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<mods:title id="C408419A6CFE81FB7FD05904C863FCD7">New gammaroid family, genera and species from subterranean waters of Japan, and their phylogenetic relationships (Crustacea: Amphipoda)</mods:title>
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<mods:namePart id="BF4AAF371464E5AA53F81B1EF734F853">Tomikawa, Ko</mods:namePart>
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<mods:namePart id="0B2223BEC201C6C69E801954320A2F7A">Morino, Hiroshi</mods:namePart>
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<treatment id="D92D36364A38FFEB73A0F996FEF0DE97" ID-DOI="http://doi.org/10.5281/zenodo.5114050" ID-GBIF-Taxon="183630800" ID-Zenodo-Dep="5114050" LSID="urn:lsid:zoobank.org:act:69D9B779-7C99-4564-BDC6-4D0960E0DC83" httpUri="http://treatment.plazi.org/id/D92D36364A38FFEB73A0F996FEF0DE97" lastPageId="10" lastPageNumber="653" pageId="9" pageNumber="652">
<subSubSection id="199ED4AB4A38FFE873A0F996FAE1DED3" box="[920,1349,1650,1674]" pageId="9" pageNumber="652" type="nomenclature">
<paragraph id="513B87204A38FFE873A0F996FAE1DED3" blockId="9.[920,1349,1650,1674]" box="[920,1349,1650,1674]" pageId="9" pageNumber="652">
<heading id="0A73304C4A38FFE873A0F996FAE1DED3" box="[920,1349,1650,1674]" centered="true" fontSize="9" level="2" pageId="9" pageNumber="652" reason="3">
<taxonomicName id="9684FCA34A38FFE873A0F996FB60DED3" ID-CoL="7PQMX" authority="TOMIKAWA" authorityName="Tomikawa &amp; Kobayashi &amp; Morino &amp; Mawatari" authorityYear="2007" box="[920,1220,1650,1674]" class="Malacostraca" family="Mesogammaridae" genus="Octopupilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="9" pageNumber="652" phylum="Arthropoda" rank="genus" status="gen. nov.">OCTOPUPILLA TOMIKAWA</taxonomicName>
<taxonomicNameLabel id="78C3E6494A38FFE874F4F992FAE1DED3" box="[1228,1349,1650,1674]" pageId="9" pageNumber="652" rank="genus">GEN. NOV.</taxonomicNameLabel>
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<subSubSection id="199ED4AB4A38FFEB7302F947FE8FDCAB" lastPageId="10" lastPageNumber="653" pageId="9" pageNumber="652" type="diagnosis">
<paragraph id="513B87204A38FFEB7302F947FE8FDCAB" blockId="9.[826,1442,1699,1905]" lastBlockId="10.[142,758,1214,1266]" lastPageId="10" lastPageNumber="653" pageId="9" pageNumber="652">Diagnosis: Eyes vestigial, represented by eight pigment spots; dorsal margin of urosomite 3 with robust setae; inner plate of maxilla 2 with facial setae in oblique row; carpus of gnathopods 1 and 2 not lobate; propodus of gnathopods 1 and 2 elongate; peduncle of pleopods with marginal setae; epimeral plates 2 and 3 with robust ventromarginal setae; uropod 3 magniramous (length of inner ramus about 0.7 × length of outer ramus).</paragraph>
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<caption id="05FBD7A84A3BFFEB70B6FBB0FB06DCDD" ID-DOI="http://doi.org/10.5281/zenodo.4634863" ID-Zenodo-Dep="4634863" httpUri="https://zenodo.org/record/4634863/files/figure.png" pageId="10" pageNumber="653" startId="10.[142,220,1108,1127]" targetBox="[142,1422,194,1075]" targetPageId="10">
<paragraph id="513B87204A3BFFEB70B6FBB0FB06DCDD" blockId="10.[142,1422,1108,1157]" pageId="10" pageNumber="653">
<emphasis id="63F05B324A3BFFEB70B6FBB0FF52DC3E" bold="true" box="[142,246,1108,1127]" pageId="10" pageNumber="653">Figure 6.</emphasis>
Scanning electron micrographs of
<taxonomicName id="9684FCA34A3BFFEB725EFBB0FC6ADC3E" authority="(Ueno, 1955)" baseAuthorityName="Ueno" baseAuthorityYear="1955" box="[614,974,1108,1127]" class="Malacostraca" family="Mesogammaridae" genus="Eoniphargus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="10" pageNumber="653" phylum="Arthropoda" rank="species" species="kojimai">
<emphasis id="63F05B324A3BFFEB725EFBB0FCE6DC3E" box="[614,834,1108,1127]" italics="true" pageId="10" pageNumber="653">Eoniphargus kojimai</emphasis>
(
<bibRefCitation id="3515FAD14A3BFFEB7368FBB0FC63DC3E" author="Ueno M" box="[848,967,1108,1127]" pageId="10" pageNumber="653" pagination="146 - 152" refId="ref13734" refString="Ueno M. 1955. Occurrence of a freshwater gammarid (Amphipoda) of the Niphargus group in Japan. Bulletin of the Biogeographical Society of Japan 16: 146 - 152." type="journal article" year="1955">Uéno, 1955</bibRefCitation>
)
</taxonomicName>
, male, 4.3 mm; female, 4.4 mm. A, antenna 2; B, calceoli on antenna 2; C, right mandible; D, molar of right mandible. AB, male; CD, female.
</paragraph>
</caption>
<subSubSection id="199ED4AB4A3BFFEB70B6FAEFFEA7DD67" pageId="10" pageNumber="653" type="type_taxon">
<paragraph id="513B87204A3BFFEB70B6FAEFFEA7DD67" blockId="10.[142,758,1290,1342]" pageId="10" pageNumber="653">
Type species:
<taxonomicName id="9684FCA34A3BFFEB7112FAEEFDC9DD79" ID-CoL="48KJB" authority="Tomikawa" authorityName="Tomikawa &amp; Kobayashi &amp; Morino &amp; Mawatari" authorityYear="2007" box="[298,621,1290,1312]" class="Malacostraca" family="Mesogammaridae" genus="Octopupilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="10" pageNumber="653" phylum="Arthropoda" rank="species" species="felix" status="sp. nov.">
<emphasis id="63F05B324A3BFFEB7112FAEEFE48DD46" box="[298,492,1290,1311]" italics="true" pageId="10" pageNumber="653">Octopupilla felix</emphasis>
Tomikawa
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<taxonomicNameLabel id="78C3E6494A3BFFEB7241FAEFFD74DD79" box="[633,720,1291,1312]" pageId="10" pageNumber="653" rank="species">sp. nov.</taxonomicNameLabel>
by monotypy.
</paragraph>
</subSubSection>
<subSubSection id="199ED4AB4A3BFFEB70B6FAB3FDC2DDF3" pageId="10" pageNumber="653" type="etymology">
<paragraph id="513B87204A3BFFEB70B6FAB3FDC2DDF3" blockId="10.[142,757,1367,1450]" pageId="10" pageNumber="653">Etymology: The genus name is from the Latin octo (eight) and pupilla (pupil) referring to the vestigial eyes represented by eight pigment spots.</paragraph>
</subSubSection>
<subSubSection id="199ED4AB4A3BFFEB70B6FA26FEF0DE97" pageId="10" pageNumber="653" type="discussion">
<paragraph id="513B87204A3BFFEB70B6FA26FEF0DE97" blockId="10.[142,758,1474,1742]" pageId="10" pageNumber="653">
Remarks:
<taxonomicName id="9684FCA34A3BFFEB70C7FA26FE20DD8E" authorityName="Tomikawa &amp; Kobayashi &amp; Morino &amp; Mawatari" authorityYear="2007" box="[255,388,1474,1495]" class="Malacostraca" family="Mesogammaridae" genus="Octopupilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="10" pageNumber="653" phylum="Arthropoda" rank="genus">
<emphasis id="63F05B324A3BFFEB70C7FA26FE20DD8E" box="[255,388,1474,1495]" italics="true" pageId="10" pageNumber="653">Octopupilla</emphasis>
</taxonomicName>
is distinguished from
<taxonomicName id="9684FCA34A3BFFEB72BBFA26FF11DDAE" authorityName=": Straskraba" authorityYear="1964" class="Malacostraca" family="Mesogammaridae" genus="Eoniphargus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="10" pageNumber="653" phylum="Arthropoda" rank="genus">
<emphasis id="63F05B324A3BFFEB72BBFA26FF11DDAE" italics="true" pageId="10" pageNumber="653">Eoniphargus</emphasis>
</taxonomicName>
by the following characters (
<taxonomicName id="9684FCA34A3BFFEB7201FA05FD68DDAF" authorityName=": Straskraba" authorityYear="1964" box="[569,716,1505,1526]" class="Malacostraca" family="Mesogammaridae" genus="Eoniphargus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="10" pageNumber="653" phylum="Arthropoda" rank="genus">
<emphasis id="63F05B324A3BFFEB7201FA05FD68DDAF" box="[569,716,1505,1526]" italics="true" pageId="10" pageNumber="653">Eoniphargus</emphasis>
</taxonomicName>
in parentheses): eyes are vestigial, represented by eight pigment spots (absent); the dorsal margin of the urosomite 3 has robust setae (lacking); epimeral plates 2 and 3 with robust ventromarginal setae (setae lacking); uropod 3 is magniramous (parviramus); and the inner ramus of uropod 3 is 0.7 times as long as the outer (0.3 times).
</paragraph>
</subSubSection>
</treatment>
</document>