gridrow="10" pageId="53" pageNumber="54">
<th id="76DF73472F58002802E4FABDE1BA2B7E" box="[187,432,1292,1314]" gridcol="0" gridrow="10" pageId="53" pageNumber="54">male tepal no.</th>
<td id="76DF73472F5800280396FABDE2542B7E" box="[457,606,1292,1314]" gridcol="1" gridrow="10" pageId="53" pageNumber="54">4</td>
<td id="76DF73472F58002800CCFABDE3312B7E" box="[659,827,1292,1314]" gridcol="2" gridrow="10" pageId="53" pageNumber="54">4</td>
<td id="76DF73472F580028012EFABDE4062B7E" box="[881,1036,1292,1314]" gridcol="3" gridrow="10" pageId="53" pageNumber="54">4</td>
<td id="76DF73472F5800280671FABDE4DC2B7E" box="[1070,1238,1292,1314]" gridcol="4" gridrow="10" pageId="53" pageNumber="54">4</td>
<td id="76DF73472F58002806A9FABDE5812B7E" box="[1270,1419,1292,1314]" gridcol="5" gridrow="10" pageId="53" pageNumber="54">4</td>
<td id="76DF73472F58002807EFFABDE66B2B7E" box="[1456,1633,1292,1314]" gridcol="6" gridrow="10" pageId="53" pageNumber="54">4</td>
<td id="76DF73472F58002804C4FABDE78E2B7E" box="[1691,1924,1292,1314]" gridcol="7" gridrow="10" pageId="53" pageNumber="54">2</td>
</tr>
<tr id="350E1A3B2F58002802E4FAF6E78E2B01" box="[187,1924,1351,1373]" gridrow="11" pageId="53" pageNumber="54">
<th id="76DF73472F58002802E4FAF6E1BA2B01" box="[187,432,1351,1373]" gridcol="0" gridrow="11" pageId="53" pageNumber="54">female tepal no.</th>
<td id="76DF73472F5800280396FAF6E2542B01" box="[457,606,1351,1373]" gridcol="1" gridrow="11" pageId="53" pageNumber="54">4–6</td>
<td id="76DF73472F58002800CCFAF6E3312B01" box="[659,827,1351,1373]" gridcol="2" gridrow="11" pageId="53" pageNumber="54">5</td>
<td id="76DF73472F580028012EFAF6E4062B01" box="[881,1036,1351,1373]" gridcol="3" gridrow="11" pageId="53" pageNumber="54">3</td>
<td id="76DF73472F5800280671FAF6E4DC2B01" box="[1070,1238,1351,1373]" gridcol="4" gridrow="11" pageId="53" pageNumber="54">5</td>
<td id="76DF73472F58002806A9FAF6E5812B01" box="[1270,1419,1351,1373]" gridcol="5" gridrow="11" pageId="53" pageNumber="54">5</td>
<td id="76DF73472F58002807EFFAF6E66B2B01" box="[1456,1633,1351,1373]" gridcol="6" gridrow="11" pageId="53" pageNumber="54">4 to 5</td>
<td id="76DF73472F58002804C4FAF6E78E2B01" box="[1691,1924,1351,1373]" gridcol="7" gridrow="11" pageId="53" pageNumber="54">2</td>
</tr>
<tr id="350E1A3B2F58002802E4FA30E78E2BCB" box="[187,1924,1409,1431]" gridrow="12" pageId="53" pageNumber="54">
<th id="76DF73472F58002802E4FA30E1BA2BCB" box="[187,432,1409,1431]" gridcol="0" gridrow="12" pageId="53" pageNumber="54">stamen no.</th>
<td id="76DF73472F5800280396FA30E2542BCB" box="[457,606,1409,1431]" gridcol="1" gridrow="12" pageId="53" pageNumber="54">60– 90</td>
<td id="76DF73472F58002800CCFA30E3312BCB" box="[659,827,1409,1431]" gridcol="2" gridrow="12" pageId="53" pageNumber="54">150–180</td>
<td id="76DF73472F580028012EFA30E4062BCB" box="[881,1036,1409,1431]" gridcol="3" gridrow="12" pageId="53" pageNumber="54">n/a</td>
<td id="76DF73472F5800280671FA30E4DC2BCB" box="[1070,1238,1409,1431]" gridcol="4" gridrow="12" pageId="53" pageNumber="54">120–150</td>
<td id="76DF73472F58002806A9FA30E5812BCB" box="[1270,1419,1409,1431]" gridcol="5" gridrow="12" pageId="53" pageNumber="54">80–100</td>
<td id="76DF73472F58002807EFFA30E66B2BCB" box="[1456,1633,1409,1431]" gridcol="6" gridrow="12" pageId="53" pageNumber="54">100–120</td>
<td id="76DF73472F58002804C4FA30E78E2BCB" box="[1691,1924,1409,1431]" gridcol="7" gridrow="12" pageId="53" pageNumber="54">30 –40</td>
</tr>
<tr id="350E1A3B2F58002802E4FA0DE78E2B8E" box="[187,1924,1468,1490]" gridrow="13" pageId="53" pageNumber="54">
<th id="76DF73472F58002802E4FA0DE1BA2B8E" box="[187,432,1468,1490]" gridcol="0" gridrow="13" pageId="53" pageNumber="54">styles</th>
<td id="76DF73472F5800280396FA0DE2542B8E" box="[457,606,1468,1490]" gridcol="1" gridrow="13" pageId="53" pageNumber="54">3</td>
<td id="76DF73472F58002800CCFA0DE3312B8E" box="[659,827,1468,1490]" gridcol="2" gridrow="13" pageId="53" pageNumber="54">2 –3</td>
<td id="76DF73472F580028012EFA0DE4062B8E" box="[881,1036,1468,1490]" gridcol="3" gridrow="13" pageId="53" pageNumber="54">2</td>
<td id="76DF73472F5800280671FA0DE4DC2B8E" box="[1070,1238,1468,1490]" gridcol="4" gridrow="13" pageId="53" pageNumber="54">2</td>
<td id="76DF73472F58002806A9FA0DE5812B8E" box="[1270,1419,1468,1490]" gridcol="5" gridrow="13" pageId="53" pageNumber="54">2</td>
<td id="76DF73472F58002807EFFA0DE66B2B8E" box="[1456,1633,1468,1490]" gridcol="6" gridrow="13" pageId="53" pageNumber="54">2</td>
<td id="76DF73472F58002804C4FA0DE78E2B8E" box="[1691,1924,1468,1490]" gridcol="7" gridrow="13" pageId="53" pageNumber="54">2 –3</td>
</tr>
<tr id="350E1A3B2F58002802E4FA46E78E2851" box="[187,1924,1527,1549]" gridrow="14" pageId="53" pageNumber="54">
<th id="76DF73472F58002802E4FA46E1BA2851" box="[187,432,1527,1549]" gridcol="0" gridrow="14" pageId="53" pageNumber="54">longest wing shape</th>
<td id="76DF73472F5800280396FA46E2542851" box="[457,606,1527,1549]" gridcol="1" gridrow="14" pageId="53" pageNumber="54">rounded-oblong</td>
<td id="76DF73472F58002800CCFA46E3312851" box="[659,827,1527,1549]" gridcol="2" gridrow="14" pageId="53" pageNumber="54">rounded-oblong</td>
<td id="76DF73472F580028012EFA46E4062851" box="[881,1036,1527,1549]" gridcol="3" gridrow="14" pageId="53" pageNumber="54">n/a</td>
<td id="76DF73472F5800280671FA46E4DC2851" box="[1070,1238,1527,1549]" gridcol="4" gridrow="14" pageId="53" pageNumber="54">oblong-triangular</td>
<td id="76DF73472F58002806A9FA46E5812851" box="[1270,1419,1527,1549]" gridcol="5" gridrow="14" pageId="53" pageNumber="54">triangular</td>
<td id="76DF73472F58002807EFFA46E66B2851" box="[1456,1633,1527,1549]" gridcol="6" gridrow="14" pageId="53" pageNumber="54">rounded-oblong</td>
<td id="76DF73472F58002804C4FA46E78E2851" box="[1691,1924,1527,1549]" gridcol="7" gridrow="14" pageId="53" pageNumber="54">rounded-oblong</td>
</tr>
<tr id="350E1A3B2F58002802E4F980E78E281B" box="[187,1924,1585,1607]" gridrow="15" pageId="53" pageNumber="54">
<th id="76DF73472F58002802E4F980E1BA281B" box="[187,432,1585,1607]" gridcol="0" gridrow="15" pageId="53" pageNumber="54">longest wing length (cm)</th>
<td id="76DF73472F5800280396F980E254281B" box="[457,606,1585,1607]" gridcol="1" gridrow="15" pageId="53" pageNumber="54">1–3</td>
<td id="76DF73472F58002800CCF980E331281B" box="[659,827,1585,1607]" gridcol="2" gridrow="15" pageId="53" pageNumber="54">3 –5</td>
<td id="76DF73472F580028012EF980E406281B" box="[881,1036,1585,1607]" gridcol="3" gridrow="15" pageId="53" pageNumber="54">n/a</td>
<td id="76DF73472F5800280671F980E4DC281B" box="[1070,1238,1585,1607]" gridcol="4" gridrow="15" pageId="53" pageNumber="54">1–2.5</td>
<td id="76DF73472F58002806A9F980E581281B" box="[1270,1419,1585,1607]" gridcol="5" gridrow="15" pageId="53" pageNumber="54">1–2.5</td>
<td id="76DF73472F58002807EFF980E66B281B" box="[1456,1633,1585,1607]" gridcol="6" gridrow="15" pageId="53" pageNumber="54">1–1.5</td>
<td id="76DF73472F58002804C4F980E78E281B" box="[1691,1924,1585,1607]" gridcol="7" gridrow="15" pageId="53" pageNumber="54">1.5– 2</td>
</tr>
<tr id="350E1A3B2F58002802E4F9DDE78E28DE" box="[187,1924,1644,1666]" gridrow="16" pageId="53" pageNumber="54">
<th id="76DF73472F58002802E4F9DDE1BA28DE" box="[187,432,1644,1666]" gridcol="0" gridrow="16" pageId="53" pageNumber="54">side wing shape</th>
<td id="76DF73472F5800280396F9DDE25428DE" box="[457,606,1644,1666]" gridcol="1" gridrow="16" pageId="53" pageNumber="54">thin oblong</td>
<td id="76DF73472F58002800CCF9DDE33128DE" box="[659,827,1644,1666]" gridcol="2" gridrow="16" pageId="53" pageNumber="54">thin oblong</td>
<td id="76DF73472F580028012EF9DDE40628DE" box="[881,1036,1644,1666]" gridcol="3" gridrow="16" pageId="53" pageNumber="54">n/a</td>
<td id="76DF73472F5800280671F9DDE4DC28DE" box="[1070,1238,1644,1666]" gridcol="4" gridrow="16" pageId="53" pageNumber="54">thin oblong</td>
<td id="76DF73472F58002806A9F9DDE58128DE" box="[1270,1419,1644,1666]" gridcol="5" gridrow="16" pageId="53" pageNumber="54">curved triangle</td>
<td id="76DF73472F58002807EFF9DDE66B28DE" box="[1456,1633,1644,1666]" gridcol="6" gridrow="16" pageId="53" pageNumber="54">thin oblong</td>
<td id="76DF73472F58002804C4F9DDE78E28DE" box="[1691,1924,1644,1666]" gridcol="7" gridrow="16" pageId="53" pageNumber="54">curved oblong</td>
</tr>
<tr id="350E1A3B2F58002802E4F916E78E28E1" box="[187,1924,1703,1725]" gridrow="17" pageId="53" pageNumber="54">
<th id="76DF73472F58002802E4F916E1BA28E1" box="[187,432,1703,1725]" gridcol="0" gridrow="17" pageId="53" pageNumber="54">wings extend along pedicel</th>
<td id="76DF73472F5800280396F916E25428E1" box="[457,606,1703,1725]" gridcol="1" gridrow="17" pageId="53" pageNumber="54">no</td>
<td id="76DF73472F58002800CCF916E33128E1" box="[659,827,1703,1725]" gridcol="2" gridrow="17" pageId="53" pageNumber="54">slightly</td>
<td id="76DF73472F580028012EF916E40628E1" box="[881,1036,1703,1725]" gridcol="3" gridrow="17" pageId="53" pageNumber="54">n/a</td>
<td id="76DF73472F5800280671F916E4DC28E1" box="[1070,1238,1703,1725]" gridcol="4" gridrow="17" pageId="53" pageNumber="54">no</td>
<td id="76DF73472F58002806A9F916E58128E1" box="[1270,1419,1703,1725]" gridcol="5" gridrow="17" pageId="53" pageNumber="54">yes and in front</td>
<td id="76DF73472F58002807EFF916E66B28E1" box="[1456,1633,1703,1725]" gridcol="6" gridrow="17" pageId="53" pageNumber="54">slightly</td>
<td id="76DF73472F58002804C4F916E78E28E1" box="[1691,1924,1703,1725]" gridcol="7" gridrow="17" pageId="53" pageNumber="54">yes and in front</td>
</tr>
</table>
</paragraph>
<subSubSection id="C3244BF22F5BFFE102E2FEBAE54C2F08" box="[189,1350,267,340]" pageId="54" pageNumber="55" type="etymology">
<paragraph id="8B8118792F5BFFE102E2FEBAE14E2F79" blockId="54.[189,324,267,293]" box="[189,324,267,293]" pageId="54" pageNumber="55">
<heading id="D0C9AF152F5BFFE102E2FEBAE14E2F79" bold="true" box="[189,324,267,293]" fontSize="11" level="3" pageId="54" pageNumber="55" reason="3">
<emphasis id="B94AC46B2F5BFFE102E2FEBAE14E2F79" bold="true" box="[189,324,267,293]" pageId="54" pageNumber="55">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8B8118792F5BFFE102E2FE88E54C2F08" blockId="54.[189,1350,313,340]" box="[189,1350,313,340]" pageId="54" pageNumber="55">
The epithet honours Walter N. Koelz (
<date id="FF803EB92F5BFFE10022FE88E3002F0F" box="[637,778,313,340]" pageId="54" pageNumber="55" value="1895" valueMax="1989">1895–1989</date>
), the American zoologist who collected the 
<typeStatus id="5485A6DB2F5BFFE10753FE8BE5482F08" box="[1292,1346,314,340]" pageId="54" pageNumber="55">type</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="C3244BF22F5BFFE102E2FECAE4E82C0B" pageId="54" pageNumber="55" type="materials_examined">
<paragraph id="8B8118792F5BFFE102E2FECAE0F02FC9" blockId="54.[189,250,379,405]" box="[189,250,379,405]" pageId="54" pageNumber="55">
<heading id="D0C9AF152F5BFFE102E2FECAE0F02FC9" bold="true" box="[189,250,379,405]" fontSize="11" level="3" pageId="54" pageNumber="55" reason="3">
<emphasis id="B94AC46B2F5BFFE102E2FECAE0F02FC9" bold="true" box="[189,250,379,405]" pageId="54" pageNumber="55">
<typeStatus id="5485A6DB2F5BFFE102E2FECAE0F02FC9" box="[189,250,379,405]" pageId="54" pageNumber="55">Type</typeStatus>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B8118792F5BFFE102E2FE18E46A2F98" blockId="54.[189,1120,425,452]" box="[189,1120,425,452]" pageId="54" pageNumber="55">
<collectingCountry id="F32958E92F5BFFE102E2FE18E0F62F9F" box="[189,252,425,451]" name="India" pageId="54" pageNumber="55">India</collectingCountry>
, 
<collectingRegion id="49FAD69B2F5BFFE10357FE18E1712F9F" box="[264,379,425,451]" country="India" name="Nagaland" pageId="54" pageNumber="55">Nagaland</collectingRegion>
, Takubama, 
<date id="FF803EB92F5BFFE10050FE18E2972F98" box="[527,669,425,452]" pageId="54" pageNumber="55" value="1950-09-04">4 Sep. 1950</date>
, 
<emphasis id="B94AC46B2F5BFFE100F6FE18E3362F9F" box="[681,828,425,451]" italics="true" pageId="54" pageNumber="55">Koelz 26053</emphasis>
(holo-: 
<collectionCode id="ED2F80BC2F5BFFE101C4FE18E3E12F98" box="[923,1003,425,452]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15602" lsid="urn:lsid:biocol.org:col:15602" name="University of Michigan" pageId="54" pageNumber="55" type="Herbarium">MICH</collectionCode>
1225736).
</paragraph>
<paragraph id="8B8118792F5BFFE102E2FE5BE1F32C58" blockId="54.[189,505,490,516]" box="[189,505,490,516]" pageId="54" pageNumber="55">
<heading id="D0C9AF152F5BFFE102E2FE5BE1F32C58" bold="true" box="[189,505,490,516]" fontSize="11" level="3" pageId="54" pageNumber="55" reason="3">
<emphasis id="B94AC46B2F5BFFE102E2FE5BE1F32C58" bold="true" box="[189,505,490,516]" pageId="54" pageNumber="55">Other material examined</emphasis>
</heading>
</paragraph>
<paragraph id="8B8118792F5BFFE102E2FDA8E4E82C0B" blockId="54.[189,1398,537,599]" pageId="54" pageNumber="55">
<collectingCountry id="F32958E92F5BFFE102E2FDA8E11F2C6F" box="[189,277,537,563]" name="India" pageId="54" pageNumber="55">INDIA</collectingCountry>
: 
<emphasis id="B94AC46B2F5BFFE10341FDA8E1842C6F" bold="true" box="[286,398,537,563]" pageId="54" pageNumber="55">
<collectingRegion id="49FAD69B2F5BFFE10341FDA8E1842C6F" box="[286,398,537,563]" country="India" name="Manipur" pageId="54" pageNumber="55">Manipur</collectingRegion>
</emphasis>
: Karong, 
<date id="FF803EB92F5BFFE1005FFDA8E29C2C68" box="[512,662,537,564]" pageId="54" pageNumber="55" value="1950-10-26">26 Oct. 1950</date>
, 
<emphasis id="B94AC46B2F5BFFE100FFFDA8E33A2C6F" box="[672,816,537,563]" italics="true" pageId="54" pageNumber="55">Koelz 26739</emphasis>
(
<collectionCode id="ED2F80BC2F5BFFE10161FDA8E39A2C68" box="[830,912,537,564]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15602" lsid="urn:lsid:biocol.org:col:15602" name="University of Michigan" pageId="54" pageNumber="55" type="Herbarium">MICH</collectionCode>
1225781). 
<emphasis id="B94AC46B2F5BFFE10656FDA8E4882C6F" bold="true" box="[1033,1154,537,563]" pageId="54" pageNumber="55">
<collectingRegion id="49FAD69B2F5BFFE10656FDA8E4882C6F" box="[1033,1154,537,563]" country="India" name="Nagaland" pageId="54" pageNumber="55">Nagaland</collectingRegion>
</emphasis>
: Naga Hills, Kojima, 
<date id="FF803EB92F5BFFE102E2FD8DE1522C0B" box="[189,344,572,599]" pageId="54" pageNumber="55" value="1885-10-26">26 Oct. 1885</date>
, 
<emphasis id="B94AC46B2F5BFFE1033BFD8DE21F2C0A" box="[356,533,572,598]" italics="true" pageId="54" pageNumber="55">Clarke 41163A</emphasis>
(
<collectionCode id="ED2F80BC2F5BFFE1007AFD8CE2342C0B" box="[549,574,573,599]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" pageId="54" pageNumber="55" type="Herbarium">K</collectionCode>
); Naga Hills, Zakhama, 
<date id="FF803EB92F5BFFE10102FD8CE3F22C0B" box="[861,1016,573,599]" pageId="54" pageNumber="55" value="1949-10-22">22 Oct. 1949</date>
, 
<emphasis id="B94AC46B2F5BFFE1065BFD8DE4982C0B" box="[1028,1170,572,599]" italics="true" pageId="54" pageNumber="55">Ward 18912</emphasis>
(
<collectionCode id="ED2F80BC2F5BFFE106FDFD8CE4DF2C0B" box="[1186,1237,573,599]" country="United Kingdom" name="Bristol Museum" pageId="54" pageNumber="55">BM</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="C3244BF22F5BFFE102E2FDCFE40E2BDB" pageId="54" pageNumber="55" type="description">
<paragraph id="8B8118792F5BFFE102E2FDCFE1442CC4" blockId="54.[189,334,638,664]" box="[189,334,638,664]" pageId="54" pageNumber="55">
<heading id="D0C9AF152F5BFFE102E2FDCFE1442CC4" bold="true" box="[189,334,638,664]" fontSize="11" level="3" pageId="54" pageNumber="55" reason="3">
<emphasis id="B94AC46B2F5BFFE102E2FDCFE1442CC4" bold="true" box="[189,334,638,664]" pageId="54" pageNumber="55">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8B8118792F5BFFE102E2FD1DE40E2BDB" blockId="54.[189,1399,684,1415]" pageId="54" pageNumber="55">
Rhizomatous, monoecious herb, 
<quantity id="4CC6B59C2F5BFFE10010FD1DE2C52C9B" box="[591,719,684,711]" metricMagnitude="-1" metricUnit="m" metricValue="5.75" metricValueMax="9.0" metricValueMin="2.5" pageId="54" pageNumber="55" unit="cm" value="57.5" valueMax="90.0" valueMin="25.0">25–90 cm</quantity>
high. Rhizome: 
<quantity id="4CC6B59C2F5BFFE101FCFD1DE4202C9B" box="[931,1066,684,711]" metricMagnitude="-2" metricUnit="m" metricValue="1.25" metricValueMax="1.5" metricValueMin="1.0" pageId="54" pageNumber="55" unit="mm" value="12.5" valueMax="15.0" valueMin="10.0">10–15 mm</quantity>
wide, glabrous, internodes 
<quantity id="4CC6B59C2F5BFFE102E2FD7EE1492CB6" box="[189,323,719,746]" metricMagnitude="-2" metricUnit="m" metricValue="1.75" metricValueMax="2.0" metricValueMin="1.5" pageId="54" pageNumber="55" unit="mm" value="17.5" valueMax="20.0" valueMin="15.0">15 –20 mm</quantity>
long. Stipules: ovate, 10–19 × 
<quantity id="4CC6B59C2F5BFFE100E1FD61E33F2CB6" box="[702,821,720,746]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" metricValueMax="1.2" metricValueMin="0.8" pageId="54" pageNumber="55" unit="mm" value="10.0" valueMax="12.0" valueMin="8.0">8–12 mm</quantity>
, glabrous, deciduous. Leaves: petiole 
<quantity id="4CC6B59C2F5BFFE106A4FD61E57D2CB6" box="[1275,1399,719,746]" metricMagnitude="-1" metricUnit="m" metricValue="4.0" metricValueMax="5.0" metricValueMin="3.0" pageId="54" pageNumber="55" unit="cm" value="40.0" valueMax="50.0" valueMin="30.0">30–50 cm</quantity>
long, sparsely puberulous; lamina ovate to broadly ovate, basifixed, cordate, 20–40(–50) × 
<quantity id="4CC6B59C2F5BFFE106AAFD42E5782D51" box="[1269,1394,755,781]" metricMagnitude="-1" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="54" pageNumber="55" unit="cm" value="25.0" valueMax="30.0" valueMin="20.0">20–30 cm</quantity>
, asymmetric, upper surface green, sparsely strigose all over, denser near the margin, underside green to pale green, sparsely strigose, denser on the veins, venation palmate, midrib 16 –22(– 40) cm long; margin deeply lobed, with ca 6 major lobes with further smaller lobelets, irregularly serrulate, with sparse hairs; apex acuminate. Inflorescences: cymose, axillary or terminal, few; peduncle sparsely puberulous, branching 3–4 times, primary peduncle 
<quantity id="4CC6B59C2F5BFFE100D2FC12E30F2DE1" box="[653,773,931,957]" metricMagnitude="-1" metricUnit="m" metricValue="4.5" metricValueMax="6.0" metricValueMin="3.0" pageId="54" pageNumber="55" unit="cm" value="45.0" valueMax="60.0" valueMin="30.0">30–60 cm</quantity>
long, secondary 
<quantity id="4CC6B59C2F5BFFE10198FC12E43B2DE1" box="[967,1073,931,958]" metricMagnitude="-2" metricUnit="m" metricValue="7.5" metricValueMax="10.0" metricValueMin="5.0" pageId="54" pageNumber="55" unit="cm" value="7.5" valueMax="10.0" valueMin="5.0">5–10 cm</quantity>
, tertiary 
<quantity id="4CC6B59C2F5BFFE106CBFC12E4FA2DE1" box="[1172,1264,931,957]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" metricValueMax="3.0" metricValueMin="1.0" pageId="54" pageNumber="55" unit="cm" value="2.0" valueMax="3.0" valueMin="1.0">1–3 cm</quantity>
, quaternary ca 
<quantity id="4CC6B59C2F5BFFE10280FC77E1112DBC" box="[223,283,966,992]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="54" pageNumber="55" unit="cm" value="1.0">1 cm</quantity>
, with 8– 
<specimenCount id="9D38D3F02F5BFFE103DEFC77E1FD2DBC" box="[385,503,966,992]" pageId="54" pageNumber="55" type="female">12 female</specimenCount>
and 2– 
<specimenCount id="9D38D3F02F5BFFE1000EFC77E2AB2DBC" box="[593,673,966,992]" pageId="54" pageNumber="55" type="male">6 male</specimenCount>
flowers; bracts ovate-orbicular, 9–15 × 
<quantity id="4CC6B59C2F5BFFE10627FC77E4E72DBC" box="[1144,1261,966,992]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" metricValueMax="11.0" metricValueMin="6.0" pageId="54" pageNumber="55" unit="mm" value="8.5" valueMax="11.0" valueMin="6.0">6–11 mm</quantity>
, deciduous. Male flower: pedicel 
<quantity id="4CC6B59C2F5BFFE103E5FC58E24A2A58" box="[442,576,1001,1028]" metricMagnitude="-2" metricUnit="m" metricValue="1.6" metricValueMax="2.0" metricValueMin="1.2" pageId="54" pageNumber="55" unit="mm" value="16.0" valueMax="20.0" valueMin="12.0">12–20 mm</quantity>
long, glabrous; tepals 4, outer tepals ovate, 10–14 × 
<quantity id="4CC6B59C2F5BFFE106E4FC58E5252A58" box="[1211,1327,1001,1028]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" metricValueMax="10.0" metricValueMin="7.0" pageId="54" pageNumber="55" unit="mm" value="8.5" valueMax="10.0" valueMin="7.0">7–10 mm</quantity>
, pink, glabrous, margin entire; inner tepals oblong to narrowly obovate, 7–14 × 
<quantity id="4CC6B59C2F5BFFE10619FBBDE4BE2A7B" box="[1094,1204,1036,1063]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" metricValueMax="9.0" metricValueMin="5.0" pageId="54" pageNumber="55" unit="mm" value="7.0" valueMax="9.0" valueMin="5.0">5–9 mm</quantity>
, pink, glabrous, entire; androecium with 60–90 stamens, symmetric; filaments 
<quantity id="4CC6B59C2F5BFFE101F0FB81E41C2A16" box="[943,1046,1072,1098]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="54" pageNumber="55" unit="mm" value="1.5" valueMax="2.0" valueMin="1.0">1–2 mm</quantity>
long, unequal, fused at base; anther oblong-cuneate to oblong-elliptic, 
<quantity id="4CC6B59C2F5BFFE100E4FBE2E3332A31" box="[699,825,1107,1134]" metricMagnitude="-3" metricUnit="m" metricValue="1.25" metricValueMax="1.5" metricValueMin="1.0" pageId="54" pageNumber="55" unit="mm" value="1.25" valueMax="1.5" valueMin="1.0">1–1.5 mm</quantity>
long, dehiscing through slits longer than half the length of the anther, not hooded, connective extended. Female flower: pedicel 
<quantity id="4CC6B59C2F5BFFE106F1FBC7E53A2ACC" box="[1198,1328,1142,1169]" metricMagnitude="-2" metricUnit="m" metricValue="1.35" metricValueMax="1.5" metricValueMin="1.2" pageId="54" pageNumber="55" unit="mm" value="13.5" valueMax="15.0" valueMin="12.0">12–15 mm</quantity>
long, glabrous; bracteoles present; tepals (4–)6, equal, ovate to elliptic, outer tepals 10–15 × 
<quantity id="4CC6B59C2F5BFFE106A6FB28E5782AE8" box="[1273,1394,1177,1204]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" metricValueMax="11.0" metricValueMin="8.0" pageId="54" pageNumber="55" unit="mm" value="9.5" valueMax="11.0" valueMin="8.0">8–11 mm</quantity>
, pale pink, glabrous, margin entire, inner tepals similar but smaller; ovary 2-locular, placentae bifid; capsule ellipsoid, 6–8 × 
<quantity id="4CC6B59C2F5BFFE10382FB51E24C2AA6" box="[477,582,1247,1274]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" metricValueMax="6.0" metricValueMin="3.0" pageId="54" pageNumber="55" unit="mm" value="4.5" valueMax="6.0" valueMin="3.0">3–6 mm</quantity>
, strigose along the base of the main wing otherwise glabrous, with one long oblong wing and two short triangular wings; styles 3, convoluted with ends twisted twice. Fruit: pedicel 
<quantity id="4CC6B59C2F5BFFE10347FA97E1962B1C" box="[280,412,1318,1345]" metricMagnitude="-2" metricUnit="m" metricValue="2.25" metricValueMax="2.5" metricValueMin="2.0" pageId="54" pageNumber="55" unit="mm" value="22.5" valueMax="25.0" valueMin="20.0">20 –25 mm</quantity>
long, recurved; capsule ellipsoid, 12–15 × 
<quantity id="4CC6B59C2F5BFFE101CCFA97E3F72B1C" box="[915,1021,1318,1344]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" metricValueMax="6.0" metricValueMin="4.0" pageId="54" pageNumber="55" unit="mm" value="5.0" valueMax="6.0" valueMin="4.0">4–6 mm</quantity>
, strigose near base of main wing otherwise glabrous; wings extending along the pedicel slightly, unequal; longest wing oblong-elliptic, 10 –30 × 
<quantity id="4CC6B59C2F5BFFE10373FADDE1A82BDB" box="[300,418,1388,1415]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" metricValueMax="1.5" metricValueMin="0.5" pageId="54" pageNumber="55" unit="mm" value="10.0" valueMax="15.0" valueMin="5.0">5–15 mm</quantity>
; shortest wings triangular-oblong, 3–5 × 
<quantity id="4CC6B59C2F5BFFE101D5FADDE3F52BDB" box="[906,1023,1388,1415]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" metricValueMax="1.1" metricValueMin="0.9" pageId="54" pageNumber="55" unit="mm" value="10.0" valueMax="11.0" valueMin="9.0">9–11 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="C3244BF22F5BFFE102E2FA1FE4BE28D5" pageId="54" pageNumber="55" type="distribution">
<paragraph id="8B8118792F5BFFE102E2FA1FE21C2B94" blockId="54.[189,534,1454,1480]" box="[189,534,1454,1480]" pageId="54" pageNumber="55">
<heading id="D0C9AF152F5BFFE102E2FA1FE21C2B94" bold="true" box="[189,534,1454,1480]" fontSize="11" level="3" pageId="54" pageNumber="55" reason="3">
<emphasis id="B94AC46B2F5BFFE102E2FA1FE21C2B94" bold="true" box="[189,534,1454,1480]" pageId="54" pageNumber="55">Distribution and phenology</emphasis>
</heading>
</paragraph>
<paragraph id="8B8118792F5BFFE102E2FA6DE3362845" blockId="54.[189,1399,1500,1562]" pageId="54" pageNumber="55">
Endemic to the 
<collectingRegion id="49FAD69B2F5BFFE10331FA6DE1CE2BAA" box="[366,452,1500,1526]" country="Myanmar" name="Rakhine" pageId="54" pageNumber="55">Arakan</collectingRegion>
Mountain Range, usually found growing on cliff faces; 
<quantity id="4CC6B59C2F5BFFE1061CFA6DE4E62BAB" box="[1091,1260,1500,1527]" metricMagnitude="3" metricUnit="m" metricValue="1.55" metricValueMax="2.1" metricValueMin="1.0" pageId="54" pageNumber="55" unit="m" value="1550.0" valueMax="2100.0" valueMin="1000.0">1000–2100 m</quantity>
. Flowering: September to October; fruiting: October to November.
</paragraph>
<paragraph id="8B8118792F5BFFE102E2F9F0E1BC2807" blockId="54.[189,438,1601,1627]" box="[189,438,1601,1627]" pageId="54" pageNumber="55">
<heading id="D0C9AF152F5BFFE102E2F9F0E1BC2807" bold="true" box="[189,438,1601,1627]" fontSize="11" level="3" pageId="54" pageNumber="55" reason="3">
<emphasis id="B94AC46B2F5BFFE102E2F9F0E1BC2807" bold="true" box="[189,438,1601,1627]" pageId="54" pageNumber="55">Conservation status</emphasis>
</heading>
</paragraph>
<paragraph id="8B8118792F5BFFE102E2F9C1E4BE28D5" blockId="54.[189,1204,1647,1674]" box="[189,1204,1647,1674]" pageId="54" pageNumber="55">
Data Deficient. The full distribution of 
<taxonomicName id="4C3E63FA2F5BFFE100D6F9C1E2FD28D5" authorityName="R. Camfield" authorityYear="2018" box="[649,759,1647,1673]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="54" pageNumber="55" phylum="Tracheophyta" rank="species" species="koelzii">
<emphasis id="B94AC46B2F5BFFE100D6F9C1E2FD28D5" box="[649,759,1647,1673]" italics="true" pageId="54" pageNumber="55">B. koelzii</emphasis>
</taxonomicName>
in the 
<collectingRegion id="49FAD69B2F5BFFE1011AF9DEE39128D5" box="[837,923,1647,1673]" country="Myanmar" name="Rakhine" pageId="54" pageNumber="55">Arakan</collectingRegion>
mountains is unknown.
</paragraph>
</subSubSection>
<subSubSection id="C3244BF22F5BFFE002E2F900E3EE2C90" lastPageId="55" lastPageNumber="56" pageId="54" pageNumber="55" type="discussion">
<paragraph id="8B8118792F5BFFE102E2F900E1252897" blockId="54.[189,303,1713,1739]" box="[189,303,1713,1739]" pageId="54" pageNumber="55">
<heading id="D0C9AF152F5BFFE102E2F900E1252897" bold="true" box="[189,303,1713,1739]" fontSize="11" level="3" pageId="54" pageNumber="55" reason="3">
<emphasis id="B94AC46B2F5BFFE102E2F900E1252897" bold="true" box="[189,303,1713,1739]" pageId="54" pageNumber="55">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8B8118792F5BFFE102E2F96EE3CE29DA" blockId="54.[189,1399,1759,1927]" pageId="54" pageNumber="55">
The female flowers on the material we have seen of this species had 3 styles; allied species can have either 2 or 3 styles; hence we consider there is the potential for 
<taxonomicName id="4C3E63FA2F5BFFE101E0F8B5E4242941" authorityName="R. Camfield" authorityYear="2018" box="[959,1070,1795,1821]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="54" pageNumber="55" phylum="Tracheophyta" rank="species" species="koelzii">
<emphasis id="B94AC46B2F5BFFE101E0F8B5E4242941" box="[959,1070,1795,1821]" italics="true" pageId="54" pageNumber="55">B. koelzii</emphasis>
</taxonomicName>
to also be polymorphic for style number, which may be revealed given more material. The size of the female flowers is based on measurements from unopened buds, and we would expect the mature flowers to be larger. The bulbils are seen on larger, older leaves and are quite commonly observed.
</paragraph>
<paragraph id="8B8118792F5AFFE002E2FEBAE3EE2C90" blockId="55.[189,1399,267,716]" pageId="55" pageNumber="56">
This species is allied to several others which share lacerate leaves; 
<taxonomicName id="4C3E63FA2F5AFFE00644FEBDE1E42F15" authority="S. Rajbh. (Rajbhandary et al. 2010)" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="55" pageNumber="56" phylum="Tracheophyta" rank="species" species="panchatharensis">
<emphasis id="B94AC46B2F5AFFE00644FEBDE5092F79" box="[1051,1283,267,293]" italics="true" pageId="55" pageNumber="56">B. panchatharensis</emphasis>
S.Rajbh. (
<bibRefCitation id="EFAF65882F5AFFE0029AFE9FE1EF2F15" author="Rajbhandary S. &amp; Hughes M. &amp; Shrestha K. K." box="[197,485,302,329]" pageId="55" pageNumber="56" pagination="151 - 162" refId="ref57152" refString="Rajbhandary S., Hughes M. &amp; Shrestha K. K. 2010. Three new species of Begonia sect. Platycentrum from Nepal. Gardens' Bulletin Singapore 62 (1): 151 - 162." type="journal article" year="2010">
Rajbhandary 
<emphasis id="B94AC46B2F5AFFE0033AFE9EE1902F15" box="[357,410,303,329]" italics="true" pageId="55" pageNumber="56">et al</emphasis>
. 2010
</bibRefCitation>
)
</taxonomicName>
from 
<collectingCountry id="F32958E92F5AFFE00067FE9FE2742F14" box="[568,638,302,328]" name="Nepal" pageId="55" pageNumber="56">Nepal</collectingCountry>
and 
<collectingCountry id="F32958E92F5AFFE000E6FE9FE31B2F14" box="[697,785,302,328]" name="Bhutan" pageId="55" pageNumber="56">Bhutan</collectingCountry>
, 
<taxonomicName id="4C3E63FA2F5AFFE00141FE9EE4C62F15" authority="H. Lev. (Leveille 1909)" authorityName="H. Lev. (Leveille" authorityYear="1909" box="[798,1228,302,329]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="55" pageNumber="56" phylum="Tracheophyta" rank="species" species="pedatifida">
<emphasis id="B94AC46B2F5AFFE00141FE9EE3BF2F15" box="[798,949,303,329]" italics="true" pageId="55" pageNumber="56">B. pedatifida</emphasis>
H.Lév. (
<bibRefCitation id="EFAF65882F5AFFE00641FE9FE4CE2F15" author="Leveille H." box="[1054,1220,302,329]" pageId="55" pageNumber="56" pagination="20 - 23" refId="ref56059" refString="Leveille H. 1909. Decades plantarum novarum XVI. Repertorium Specierum Novarum Regni Vegetabilis 7: 20 - 23. Available from http: // biodiversitylibrary. org / page / 265226 [accessed May 2009]." type="journal article" year="1909">Léveillé 1909</bibRefCitation>
)
</taxonomicName>
from Yunnan, 
<taxonomicName id="4C3E63FA2F5AFFE002E2FEE2E1532F30" authorityName="Rekha Morris &amp; P. D. McMillan (Morris &amp; McMillan" authorityYear="2012" box="[189,345,338,364]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="55" pageNumber="56" phylum="Tracheophyta" rank="species" species="shilendrae">
<emphasis id="B94AC46B2F5AFFE002E2FEE2E1532F30" box="[189,345,338,364]" italics="true" pageId="55" pageNumber="56">B. shilendrae</emphasis>
</taxonomicName>
from Arunachal-Pradesh, 
<taxonomicName id="4C3E63FA2F5AFFE000D2FEE2E57C2F30" authority="S. H. Huang &amp; Y. M. Shui (Huang &amp; Shui 1994)" authorityName="S. H. Huang &amp; Y. M. Shui (Huang &amp; Shui" authorityYear="1994" box="[653,1398,338,364]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="55" pageNumber="56" phylum="Tracheophyta" rank="species" species="rubropunctata">
<emphasis id="B94AC46B2F5AFFE000D2FEE2E35F2F30" box="[653,853,338,364]" italics="true" pageId="55" pageNumber="56">B. rubropunctata</emphasis>
S.H.Huang &amp; Y.M.Shui (
<bibRefCitation id="EFAF65882F5AFFE006DEFEE3E5642F30" author="Huang S. H. &amp; Shui Y. M." box="[1153,1390,338,364]" pageId="55" pageNumber="56" pagination="333 - 342" refId="ref54883" refString="Huang S. H. &amp; Shui Y. M. 1994. New taxa of Begonia from Yunnan. Acta Botanica Yunnanica 16 (4): 333 - 342." type="journal article" year="1994">Huang &amp; Shui 1994</bibRefCitation>
)
</taxonomicName>
from Yunnan, 
<taxonomicName id="4C3E63FA2F5AFFE00331FEC7E3402FCC" authority="Irmsch. (Irmscher 1951)" authorityName="Irmsch. (Irmscher" authorityYear="1951" box="[366,842,373,400]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="55" pageNumber="56" phylum="Tracheophyta" rank="species" species="macrotoma">
<emphasis id="B94AC46B2F5AFFE00331FEC7E2122FD3" box="[366,536,374,399]" italics="true" pageId="55" pageNumber="56">B. macrotoma</emphasis>
Irmsch. (
<bibRefCitation id="EFAF65882F5AFFE000D0FEC4E3482FCC" author="Irmscher E." box="[655,834,373,400]" pageId="55" pageNumber="56" pagination="35 - 45" refId="ref55254" refString="Irmscher E. 1951. Some new Chinese species of Begonia. Notes from the Royal Botanic Gardens Edinburgh 21: 35 - 45." type="journal article" year="1951">Irmscher 1951</bibRefCitation>
)
</taxonomicName>
from Yunnan and 
<taxonomicName id="4C3E63FA2F5AFFE0066AFEC7E23B2FEE" authority="K. Y. Guan &amp; D. K. Tian (Guan &amp; Tian 2000)" authorityName="K. Y. Guan &amp; D. K. Tian (Guan &amp; Tian" authorityYear="2000" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="55" pageNumber="56" phylum="Tracheophyta" rank="species" species="longialata">
<emphasis id="B94AC46B2F5AFFE0066AFEC7E4D92FD3" box="[1077,1235,373,399]" italics="true" pageId="55" pageNumber="56">B. longialata</emphasis>
K.Y.Guan &amp; D.K.Tian (
<bibRefCitation id="EFAF65882F5AFFE00361FE29E2232FEE" author="Guan K. Y. &amp; Tian D. K." box="[318,553,408,434]" pageId="55" pageNumber="56" pagination="129 - 134" refId="ref54214" refString="Guan K. Y. &amp; Tian D. K. 2000. Three new species of Begonia from Yunnan. Acta Botanica Yunnanica 22 (2): 129 - 134." type="journal article" year="2000">Guan &amp; Tian 2000</bibRefCitation>
)
</taxonomicName>
from Yunnan. 
<taxonomicName id="4C3E63FA2F5AFFE000B3FE28E3AC2FEE" authorityName="R. Camfield" authorityYear="2018" box="[748,934,408,434]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="55" pageNumber="56" phylum="Tracheophyta" rank="species" species="koelzii">
<emphasis id="B94AC46B2F5AFFE000B3FE28E3AC2FEE" box="[748,934,408,434]" italics="true" pageId="55" pageNumber="56">Begonia koelzii</emphasis>
</taxonomicName>
is distinct from these in a number of vegetative and floral characters (
<tableCitation id="C6BC2DC22F5AFFE00063FE0AE29F2F89" box="[572,661,443,469]" captionStart="Table 1" captionStartId="53.[183,252,558,586]" captionText="Table 1. Comparison of the key vegetative and floral characters of the new species B. koelzii R.Camfield sp. nov. and several similar species. Data collected from herbarium specimens at E, HAST and K, species protologues and the Flora of China (Gu et al. 2007)." pageId="55" pageNumber="56">Table 1</tableCitation>
). 
<taxonomicName id="4C3E63FA2F5AFFE000F1FE0DE3972F89" authorityName="Irmsch. (Irmscher" authorityYear="1951" box="[686,925,444,469]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="55" pageNumber="56" phylum="Tracheophyta" rank="species" species="macrotoma">
<emphasis id="B94AC46B2F5AFFE000F1FE0DE3972F89" box="[686,925,444,469]" italics="true" pageId="55" pageNumber="56">Begonia macrotoma</emphasis>
</taxonomicName>
is recorded from Northeast 
<collectingCountry id="F32958E92F5AFFE006B6FE0AE52F2F89" box="[1257,1317,443,469]" name="India" pageId="55" pageNumber="56">India</collectingCountry>
by 
<bibRefCitation id="EFAF65882F5AFFE0070CFE0AE1592FA5" author="Gu C. &amp; Peng C. &amp; Turland N. J." pageId="55" pageNumber="56" pagination="153 - 207" refId="ref54157" refString="Gu C., Peng C. - I &amp; Turland N. J. 2007. Begoniaceae. In: Wu Z. Y., Raven P. H. &amp; Hong D. Y. (eds) Flora of China 13: 153 - 207. Missouri Botanical Garden Press, Missouri." type="journal article" year="2007">
Gu 
<emphasis id="B94AC46B2F5AFFE002E2FE6EE0FE2FA5" box="[189,244,479,505]" italics="true" pageId="55" pageNumber="56">et al</emphasis>
. (2007)
</bibRefCitation>
but no specimens were seen during this revision to confirm this. 
<taxonomicName id="4C3E63FA2F5AFFE0062BFE6EE5532FA5" authorityName="K. Y. Guan &amp; D. K. Tian (Guan &amp; Tian" authorityYear="2000" box="[1140,1369,479,505]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="55" pageNumber="56" phylum="Tracheophyta" rank="species" species="longialata">
<emphasis id="B94AC46B2F5AFFE0062BFE6EE5532FA5" box="[1140,1369,479,505]" italics="true" pageId="55" pageNumber="56">Begonia longialata</emphasis>
</taxonomicName>
is reported by 
<bibRefCitation id="EFAF65882F5AFFE00314FDB3E20C2C40" author="Gu C. &amp; Peng C. &amp; Turland N. J." box="[331,518,514,540]" pageId="55" pageNumber="56" pagination="153 - 207" refId="ref54157" refString="Gu C., Peng C. - I &amp; Turland N. J. 2007. Begoniaceae. In: Wu Z. Y., Raven P. H. &amp; Hong D. Y. (eds) Flora of China 13: 153 - 207. Missouri Botanical Garden Press, Missouri." type="journal article" year="2007">
Gu 
<emphasis id="B94AC46B2F5AFFE00329FDB2E1A32C40" box="[374,425,514,540]" italics="true" pageId="55" pageNumber="56">et al</emphasis>
. (2007)
</bibRefCitation>
as endemic to Yunnan; however, photographs have been taken in 
<collectingRegion id="49FAD69B2F5AFFE00754FDB3E5782C40" box="[1291,1394,514,540]" country="India" name="Manipur" pageId="55" pageNumber="56">Manipur</collectingRegion>
, on route to Khayang from Ukhrul (
<bibRefCitation id="EFAF65882F5AFFE00004FD94E2E12C63" author="Macer N." box="[603,747,549,575]" pageId="55" pageNumber="56" refId="ref56268" refString="Macer N. 2013. No Whiteys since Kingdon-Ward - Planthunting in darkest Manipur, November 2012. Available from http: // www. panglobalplants. com / 2013 / 02 / 13 / no-whiteys-since-kingdon-ward-planthunting-indarkest-manipur / [accessed Nov. 2014]." type="url" year="2013">Macer 2013</bibRefCitation>
) of a caulescent plant with lacerate leaves which could potentially represent it in our study region; however, no herbarium material has been seen. 
<taxonomicName id="4C3E63FA2F5AFFE0074AFDF8E1332CD9" authorityName="Rekha Morris &amp; P. D. McMillan (Morris &amp; McMillan" authorityYear="2012" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="55" pageNumber="56" phylum="Tracheophyta" rank="species" species="shilendrae">
<emphasis id="B94AC46B2F5AFFE0074AFDF8E1332CD9" italics="true" pageId="55" pageNumber="56">Begonia shilendrae</emphasis>
</taxonomicName>
is the only other allied species currently known to occur within the study area, but their distributions do not overlap; 
<taxonomicName id="4C3E63FA2F5AFFE00055FD3EE2AF2CF5" authorityName="Rekha Morris &amp; P. D. McMillan (Morris &amp; McMillan" authorityYear="2012" box="[522,677,655,681]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="55" pageNumber="56" phylum="Tracheophyta" rank="species" species="shilendrae">
<emphasis id="B94AC46B2F5AFFE00055FD3EE2AF2CF5" box="[522,677,655,681]" italics="true" pageId="55" pageNumber="56">B. shilendrae</emphasis>
</taxonomicName>
is known only from the 
<typeStatus id="5485A6DB2F5AFFE001E2FD3EE3FA2CF5" box="[957,1008,655,681]" pageId="55" pageNumber="56">type</typeStatus>
locality of Itanagar in Arunachal- Pradesh, whereas 
<taxonomicName id="4C3E63FA2F5AFFE003CFFD02E1F42C90" authorityName="R. Camfield" authorityYear="2018" box="[400,510,690,716]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="55" pageNumber="56" phylum="Tracheophyta" rank="species" species="koelzii">
<emphasis id="B94AC46B2F5AFFE003CFFD02E1F42C90" box="[400,510,690,716]" italics="true" pageId="55" pageNumber="56">B. koelzii</emphasis>
</taxonomicName>
is recorded from 
<collectingRegion id="49FAD69B2F5AFFE00091FD03E3392C90" box="[718,819,690,716]" country="India" name="Manipur" pageId="55" pageNumber="56">Manipur</collectingRegion>
and 
<collectingRegion id="49FAD69B2F5AFFE00133FD03E3EA2C90" box="[876,992,690,716]" country="India" name="Nagaland" pageId="55" pageNumber="56">Nagaland</collectingRegion>
.
</paragraph>
</subSubSection>
</treatment>
</document>