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<document ID-CLB-Dataset="24980" ID-DOI="http://dx.doi.org/10.3897/zookeys.885.38655" ID-GBIF-Dataset="8e14d288-ddb5-485a-bd43-0cd3f46754ff" ID-PMC="PMC6848236" ID-Pensoft-Pub="1313-2970-885-51" ID-Pensoft-UUID="BCEA8CB1014D5D329A2E4B2CF99881FF" ID-PubMed="31736618" ID-ZooBank="BA152050AF7344CA8CED6D30F963CBC9" ModsDocID="1313-2970-885-51" checkinTime="1572912880629" checkinUser="pensoft" docAuthor="Zlatkov, Boyan &amp; Huemer, Peter" docDate="2019" docId="2C69A24D39965AF398D52A8EF7FE9022" docLanguage="en" docName="ZooKeys 885: 51-87" docOrigin="ZooKeys 885" docSource="http://dx.doi.org/10.3897/zookeys.885.38655" docTitle="Clepsis acclivana Zerny 1933, stat. rev." docType="treatment" docVersion="6" id="BCEA8CB1014D5D329A2E4B2CF99881FF" lastPageNumber="51" masterDocId="BCEA8CB1014D5D329A2E4B2CF99881FF" masterDocTitle="Remarkable confusion in some Western Palearctic Clepsis leads to a revised taxonomic concept (Lepidoptera, Tortricidae)" masterLastPageNumber="87" masterPageNumber="51" pageNumber="51" updateTime="1732932338753" updateUser="ExternalLinkService">
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<mods:title id="B35558E331CB6132E005D9A0BACE26B3">Remarkable confusion in some Western Palearctic Clepsis leads to a revised taxonomic concept (Lepidoptera, Tortricidae)</mods:title>
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<mods:namePart id="9D29DBBF8FE330371E3BF67A3021C567">Zlatkov, Boyan</mods:namePart>
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<mods:namePart id="4F0E1B52816473E26D51D8EC42DEF138">Huemer, Peter</mods:namePart>
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<mods:date id="4573F276D10D2965B9C26D6C9E068D35">2019</mods:date>
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<mods:classification id="F70A496FD28330BF23F141DC574EFB7A">journal article</mods:classification>
<mods:identifier id="A0EE483E55F930BDEF5A58CE061C71EC" type="DOI">http://dx.doi.org/10.3897/zookeys.885.38655</mods:identifier>
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<treatment id="2C69A24D39965AF398D52A8EF7FE9022" ID-GBIF-Taxon="160133797" LSID="urn:lsid:plazi:treatment:2C69A24D39965AF398D52A8EF7FE9022" httpUri="http://treatment.plazi.org/id/2C69A24D39965AF398D52A8EF7FE9022" lastPageNumber="51" pageId="0" pageNumber="51" scope_family="Tortricidae" scope_order="Lepidoptera">
<subSubSection id="50A93CD472487606D0A40CDCF92E7A2A" pageId="0" pageNumber="51" type="nomenclature">
<paragraph id="A1B880CF5AE8CBE31601B9ED9C347A45" pageId="0" pageNumber="51">
<taxonomicName id="42B4904C7AF6B43D3384A750D486FA4F" ID-CoL="P5TZ" LSID="2C69A24D-3996-5AF3-98D5-2A8EF7FE9022" authority="Zerny, 1933" authorityName="Zerny" authorityYear="1933" class="Insecta" family="Tortricidae" genus="Clepsis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Clepsis acclivana" order="Lepidoptera" pageId="0" pageNumber="51" phylum="Arthropoda" rank="species" species="acclivana">Clepsis acclivana (Zerny, 1933)</taxonomicName>
<taxonomicNameLabel id="6DBAA37B9A0E9999EF15BB91635F0603" pageId="0" pageNumber="51">stat. rev.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="6F042D5FC09267F41FFC500CFC99ECDB" pageId="0" pageNumber="51" type="reference_group">
<paragraph id="EB21950E94590CE480AA454F4BD47C7D" pageId="0" pageNumber="51">
<taxonomicName id="139F90BD5B97320444E997EBA8BAC1EF" class="Insecta" family="Tortricidae" genus="Cacoecia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cacoecia acclivana" order="Lepidoptera" pageId="0" pageNumber="51" phylum="Arthropoda" rank="species" species="acclivana">Cacoecia acclivana</taxonomicName>
Zerny, 1933:108, pl. 1, fig. 11 (Lebanon)
</paragraph>
</subSubSection>
<subSubSection id="22E0C50AF3F8C52E50D1833E758D1F97" pageId="0" pageNumber="51" type="materials_examined">
<paragraph id="584F003E9A30639B784BA5CEBD79FC46" pageId="0" pageNumber="51">Material examined.</paragraph>
<paragraph id="7F3095AA3A3ED1FA0BB902C905D0BF8E" pageId="0" pageNumber="51">
Lectotype ♂ by designation of
<bibRefCitation id="22A5B3BB04FAE3B6F4BE1757BC791702" author="Razowski, J" journalOrPublisher="Acta zoologica cracoviensia" pageId="0" pageNumber="51" pagination="101 - 198" refId="B34" refString="Razowski, J, 1979. Revision of the Genus Clepsis Guenee (Lepidoptera, Tortricidae). Part I. Acta zoologica cracoviensia 23: 101 - 198" title="Revision of the Genus Clepsis Guenee (Lepidoptera, Tortricidae). Part I." volume="23" year="1979">Razowski (1979)</bibRefCitation>
, pinned, with 6 labels: &quot;Nord-Libanon /
<normalizedToken id="35BBF336667891D9551D48510D8672FD" originalValue="Becharré">Becharre</normalizedToken>
, 1400 m / 21.-28.vi.[19]31. Zerny&quot; &quot;Cacoecia / acclivana / Zerny Type!&quot; [handwritten]
<normalizedToken id="D380BC9202E732B2CDA23EB48B74CB61" originalValue="“Cacoecia”">&quot;Cacoecia&quot;</normalizedToken>
♂ / acclivana Zerny / N. Obraztsov det. 1965 / prep. No. V. 49&quot; [handwritten and printed]
<normalizedToken id="94665880939A886E3859C8E071F8E557" originalValue="“Lectotype”">&quot;Lectotype&quot;</normalizedToken>
[green label] &quot;Nat. hist. Mus. / Wien / Gen. Praep. / MV 2533&quot; [blue label] &quot;NHMW / Type fot / 2013&quot;; male genitalia on a slide with two labels:
<normalizedToken id="15E09264C80830E702C60FBE426B1DC4" originalValue="“Cacoecia”">&quot;Cacoecia&quot;</normalizedToken>
/ acclivana / Zerny / Nord-Libanon /
<normalizedToken id="7DD5B613AF5D00E2E6B648B9F279715D" originalValue="Becharré">Becharre</normalizedToken>
/ 1400 m 21.-28.vi / 1931 Zerny&quot; &quot;♂ / V. 49 / Mus. Vind. 2533 / Lectotypus&quot; [both handwritten].
</paragraph>
<paragraph id="87E280CAA6EE2C7EB2D3D3CBA26CB14C" pageId="0" pageNumber="51">LEBANON • 1 ♂; Bsharri; alt. 1400 m; 21-28 Jun. 1931; Zerny leg.; GS V. 49; NHMW 2533.</paragraph>
<paragraph id="5B1D6C3E658958418621D96EBA002E4A" pageId="0" pageNumber="51">
Paralectotype ♂, pinned, with four labels: &quot;Nord-Libanon /
<normalizedToken id="37FE2235217A8AF30F6B434C38AE0572" originalValue="Becharré">Becharre</normalizedToken>
, 1400 m / 11.-20.vi.[19]31. Zerny&quot;
<normalizedToken id="86F40E666E1BC0B5EB93F31136C0E4EF" originalValue="“Lectoparatype”">&quot;Lectoparatype&quot;</normalizedToken>
[green label] &quot;Cacoecia / acclivana / Zerny Type!&quot; [handwritten] &quot;NHMW / Gen. Prep. ♂ / No. 1/15.2.2018&quot;; male genitalia on a slide with two labels: &quot;Paralectotype /
<emphasis id="B7400B3EB748BCEF6CAE729464A8D672" italics="true" pageId="0" pageNumber="51">Cacoecia acclivana</emphasis>
/ Zerny, 1933 / Nord-Libanon,
<normalizedToken id="15E72EE1E88944DD026BC3A000C98724" originalValue="Becharré">Becharre</normalizedToken>
, 1400 / m, 11.-20.vi.[19]31, Zerny&quot; [red label] &quot;NHMW / Gen. prep. / ♂ / No. 1/15.2.2018 / B. Zlatkov 2018 Euparal&quot;.
</paragraph>
<paragraph id="BF81ED4CEE46746D2A1FEBF7FDB2B390" pageId="0" pageNumber="51">LEBANON • 1 ♂; Bsharri; alt. 1400 m; 11-20 Jun. 1931; Zerny leg.; GS 1/15.2.2018; NHMW.</paragraph>
</subSubSection>
<subSubSection id="F654544C3B0DFE3D8AB2297C91FC29FC" pageId="0" pageNumber="51" type="diagnosis">
<paragraph id="B979E2BC839301CCD22EABCBAEB41EDF" pageId="0" pageNumber="51">Diagnosis.</paragraph>
<paragraph id="CF02223448B54B70CFEFBF61A38D0D23" pageId="0" pageNumber="51">
<taxonomicName id="AE365864EB458FE9442BF70F7DA4D052" class="Insecta" family="Tortricidae" genus="Clepsis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Clepsis acclivana" order="Lepidoptera" pageId="0" pageNumber="51" phylum="Arthropoda" rank="species" species="acclivana">
<emphasis id="BB8F3B6E1AAE877A61563911FF072827" italics="true" pageId="0" pageNumber="51">Clepsis acclivana</emphasis>
</taxonomicName>
is most similar to
<taxonomicName id="280EAE84D0A07AEE00865DE417CB66BC" lsidName="C. trivia" pageId="0" pageNumber="51" rank="species" species="trivia">
<emphasis id="FD66AC4B090E156B5A3F981DDBC482AC" italics="true" pageId="0" pageNumber="51">C. trivia</emphasis>
</taxonomicName>
but the forewings are paler and wider with more distinct markings, the uncus is narrower, the median part of the gnathos is smaller and its arms are not angled, the sacculus is more curved, the labis is more massive and with shorter acanthae, and the apex of the phallic process is curved ventrolaterally instead of dorsally.
</paragraph>
</subSubSection>
<subSubSection id="B74C620DBB0BBFE8B2CA830CFDF62929" pageId="0" pageNumber="51" type="description">
<paragraph id="2A96A98BBC473B82156ED967C3F69DE7" pageId="0" pageNumber="51">Description.</paragraph>
<paragraph id="DB734AC90D6D867BF5830ADA2496AF36" pageId="0" pageNumber="51">
Adult. Sexual dimorphism unknown. Male (
<figureCitation id="D395319F37B7F21F48C05D2C4947C970" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Adults of Clepsis neglectana species group A-C C. neglectana: A lectotype of Tortrix neglectana B male, Southern Germany C Female, Germany D-G C. striolana: D lectotype of T. striolana E paralectotype of T. striolana F male, Austria, South Tyrol G female, Austria, South Tyrol H-K C. trivia: H holotype of T. trivia I-J males, Crete K female, Crete L-M C. acclivana: L lectotype of Cacoecia acclivana M paralectotype of C. acclivana N T. severana holotype O Cacoecia unifasciana var. semiana holotype. Scale bar: 5 mm, all to scale." figureDoi="10.3897/zookeys.885.38655.figure2" httpUri="https://binary.pensoft.net/fig/352380" pageId="0" pageNumber="51" tableDoi="10.3897/zookeys.885.38655.figure2">Fig. 2L, M</figureCitation>
). Head. Vertex pale fulvous, frons, palps and antennae with ochreous scales. Antennae with numerous sensilla trichodea as long as width of flagellomeres. Thorax dorsally pale fulvous, ventrally creamy, legs pale brown, tegula pale fulvous. Forewing with length of 7.8 mm (in both specimens), costa basally convex, apically straight, with costal fold extending from base to 0.4
<normalizedToken id="9870A24B5E0ACD8A9675E8D5D40DDEC1" originalValue="×">x</normalizedToken>
length of costal margin (
<figureCitation id="18F4DA7BF2062AF1F6C9F5A2AE8D6D08" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Costal folds of males of Clepsis spp. A C. neglectana, Germany B C. striolana, Austria C C. trivia, Crete D C. acclivana, lectotype of Cacoecia acclivana E C. consimilana, neotype of Tortrix consimilana F C. eatoniana, Spain. Scale bar: 1 mm, all to scale." figureDoi="10.3897/zookeys.885.38655.figure3" httpUri="https://binary.pensoft.net/fig/352381" pageId="0" pageNumber="51" tableDoi="10.3897/zookeys.885.38655.figure3">Fig. 3D</figureCitation>
). Upperside background pale yellowish with fulvous reticulate pattern. Markings ill-defined, consisting of brown and ferruginous scales: basal blotch faint, with remnants only at costa as dark line; median fascia narrow, more prominent at dorsum; subapical blotch reduced, dash-like. Cilia concolourous with background. Underside pale grey-brown, costal and terminal areas creamy with some reticulate pattern. Hindwings upperside monochrome pale grey, cilia concolourous, underside whitish. Abdomen pale grey. Male genitalia (
<figureCitation id="5B352193D3E9A60EF7E83B7E1BDDB573" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Male genitalia of the Clepsis neglectana species group (some without phallus) A-B C. acclivana: A Cacoecia acclivana lectotype B C. acclivana paralectotype, right valva intentionally not cleaned from scales C Clepsis semiana, holotype of Cacoecia unifasciana var. semiana. Scale bar: 500 μm, all to scale." figureDoi="10.3897/zookeys.885.38655.figure10" httpUri="https://binary.pensoft.net/fig/352388" pageId="0" pageNumber="51" tableDoi="10.3897/zookeys.885.38655.figure10">Fig. 10A, B</figureCitation>
). Uncus round apically, with parallel lateral margins, gnathos relatively small, socius membranous. Valvae pointed laterally or ventrolaterally when mounted on slide. Costal sclerite of valva very wide, with short wide labis covered with small acanthae (
<figureCitation id="E770BA8E2E2F134257D0EC31FE653DC3" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Labides of Clepsis spp. A, B C. neglectana: A Southern Germany B Finland C C. striolana, Austria, North Tyrol D C. trivia, Crete E C. acclivana, paralectotype of Cacoecia acclivana F, G C. consimilana: F Germany, neotype of Tortrix consimilana G Italy H C. eatoniana, Spain. Scale bar: 100 μm, all to scale." figureDoi="10.3897/zookeys.885.38655.figure5" httpUri="https://binary.pensoft.net/fig/352383" pageId="0" pageNumber="51" tableDoi="10.3897/zookeys.885.38655.figure5">Fig. 5E</figureCitation>
). Basal and apical parts of sacculus with equal length forming angle of ca. 140°, saccular process almost right-angled, relatively large. Membranous part of valva with protuberance devoid of tuft of setae but has deciduous scales; its terminal part with nearly symmetrical dorsal and ventral curvature, brachiola prominent, pointed laterally. Posterior part of phallus slightly sinuate, with lateral process as long as 0.29
<normalizedToken id="4212A05F5DB1DC29B908A679C0753431" originalValue="×">x</normalizedToken>
distance between anterior opening and tip of phallus, apically bent ventrolaterally. Anterior and posterior part of phallus form angle of 130°. Caulis large, diverging widely from coecum. Vesica bent at ca. 110° dorsad, with basal widening and terminal diverticulum dorsally, slightly pointed to right. Three sockets of ventroapically located deciduous cornuti adjacent to gonopore are detectable (
<figureCitation id="1701A819AA4B75B362D1E12F6A3395E9" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Phalli with vesicae everted of the Clepsis neglectana species group A C. neglectana, Germany B C. striolana, Austria C C. trivia, Crete D C. acclivana, paralectotype of Cacoecia acclivana. The phallus of each species is shown in three aspects from top to bottom: sclerotised phallic tube (without vesica) in dorsal view (viewpoint marked with black arrow), vesica in dorsal view (viewpoint marked with white arrow) and whole phallus in left view. 1 cornutus, 2 diverticulum, 3 location of gonopore, 4 phallic process, 5 caulis. Scale bar: 250 μm." figureDoi="10.3897/zookeys.885.38655.figure6" httpUri="https://binary.pensoft.net/fig/352384" pageId="0" pageNumber="51" tableDoi="10.3897/zookeys.885.38655.figure6">Fig. 6D</figureCitation>
).
</paragraph>
<caption id="B4E4808A3EBE0831D54A2DE935E5EA3F" doi="10.3897/zookeys.885.38655.figure10" httpUri="https://binary.pensoft.net/fig/352388" pageId="0" pageNumber="51" start="Figure 10" startId="F10">
<paragraph id="56896D1D13CD33489F4AB2BAA6FECD5C" pageId="0" pageNumber="51">
<emphasis id="9AF92BF7682C0D6DCC9D022DCF15E3A8" bold="true" pageId="0" pageNumber="51">Figure 10.</emphasis>
Male genitalia of the
<taxonomicName id="CDED6E92CB88A569669C753BF6A044F7" class="Insecta" family="Tortricidae" genus="Clepsis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Clepsis neglectana" order="Lepidoptera" pageId="0" pageNumber="51" phylum="Arthropoda" rank="species" species="neglectana">
<emphasis id="4909C1726966E929A1F89B6D2D7A96B5" italics="true" pageId="0" pageNumber="51">Clepsis neglectana</emphasis>
</taxonomicName>
species group (some without phallus)
<emphasis id="9306EA7C49D2CB1508A93FBB0DF81D39" bold="true" pageId="0" pageNumber="51">
<normalizedToken id="4C05444B903F60ED518193293F6B3905" originalValue="AB">A-B</normalizedToken>
</emphasis>
<taxonomicName id="D6B0F180519F6EA40A9D0DB8223ED0AD" lsidName="C. acclivana" pageId="0" pageNumber="51" rank="species" species="acclivana">
<emphasis id="FB5AEB50EF4F64C4486EEA74081609CA" italics="true" pageId="0" pageNumber="51">C. acclivana</emphasis>
</taxonomicName>
:
<emphasis id="8C43079021FF138BD6999832FA6C5809" bold="true" pageId="0" pageNumber="51">A</emphasis>
<taxonomicName id="0F4FE7775D1B98437F5180B2A7FE37BE" class="Insecta" family="Tortricidae" genus="Cacoecia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cacoecia acclivana" order="Lepidoptera" pageId="0" pageNumber="51" phylum="Arthropoda" rank="species" species="acclivana">
<emphasis id="C7D10E9D94D9A925938BE0C19C265B81" italics="true" pageId="0" pageNumber="51">Cacoecia acclivana</emphasis>
</taxonomicName>
lectotype
<emphasis id="1176E1A3BE4292975EB415415A5936E1" bold="true" pageId="0" pageNumber="51">B</emphasis>
<taxonomicName id="FD5E55FDFD5D918EA5BDFB832FDF768A" lsidName="C. acclivana" pageId="0" pageNumber="51" rank="species" species="acclivana">
<emphasis id="FA39B5DDB4015D2D295E8837EEFCA8C3" italics="true" pageId="0" pageNumber="51">C. acclivana</emphasis>
</taxonomicName>
paralectotype, right valva intentionally not cleaned from scales
<emphasis id="75B38B163CA19AC9B16457D935D0F597" bold="true" pageId="0" pageNumber="51">C</emphasis>
<taxonomicName id="7A74715B1895F2983D8973571527B381" class="Insecta" family="Tortricidae" genus="Clepsis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Clepsis semiana" order="Lepidoptera" pageId="0" pageNumber="51" phylum="Arthropoda" rank="species" species="semiana">
<emphasis id="4CD697DD74ED7DB2335A9FE635204348" italics="true" pageId="0" pageNumber="51">Clepsis semiana</emphasis>
</taxonomicName>
, holotype of
<taxonomicName id="55331AC9AD607FBFAB0A8079033F98D4" class="Insecta" family="Tortricidae" genus="Cacoecia" higherTaxonomySource="CoL" infraspecific-rank="var." kingdom="Animalia" lsidName="Cacoecia unifasciana var. semiana" order="Lepidoptera" pageId="0" pageNumber="51" phylum="Arthropoda" rank="variety" species="unifasciana" variety="semiana">Cacoecia unifasciana var. semiana</taxonomicName>
. Scale bar: 500
<normalizedToken id="C9C62A9DD3AAC508FE10E0A4E50AECB2" originalValue="μm">μm</normalizedToken>
, all to scale.
</paragraph>
</caption>
<paragraph id="90182FC7F466E2EB79FAA0DECC9D24A8" pageId="0" pageNumber="51">Female unknown.</paragraph>
<paragraph id="54F961463FB4FC8A32150D2EF966C8A9" pageId="0" pageNumber="51">Preimaginal stages unknown.</paragraph>
</subSubSection>
<subSubSection id="F1984BFA1D4D0BD403907712CEEC43ED" pageId="0" pageNumber="51" type="molecular data">
<paragraph id="7708EA9D71F70167F9244163FC2C9E2A" pageId="0" pageNumber="51">Molecular data.</paragraph>
<paragraph id="93E7C48437C99E2CF2B3EAAF58A1907B" pageId="0" pageNumber="51">Unknown.</paragraph>
</subSubSection>
<subSubSection id="24A1DFCAF3EB656CBAC168F816E2A39B" pageId="0" pageNumber="51" type="distribution">
<paragraph id="76B4B8E79807CA22C4DBC87FACC7F0CB" pageId="0" pageNumber="51">Distribution.</paragraph>
<paragraph id="F8DFD7AA879357C7BC06DC2F2B65ED22" pageId="0" pageNumber="51">
Known from the type locality only: Lebanon (
<figureCitation id="B92C5036D86D9BEF9FDDF6FAFD5D3901" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Distribution map of examined material of Clepsis neglectana and C. consimilana species groups. Map created with SimpleMappr (http: // www. simplemappr. net)." figureDoi="10.3897/zookeys.885.38655.figure17" httpUri="https://binary.pensoft.net/fig/352395" pageId="0" pageNumber="51" tableDoi="10.3897/zookeys.885.38655.figure17">Fig. 17</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="108C63E8AD74C66DCC6C4E25A5FCACAF" pageId="0" pageNumber="51" type="ecology">
<paragraph id="E2AAE79B583E02D0B72A143C473C6D39" pageId="0" pageNumber="51">Ecology.</paragraph>
<paragraph id="67C16F9F1D03474FF78FD6E61498003A" pageId="0" pageNumber="51">Not known.</paragraph>
</subSubSection>
<subSubSection id="6CC8E260E661507E652877EEFDB41EA5" pageId="0" pageNumber="51" type="remarks">
<paragraph id="1583BDA574B70A717137E4724BF80212" pageId="0" pageNumber="51">Remarks.</paragraph>
<paragraph id="52CE6BC6830155187028BCAB39731A51" pageId="0" pageNumber="51">
Comparison of the wing pattern and genitalia of the lectotype and paralectotype of
<taxonomicName id="F47E845EAF45AD87357EF74C39298CA2" lsidName="C. acclivana" pageId="0" pageNumber="51" rank="species" species="acclivana">
<emphasis id="EE69DB9452BEB1CFD4A4B7B75BB2C838" italics="true" pageId="0" pageNumber="51">C. acclivana</emphasis>
</taxonomicName>
with the species considered above confirmed the assumption that it is a distinct species. The barcoding distance to the other species was not studied because of lack of fresh material, but the great similarity in all morphological characters of the two specimens convinced us that they can represent a distinct species, therefore we resurrect the initial status of the taxon
<taxonomicName id="6CCE45C43066C946A6F8F4930F3CB9C0" lsidName="acclivana" pageId="0" pageNumber="51" rank="species" species="acclivana">
<emphasis id="50C17E5B51D8EF1A95CFFB40111011E5" italics="true" pageId="0" pageNumber="51">acclivana</emphasis>
</taxonomicName>
synonymised by
<bibRefCitation id="6303623072D43970D5879B7C6A44FEB9" author="Razowski, J" journalOrPublisher="Acta zoologica cracoviensia" pageId="0" pageNumber="51" pagination="101 - 198" refId="B34" refString="Razowski, J, 1979. Revision of the Genus Clepsis Guenee (Lepidoptera, Tortricidae). Part I. Acta zoologica cracoviensia 23: 101 - 198" title="Revision of the Genus Clepsis Guenee (Lepidoptera, Tortricidae). Part I." volume="23" year="1979">Razowski (1979)</bibRefCitation>
with
<taxonomicName id="77B4C9F846D5C0D3A63CF7A6660A4552" lsidName="C. neglectana" pageId="0" pageNumber="51" rank="species" species="neglectana">
<emphasis id="610EC6C9517FF160ED8F81744D1F560C" italics="true" pageId="0" pageNumber="51">C. neglectana</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="F625EF0385829E84E5ABB78B44FC0E73" pageId="0" pageNumber="51" type="taxa incertae sedis">
<paragraph id="EC6F678ED34C469CAEB890B0DF775CA7" pageId="0" pageNumber="51">Taxa incertae sedis.</paragraph>
<paragraph id="406082EA3E7D29E7097AE5506A7A934C" pageId="0" pageNumber="51">
Without further morphological or genetic support the following species cannot be interpreted with certainty, but the synonymy with
<taxonomicName id="E2A028C775D3DE7BD783FEE7581A0FD6" lsidName="C. neglectana" pageId="0" pageNumber="51" rank="species" species="neglectana">
<emphasis id="EE858B0B5E5057C8F0E853995CDC7F36" italics="true" pageId="0" pageNumber="51">C. neglectana</emphasis>
</taxonomicName>
,
<taxonomicName id="6D09B6B33F04DEA2F566D0FE96DE68DF" lsidName="C. acclivana" pageId="0" pageNumber="51" rank="species" species="acclivana">
<emphasis id="5D8700A69EF168F23BE4A31DBC73D023" italics="true" pageId="0" pageNumber="51">C. acclivana</emphasis>
</taxonomicName>
or
<taxonomicName id="C91272943BD00A1BE4DE204A4249461F" lsidName="C. trivia" pageId="0" pageNumber="51" rank="species" species="trivia">
<emphasis id="3259321A5AD660CCD515D73C000EB4E1" italics="true" pageId="0" pageNumber="51">C. trivia</emphasis>
</taxonomicName>
does not seem justified for now. They demonstrate some morphological gap; at least the differences between them are not smaller than the differences with the above mentioned species of the group. Additional material and genetic study is necessary to solve their real status.
</paragraph>
</subSubSection>
</treatment>
</document>