157 lines
18 KiB
XML
157 lines
18 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.379.6457" ID-GBIF-Dataset="11de03c0-97c4-47b9-812b-b8e99890bb88" ID-PMC="PMC3935226" ID-Pensoft-Pub="1313-2970-379-1" ID-PubMed="24574855" ID-ZBK="237EB0D412AF485689C55E2AA52C4CEA" ModsDocAuthor="" ModsDocDate="2014" ModsDocID="1313-2970-379-1" ModsDocOrigin="ZooKeys 379" ModsDocTitle="Revision of the genus Reichardtiolus Kryzhanovskij, 1959 (Coleoptera, Histeridae, Saprininae)" checkinTime="1451246330986" checkinUser="pensoft" docAuthor="Lackner, Tomas" docDate="2014" docId="99EB118D2F61F74BC11CC8FB10AC94A0" docLanguage="en" docName="ZooKeys 379: 1-27" docOrigin="ZooKeys 379" docSource="http://dx.doi.org/10.3897/zookeys.379.6457" docTitle="Reichardtiolus aldhaferi Lackner, 2014, sp. n." docType="treatment" docUuid="5DBC0C28-18FC-40FA-92B4-21222C33DE98" docUuidSource="ZooBank" docVersion="4" lastPageNumber="11" masterDocId="FFC4FF952E63FFB76559FFBF62144D69" masterDocTitle="Revision of the genus Reichardtiolus Kryzhanovskij, 1959 (Coleoptera, Histeridae, Saprininae)" masterLastPageNumber="27" masterPageNumber="1" pageNumber="10" updateTime="1668157689324" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Revision of the genus Reichardtiolus Kryzhanovskij, 1959 (Coleoptera, Histeridae, Saprininae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Lackner, Tomas</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2014</mods:date>
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<mods:detail type="volume">
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<mods:number>379</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>27</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.379.6457</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.379.6457</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-379-1</mods:identifier>
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<mods:identifier type="ZBK">237EB0D412AF485689C55E2AA52C4CEA</mods:identifier>
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<mods:identifier type="ZooBank">237EB0D412AF485689C55E2AA52C4CEA</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="152050881" LSID="urn:lsid:zoobank.org:act:5DBC0C28-18FC-40FA-92B4-21222C33DE98" httpUri="http://treatment.plazi.org/id/99EB118D2F61F74BC11CC8FB10AC94A0" lastPageId="10" lastPageNumber="11" pageId="9" pageNumber="10">
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<subSubSection pageId="9" pageNumber="10" type="nomenclature">
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<paragraph pageId="9" pageNumber="10">
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<taxonomicName LSID="http://zoobank.org/5DBC0C28-18FC-40FA-92B4-21222C33DE98" class="Insecta" family="Histeridae" genus="Reichardtiolus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Reichardtiolus aldhaferi" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="aldhaferi">Reichardtiolus aldhaferi</taxonomicName>
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<taxonomicNameLabel pageId="9" pageNumber="10">sp. n.</taxonomicNameLabel>
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Figs 34-47
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</paragraph>
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</subSubSection>
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<subSubSection pageId="9" pageNumber="10" type="type locality">
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<paragraph pageId="9" pageNumber="10">Type locality.</paragraph>
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<paragraph pageId="9" pageNumber="10">Saudi Arabia, environs of Riyadh, Rhodet Khorim.</paragraph>
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</subSubSection>
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<subSubSection pageId="9" pageNumber="10" type="type material examined">
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<paragraph pageId="9" pageNumber="10">Type material examined.</paragraph>
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<paragraph pageId="9" pageNumber="10">
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Holotype, male, side-mounted on a triangular point with male genitalia extracted, dismembered and glued to the same mounting-point as the specimen, with following labels:
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<normalizedToken originalValue="“♂”">"♂"</normalizedToken>
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(printed); followed by: "Saudi Arabia, Rhodet Khorim / 25°25.943'N, 47°13.863', Alt. / 572m 5.ii.2012 HP (B)" (printed, black-margined label); followed by: "
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<taxonomicName class="Insecta" family="Histeridae" genus="Reichardtiolus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Reichardtiolus aldhaferi" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="aldhaferi">Reichardtiolus aldhaferi</taxonomicName>
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/ sp. n. Det. T. Lackner / 2013 HOLOTYPE" (red label, printed) (KSMA). Paratypes: 3 ♂♂ & 1 ♀, idem as Holotype (1 ♂ and 1 ♀ are sputter-coated with gold); 2 ♀♀, with following labels:
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<normalizedToken originalValue="“♀”">"♀"</normalizedToken>
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(printed), followed by: "Saudi Arabia, Rhodet Khorim / N: 25°22'58" / E:47°16'44" / 08.i.2012 Light Trap (A) (printed, black-margined label); 1 ♀, with following labels:
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<normalizedToken originalValue="“♀”">"♀"</normalizedToken>
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(printed), followed by: "Saudi Arabia Rhodet Khorim / N: 25°25'94"/ E: 47°13'86" / 25.xii.2011 Light Trap (B) (printed, black-margined label); 1 ♀, with following labels:
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<normalizedToken originalValue="“♀”">"♀"</normalizedToken>
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(printed), followed by: "Saudi Arabia Kharah, Al / Mozahmiah 30km W.Riyadh / 24.ii.2011/LT /
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<geoCoordinate direction="north" orientation="latitude" precision="15" value="28.3925">28°23'33"N</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="15" value="46.244167">46°14'39"E</geoCoordinate>
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/ Al Dhafer, H.; Kondratieff,B.; / Fadl, H.&Al Gharbawi, A. (printed-written, black-margined label); 1 ♀, with following labels:
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<normalizedToken originalValue="“♀”">"♀"</normalizedToken>
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(printed), followed by: KSA: Riyadh: Dirab / 20.i.1986 LT (written). All exs. KSMA except for 1 ♂ from Rhodet Khorim, 5.ii.2012 and 1 ♀, ibid, but 25.xii.2011 in coll. TLAN.
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="10" lastPageNumber="11" pageId="9" pageNumber="10" type="diagnostic description">
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<paragraph pageId="9" pageNumber="10">Diagnostic description.</paragraph>
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<paragraph lastPageId="10" lastPageNumber="11" pageId="9" pageNumber="10">
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Body size: PEL: 2.50-3.25 mm; APW: 0.85-1.15 mm; PPW: 1.80-2.25 mm; EW: 2.00-2.50 mm; EL: 1.50-2.00 mm. Body darker than that of
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<taxonomicName class="Insecta" family="Histeridae" genus="Reichardtiolus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Reichardtiolus duriculus" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="duriculus">
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<pageBreakToken pageId="10" pageNumber="11" start="start">Reichardtiolus</pageBreakToken>
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duriculus
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</taxonomicName>
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, otherwise similar to it. Legs and antennae darker than those of
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<taxonomicName class="Insecta" family="Histeridae" genus="Reichardtiolus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Reichardtiolus duriculus" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="duriculus">Reichardtiolus duriculus</taxonomicName>
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; mouthparts similar except mentum, which is on its anterior margin more emarginated than that of
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<taxonomicName class="Insecta" family="Histeridae" genus="Reichardtiolus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Reichardtiolus duriculus" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="duriculus">Reichardtiolus duriculus</taxonomicName>
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(compare Figs 4 and 35). Clypeus anteriorly elevated (Fig. 34), with slight median depression, rugosely punctate; frons (Fig. 34) coarsely and densely punctate, medially rugulose-lacunose, with shallow depressions; frontal and supraorbital striae and eyes as in
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<taxonomicName class="Insecta" family="Histeridae" genus="Reichardtiolus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Reichardtiolus duriculus" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="duriculus">Reichardtiolus duriculus</taxonomicName>
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. Pronotum slightly less acutely narrowing apically than that of
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<taxonomicName class="Insecta" family="Histeridae" genus="Reichardtiolus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Reichardtiolus duriculus" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="duriculus">Reichardtiolus duriculus</taxonomicName>
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; punctuation on pronotal disk sparser than that of
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<taxonomicName class="Insecta" family="Histeridae" genus="Reichardtiolus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Reichardtiolus duriculus" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="duriculus">Reichardtiolus duriculus</taxonomicName>
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. Elytra similar to those of
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<taxonomicName class="Insecta" family="Histeridae" genus="Reichardtiolus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Reichardtiolus duriculus" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="duriculus">Reichardtiolus duriculus</taxonomicName>
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, but dorsal elytral striae weaker, occasionally striae 3-4 shortened apically, only half as long as striae 1-2 or even evanescent; between 4th dorsal elytral and sutural striae in several specimens punctures scratch-like and surface with variously deep longitudinal wrinkles; rarely with shallow depression between the bases of 4th and sutural elytral striae. Punctuation of elytral disk sparser than that of
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<taxonomicName class="Insecta" family="Histeridae" genus="Reichardtiolus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Reichardtiolus duriculus" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="duriculus">Reichardtiolus duriculus</taxonomicName>
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, punctures separated by several times their diameter; in fourth elytral interval occasionally scratch-like. Propygidium and pygidium similar to those of
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<taxonomicName class="Insecta" family="Histeridae" genus="Reichardtiolus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Reichardtiolus duriculus" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="duriculus">Reichardtiolus duriculus</taxonomicName>
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, but punctuation denser and coarser in
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<taxonomicName class="Insecta" family="Histeridae" genus="Reichardtiolus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Reichardtiolus aldhaferi" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="aldhaferi">Reichardtiolus aldhaferi</taxonomicName>
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, although not as dense as in
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<taxonomicName class="Insecta" family="Histeridae" genus="Reichardtiolus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Reichardtiolus sphingis" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="sphingis">Reichardtiolus sphingis</taxonomicName>
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(compare Figs 12, 13 and 37). Structure of prosternal process similar to that of
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<taxonomicName class="Insecta" family="Histeridae" genus="Reichardtiolus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Reichardtiolus duriculus" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="duriculus">Reichardtiolus duriculus</taxonomicName>
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, but prosternal keel laterally more compressed and setose (compare Figs 10 and 36); carinal prosternal striae occasionally very approximate, medially almost united and difficult to discern; prosternal foveae smaller than those of
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<taxonomicName class="Insecta" family="Histeridae" genus="Reichardtiolus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Reichardtiolus duriculus" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="duriculus">Reichardtiolus duriculus</taxonomicName>
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. Mesoventrite sub-square, trapezoidal, punctuation sparse, punctures separated by several times their own diameter; marginal mesoventral stria always complete anteriorly, almost straight; meso-metaventral sutural stria absent, suture distinct. Metaventrite, metepisternum and abdominal ventrites similar to those of
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<taxonomicName class="Insecta" family="Histeridae" genus="Reichardtiolus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Reichardtiolus duriculus" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="duriculus">Reichardtiolus duriculus</taxonomicName>
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. Legs as in
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<taxonomicName class="Insecta" family="Histeridae" genus="Reichardtiolus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Reichardtiolus duriculus" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="duriculus">Reichardtiolus duriculus</taxonomicName>
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; except denticles of mesotibia that are sparser, thinner and shorter. Male genitalia: 8th sternite (Figs 38-39) strongly sclerotized laterally, apically with pseudopores and a row of short setae and small velum covered with minute setae; 8th tergite (Fig. 39) deeply emarginated apically, on basal half with prominent pores; 8th sternite and tergite fused laterally (Fig. 40). 9th tergite (Fig. 41) well sclerotized along margins, laterally without projection (Fig. 42), apically with two bisinuate strongly sclerotized lines visible from dorsal view, apical half covered with pseudopores, sclerotization of tergite medially divided, two parts held together by weakly sclerotized part; 10th tergite basally faintly inwardly arcuate (Fig. 41). Tips of apical end of spiculum gastrale (Fig. 43) without strongly sclerotized parts, apical end strongly inwardly arcuate, basal end outwardly arcuate. Aedeagus (Figs 45-46) similar to that of
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<taxonomicName class="Insecta" family="Histeridae" genus="Reichardtiolus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Reichardtiolus duriculus" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="duriculus">Reichardtiolus duriculus</taxonomicName>
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, but laterally more curved and medially thickened (compare Figs 23 and 46).
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</paragraph>
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<caption pageId="10" pageNumber="11">
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<paragraph pageId="10" pageNumber="11">
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Figures 34-37. 34
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<taxonomicName class="Insecta" family="Histeridae" genus="Reichardtiolus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Reichardtiolus aldhaferi" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="aldhaferi">Reichardtiolus aldhaferi</taxonomicName>
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sp. n., head, dorsal view 35
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<taxonomicName class="Insecta" family="Histeridae" genus="Reichardtiolus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Reichardtiolus aldhaferi" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="aldhaferi">Reichardtiolus aldhaferi</taxonomicName>
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sp. n., mentum, ventral view 36
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<taxonomicName class="Insecta" family="Histeridae" genus="Reichardtiolus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Reichardtiolus aldhaferi" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="aldhaferi">Reichardtiolus aldhaferi</taxonomicName>
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sp. n., prosternum 37
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<taxonomicName class="Insecta" family="Histeridae" genus="Reichardtiolus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Reichardtiolus aldhaferi" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="aldhaferi">Reichardtiolus aldhaferi</taxonomicName>
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sp. n., pygidium.
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</paragraph>
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</caption>
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<caption pageId="10" pageNumber="11">
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<paragraph pageId="10" pageNumber="11">
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Figures 38-47. 38
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<taxonomicName class="Insecta" family="Histeridae" genus="Reichardtiolus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Reichardtiolus aldhaferi" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="aldhaferi">Reichardtiolus aldhaferi</taxonomicName>
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sp. n., 8th sternite and tergite, ventral view 39 ditto, ventral view 40 ditto, lateral view 41
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<taxonomicName class="Insecta" family="Histeridae" genus="Reichardtiolus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Reichardtiolus aldhaferi" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="aldhaferi">Reichardtiolus aldhaferi</taxonomicName>
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sp. n., 9th + 10th tergites, dorsal view 42 ditto, lateral view 43
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<taxonomicName class="Insecta" family="Histeridae" genus="Reichardtiolus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Reichardtiolus aldhaferi" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="aldhaferi">Reichardtiolus aldhaferi</taxonomicName>
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sp. n., spiculum gastrale, ventral view 44 ditto, lateral view 45
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<taxonomicName class="Insecta" family="Histeridae" genus="Reichardtiolus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Reichardtiolus aldhaferi" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="aldhaferi">Reichardtiolus aldhaferi</taxonomicName>
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sp. n., aedeagus, dorsal view 46 ditto, lateral view 47
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<taxonomicName class="Insecta" family="Histeridae" genus="Reichardtiolus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Reichardtiolus aldhaferi" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="aldhaferi">Reichardtiolus aldhaferi</taxonomicName>
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sp. n., apex of aedeagus, frontal view.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="10" pageNumber="11" type="differential diagnosis">
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<paragraph pageId="10" pageNumber="11">Differential diagnosis.</paragraph>
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<paragraph pageId="10" pageNumber="11">As with preceding species.</paragraph>
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</subSubSection>
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<subSubSection pageId="10" pageNumber="11" type="biology">
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<paragraph pageId="10" pageNumber="11">Biology.</paragraph>
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<paragraph pageId="10" pageNumber="11">Unknown, presumably similar to the congeners, the examined specimens were collected at light in winter months.</paragraph>
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</subSubSection>
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<subSubSection pageId="10" pageNumber="11" type="distribution">
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<paragraph pageId="10" pageNumber="11">Distribution.</paragraph>
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<paragraph pageId="10" pageNumber="11">Saudi Arabia, environs of Riyadh (Fig. 72).</paragraph>
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</subSubSection>
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<subSubSection pageId="10" pageNumber="11" type="etymology">
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<paragraph pageId="10" pageNumber="11">Etymology.</paragraph>
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<paragraph pageId="10" pageNumber="11">Patronymic, named after the head of the entomology department at KSMA, H. M. Al Dhafer.</paragraph>
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</subSubSection>
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</treatment>
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</document> |