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<document id="2270D36CB2FA9E4FE2A03CE6BE4D710F" ID-CLB-Dataset="5852" ID-DOI="10.1206/3867.1" ID-GBIF-Dataset="1d6299e9-5c62-4abd-be37-59efcb217cbf" ID-ISSN="0003-0082" ID-Zenodo-Dep="4598386" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1615481346733" checkinUser="felipe" docAuthor="Velazco, Paúl M. &amp; Novacek, Michael J." docDate="2016" docId="0386879BFB149C68FE47353A8B73FA3A" docLanguage="en" docName="AmMusNovit.2016.3867.1-42.pdf" docOrigin="American Museum Novitates 2016 (3867)" docSource="http://www.bioone.org/doi/10.1206/3867.1" docStyle="DocumentStyle:26C3580A17D4BBE58BD8D8AC6AE09AC4.6:AmMusNovit.2010-.journal_article" docStyleId="26C3580A17D4BBE58BD8D8AC6AE09AC4" docStyleName="AmMusNovit.2010-.journal_article" docStyleVersion="6" docTitle="Palaeictops matthewi Novacek 1977" docType="treatment" docVersion="6" lastPageNumber="17" masterDocId="FFBFFFE3FB049C78FF8C346B881CFF9C" masterDocTitle="Systematics of the genus Palaeictops Matthew, 1899 (Mammalia: Leptictidae), with the description of two new species from the Middle Eocene of Utah and Wyoming" masterLastPageNumber="42" masterPageNumber="1" pageNumber="17" updateTime="1698934683797" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="E3B93CFA0019D11DF6965DEC0CEE9562">Systematics of the genus Palaeictops Matthew, 1899 (Mammalia: Leptictidae), with the description of two new species from the Middle Eocene of Utah and Wyoming</mods:title>
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<mods:namePart id="DFAD9142C0CEB0272E5CC80E50A17657">Velazco, Paúl M.</mods:namePart>
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<treatment id="0386879BFB149C68FE47353A8B73FA3A" ID-DOI="http://doi.org/10.5281/zenodo.5056595" ID-GBIF-Taxon="183232359" ID-Zenodo-Dep="5056595" LSID="urn:lsid:plazi:treatment:0386879BFB149C68FE47353A8B73FA3A" httpUri="http://treatment.plazi.org/id/0386879BFB149C68FE47353A8B73FA3A" lastPageNumber="17" pageId="16" pageNumber="17">
<subSubSection id="C3356506FB149C68FE47353A8B68FEF7" box="[459,884,337,363]" pageId="16" pageNumber="17" type="nomenclature">
<paragraph id="8B90368DFB149C68FE47353A8B68FEF7" blockId="16.[120,1228,225,363]" box="[459,884,337,363]" pageId="16" pageNumber="17">
<taxonomicName id="4C2F4D0EFB149C68FE47353A8B68FEF7" authority="Novacek, 1977" authorityName="Novacek" authorityYear="1977" box="[459,884,337,363]" class="Mammalia" family="Leptictidae" genus="Palaeictops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pilosa" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="matthewi">
<emphasis id="B95BEA9FFB149C68FE47353A8ADCFEF7" box="[459,704,337,363]" italics="true" pageId="16" pageNumber="17">Palaeictops matthewi</emphasis>
<bibRefCitation id="EFBE4B7CFB149C68FD44353A8B68FEF7" author="Novacek, M. J." box="[712,884,337,363]" pageId="16" pageNumber="17" pagination="1 - 42" refId="ref15221" refString="Novacek, M. J. 1977. A review of Paleocene and Eocene Leptictidae (Eutheria, Mammalia) from North America. PaleoBios 24: 1 - 42." type="journal article" year="1977">Novacek, 1977</bibRefCitation>
</taxonomicName>
</paragraph>
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<subSubSection id="C3356506FB149C68FDA735E28B09FE3F" box="[555,789,393,419]" pageId="16" pageNumber="17" type="description">
<paragraph id="8B90368DFB149C68FDA735E28B09FE3F" blockId="16.[555,789,393,419]" box="[555,789,393,419]" pageId="16" pageNumber="17">
<heading id="D0D881E1FB149C68FDA735E28B09FE3F" box="[555,789,393,419]" centered="true" fontSize="36" level="3" pageId="16" pageNumber="17" reason="7">
<figureCitation id="13142A08FB149C68FDA735E28AA6FE3F" box="[555,698,393,419]" captionStart-0="FIG" captionStart-1="FIG" captionStart-2="FIG" captionStartId-0="18.[121,161,1272,1293]" captionStartId-1="19.[120,160,840,861]" captionStartId-2="20.[121,162,837,858]" captionTargetBox-0="[120,1224,224,1246]" captionTargetBox-1="[319,1025,224,812]" captionTargetBox-2="[316,1029,224,810]" captionTargetId-0="figure-97@18.[120,1224,224,1246]" captionTargetId-1="figure-344@19.[319,1025,224,812]" captionTargetId-2="figure-373@20.[316,1029,224,810]" captionTargetPageId-0="18" captionTargetPageId-1="19" captionTargetPageId-2="20" captionText-0="FIG.8. The skull of Palaeictops matthewi FMNH P26904 (holotype), containing P2–P5 (broken), M1–M2 (roots), and M3 (broken) on the left side; the right side retains roots of C–P1, P2 (broken), P4–P5, and M1–M3.A, (opposite page) dorsal (top) and ventral (bottom) views and B, (above) right lateral (top) and left lateral (bottom) views.Scale bar = 1 cm." captionText-1="FIG. 9. Right ramus of Palaeictops matthewi FMNH P26904 (holotype), containing c–p1 (broken), p2–p5, m1– m2 (broken), and root of i2, in (clockwise from upper left) medial, occlusal, and lateral views. Scale bar = 1 cm." captionText-2="FIG. 10. Left ramus of Palaeictops matthewi FMNH P26904 (holotype), containing c (broken), p1, p2 (broken), p4–p5, m1–m3 (broken), and root of i2, in (clockwise from upper left) medial, occlusal, and lateral views. Scale bar = 1 cm." figureDoi-0="http://doi.org/10.5281/zenodo.4598405" figureDoi-1="http://doi.org/10.5281/zenodo.4598407" figureDoi-2="http://doi.org/10.5281/zenodo.4598409" httpUri-0="https://zenodo.org/record/4598405/files/figure.png" httpUri-1="https://zenodo.org/record/4598407/files/figure.png" httpUri-2="https://zenodo.org/record/4598409/files/figure.png" pageId="16" pageNumber="17">Figure 8–10</figureCitation>
, table 1
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3356506FB149C68FF2435AA8B3AFD7D" pageId="16" pageNumber="17" type="materials_examined">
<paragraph id="8B90368DFB149C68FF2435AA8A65FD0A" blockId="16.[120,1226,449,1447]" pageId="16" pageNumber="17">
<typeStatus id="5494882FFB149C68FF2435AA8934FE47" box="[168,296,449,475]" pageId="16" pageNumber="17" type="holotype">HOLOTYPE</typeStatus>
:
<collectionCode id="ED3EAE48FB149C68FEBB35A98991FE40" box="[311,397,450,476]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="16" pageNumber="17" type="Museum">FMNH</collectionCode>
P26904 (fig. 8A–B), consisting of a damaged skull that retains P2–P5 (broken), M1–M2 (roots), and M3 (broken) on the left side; the right side retains roots of C– P1, P2 (broken), P4–P5, and M1–M3. Right ramus with c–p1 (broken), p2–p5, m1–m2 (broken), and root of i2 (fig. 9). Left ramus with c (broken), p1, p2 (broken), p4–p5, m1–m3 (broken), and root of i2 (fig. 10). Partial skeleton including lumbar and caudal vertebrae, pelvis, femur, tibia, carpals, tarsals, and phalanges.
</paragraph>
<paragraph id="8B90368DFB149C68FF2436C98B3AFD7D" blockId="16.[120,1226,449,1447]" pageId="16" pageNumber="17">
TYPE LOCALITY: Exposure near top of Buzzard Pass, Mesa Co.,
<collectingRegion id="49EBF86FFB149C68FCF336CA8BF3FD27" box="[895,1007,673,699]" country="United States of America" name="Colorado" pageId="16" pageNumber="17">Colorado</collectingRegion>
. De Beque Formation, Rifle Member, Lower Eocene (Wasatchian NALMA).
</paragraph>
</subSubSection>
<subSubSection id="C3356506FB149C68FF24368788E6FB97" pageId="16" pageNumber="17" type="diagnosis">
<paragraph id="8B90368DFB149C68FF24368788E6FB97" blockId="16.[120,1226,449,1447]" pageId="16" pageNumber="17">
DIAGNOSIS (modified from
<bibRefCitation id="EFBE4B7CFB149C68FE7A36878AB7FC9A" author="Novacek, M. J." box="[502,683,748,774]" pageId="16" pageNumber="17" pagination="1 - 42" refId="ref15221" refString="Novacek, M. J. 1977. A review of Paleocene and Eocene Leptictidae (Eutheria, Mammalia) from North America. PaleoBios 24: 1 - 42." type="journal article" year="1977">Novacek, 1977</bibRefCitation>
): Similar to
<taxonomicName id="4C2F4D0EFB149C68FCCC36878C4CFC9A" box="[832,1104,748,774]" class="Mammalia" family="Leptictidae" genus="Palaeictops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pilosa" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="multicuspis">
<emphasis id="B95BEA9FFB149C68FCCC36878C4CFC9A" box="[832,1104,748,774]" italics="true" pageId="16" pageNumber="17">Palaeictops multicuspis</emphasis>
</taxonomicName>
in having tall, trenchant p4 but differs from
<taxonomicName id="4C2F4D0EFB149C68FD9D37798AADFCB7" box="[529,689,785,811]" class="Mammalia" family="Leptictidae" genus="Palaeictops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pilosa" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="multicuspis">
<emphasis id="B95BEA9FFB149C68FD9D37798AADFCB7" box="[529,689,785,811]" italics="true" pageId="16" pageNumber="17">P. multicuspis</emphasis>
</taxonomicName>
in slightly smaller size and in having a more slender ramus; p2 with only one main, trenchant cusp and a basal posterior cuspule; p4 with anterior main cusp larger than posterior main cusp and a short transverse ridge on the posterior heel separated from the rest of the crown by a transverse trough (figs. 9, 10). P4 trenchant tooth, taller than P5 with a very prominent paracone, smaller metacone and low protocone (fig. 8B). Species has postcranial features used to diagnose
<taxonomicName id="4C2F4D0EFB149C68FC4D37A78C58FC7A" authorityName="Matthew" authorityYear="1899" box="[961,1092,972,998]" class="Mammalia" family="Leptictidae" genus="Palaeictops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pilosa" pageId="16" pageNumber="17" phylum="Chordata" rank="genus">
<emphasis id="B95BEA9FFB149C68FC4D37A78C58FC7A" box="[961,1092,972,998]" italics="true" pageId="16" pageNumber="17">Palaeictops</emphasis>
</taxonomicName>
(see above diagnosis).
</paragraph>
</subSubSection>
<subSubSection id="C3356506FB149C68FF24307C8BF5FB3D" pageId="16" pageNumber="17" type="distribution">
<paragraph id="8B90368DFB149C68FF24307C88EDFBCA" blockId="16.[120,1226,449,1447]" pageId="16" pageNumber="17">
DISTRIBUTION: De Beque and Huerfano formations,
<collectingRegion id="49EBF86FFB149C68FC9C307C8B9CFBAD" box="[784,896,1047,1073]" country="United States of America" name="Colorado" pageId="16" pageNumber="17">Colorado</collectingRegion>
, Lower Eocene (Wasatchian NALMA).
</paragraph>
<paragraph id="8B90368DFB149C68FF2430098BF5FB3D" blockId="16.[120,1226,449,1447]" pageId="16" pageNumber="17">
REFERRED SPECIMENS: AMNH 17555, skull fragment, partial lower jaws and partial skeleton. Garcia Canon region,
<collectingRegion id="49EBF86FFB149C68FE4D30EC8A2EFB3D" box="[449,562,1159,1185]" country="United States of America" name="Colorado" pageId="16" pageNumber="17">Colorado</collectingRegion>
. Lower beds of Huerfano Formation.
</paragraph>
</subSubSection>
<subSubSection id="C3356506FB149C68FF2430C78B73FA3A" pageId="16" pageNumber="17" type="discussion">
<paragraph id="8B90368DFB149C68FF2430C78B73FA3A" blockId="16.[120,1226,449,1447]" pageId="16" pageNumber="17">
REMARKS:
<bibRefCitation id="EFBE4B7CFB149C68FEAB30C68AD8FB5A" author="Bown, T. M. &amp; D. Schankler" box="[295,708,1196,1223]" pageId="16" pageNumber="17" pagination="1 - 73" refId="ref13587" refString="Bown, T. M., and D. Schankler. 1982. A review of the Proteutheria and Insectivora of the Willwood Formation (Lower Eocene), Bighorn Basin, Wyoming. Geological Survey Bulletin 1523: 1 - 73." type="journal article" year="1982">Bown and Schankler (1982: 11–12)</bibRefCitation>
stated that
<taxonomicName id="4C2F4D0EFB149C68FCC030C78C5CFB5A" authorityName="Novacek" authorityYear="1977" box="[844,1088,1196,1222]" class="Mammalia" family="Leptictidae" genus="Palaeictops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pilosa" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="matthewi">
<emphasis id="B95BEA9FFB149C68FCC030C78C5CFB5A" box="[844,1088,1196,1222]" italics="true" pageId="16" pageNumber="17">Palaeictops matthewi</emphasis>
</taxonomicName>
is probably conspecific with
<taxonomicName id="4C2F4D0EFB149C68FEB230B989C6FB77" box="[318,474,1233,1259]" class="Mammalia" family="Leptictidae" genus="Palaeictops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pilosa" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="multicuspis">
<emphasis id="B95BEA9FFB149C68FEB230B989C6FB77" box="[318,474,1233,1259]" italics="true" pageId="16" pageNumber="17">P. multicuspis</emphasis>
</taxonomicName>
, because differences in size and p2 morphology are trivial. Synonymy is, however, unwarranted. The jaw proportions and p2 and p
<quantity id="4CD79B68FB149C68FC09309C8BAEFA8D" box="[901,946,1271,1297]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="16" pageNumber="17" unit="in" value="4.0">4 in</quantity>
these species are clearly distinct as described in the diagnosis of each species (this paper).
<bibRefCitation id="EFBE4B7CFB149C68FC0C31768CD4FAAA" author="Bown, T. M. &amp; D. Schankler" box="[896,1224,1308,1335]" pageId="16" pageNumber="17" pagination="1 - 73" refId="ref13587" refString="Bown, T. M., and D. Schankler. 1982. A review of the Proteutheria and Insectivora of the Willwood Formation (Lower Eocene), Bighorn Basin, Wyoming. Geological Survey Bulletin 1523: 1 - 73." type="journal article" year="1982">Bown and Schankler (1982)</bibRefCitation>
maintained that such characters might be insignificant and subject to intraspecific variation. There is, however, no sample evidence supporting this assessment and, in absence of data to the contrary,
<taxonomicName id="4C2F4D0EFB149C68FE9B31E68983FA3A" authorityName="Novacek" authorityYear="1977" box="[279,415,1420,1446]" class="Mammalia" family="Leptictidae" genus="Palaeictops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pilosa" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="matthewi">
<emphasis id="B95BEA9FFB149C68FE9B31E68983FA3A" box="[279,415,1420,1446]" italics="true" pageId="16" pageNumber="17">P. matthewi</emphasis>
</taxonomicName>
is here recognized as diagnosed above.
</paragraph>
</subSubSection>
</treatment>
</document>
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