treatments-xml/data/30/76/88/3076884BFFD61021FF1EF8D5471CF814.xml

<document id="1F62787506C731E82C12C719F1008446" ID-CLB-Dataset="27969" ID-DOI="10.11646/zootaxa.4521.1.7" ID-GBIF-Dataset="d78d1897-cc4e-4bde-b97d-0c5b8021451d" ID-ISSN="1175-5326" ID-Zenodo-Dep="2609721" ID-ZooBank="13AD67F2-D182-47F5-8664-5BF46B6102B0" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1553623459469" checkinUser="plazi" docAuthor="Cunha, Carlo M. &amp; Simone, Luiz Ricardo L." docDate="2018" docId="3076884BFFD61021FF1EF8D5471CF814" docLanguage="en" docName="zootaxa.4521.1.7.pdf" docOrigin="Zootaxa 4521 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Rapturella atlas Cunha &amp; Simone 2018, n. sp." docType="treatment" docUuid="B53DED70-F324-4F4A-9B87-0CB1D59D38FF" docUuidSource="ZooBank" docVersion="9" lastPageNumber="126" masterDocId="CC4FF033FFD61020FF89FFC04311FF83" masterDocTitle="A new species of the genus Rapturella (Gastropoda: Acteonidae) from southeast Brazil" masterLastPageNumber="128" masterPageNumber="125" pageNumber="125" updateTime="1698713667879" updateUser="ExternalLinkService">
<mods:mods id="6ADC82F0DF166BE56FC9640929CCA92B" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="7B906578C9A47BA4D61B345A1078E919">
<mods:title id="49898501148368B4459F411E00663C5E">A new species of the genus Rapturella (Gastropoda: Acteonidae) from southeast Brazil</mods:title>
</mods:titleInfo>
<mods:name id="C7E6DBA83917296FD41B22C49DD687B3" type="personal">
<mods:role id="4E225FD7DCE7A60A1195980853053785">
<mods:roleTerm id="96E50E4A60820686A97969CFD7263DC2">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="3141E286BC50A4E5DB5DBF9B44C8B0CB">Cunha, Carlo M.</mods:namePart>
</mods:name>
<mods:name id="34901BB7B0A8F966342CFEB09E8ACA55" type="personal">
<mods:role id="224A4606FB861AB57FBFCC1DE2354D52">
<mods:roleTerm id="2497A2D55BBFF83A2231D02FF3936A21">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="436CB47DA68448FEE77D6814B9A110B8">Simone, Luiz Ricardo L.</mods:namePart>
</mods:name>
<mods:typeOfResource id="BD73E471199645DBDF2688A6C1F4407A">text</mods:typeOfResource>
<mods:relatedItem id="6D1974EB6187A9E808F6B3B09F6C4DDB" type="host">
<mods:titleInfo id="0575318080CF204A6F26071D028D4623">
<mods:title id="FDF09C3EE1B43AE9C96F1BDF3B1791DF">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="36D0A73E622D49B2EBD1A0906BC1B6D3">
<mods:date id="BC2064C11650EE60131C5A043A22A27D">2018</mods:date>
<mods:detail id="8F8F7A3CD2F514EF98887A5A2E293684" type="pubDate">
<mods:number id="0D398CFD4F8B0D41B4D70D4D45C8E894">2018-11-13</mods:number>
</mods:detail>
<mods:detail id="52825144D363DE95C2E48EBC7A942789" type="volume">
<mods:number id="09705EEFEFDC91C3E08C9EB6188EDFFA">4521</mods:number>
</mods:detail>
<mods:detail id="1E23712BBE361B80E4592C8FCE5EEA61" type="issue">
<mods:number id="2F6340AE74FEC4085D5A60A9509F6598">1</mods:number>
</mods:detail>
<mods:extent id="FE66BA85EE4F65E6BA110319431B0052" unit="page">
<mods:start id="BD3EEF6CE56DA1D8C2661B43DC05FCC7">125</mods:start>
<mods:end id="DE88BA3B475E31EA47FB0B075E9949C9">128</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="F465C5E4EF8CDA544F65763E1BB064EC">journal article</mods:classification>
<mods:identifier id="3FC64E068EAA12B6FC0EDD82202DD97B" type="CLB-Dataset">27969</mods:identifier>
<mods:identifier id="CC8BAF20474B66B337D7F1831A521863" type="DOI">10.11646/zootaxa.4521.1.7</mods:identifier>
<mods:identifier id="5A159DCEC0EEB91A66BB4A77485E154F" type="GBIF-Dataset">d78d1897-cc4e-4bde-b97d-0c5b8021451d</mods:identifier>
<mods:identifier id="DB3AA6B30F74090575F95D5874118DA1" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="0BC26AEED24214AF3D9D3EF717E01E02" type="Zenodo-Dep">2609721</mods:identifier>
<mods:identifier id="74D6E7455B7C39EE7A660334CC0C53A6" type="ZooBank">13AD67F2-D182-47F5-8664-5BF46B6102B0</mods:identifier>
</mods:mods>
<treatment id="3076884BFFD61021FF1EF8D5471CF814" ID-DOI="http://doi.org/10.5281/zenodo.5971827" ID-GBIF-Taxon="154498698" ID-Zenodo-Dep="5971827" LSID="urn:lsid:zoobank.org:act:B53DED70-F324-4F4A-9B87-0CB1D59D38FF" httpUri="http://treatment.plazi.org/id/3076884BFFD61021FF1EF8D5471CF814" lastPageId="1" lastPageNumber="126" pageId="0" pageNumber="125">
<subSubSection id="F0C56AD6FFD61020FF1EF8D54135F8B3" box="[151,548,1813,1840]" pageId="0" pageNumber="125" type="nomenclature">
<paragraph id="B860395DFFD61020FF1EF8D54135F8B3" blockId="0.[151,1436,1813,2010]" box="[151,548,1813,1840]" pageId="0" pageNumber="125">
<heading id="E3288E31FFD61020FF1EF8D54135F8B3" bold="true" box="[151,548,1813,1840]" fontSize="11" level="1" pageId="0" pageNumber="125" reason="1">
<emphasis id="8AABE54FFFD61020FF1EF8D54135F8B3" bold="true" box="[151,548,1813,1840]" pageId="0" pageNumber="125">
<taxonomicName id="7FDF42DEFFD61020FF1EF8D54249F8AC" ID-CoL="4RL3M" authority="Cunha &amp; Simone, 2018" authorityName="Cunha &amp; Simone" authorityYear="2018" box="[151,344,1813,1839]" class="Gastropoda" family="Acteonidae" genus="Rapturella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Heterostropha" pageId="0" pageNumber="125" phylum="Mollusca" rank="species" species="atlas" status="sp. nov.">
<emphasis id="8AABE54FFFD61020FF1EF8D54249F8AC" bold="true" box="[151,344,1813,1839]" italics="true" pageId="0" pageNumber="125">Rapturella atlas</emphasis>
</taxonomicName>
<taxonomicNameLabel id="91985834FFD61020FED6F8D642B0F8B3" box="[351,417,1814,1840]" pageId="0" pageNumber="125" rank="species">n. sp.</taxonomicNameLabel>
(
<figureCitation id="20E425D8FFD61020FE39F8D6410BF8B3" box="[432,538,1814,1840]" captionStart="FIGURE 1" captionStartId="2.[151,244,1481,1502]" captionTargetBox="[151,1436,402,1457]" captionTargetId="figure@2.[151,1436,402,1457]" captionTargetPageId="2" captionText="FIGURE 1. Rapturella atlas n. sp. A, Holotype, MZSP 91018 (L 2.8 mm), apertural view; B, same, dorsal view; C, paratype, MZSP 107496 (L 3.1 mm), apertural view (SEM); D, same, detail of ornamentation on the last whorl; E, same, apical view; F, same, detail of protoconch, arrows indicate start of teleoconch. Scale bars: D. 100 µm, F. 200 µm." httpUri="https://zenodo.org/record/2609723/files/figure.png" pageId="0" pageNumber="125">Figure 1</figureCitation>
)
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="F0C56AD6FFD61021FF1EF8F84040FF52" lastPageId="1" lastPageNumber="126" pageId="0" pageNumber="125" type="materials_examined">
<paragraph id="B860395DFFD61020FF1EF8F842C7F859" blockId="0.[151,1436,1813,2010]" pageId="0" pageNumber="125">
<emphasis id="8AABE54FFFD61020FF1EF8F84252F8CC" bold="true" box="[151,323,1848,1871]" pageId="0" pageNumber="125">Type material.</emphasis>
<materialsCitation id="08B73300FFD61020FEC6F8F841FAF8CC" ID-GBIF-Occurrence="2028520922" box="[335,747,1848,1871]" collectionCode="MZSP" country="Brazil" pageId="0" pageNumber="125" specimenCode="MZSP 91018" specimenCount="1" typeStatus="holotype">
<typeStatus id="676487FFFFD61020FEC6F8F842A3F8CC" box="[335,434,1848,1871]" pageId="0" pageNumber="125" type="holotype">Holotype</typeStatus>
<specimenCode id="E8799126FFD61020FE37F8F84145F8CC" box="[446,596,1848,1871]" collectionCode="MZSP" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="0" pageNumber="125" type="Museum">MZSP 91018</specimenCode>
(
<figureCitation id="20E425D8FFD61020FDE1F8F841B8F8CC" box="[616,681,1848,1871]" captionStart="FIGURE 1" captionStartId="2.[151,244,1481,1502]" captionTargetBox="[151,1436,402,1457]" captionTargetId="figure@2.[151,1436,402,1457]" captionTargetPageId="2" captionText="FIGURE 1. Rapturella atlas n. sp. A, Holotype, MZSP 91018 (L 2.8 mm), apertural view; B, same, dorsal view; C, paratype, MZSP 107496 (L 3.1 mm), apertural view (SEM); D, same, detail of ornamentation on the last whorl; E, same, apical view; F, same, detail of protoconch, arrows indicate start of teleoconch. Scale bars: D. 100 µm, F. 200 µm." httpUri="https://zenodo.org/record/2609723/files/figure.png" pageId="0" pageNumber="125">Fig. 1</figureCitation>
A–B)
</materialsCitation>
. 
<materialsCitation id="08B73300FFD61020FD72F8F842C3F859" ID-GBIF-Occurrence="2028520920" collectingDate="1997-11-20" collectingDateMax="2005-06-18" collectingDateMin="1997-11-20" collectionCode="MZSP, MNHN, MNRJ" country="Brazil" elevation="155" latitude="-25.24" location="Sao Sebastiao" longLatPrecision="124" longitude="-45.051666" municipality="Cape Sao Tome" pageId="0" pageNumber="125" specimenCode="MZSP 107499, 1, MZSP 107496, MZSP 107497, MNHN 25271, 2, MNRJ 30855, MZSP 107498, MZSP 107500, 4" specimenCount="1" stateProvince="Rio de Janeiro" typeStatus="paratype">
<typeStatus id="676487FFFFD61020FD72F8F84076F8CC" box="[763,871,1848,1871]" pageId="0" pageNumber="125" type="paratype">Paratypes</typeStatus>
: 
<collectingCountry id="C0C879CDFFD61020FCFEF8F840CBF8CC" box="[887,986,1848,1871]" name="Brazil" pageId="0" pageNumber="125">BRAZIL</collectingCountry>
, 
<collectingRegion id="7A1BF7BFFFD61020FC63F8F84781F8CC" box="[1002,1168,1848,1871]" country="Brazil" name="Rio de Janeiro" pageId="0" pageNumber="125">Rio de Janeiro</collectingRegion>
state, 
<collectingMunicipality id="5804A327FFD61020FB68F8F84689F8CC" box="[1249,1432,1848,1871]" pageId="0" pageNumber="125">Cape São Tomé</collectingMunicipality>
, 
<geoCoordinate id="DDEB5F9AFFD61020FF1EF89B4206F8F1" box="[151,279,1883,1906]" direction="south" orientation="latitude" pageId="0" pageNumber="125" precision="9" value="-21.972">21°58.32' S</geoCoordinate>
<geoCoordinate id="DDEB5F9AFFD61020FEA8F89B42BDF8F1" box="[289,428,1883,1906]" direction="west" orientation="longitude" pageId="0" pageNumber="125" precision="9" value="-40.03417">40°02.05' W</geoCoordinate>
, 
<quantity id="7F2794B8FFD61020FE33F89B4115F8F1" box="[442,516,1883,1906]" metricMagnitude="2" metricUnit="m" metricValue="1.55" pageId="0" pageNumber="125" unit="m" value="155.0">
<elevation id="33F2DE6EFFD61020FE33F89B4115F8F1" box="[442,516,1883,1906]" metricMagnitude="2" metricUnit="m" metricValue="1.55" pageId="0" pageNumber="125" unit="m" value="155.0">155 m</elevation>
</quantity>
, 
<specimenCode id="E8799126FFD61020FD9AF89B41A6F8F1" box="[531,695,1883,1906]" collectionCode="MZSP" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="0" pageNumber="125" type="Museum">MZSP 107499</specimenCode>
, 
<specimenCode id="E8799126FFD61020FD4CF89B41C3F8F1" box="[709,722,1883,1906]" collectionCode="MZSP" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="0" pageNumber="125" type="Museum">1</specimenCode>
shell (
<date id="CC611F9DFFD61020FCAAF89B4088F8F1" box="[803,921,1883,1906]" pageId="0" pageNumber="125" value="2005-06-18">
<collectingDate id="DC25E675FFD61020FCAAF89B4088F8F1" box="[803,921,1883,1906]" pageId="0" pageNumber="125" value="2005-06-18">18.vi.2005</collectingDate>
</date>
); 
<collectingRegion id="7A1BF7BFFFD61020FC39F89B4731F8F1" box="[944,1056,1883,1906]" country="Brazil" name="Sao Paulo" pageId="0" pageNumber="125">São Paulo</collectingRegion>
state, 
<location id="BD006F86FFD61020FBE4F89B4618F8F1" LSID="urn:lsid:plazi:treatment:3076884BFFD61021FF1EF8D5471CF814:BD006F86FFD61020FBE4F89B4618F8F1" box="[1133,1289,1883,1906]" country="Brazil" latitude="-25.24" longLatPrecision="124" longitude="-45.051666" municipality="Cape Sao Tome" name="Sao Sebastiao" pageId="0" pageNumber="125" stateProvince="Rio de Janeiro">São Sebastião</location>
, 24°44'69&quot; S 44°44'965&quot; W, 
<quantity id="7F2794B8FFD61020FEB1F8BE426CF816" box="[312,381,1918,1941]" metricMagnitude="2" metricUnit="m" metricValue="1.53" pageId="0" pageNumber="125" unit="m" value="153.0">
<elevation id="33F2DE6EFFD61020FEB1F8BE426CF816" box="[312,381,1918,1941]" metricMagnitude="2" metricUnit="m" metricValue="1.53" pageId="0" pageNumber="125" unit="m" value="153.0">153 m</elevation>
</quantity>
, 
<specimenCode id="E8799126FFD61020FE01F8BE4132F816" box="[392,547,1918,1941]" collectionCode="MZSP" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="0" pageNumber="125" type="Museum">MZSP 107496</specimenCode>
(
<figureCitation id="20E425D8FFD61020FDB8F8BE417DF816" box="[561,620,1918,1941]" captionStart="FIGURE 1" captionStartId="2.[151,244,1481,1502]" captionTargetBox="[151,1436,402,1457]" captionTargetId="figure@2.[151,1436,402,1457]" captionTargetPageId="2" captionText="FIGURE 1. Rapturella atlas n. sp. A, Holotype, MZSP 91018 (L 2.8 mm), apertural view; B, same, dorsal view; C, paratype, MZSP 107496 (L 3.1 mm), apertural view (SEM); D, same, detail of ornamentation on the last whorl; E, same, apical view; F, same, detail of protoconch, arrows indicate start of teleoconch. Scale bars: D. 100 µm, F. 200 µm." httpUri="https://zenodo.org/record/2609723/files/figure.png" pageId="0" pageNumber="125">Fig. 1</figureCitation>
C–F), 1 shell, 
<specimenCode id="E8799126FFD61020FC8FF8BE40B4F816" box="[774,933,1918,1941]" collectionCode="MZSP" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="0" pageNumber="125" type="Museum">MZSP 107497</specimenCode>
, 
<specimenCode id="E8799126FFD61020FC27F8BE40AAF816" box="[942,955,1918,1941]" collectionCode="MZSP" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="0" pageNumber="125" type="Museum">1</specimenCode>
shell, 
<specimenCode id="E8799126FFD61020FB88F8BE478FF816" box="[1025,1182,1918,1941]" collectionCode="MNHN" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="0" pageNumber="125">MNHN 25271</specimenCode>
, 
<specimenCode id="E8799126FFD61020FB21F8BE47A4F816" box="[1192,1205,1918,1941]" collectionCode="MNHN" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="0" pageNumber="125">2</specimenCode>
shells, 
<specimenCode id="E8799126FFD61020FA8CF8BE4686F816" box="[1285,1431,1918,1941]" collectionCode="MNRJ" country="Brazil" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="0" pageNumber="125">MNRJ 30855</specimenCode>
, 
<specimenCode id="E8799126FFD61020FF1EF86043B5F834" box="[151,164,1952,1975]" collectionCode="MNRJ" country="Brazil" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="0" pageNumber="125">2</specimenCode>
shells; 
<geoCoordinate id="DDEB5F9AFFD61020FF75F860427CF834" box="[252,365,1952,1975]" direction="south" orientation="latitude" pageId="0" pageNumber="125" precision="92" value="-24.713333">24°42.8' S</geoCoordinate>
<geoCoordinate id="DDEB5F9AFFD61020FEFEF86042E2F834" box="[375,499,1952,1975]" direction="west" orientation="longitude" pageId="0" pageNumber="125" precision="92" value="-44.715">44°42.9' W</geoCoordinate>
, 
<quantity id="7F2794B8FFD61020FD88F8604158F834" box="[513,585,1952,1975]" metricMagnitude="2" metricUnit="m" metricValue="1.6199999999999999" pageId="0" pageNumber="125" unit="m" value="162.0">
<elevation id="33F2DE6EFFD61020FD88F8604158F834" box="[513,585,1952,1975]" metricMagnitude="2" metricUnit="m" metricValue="1.6199999999999999" pageId="0" pageNumber="125" unit="m" value="162.0">162 m</elevation>
</quantity>
, 
<specimenCode id="E8799126FFD61020FDD1F86041E8F834" box="[600,761,1952,1975]" collectionCode="MZSP" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="0" pageNumber="125" type="Museum">MZSP 107498</specimenCode>
, 
<specimenCode id="E8799126FFD61020FC8EF8604005F834" box="[775,788,1952,1975]" collectionCode="MZSP" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="0" pageNumber="125" type="Museum">2</specimenCode>
shells (
<date id="CC611F9DFFD61020FCE4F86040F2F834" box="[877,995,1952,1975]" pageId="0" pageNumber="125" value="1997-11-20">
<collectingDate id="DC25E675FFD61020FCE4F86040F2F834" box="[877,995,1952,1975]" pageId="0" pageNumber="125" value="1997-11-20">20.xi.1997</collectingDate>
</date>
); 
<geoCoordinate id="DDEB5F9AFFD61020FC73F860477AF834" box="[1018,1131,1952,1975]" direction="south" orientation="latitude" pageId="0" pageNumber="125" precision="92" value="-25.24">25°14.4' S</geoCoordinate>
<geoCoordinate id="DDEB5F9AFFD61020FBFCF86047E3F834" box="[1141,1266,1952,1975]" direction="west" orientation="longitude" pageId="0" pageNumber="125" precision="92" value="-45.051666">45°03.1' W</geoCoordinate>
, 
<quantity id="7F2794B8FFD61020FB76F8604659F834" box="[1279,1352,1952,1975]" metricMagnitude="2" metricUnit="m" metricValue="1.47" pageId="0" pageNumber="125" unit="m" value="147.0">
<elevation id="33F2DE6EFFD61020FB76F8604659F834" box="[1279,1352,1952,1975]" metricMagnitude="2" metricUnit="m" metricValue="1.47" pageId="0" pageNumber="125" unit="m" value="147.0">147 m</elevation>
</quantity>
, 
<specimenCode id="E8799126FFD61020FADFF86043FBF859" collectionCode="MZSP" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="0" pageNumber="125" type="Museum">MZSP 107500</specimenCode>
, 
<specimenCode id="E8799126FFD61020FF7DF8034210F859" box="[244,257,1987,2010]" collectionCode="MZSP" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="0" pageNumber="125" type="Museum">4</specimenCode>
shells (
<date id="CC611F9DFFD61020FEDDF80342DBF859" box="[340,458,1987,2010]" pageId="0" pageNumber="125" value="1997-11-21">
<collectingDate id="DC25E675FFD61020FEDDF80342DBF859" box="[340,458,1987,2010]" pageId="0" pageNumber="125" value="1997-11-21">21.xi.1997</collectingDate>
</date>
)
</materialsCitation>
.
</paragraph>
<paragraph id="B860395DFFD71021FF4EFF584040FF52" blockId="1.[151,1436,152,1943]" pageId="1" pageNumber="126">
<materialsCitation id="08B73300FFD71021FF4EFF58405CFF52" ID-GBIF-Occurrence="2028520921" collectingDate="2005-06-29" collectorName="Project Frade Development &amp; C. H. Caetano" country="Brazil" elevation="162" latitude="-22.0118" location="Cape Sao Tome" longLatPrecision="1" longitude="-40.05633" pageId="1" pageNumber="126" specimenCount="1" stateProvince="Rio de Janeiro" typeStatus="holotype">
<emphasis id="8AABE54FFFD71021FF4EFF584270FF2C" bold="true" box="[199,353,152,175]" pageId="1" pageNumber="126">
<typeStatus id="676487FFFFD71021FF4EFF5843EEFF2C" box="[199,255,152,175]" pageId="1" pageNumber="126">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="C0C879CDFFD71021FEE2FF5842C1FF2C" box="[363,464,152,175]" name="Brazil" pageId="1" pageNumber="126">BRAZIL</collectingCountry>
: 
<collectingRegion id="7A1BF7BFFFD71021FE55FF584191FF2C" box="[476,640,152,175]" country="Brazil" name="Rio de Janeiro" pageId="1" pageNumber="126">Rio de Janeiro</collectingRegion>
, off 
<location id="BD006F86FFD71021FD3FFF584079FF2C" LSID="urn:lsid:plazi:treatment:3076884BFFD61021FF1EF8D5471CF814:BD006F86FFD71021FD3FFF584079FF2C" box="[694,872,152,175]" country="Brazil" latitude="-22.0118" longLatPrecision="1" longitude="-40.05633" name="Cape Sao Tome" pageId="1" pageNumber="126" stateProvince="Rio de Janeiro">Cape São Tomé</location>
, 
<geoCoordinate id="DDEB5F9AFFD71021FCFCFF584713FF2C" box="[885,1026,152,175]" direction="south" orientation="latitude" pageId="1" pageNumber="126" precision="1" value="-22.0118">22°00.708' S</geoCoordinate>
<geoCoordinate id="DDEB5F9AFFD71021FB82FF5847B2FF2C" box="[1035,1187,152,175]" direction="west" orientation="longitude" pageId="1" pageNumber="126" precision="1" value="-40.05633">40°03.380' W</geoCoordinate>
, 
<quantity id="7F2794B8FFD71021FB39FF5847E6FF2C" box="[1200,1271,152,175]" metricMagnitude="2" metricUnit="m" metricValue="1.6199999999999999" pageId="1" pageNumber="126" unit="m" value="162.0">
<elevation id="33F2DE6EFFD71021FB39FF5847E6FF2C" box="[1200,1271,152,175]" metricMagnitude="2" metricUnit="m" metricValue="1.6199999999999999" pageId="1" pageNumber="126" unit="m" value="162.0">162 m</elevation>
</quantity>
(
<collectorName id="152A5C8BFFD71021FA81FF584238FF52" pageId="1" pageNumber="126">Project Frade Development</collectorName>
, sta BC_A_E39_1, 
<collectorName id="152A5C8BFFD71021FE75FF7A419DFF52" box="[508,652,186,209]" pageId="1" pageNumber="126">C.H. Caetano</collectorName>
coll., 
<date id="CC611F9DFFD71021FD46FF7A4057FF52" box="[719,838,186,209]" pageId="1" pageNumber="126" value="2005-06-29">
<collectingDate id="DC25E675FFD71021FD46FF7A4057FF52" box="[719,838,186,209]" pageId="1" pageNumber="126" value="2005-06-29">29.vi.2005</collectingDate>
</date>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="F0C56AD6FFD71021FF4EFF1D4743FE94" pageId="1" pageNumber="126" type="etymology">
<paragraph id="B860395DFFD71021FF4EFF1D4743FE94" blockId="1.[151,1436,152,1943]" pageId="1" pageNumber="126">
<emphasis id="8AABE54FFFD71021FF4EFF1D4256FF77" bold="true" box="[199,327,221,244]" pageId="1" pageNumber="126">Etymology.</emphasis>
In honors of Atlas, the nick name of Frank Fontaine, the character of the science-fiction video game series BioShock: an allusion to the hidden identity of the species. The noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="F0C56AD6FFD71021FF4EFEE240F3FEFC" pageId="1" pageNumber="126" type="diagnosis">
<paragraph id="B860395DFFD71021FF4EFEE240F3FEFC" blockId="1.[151,1436,152,1943]" pageId="1" pageNumber="126">
<emphasis id="8AABE54FFFD71021FF4EFEE2422DFEBA" bold="true" box="[199,316,290,313]" pageId="1" pageNumber="126">Diagnosis.</emphasis>
Small shell with walls strongly thickened. Umbilicus deep and narrow. Spiral grooves on teleoconch separated by regularly spaced intervals, each interval about 10 times wider than the grooves; last whorl has about 11 spiral grooves, but no subsutural grooves. Palatal region has no thickened area.
</paragraph>
</subSubSection>
<subSubSection id="F0C56AD6FFD71021FF4EFE4A4259FC2A" pageId="1" pageNumber="126" type="description">
<paragraph id="B860395DFFD71021FF4EFE4A407DFD7F" blockId="1.[151,1436,152,1943]" pageId="1" pageNumber="126">
<emphasis id="8AABE54FFFD71021FF4EFE4A4243FE22" bold="true" box="[199,338,394,417]" pageId="1" pageNumber="126">Description.</emphasis>
Shell oval, profile rounded, maximum length (L) 
<quantity id="7F2794B8FFD71021FCF6FE4A40C7FE22" box="[895,982,394,417]" metricMagnitude="-3" metricUnit="m" metricValue="3.1" pageId="1" pageNumber="126" unit="mm" value="3.1">3.1 mm</quantity>
; about 1.3 times longer than wide. Walls thickened. Color white; spire short, length about 1/4 of total length (
<figureCitation id="20E425D8FFD71021FC07FE6D4719FE47" box="[910,1032,429,452]" captionStart="FIGURE 1" captionStartId="2.[151,244,1481,1502]" captionTargetBox="[151,1436,402,1457]" captionTargetId="figure@2.[151,1436,402,1457]" captionTargetPageId="2" captionText="FIGURE 1. Rapturella atlas n. sp. A, Holotype, MZSP 91018 (L 2.8 mm), apertural view; B, same, dorsal view; C, paratype, MZSP 107496 (L 3.1 mm), apertural view (SEM); D, same, detail of ornamentation on the last whorl; E, same, apical view; F, same, detail of protoconch, arrows indicate start of teleoconch. Scale bars: D. 100 µm, F. 200 µm." httpUri="https://zenodo.org/record/2609723/files/figure.png" pageId="1" pageNumber="126">Fig. 1A, B</figureCitation>
). Protoconch small, glossy, 1 whorl; width about 
<quantity id="7F2794B8FFD71021FEAAFE10426DFE64" box="[291,380,464,487]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="1" pageNumber="126" unit="mm" value="0.4">0.4 mm</quantity>
; separated from teleoconch by narrow orthocline furrow (
<figureCitation id="20E425D8FFD71021FB89FE104745FE64" box="[1024,1108,464,487]" captionStart="FIGURE 1" captionStartId="2.[151,244,1481,1502]" captionTargetBox="[151,1436,402,1457]" captionTargetId="figure@2.[151,1436,402,1457]" captionTargetPageId="2" captionText="FIGURE 1. Rapturella atlas n. sp. A, Holotype, MZSP 91018 (L 2.8 mm), apertural view; B, same, dorsal view; C, paratype, MZSP 107496 (L 3.1 mm), apertural view (SEM); D, same, detail of ornamentation on the last whorl; E, same, apical view; F, same, detail of protoconch, arrows indicate start of teleoconch. Scale bars: D. 100 µm, F. 200 µm." httpUri="https://zenodo.org/record/2609723/files/figure.png" pageId="1" pageNumber="126">Fig. 1F</figureCitation>
: arrows). Teleoconch up to 4 whorls, each whorl convex (
<figureCitation id="20E425D8FFD71021FE56FE324129FD8A" box="[479,568,498,521]" captionStart="FIGURE 1" captionStartId="2.[151,244,1481,1502]" captionTargetBox="[151,1436,402,1457]" captionTargetId="figure@2.[151,1436,402,1457]" captionTargetPageId="2" captionText="FIGURE 1. Rapturella atlas n. sp. A, Holotype, MZSP 91018 (L 2.8 mm), apertural view; B, same, dorsal view; C, paratype, MZSP 107496 (L 3.1 mm), apertural view (SEM); D, same, detail of ornamentation on the last whorl; E, same, apical view; F, same, detail of protoconch, arrows indicate start of teleoconch. Scale bars: D. 100 µm, F. 200 µm." httpUri="https://zenodo.org/record/2609723/files/figure.png" pageId="1" pageNumber="126">Fig. 1C</figureCitation>
). Surface smooth, glossy, sculptured with narrow punctuated spiral grooves distributed on entire teleoconch; punctuations very small, oval (
<figureCitation id="20E425D8FFD71021FCC7FDD540B5FDAF" box="[846,932,533,556]" captionStart="FIGURE 1" captionStartId="2.[151,244,1481,1502]" captionTargetBox="[151,1436,402,1457]" captionTargetId="figure@2.[151,1436,402,1457]" captionTargetPageId="2" captionText="FIGURE 1. Rapturella atlas n. sp. A, Holotype, MZSP 91018 (L 2.8 mm), apertural view; B, same, dorsal view; C, paratype, MZSP 107496 (L 3.1 mm), apertural view (SEM); D, same, detail of ornamentation on the last whorl; E, same, apical view; F, same, detail of protoconch, arrows indicate start of teleoconch. Scale bars: D. 100 µm, F. 200 µm." httpUri="https://zenodo.org/record/2609723/files/figure.png" pageId="1" pageNumber="126">Fig. 1D</figureCitation>
); grooves regularly spaced, separated by gaps about 10 times wider than grooves; last whorl with about 11 spiral grooves, gradually becoming deeper and closer to each other towards umbilical area; 3 grooves on anterior half and 8 on posterior half; umbilicus rimate, partially covered by extension of columellar margin. Aperture about 2/3 of total length, antero-posteriorly elongated, length about 1.5 times width; posterior end pointed, anterior (siphonal) rounded. Inner lip concave; superior half convex, rounded, with thin callus, about 1.3 times length of inferior half; inferior half slightly concave, with shallow edge, and small thickened columellar fold in posterior region. Outer lip thick, with sharp edge.
</paragraph>
<paragraph id="B860395DFFD71021FF4EFCC84259FC2A" blockId="1.[151,1436,152,1943]" pageId="1" pageNumber="126">
<emphasis id="8AABE54FFFD71021FF4EFCC8427FFC9C" bold="true" box="[199,366,776,799]" pageId="1" pageNumber="126">Measurements</emphasis>
(length by width in mm). 
<typeStatus id="676487FFFFD71021FD01FCC841FEFC9C" box="[648,751,776,799]" pageId="1" pageNumber="126" type="holotype">Holotype</typeStatus>
: 2.8 by 1.6 (
<figureCitation id="20E425D8FFD71021FCFDFCC840BEFC9C" box="[884,943,776,799]" captionStart="FIGURE 1" captionStartId="2.[151,244,1481,1502]" captionTargetBox="[151,1436,402,1457]" captionTargetId="figure@2.[151,1436,402,1457]" captionTargetPageId="2" captionText="FIGURE 1. Rapturella atlas n. sp. A, Holotype, MZSP 91018 (L 2.8 mm), apertural view; B, same, dorsal view; C, paratype, MZSP 107496 (L 3.1 mm), apertural view (SEM); D, same, detail of ornamentation on the last whorl; E, same, apical view; F, same, detail of protoconch, arrows indicate start of teleoconch. Scale bars: D. 100 µm, F. 200 µm." httpUri="https://zenodo.org/record/2609723/files/figure.png" pageId="1" pageNumber="126">Fig. 1</figureCitation>
A–B). 
<typeStatus id="676487FFFFD71021FC74FCC84778FC9C" box="[1021,1129,776,799]" pageId="1" pageNumber="126" type="paratype">Paratypes</typeStatus>
: MZSP 107496 (
<figureCitation id="20E425D8FFD71021FA96FCC8464BFC9C" box="[1311,1370,776,799]" captionStart="FIGURE 1" captionStartId="2.[151,244,1481,1502]" captionTargetBox="[151,1436,402,1457]" captionTargetId="figure@2.[151,1436,402,1457]" captionTargetPageId="2" captionText="FIGURE 1. Rapturella atlas n. sp. A, Holotype, MZSP 91018 (L 2.8 mm), apertural view; B, same, dorsal view; C, paratype, MZSP 107496 (L 3.1 mm), apertural view (SEM); D, same, detail of ornamentation on the last whorl; E, same, apical view; F, same, detail of protoconch, arrows indicate start of teleoconch. Scale bars: D. 100 µm, F. 200 µm." httpUri="https://zenodo.org/record/2609723/files/figure.png" pageId="1" pageNumber="126">Fig. 1</figureCitation>
C–F), 1 shell, 3.1 by 2.2; MZSP 107497, 1 shell, 2.6 by 1.7; MZSP 107498, 1 shell, 2.7 by 1.8; MZSP 107500, 3 shells, (#1) 2.3 by 1.5, (#2) 1.7 by 1.3, (#3) 2.3 by 1.6; MZSP 138843, 1 shell, 2.7 by 1.8; MNHN 25271, 2 shells, (#1), 2.0 by 1.4, (#2) 2.5 by 1.7; MNRJ 30855, 2 shells, (#1) 1.8 by 1.3; (#2) 2.5 by 1.7; MZSP 90698, 1 shell, 3.3 by 2.2; MZSP 91018, 1 shell, 2.6 by 1.7.
</paragraph>
</subSubSection>
<subSubSection id="F0C56AD6FFD71021FF4EFC7541C3FC6C" pageId="1" pageNumber="126" type="distribution">
<paragraph id="B860395DFFD71021FF4EFC7541CDFC4F" blockId="1.[151,1436,152,1943]" box="[199,732,949,972]" pageId="1" pageNumber="126">
<emphasis id="8AABE54FFFD71021FF4EFC754248FC4F" bold="true" box="[199,345,949,972]" pageId="1" pageNumber="126">Distribution.</emphasis>
<collectingRegion id="7A1BF7BFFFD71021FED6FC7542E8FC4F" box="[351,505,949,972]" country="Brazil" name="Rio de Janeiro" pageId="1" pageNumber="126">Rio de Janeiro</collectingRegion>
to 
<collectingRegion id="7A1BF7BFFFD71021FD92FC754198FC4F" box="[539,649,949,972]" country="Brazil" name="Sao Paulo" pageId="1" pageNumber="126">São Paulo</collectingRegion>
, 
<collectingCountry id="C0C879CDFFD71021FD1DFC7541C9FC4F" box="[660,728,949,972]" name="Brazil" pageId="1" pageNumber="126">Brazil</collectingCountry>
.
</paragraph>
<paragraph id="B860395DFFD71021FF4EFC1841C3FC6C" blockId="1.[151,1436,152,1943]" box="[199,722,984,1007]" pageId="1" pageNumber="126">
<emphasis id="8AABE54FFFD71021FF4EFC184234FC6C" bold="true" box="[199,293,984,1007]" pageId="1" pageNumber="126">Habitat.</emphasis>
Muddy sand bottom, 
<quantity id="7F2794B8FFD71021FD99FC184199FC6C" box="[528,648,984,1007]" metricMagnitude="2" metricUnit="m" metricValue="1.545" metricValueMax="1.6199999999999999" metricValueMin="1.47" pageId="1" pageNumber="126" unit="m" value="154.5" valueMax="162.0" valueMin="147.0">147–162 m</quantity>
depth.
</paragraph>
</subSubSection>
<subSubSection id="F0C56AD6FFD71021FF4EFC3A42E9FB92" box="[199,504,1018,1041]" pageId="1" pageNumber="126" type="materials_examined">
<paragraph id="B860395DFFD71021FF4EFC3A42E9FB92" blockId="1.[151,1436,152,1943]" box="[199,504,1018,1041]" pageId="1" pageNumber="126">
<emphasis id="8AABE54FFFD71021FF4EFC3A42BAFB92" bold="true" box="[199,427,1018,1041]" pageId="1" pageNumber="126">Material Examined.</emphasis>
<typeStatus id="676487FFFFD71021FE3BFC3A42E4FB92" box="[434,501,1018,1041]" pageId="1" pageNumber="126">Types</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="F0C56AD6FFD71021FF4EFBDD465AFB1F" pageId="1" pageNumber="126" type="discussion">
<paragraph id="B860395DFFD71021FF4EFBDD465AFB1F" blockId="1.[151,1436,152,1943]" pageId="1" pageNumber="126">
<emphasis id="8AABE54FFFD71021FF4EFBDD4224FBB7" bold="true" box="[199,309,1053,1076]" pageId="1" pageNumber="126">Remarks.</emphasis>
<taxonomicName id="7FDF42DEFFD71021FEB5FBDC42F9FBB7" authority="Cunha &amp; Simone, 2018" authorityName="Cunha &amp; Simone" authorityYear="2018" box="[316,488,1052,1076]" class="Gastropoda" family="Acteonidae" genus="Rapturella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Heterostropha" pageId="1" pageNumber="126" phylum="Mollusca" rank="species" species="atlas" status="sp. nov.">
<emphasis id="8AABE54FFFD71021FEB5FBDC42F9FBB7" box="[316,488,1052,1076]" italics="true" pageId="1" pageNumber="126">Rapturella atlas</emphasis>
</taxonomicName>
<emphasis id="8AABE54FFFD71021FE78FBDD413FFBB7" bold="true" box="[497,558,1053,1076]" pageId="1" pageNumber="126">
<taxonomicNameLabel id="91985834FFD71021FE78FBDD413FFBB7" box="[497,558,1053,1076]" pageId="1" pageNumber="126" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
fits in the genus 
<taxonomicName id="7FDF42DEFFD71021FD60FBDC404BFBB7" authorityName="Salvador &amp; Cunha" authorityYear="2016" box="[745,858,1052,1076]" class="Gastropoda" family="Acteonidae" genus="Rapturella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Heterostropha" pageId="1" pageNumber="126" phylum="Mollusca" rank="genus">
<emphasis id="8AABE54FFFD71021FD60FBDC404BFBB7" box="[745,858,1052,1076]" italics="true" pageId="1" pageNumber="126">Rapturella</emphasis>
</taxonomicName>
because it presents most of the diagnostic features of the genus including a minute shell size with thick walls, rounded spire whorls with a subsutural shelf, a teleoconch sculptured by spiral grooves composed of rounded punctae, and the characteristic aperture with a rounded trapezoidal shape, a columellar region thickened with a weak fold, a narrow anal canal, and a rounded siphonal canal region.
</paragraph>
</subSubSection>
<subSubSection id="F0C56AD6FFD71021FF4EFB6742C5F88F" pageId="1" pageNumber="126" type="reference_group">
<paragraph id="B860395DFFD71021FF4EFB6740D3FA74" blockId="1.[151,1436,152,1943]" pageId="1" pageNumber="126">
<taxonomicName id="7FDF42DEFFD71021FF4EFB674264FB3C" ID-CoL="4RL3M" authority="Cunha &amp; Simone, 2018" authorityName="Cunha &amp; Simone" authorityYear="2018" box="[199,373,1191,1215]" class="Gastropoda" family="Acteonidae" genus="Rapturella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Heterostropha" pageId="1" pageNumber="126" phylum="Mollusca" rank="species" species="atlas" status="sp. nov.">
<emphasis id="8AABE54FFFD71021FF4EFB674264FB3C" box="[199,373,1191,1215]" italics="true" pageId="1" pageNumber="126">Rapturella atlas</emphasis>
</taxonomicName>
<emphasis id="8AABE54FFFD71021FEF7FB6842ACFB3C" bold="true" box="[382,445,1192,1215]" pageId="1" pageNumber="126">
<taxonomicNameLabel id="91985834FFD71021FEF7FB6842ACFB3C" box="[382,445,1192,1215]" pageId="1" pageNumber="126" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
most resembles 
<taxonomicName id="7FDF42DEFFD71021FDFEFB6841DEFB3C" ID-CoL="4RL3P" authorityName="Salvador &amp; Cunha" authorityYear="2016" box="[631,719,1192,1215]" class="Gastropoda" family="Acteonidae" genus="Rapturella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Heterostropha" pageId="1" pageNumber="126" phylum="Mollusca" rank="species" species="ryani">
<emphasis id="8AABE54FFFD71021FDFEFB6841DEFB3C" box="[631,719,1192,1215]" italics="true" pageId="1" pageNumber="126">R. ryani</emphasis>
</taxonomicName>
, from the 
<collectingRegion id="7A1BF7BFFFD71021FCCFFB684082FB3C" box="[838,915,1192,1215]" country="United States of America" name="Florida" pageId="1" pageNumber="126">Florida</collectingRegion>
Keys, 
<collectingCountry id="C0C879CDFFD71021FC6AFB68470DFB3C" box="[995,1052,1192,1215]" name="United States of America" pageId="1" pageNumber="126">USA</collectingCountry>
. It differs by having a smaller size (length up to 
<quantity id="7F2794B8FFD71021FEAEFB0A426AFB62" box="[295,379,1226,1249]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="1" pageNumber="126" unit="mm" value="2.5">2.5 mm</quantity>
, while 
<taxonomicName id="7FDF42DEFFD71021FE41FB0A410FFB62" ID-CoL="4RL3P" authorityName="Salvador &amp; Cunha" authorityYear="2016" box="[456,542,1226,1249]" class="Gastropoda" family="Acteonidae" genus="Rapturella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Heterostropha" pageId="1" pageNumber="126" phylum="Mollusca" rank="species" species="ryani">
<emphasis id="8AABE54FFFD71021FE41FB0A410FFB62" box="[456,542,1226,1249]" italics="true" pageId="1" pageNumber="126">R. ryani</emphasis>
</taxonomicName>
easily reaches 
<quantity id="7F2794B8FFD71021FD48FB0A4012FB62" box="[705,771,1226,1249]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="1" pageNumber="126" unit="mm" value="4.0">4 mm</quantity>
); thicker walls forming a more solid shell; 3 spiral grooves on the penultimate whorl (5 more closely spaced grooves on 
<taxonomicName id="7FDF42DEFFD71021FC99FB2D4077FA87" ID-CoL="4RL3P" authorityName="Salvador &amp; Cunha" authorityYear="2016" box="[784,870,1261,1284]" class="Gastropoda" family="Acteonidae" genus="Rapturella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Heterostropha" pageId="1" pageNumber="126" phylum="Mollusca" rank="species" species="ryani">
<emphasis id="8AABE54FFFD71021FC99FB2D4077FA87" box="[784,870,1261,1284]" italics="true" pageId="1" pageNumber="126">R. ryani</emphasis>
</taxonomicName>
), about 3 grooves in the periumbilical region (about 10 more closely spaced grooves on 
<taxonomicName id="7FDF42DEFFD71021FD9FFAD0417DFAA4" ID-CoL="4RL3P" authorityName="Salvador &amp; Cunha" authorityYear="2016" box="[534,620,1296,1319]" class="Gastropoda" family="Acteonidae" genus="Rapturella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Heterostropha" pageId="1" pageNumber="126" phylum="Mollusca" rank="species" species="ryani">
<emphasis id="8AABE54FFFD71021FD9FFAD0417DFAA4" box="[534,620,1296,1319]" italics="true" pageId="1" pageNumber="126">R. ryani</emphasis>
</taxonomicName>
), and a smaller protoconch (
<quantity id="7F2794B8FFD71021FC13FAD040EDFAA4" box="[922,1020,1296,1319]" metricMagnitude="-4" metricUnit="m" metricValue="4.2" pageId="1" pageNumber="126" unit="mm" value="0.42">0.42 mm</quantity>
, against 
<quantity id="7F2794B8FFD71021FBD1FAD047A6FAA4" box="[1112,1207,1296,1319]" metricMagnitude="-4" metricUnit="m" metricValue="4.699999999999999" pageId="1" pageNumber="126" unit="mm" value="0.47">0.47 mm</quantity>
of 
<taxonomicName id="7FDF42DEFFD71021FB50FAD0463FFAA4" ID-CoL="4RL3P" authorityName="Salvador &amp; Cunha" authorityYear="2016" box="[1241,1326,1296,1319]" class="Gastropoda" family="Acteonidae" genus="Rapturella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Heterostropha" pageId="1" pageNumber="126" phylum="Mollusca" rank="species" species="ryani">
<emphasis id="8AABE54FFFD71021FB50FAD0463FFAA4" box="[1241,1326,1296,1319]" italics="true" pageId="1" pageNumber="126">R. ryani</emphasis>
</taxonomicName>
). Despite it being a smaller species, the smaller protoconch of 
<taxonomicName id="7FDF42DEFFD71021FD44FAF24031FACA" ID-CoL="4RL3M" authorityName="Cunha &amp; Simone" authorityYear="2018" box="[717,800,1329,1353]" class="Gastropoda" family="Acteonidae" genus="Rapturella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Heterostropha" pageId="1" pageNumber="126" phylum="Mollusca" rank="species" species="atlas">
<emphasis id="8AABE54FFFD71021FD44FAF24031FACA" box="[717,800,1329,1353]" italics="true" pageId="1" pageNumber="126">R. atlas</emphasis>
</taxonomicName>
is a relevant developmental parameter. 
<taxonomicName id="7FDF42DEFFD71021FB5AFAF1466EFACA" ID-CoL="4RL3M" authority="Cunha &amp; Simone, 2018" authorityName="Cunha &amp; Simone" authorityYear="2018" box="[1235,1407,1329,1353]" class="Gastropoda" family="Acteonidae" genus="Rapturella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Heterostropha" pageId="1" pageNumber="126" phylum="Mollusca" rank="species" species="atlas" status="sp. nov.">
<emphasis id="8AABE54FFFD71021FB5AFAF1466EFACA" box="[1235,1407,1329,1353]" italics="true" pageId="1" pageNumber="126">Rapturella atlas</emphasis>
</taxonomicName>
<emphasis id="8AABE54FFFD71021FA0FFAF243A6FAEF" bold="true" pageId="1" pageNumber="126">
<taxonomicNameLabel id="91985834FFD71021FA0FFAF243A6FAEF" pageId="1" pageNumber="126" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
differs from 
<taxonomicName id="7FDF42DEFFD71021FECBFA95417AFAEF" ID-CoL="57FV3" authority="(Forbes, 1884)" authorityName="Forbes" authorityYear="1884" baseAuthorityName="Forbes" baseAuthorityYear="1884" box="[322,619,1364,1388]" class="Gastropoda" family="Acteonidae" genus="Tornatella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Heterostropha" pageId="1" pageNumber="126" phylum="Mollusca" rank="species" species="globulina">
<emphasis id="8AABE54FFFD71021FECBFA9542D4FAEF" box="[322,453,1364,1388]" italics="true" pageId="1" pageNumber="126">R. globulina</emphasis>
(Forbes, 1884)
</taxonomicName>
, from the Eastern Atlantic (
<bibRefCitation id="DC4E44ACFFD71021FC06FA95478DFAEF" author="Salvador, R. B. &amp; Cunha, C. M." box="[911,1180,1365,1388]" pageId="1" pageNumber="126" pagination="472 - 483" refId="ref2812" refString="Salvador, R. B. &amp; Cunha, C. M. (2016) Taxonomic revision of the fossil genera Bulimactaeon, Hemiauricula (= Liocarenus) and Nucleopsis, with description of a new Recent genus and species (Gastropoda: Heterobranchia: Acteonidae). Journal of Molluscan Studies, 82 (3), 472 - 483. https: // doi. org / 10.1093 / mollus / eyw 010" type="journal article" year="2016">
<collectingCountry id="C0C879CDFFD71021FC06FA9540FEFAEF" box="[911,1007,1365,1388]" name="El Salvador" pageId="1" pageNumber="126">Salvador</collectingCountry>
&amp; 
<collectingCountry id="C0C879CDFFD71021FB99FA954749FAEF" box="[1040,1112,1365,1388]" name="Saint Helena, Ascension and Tristan da Cunha" pageId="1" pageNumber="126">Cunha</collectingCountry>
, 2016
</bibRefCitation>
), by its slightly rounded whorls forming a straighter profile (whorls of 
<taxonomicName id="7FDF42DEFFD71021FD1CFAB8400AFA0C" ID-CoL="57FV3" baseAuthorityName="Forbes" baseAuthorityYear="1884" box="[661,795,1399,1423]" class="Gastropoda" family="Acteonidae" genus="Tornatella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Heterostropha" pageId="1" pageNumber="126" phylum="Mollusca" rank="species" species="globulina">
<emphasis id="8AABE54FFFD71021FD1CFAB8400AFA0C" box="[661,795,1399,1423]" italics="true" pageId="1" pageNumber="126">R. globulina</emphasis>
</taxonomicName>
are more rounded), the small subsutural shoulder on each whorl (absent on 
<taxonomicName id="7FDF42DEFFD71021FEDBFA5A42C7FA32" ID-CoL="57FV3" baseAuthorityName="Forbes" baseAuthorityYear="1884" box="[338,470,1433,1457]" class="Gastropoda" family="Acteonidae" genus="Tornatella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Heterostropha" pageId="1" pageNumber="126" phylum="Mollusca" rank="species" species="globulina">
<emphasis id="8AABE54FFFD71021FEDBFA5A42C7FA32" box="[338,470,1433,1457]" italics="true" pageId="1" pageNumber="126">R. globulina</emphasis>
</taxonomicName>
), and by the more widely spaced spiral grooves on its shell surface. Additionally, 
<taxonomicName id="7FDF42DEFFD71021FAC3FA5A468DFA32" ID-CoL="4RL3M" authority="Cunha &amp; Simone, 2018" authorityName="Cunha &amp; Simone" authorityYear="2018" box="[1354,1436,1433,1457]" class="Gastropoda" family="Acteonidae" genus="Rapturella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Heterostropha" pageId="1" pageNumber="126" phylum="Mollusca" rank="species" species="atlas" status="sp. nov.">
<emphasis id="8AABE54FFFD71021FAC3FA5A468DFA32" box="[1354,1436,1433,1457]" italics="true" pageId="1" pageNumber="126">R. atlas</emphasis>
</taxonomicName>
<emphasis id="8AABE54FFFD71021FF1EFA7D43C5FA57" bold="true" box="[151,212,1469,1492]" pageId="1" pageNumber="126">
<taxonomicNameLabel id="91985834FFD71021FF1EFA7D43C5FA57" box="[151,212,1469,1492]" pageId="1" pageNumber="126" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
differs from its congeners in lacking two closely spaced, punctate, subsutural grooves (
<bibRefCitation id="DC4E44ACFFD71021FB05FA7D43EEFA74" author="Salvador, R. B. &amp; Cunha, C. M." pageId="1" pageNumber="126" pagination="472 - 483" refId="ref2812" refString="Salvador, R. B. &amp; Cunha, C. M. (2016) Taxonomic revision of the fossil genera Bulimactaeon, Hemiauricula (= Liocarenus) and Nucleopsis, with description of a new Recent genus and species (Gastropoda: Heterobranchia: Acteonidae). Journal of Molluscan Studies, 82 (3), 472 - 483. https: // doi. org / 10.1093 / mollus / eyw 010" type="journal article" year="2016">
<collectingCountry id="C0C879CDFFD71021FB05FA7D47FDFA57" box="[1164,1260,1469,1492]" name="El Salvador" pageId="1" pageNumber="126">Salvador</collectingCountry>
&amp; 
<collectingCountry id="C0C879CDFFD71021FA98FA7D4647FA57" box="[1297,1366,1469,1492]" name="Saint Helena, Ascension and Tristan da Cunha" pageId="1" pageNumber="126">Cunha</collectingCountry>
2016: fig. 4C, F
</bibRefCitation>
), and in lacking a faint tooth-like thickening on the palatal region.
</paragraph>
<paragraph id="B860395DFFD71021FF4EF9C242C5F88F" blockId="1.[151,1436,152,1943]" pageId="1" pageNumber="126">
This discovery expands the geographic distribution of the genus 
<taxonomicName id="7FDF42DEFFD71021FC07F9C140EEF99A" ID-CoL="7PFKR" authorityName="Salvador &amp; Cunha" authorityYear="2016" box="[910,1023,1537,1561]" class="Gastropoda" family="Acteonidae" genus="Rapturella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Heterostropha" pageId="1" pageNumber="126" phylum="Mollusca" rank="genus">
<emphasis id="8AABE54FFFD71021FC07F9C140EEF99A" box="[910,1023,1537,1561]" italics="true" pageId="1" pageNumber="126">Rapturella</emphasis>
</taxonomicName>
southwards to the southwest Atlantic region off the Brazilian coast. However, the distribution of the genus remains restricted, like that of the genera 
<taxonomicName id="7FDF42DEFFD71021FF1EF987413CF9DC" ID-CoL="7NVNL" authority="Repetto &amp; Bianco, 2012" authorityName="Repetto &amp; Bianco" authorityYear="2012" box="[151,557,1607,1631]" class="Gastropoda" family="Acteonidae" genus="Callostracon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Heterostropha" pageId="1" pageNumber="126" phylum="Mollusca" rank="genus">
<emphasis id="8AABE54FFFD71021FF1EF9874233F9DC" box="[151,290,1607,1631]" italics="true" pageId="1" pageNumber="126">Callostracon</emphasis>
Repetto &amp; Bianco, 2012
</taxonomicName>
and 
<taxonomicName id="7FDF42DEFFD71021FDEBF9874073F9DC" ID-CoL="7P9FM" authority="Marcus, 1974" authorityName="Marcus" authorityYear="1974" box="[610,866,1607,1631]" class="Gastropoda" family="Acteonidae" genus="Mysouffa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Heterostropha" pageId="1" pageNumber="126" phylum="Mollusca" rank="genus">
<emphasis id="8AABE54FFFD71021FDEBF98741D4F9DC" box="[610,709,1607,1631]" italics="true" pageId="1" pageNumber="126">Mysouffa</emphasis>
<bibRefCitation id="DC4E44ACFFD71021FD44F9884073F9DC" author="Marcus E." box="[717,866,1608,1631]" pageId="1" pageNumber="126" pagination="300 - 371" refId="ref2715" refString="Marcus E. (1974) On some Cephalaspidea (Gastropoda: Opisthobranchia) from the western and middle Atlantic warm waters. Bulletin of Marine Science, 24, 300 - 371." type="journal article" year="1974">Marcus, 1974</bibRefCitation>
</taxonomicName>
, to the Atlantic and Mediterranean. This conforms to the heterogenic distribution of acteonids, with most genera confined to specific regions such as the subarctic (e.g. 
<taxonomicName id="7FDF42DEFFD71021FF1EF94D42ADF927" ID-CoL="7P9KF" authority="Thiele, 1912" authorityName="Thiele" authorityYear="1912" box="[151,444,1677,1700]" class="Gastropoda" family="Acteonidae" genus="Neactaeonina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Heterostropha" pageId="1" pageNumber="126" phylum="Mollusca" rank="genus">
<emphasis id="8AABE54FFFD71021FF1EF94D423AF927" box="[151,299,1677,1700]" italics="true" pageId="1" pageNumber="126">Neactaeonina</emphasis>
Thiele, 1912
</taxonomicName>
), the Pacific (e.g. 
<taxonomicName id="7FDF42DEFFD71021FDF7F94C40B0F927" ID-CoL="7PDPR" authority="Thiele, 1925" authorityName="Thiele" authorityYear="1925" box="[638,929,1676,1700]" class="Gastropoda" family="Acteonidae" genus="Pseudactaeon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Heterostropha" pageId="1" pageNumber="126" phylum="Mollusca" rank="genus">
<emphasis id="8AABE54FFFD71021FDF7F94C4005F927" box="[638,788,1676,1700]" italics="true" pageId="1" pageNumber="126">Pseudactaeon</emphasis>
Thiele, 1925
</taxonomicName>
and 
<taxonomicName id="7FDF42DEFFD71021FC5FF94D47E0F927" ID-CoL="7P8K9" authority="Rudman, 1971" authorityName="Rudman" authorityYear="1971" box="[982,1265,1677,1700]" class="Gastropoda" family="Acteonidae" genus="Maxacteon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Heterostropha" pageId="1" pageNumber="126" phylum="Mollusca" rank="genus">
<emphasis id="8AABE54FFFD71021FC5FF94D475DF927" box="[982,1100,1677,1700]" italics="true" pageId="1" pageNumber="126">Maxacteon</emphasis>
<bibRefCitation id="DC4E44ACFFD71021FBDDF94D47E0F927" author="Rudman W. B." box="[1108,1265,1677,1700]" pageId="1" pageNumber="126" pagination="205 - 214" refId="ref2778" refString="Rudman W. B. (1971) The family Acteonidae (Opisthobranchia, Gastropoda) in New Zealand. Journal of the Malacological Society of Australia, 2, 205 - 214." type="journal article" year="1971">Rudman, 1971</bibRefCitation>
</taxonomicName>
) or the Western Atlantic (e.g. 
<taxonomicName id="7FDF42DEFFD71021FEA5F96F4123F944" ID-CoL="7PBFW" authority="Dall, 1889" authorityName="Dall" authorityYear="1889" box="[300,562,1711,1735]" class="Gastropoda" family="Acteonidae" genus="Ovulactaeon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Heterostropha" pageId="1" pageNumber="126" phylum="Mollusca" rank="genus">
<emphasis id="8AABE54FFFD71021FEA5F96F42A5F944" box="[300,436,1711,1735]" italics="true" pageId="1" pageNumber="126">Ovulactaeon</emphasis>
Dall, 1889
</taxonomicName>
). Only the genus 
<taxonomicName id="7FDF42DEFFD71021FD7AF96F402CF945" ID-CoL="7NMTH" authorityName="Montfort" authorityYear="1810" box="[755,829,1711,1734]" class="Gastropoda" family="Acteonidae" genus="Acteon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Heterostropha" pageId="1" pageNumber="126" phylum="Mollusca" rank="genus">
<emphasis id="8AABE54FFFD71021FD7AF96F402CF945" box="[755,829,1711,1734]" italics="true" pageId="1" pageNumber="126">Acteon</emphasis>
</taxonomicName>
has a more widespread range, but, with species of such diverse shell shapes (e.g. 
<bibRefCitation id="DC4E44ACFFD71021FE27F9124158F96A" author="Rudman W. B." box="[430,585,1746,1769]" pageId="1" pageNumber="126" pagination="205 - 214" refId="ref2778" refString="Rudman W. B. (1971) The family Acteonidae (Opisthobranchia, Gastropoda) in New Zealand. Journal of the Malacological Society of Australia, 2, 205 - 214." type="journal article" year="1971">Rudman 1971</bibRefCitation>
; 
<bibRefCitation id="DC4E44ACFFD71021FDDFF91241F6F96A" author="Marcus E." box="[598,743,1746,1769]" pageId="1" pageNumber="126" pagination="300 - 371" refId="ref2715" refString="Marcus E. (1974) On some Cephalaspidea (Gastropoda: Opisthobranchia) from the western and middle Atlantic warm waters. Bulletin of Marine Science, 24, 300 - 371." type="journal article" year="1974">Marcus 1974</bibRefCitation>
; 
<bibRefCitation id="DC4E44ACFFD71021FD7AF912406AF96A" author="Valdes A." box="[755,891,1746,1769]" pageId="1" pageNumber="126" pagination="587 - 792" refId="ref2995" refString="Valdes A. (2008) Deep-sea &quot; cephalaspidean &quot; heterobranchs (Gastropoda) from the tropical southwest Pacific. In: Heros, V., Cowie, R. H. &amp; Bouchet, P. (Eds.), Tropical Deep-Sea Benthos. Vol. 25. Memoires du Museum National d'Histoire Naturelle, 196, pp. 587 - 792." type="journal article" year="2008">Valdés 2008</bibRefCitation>
), further taxonomic investigation may result in its segregation into more genera.
</paragraph>
</subSubSection>
<subSubSection id="F0C56AD6FFD71021FF4EF8D8471CF814" pageId="1" pageNumber="126" type="description">
<paragraph id="B860395DFFD71021FF4EF8D8471CF814" blockId="1.[151,1436,152,1943]" pageId="1" pageNumber="126">
The new species, as well as the genus 
<taxonomicName id="7FDF42DEFFD71021FDF6F8D741E1F8AC" authorityName="Salvador &amp; Cunha" authorityYear="2016" box="[639,752,1815,1839]" class="Gastropoda" family="Acteonidae" genus="Rapturella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Heterostropha" pageId="1" pageNumber="126" phylum="Mollusca" rank="genus">
<emphasis id="8AABE54FFFD71021FDF6F8D741E1F8AC" box="[639,752,1815,1839]" italics="true" pageId="1" pageNumber="126">Rapturella</emphasis>
</taxonomicName>
, are recognized only by a combination of few conchological characters. The lack of soft tissue samples inhibits any comparative morphological evaluations or molecular analyses. Thus, the putative relationships of the genus with other acteonids and the classification of the species, which we here assign to 
<taxonomicName id="7FDF42DEFFD71021FF75F8BF427CF814" authorityName="Salvador &amp; Cunha" authorityYear="2016" box="[252,365,1919,1943]" class="Gastropoda" family="Acteonidae" genus="Rapturella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Heterostropha" pageId="1" pageNumber="126" phylum="Mollusca" rank="genus">
<emphasis id="8AABE54FFFD71021FF75F8BF427CF814" box="[252,365,1919,1943]" italics="true" pageId="1" pageNumber="126">Rapturella</emphasis>
</taxonomicName>
, need to be tested once new quality samples become available.
</paragraph>
</subSubSection>
</treatment>
</document>