354 lines
43 KiB
XML
354 lines
43 KiB
XML
<document id="706BB1F272EE62CBC3A39D2E4BAD238D" ID-CLB-Dataset="8059" ID-DOI="10.11646/zootaxa.4933.2.1" ID-GBIF-Dataset="1a2ba1ca-028f-4b95-8dda-5ddec4e63524" ID-ISSN="1175-5326" ID-Zenodo-Dep="4550146" ID-ZooBank="5D1D20A5-0F44-4AEF-AF5F-A758FC37D076" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1613716226573" checkinUser="plazi" docAuthor="Anistratenko, Vitaliy V., Neubauer, Thomas A., Anistratenko, Olga Yu., Kijashko, Pavel V. & Wesselingh, Frank P." docDate="2021" docId="03BF87A3C84AFFBBFF3DFF49FE5BA167" docLanguage="en" docName="zootaxa.4933.2.1.pdf" docOrigin="Zootaxa 4933 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Andrusovia antecessor Anistratenko & Neubauer & Anistratenko & Kijashko & Wesselingh 2021, sp. nov." docType="treatment" docUuid="528249A2-5494-4E14-BA1F-4AB37949EA0D" docUuidSource="ZooBank" docVersion="10" lastPageNumber="163" masterDocId="FF86FFDBC841FFB7FFAAFFDEFFA5A577" masterDocTitle="A revision of the Pontocaspian gastropods of the subfamily Caspiinae (Caenogastropoda: Hydrobiidae)" masterLastPageNumber="197" masterPageNumber="151" pageNumber="162" updateTime="1698930299740" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="9F235B0C0EF7AA25BA9E122D51FBC394">A revision of the Pontocaspian gastropods of the subfamily Caspiinae (Caenogastropoda: Hydrobiidae)</mods:title>
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<mods:namePart id="CD41BFD87C00ED6A337253DBE61D40B4">Anistratenko, Vitaliy V.</mods:namePart>
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<mods:affiliation id="0C28979D797C5E6E808C990E24917977">Department of Invertebrate Fauna and Systematics, Schmalhausen Institute of Zoology, National Academy of Sciences of Ukraine, B. Khmelnytsky Str. 15, 01030 Kiev, Ukraine.</mods:affiliation>
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<mods:namePart id="2A246401F57C771730B7CDF7BBDC5250">Neubauer, Thomas A.</mods:namePart>
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<mods:affiliation id="0FE001598F728691005E9C648213E91B">Department of Animal Ecology and Systematics, Justus Liebig University, Heinrich-Buff-Ring 26 - 32 IFZ, 35392 Giessen, Germany. tneub @ zo. jlug. de; https: // orcid. org / 0000 - 0002 - 1398 - 9941 & Naturalis Biodiversity Center, Darwinweg 2, 2333 CR Leiden, The Netherlands.</mods:affiliation>
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<mods:namePart id="E43A6EBB02F395372DFC401987C0B32B">Anistratenko, Olga Yu.</mods:namePart>
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<mods:affiliation id="DE3CE9EA692E3829EA2BAC7C5305B396">Department of Invertebrate Fauna and Systematics, Schmalhausen Institute of Zoology, National Academy of Sciences of Ukraine, B. Khmelnytsky Str. 15, 01030 Kiev, Ukraine. & Department of Cainozoic Deposits, Institute of Geological Sciences, National Academy of Sciences of Ukraine, O. Gontchar Str. 55 b, 01054 Kiev, Ukraine.</mods:affiliation>
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<mods:namePart id="0D897ABE7DB6080B8E7153B1C9F6B889">Kijashko, Pavel V.</mods:namePart>
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<mods:affiliation id="5D50935C7ADE25539E58225849B9A8F2">Zoological Institute, Russian Academy of Sciences, Universitetskaya Emb. 1, 199034 Saint Petersburg, Russia.</mods:affiliation>
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<mods:affiliation id="484387FFA63812EEA83D3533271C507A">Naturalis Biodiversity Center, Darwinweg 2, 2333 CR Leiden, The Netherlands. & Department of Earth Sciences, Utrecht University, Budapestlaan 17, 3584 CD Utrecht, The Netherlands</mods:affiliation>
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<mods:date id="796FEFFC43FC043F3FC3EE97F7324AA2">2021</mods:date>
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<treatment id="03BF87A3C84AFFBBFF3DFF49FE5BA167" ID-DOI="http://doi.org/10.5281/zenodo.4559928" ID-GBIF-Taxon="179906632" ID-Zenodo-Dep="4559928" LSID="urn:lsid:zoobank.org:act:528249A2-5494-4E14-BA1F-4AB37949EA0D" httpUri="http://treatment.plazi.org/id/03BF87A3C84AFFBBFF3DFF49FE5BA167" lastPageId="12" lastPageNumber="163" pageId="11" pageNumber="162">
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<emphasis id="B962EAA7C84AFFBCFF3DFF49FDB8A5C6" bold="true" box="[151,541,151,178]" pageId="11" pageNumber="162">
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†
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<taxonomicName id="4C164D36C84AFFBCFF0CFF49FE1DA5C6" authority="Anistratenko & Neubauer & Anistratenko & Kijashko & Wesselingh, 2021" authorityName="Anistratenko & Neubauer & Anistratenko & Kijashko & Wesselingh" authorityYear="2021" box="[166,440,151,178]" class="Kinetofragminophora" family="Corynophryidae" genus="Andrusovia" kingdom="Protozoa" order="Suctorida" pageId="11" pageNumber="162" phylum="Ciliophora" rank="species" species="antecessor" status="sp. nov.">
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<emphasis id="B962EAA7C84AFFBCFF0CFF49FE1DA5C6" bold="true" box="[166,440,151,178]" italics="true" pageId="11" pageNumber="162">Andrusovia antecessor</emphasis>
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urn:lsid:zoobank.org:act:
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<paragraph id="8BA936B5C84AFFBCFF3DFEFAFA88A44B" blockId="11.[151,1325,291,316]" box="[151,1325,291,316]" pageId="11" pageNumber="162">
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<taxonomicName id="4C164D36C84AFFBCFF3DFEFAFD94A44B" authority="Starobogatov, 2000" authorityName="Starobogatov" authorityYear="2000" box="[151,561,292,316]" class="Kinetofragminophora" family="Corynophryidae" genus="Andrusovia" kingdom="Protozoa" order="Suctorida" pageId="11" pageNumber="162" phylum="Ciliophora" rank="species" species="brusinai">
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<emphasis id="B962EAA7C84AFFBCFF3DFEFAFEC3A44B" box="[151,358,292,316]" italics="true" pageId="11" pageNumber="162">Andrusovia brusinai</emphasis>
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<bibRefCitation id="EF874B44C84AFFBCFEC7FEFAFD94A44B" author="Starobogatov, Ya. I." box="[365,561,292,316]" pageId="11" pageNumber="162" pagination="37 - 42" refId="ref33581" refString="Starobogatov, Ya. I. (2000) Caspian endemic genus Andrusovia (Gastropoda Pectinibranchia Horatiidae). Ruthenica, 10 (1), 37 - 42." type="journal article" year="2000">Starobogatov, 2000</bibRefCitation>
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</taxonomicName>
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—
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<treatmentCitation id="0AB710A4C84AFFBCFDE1FEFAFCCAA44C" author="Neubauer, T. A. & van de Velde, S. & Yanina, T. A. & Wesselingh, F. P." box="[587,879,291,316]" page="54" pageId="11" pageNumber="162" year="2018">
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<bibRefCitation id="EF874B44C84AFFBCFDE1FEFAFCCAA44C" author="Neubauer, T. A. & van de Velde, S. & Yanina, T. A. & Wesselingh, F. P." box="[587,879,291,316]" pageId="11" pageNumber="162" pagination="43 - 103" refId="ref32549" refString="Neubauer, T. A., van de Velde, S., Yanina, T. A. & Wesselingh, F. P. (2018) A late Pleistocene gastropod fauna from the northern Caspian Sea with implications for Pontocaspian gastropod taxonomy. ZooKeys, 770, 43 - 103. https: // doi. org / 10.3897 / zookeys. 770.25365" type="journal article" year="2018">
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Neubauer
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<emphasis id="B962EAA7C84AFFBCFD1EFEFAFD44A44B" box="[692,737,292,316]" italics="true" pageId="11" pageNumber="162">et al</emphasis>
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. 2018: 54–56
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</bibRefCitation>
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</treatmentCitation>
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, fig. 6F–K, M, N [non
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<bibRefCitation id="EF874B44C84AFFBCFBF3FEFAFA86A44B" author="Starobogatov, Ya. I." box="[1113,1315,292,316]" pageId="11" pageNumber="162" pagination="37 - 42" refId="ref33581" refString="Starobogatov, Ya. I. (2000) Caspian endemic genus Andrusovia (Gastropoda Pectinibranchia Horatiidae). Ruthenica, 10 (1), 37 - 42." type="journal article" year="2000">Starobogatov, 2000</bibRefCitation>
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].
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<emphasis id="B962EAA7C84AFFBCFF3DFEB4FEE2A4F4" bold="true" box="[151,327,361,388]" pageId="11" pageNumber="162">Type material.</emphasis>
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<materialsCitation id="3B7E3CE8C84AFFBCFEE5FEB4FB77A4F2" ID-GBIF-Occurrence="3042095329" box="[335,1234,362,389]" collectionCode="RGM" pageId="11" pageNumber="162" specimenCode="RGM 1309839" specimenCount="1" typeStatus="holotype">
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<typeStatus id="54AD8817C84AFFBCFEE5FEB4FE19A4F3" box="[335,444,362,388]" pageId="11" pageNumber="162" type="holotype">Holotype</typeStatus>
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:
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<specimenCode id="DBB09ECEC84AFFBCFE6DFEB4FDD3A4F3" box="[455,630,362,388]" collectionCode="RGM" country="Netherlands" httpUri="http://grbio.org/cool/47nz-wbm0" name="National Museum of Natural History, Naturalis" pageId="11" pageNumber="162" type="Museum">RGM 1309839</specimenCode>
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: 1.81 x
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<quantity id="4CEE9B50C84AFFBCFD78FEB4FC9DA4F3" box="[722,824,362,388]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="11" pageNumber="162" unit="mm" value="1.8">1.80 mm</quantity>
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(shell height x width;
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<figureCitation id="132D2A30C84AFFBCFB96FEB4FB6CA4F2" box="[1084,1225,362,389]" captionStart="FIGURE 4" captionStartId="11.[151,250,1932,1956]" captionTargetBox="[151,1436,736,1909]" captionTargetId="figure-167@11.[151,1436,736,1909]" captionTargetPageId="11" captionText="FIGURE 4. Andrusovia antecessor sp. nov. from the early Late Pleistocene of Selitrennoye, Astrakhan region, Russia (locality 1). (a, d, g) Paratype (RGM 1309840). (b, c, h) Holotype (RGM 1309839). (e, f, i) Paratype (LV 201509). Scale bars = 500 μm (a–f), 100 μm (g–i)." figureDoi="http://doi.org/10.5281/zenodo.4550158" httpUri="https://zenodo.org/record/4550158/files/figure.png" pageId="11" pageNumber="162">Fig. 4b, c, h</figureCitation>
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)
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</materialsCitation>
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.
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<materialsCitation id="3B7E3CE8C84AFFBCFB75FEB4FDFEA4DF" ID-GBIF-Occurrence="3042095342" collectionCode="RGM" pageId="11" pageNumber="162" specimenCode="RGM 1309840" specimenCount="1" typeStatus="paratype">
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<typeStatus id="54AD8817C84AFFBCFB75FEB4FAF5A4F3" box="[1247,1360,362,388]" pageId="11" pageNumber="162" type="paratype">Paratypes</typeStatus>
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:
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<specimenCode id="DBB09ECEC84AFFBCFAF6FEB4FF58A4DF" collectionCode="RGM" country="Netherlands" httpUri="http://grbio.org/cool/47nz-wbm0" name="National Museum of Natural History, Naturalis" pageId="11" pageNumber="162" type="Museum">RGM 1309840</specimenCode>
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: 1.52 x
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<quantity id="4CEE9B50C84AFFBCFEFFFE50FE1CA4DF" box="[341,441,398,424]" metricMagnitude="-3" metricUnit="m" metricValue="1.44" pageId="11" pageNumber="162" unit="mm" value="1.44">1.44 mm</quantity>
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(
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<figureCitation id="132D2A30C84AFFBCFE62FE50FDF4A4DF" box="[456,593,398,424]" captionStart="FIGURE 4" captionStartId="11.[151,250,1932,1956]" captionTargetBox="[151,1436,736,1909]" captionTargetId="figure-167@11.[151,1436,736,1909]" captionTargetPageId="11" captionText="FIGURE 4. Andrusovia antecessor sp. nov. from the early Late Pleistocene of Selitrennoye, Astrakhan region, Russia (locality 1). (a, d, g) Paratype (RGM 1309840). (b, c, h) Holotype (RGM 1309839). (e, f, i) Paratype (LV 201509). Scale bars = 500 μm (a–f), 100 μm (g–i)." figureDoi="http://doi.org/10.5281/zenodo.4550158" httpUri="https://zenodo.org/record/4550158/files/figure.png" pageId="11" pageNumber="162">Fig. 4a, d, g</figureCitation>
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)
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</materialsCitation>
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;
|
||
<materialsCitation id="3B7E3CE8C84AFFBCFDCDFE50FB34A4DE" ID-GBIF-Occurrence="3042095334" accessionNumber="LV 201509" box="[615,1169,398,425]" collectionCode="MSU" pageId="11" pageNumber="162" specimenCount="1" typeStatus="paratype">
|
||
LV 201509 (
|
||
<collectionCode id="ED07AE70C84AFFBCFD59FE50FC93A4DF" box="[755,822,398,424]" country="USA" httpUri="http://grbio.org/cool/53w1-8r3k" name="Michigan State University Museum" pageId="11" pageNumber="162" type="University or college">MSU</collectionCode>
|
||
): 1.54 x
|
||
<quantity id="4CEE9B50C84AFFBCFC3CFE50FC5FA4DF" box="[918,1018,398,425]" metricMagnitude="-3" metricUnit="m" metricValue="1.55" pageId="11" pageNumber="162" unit="mm" value="1.55">1.55 mm</quantity>
|
||
(
|
||
<figureCitation id="132D2A30C84AFFBCFBA3FE50FB23A4DE" box="[1033,1158,398,425]" captionStart="FIGURE 4" captionStartId="11.[151,250,1932,1956]" captionTargetBox="[151,1436,736,1909]" captionTargetId="figure-167@11.[151,1436,736,1909]" captionTargetPageId="11" captionText="FIGURE 4. Andrusovia antecessor sp. nov. from the early Late Pleistocene of Selitrennoye, Astrakhan region, Russia (locality 1). (a, d, g) Paratype (RGM 1309840). (b, c, h) Holotype (RGM 1309839). (e, f, i) Paratype (LV 201509). Scale bars = 500 μm (a–f), 100 μm (g–i)." figureDoi="http://doi.org/10.5281/zenodo.4550158" httpUri="https://zenodo.org/record/4550158/files/figure.png" pageId="11" pageNumber="162">Fig. 4e, f, i</figureCitation>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BA936B5C84AFFBCFF6DFE6CFD71A487" blockId="11.[151,1437,361,712]" pageId="11" pageNumber="162">
|
||
<materialsCitation id="3B7E3CE8C84AFFBCFF6DFE6CFD75A487" ID-GBIF-Occurrence="3042095303" collectionCode="WGS" country="Russia" latitude="47.172554" location="Selitrennoye" longLatPrecision="1" longitude="47.44039" municipality="northern Caspian Basin" pageId="11" pageNumber="162" specimenCode="WGS84" specimenCount="1" stateProvince="Astrakhan'" typeStatus="holotype">
|
||
<emphasis id="B962EAA7C84AFFBCFF6DFE6CFEC0A4BC" bold="true" box="[199,357,433,460]" pageId="11" pageNumber="162">
|
||
<typeStatus id="54AD8817C84AFFBCFF6DFE6CFEA7A4BB" box="[199,258,434,460]" pageId="11" pageNumber="162">Type</typeStatus>
|
||
locality.
|
||
</emphasis>
|
||
<location id="8EC9606EC84AFFBCFEC6FE6CFE59A4BB" LSID="urn:lsid:plazi:treatment:03BF87A3C84AFFBBFF3DFF49FE5BA167:8EC9606EC84AFFBCFEC6FE6CFE59A4BB" box="[364,508,434,460]" country="Russia" latitude="47.172554" longLatPrecision="1" longitude="47.44039" municipality="northern Caspian Basin" name="Selitrennoye" pageId="11" pageNumber="162" stateProvince="Astrakhan'">Selitrennoye</location>
|
||
,
|
||
<collectingRegion id="49D2F857C84AFFBCFDACFE6CFD6FA4BB" box="[518,714,434,460]" country="Russia" name="Astrakhan'" pageId="11" pageNumber="162">Astrakhan region</collectingRegion>
|
||
,
|
||
<collectingCountry id="F3017625C84AFFBCFD7CFE6CFC84A4BB" box="[726,801,434,460]" name="Russia" pageId="11" pageNumber="162">Russia</collectingCountry>
|
||
(
|
||
<tableCitation id="C694030EC84AFFBCFC84FE6CFC26A4BB" box="[814,899,434,460]" captionStart="TABLE 1" captionStartId="3.[151,239,473,499]" captionTargetPageId="3" captionText="TABLE 1. Locality information of the studied material. Coordinates of collecting sites are given based on data in museum catalogue cards, labels and corresponding publications. Asterisks mark localities with approximate geographic coordinates based on indirect data provided in the ZIN catalogue. Abbreviations: q.—quadrant, st.—station." httpUri="http://table.plazi.org/id/DF69663DC842FFB4FF3DFE07F8C6A760" pageId="11" pageNumber="162" tableUuid="DF69663DC842FFB4FF3DFE07F8C6A760">Table 1</tableCitation>
|
||
, locality 1);
|
||
<collectingMunicipality id="6BCDACCFC84AFFBCFBB8FE6CFAB9A4BB" box="[1042,1308,434,460]" pageId="11" pageNumber="162">northern Caspian Basin</collectingMunicipality>
|
||
; GPS coordinates:
|
||
<geoCoordinate id="EE225072C84AFFBCFF5FFE08FE01A487" box="[245,420,470,496]" degrees="47" direction="north" minutes="10" orientation="latitude" pageId="11" pageNumber="162" precision="1" seconds="21.19" value="47.172554">47°10’21.19”N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE225072C84AFFBCFE05FE08FDFCA486" box="[431,601,470,497]" degrees="47" direction="east" minutes="26" orientation="longitude" pageId="11" pageNumber="162" precision="1" seconds="25.41" value="47.44039">47°26’25.41”E</geoCoordinate>
|
||
(
|
||
<specimenCode id="DBB09ECEC84AFFBCFDC2FE08FD6DA487" box="[616,712,470,496]" collectionCode="WGS" pageId="11" pageNumber="162">WGS84</specimenCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BA936B5C84AFFBCFF6DFE27FBE9A763" blockId="11.[151,1437,361,712]" box="[199,1100,505,532]" pageId="11" pageNumber="162">
|
||
<emphasis id="B962EAA7C84AFFBCFF6DFE27FF58A764" bold="true" box="[199,253,505,531]" pageId="11" pageNumber="162">Age.</emphasis>
|
||
Hyrcanian, early Late Pleistocene, c. 107 ± 7 ka (
|
||
<bibRefCitation id="EF874B44C84AFFBCFC86FE24FBE5A763" author="van de Velde, S. & Yanina, T. A. & Neubauer, T. A. & Wesselingh, F. P." box="[812,1088,506,532]" pageId="11" pageNumber="162" pagination="1227 - 1239" refId="ref34097" refString="van de Velde, S., Yanina, T. A., Neubauer, T. A. & Wesselingh, F. P. (2020) The Late Pleistocene mollusk fauna of Selitrennoye (Astrakhan province, Russia): A natural baseline for endemic Caspian Sea faunas. Journal of Great Lakes Research, 46 (5), 1227 - 1239. https: // doi. org / 10.1016 / j. jglr. 2019.04.001" type="journal article" year="2020">
|
||
van de Velde
|
||
<emphasis id="B962EAA7C84AFFBCFC6EFE25FC53A763" box="[964,1014,506,532]" italics="true" pageId="11" pageNumber="162">et al</emphasis>
|
||
. 2020
|
||
</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BA936B5C84AFFBCFF6DFDC3FD4FA74E" blockId="11.[151,1437,361,712]" box="[199,746,541,569]" pageId="11" pageNumber="162">
|
||
<emphasis id="B962EAA7C84AFFBCFF6DFDC3FE24A740" bold="true" box="[199,385,541,567]" pageId="11" pageNumber="162">Other material.</emphasis>
|
||
<materialsCitation id="3B7E3CE8C84AFFBCFE22FDC0FD43A74E" ID-GBIF-Occurrence="3042095335" box="[392,742,542,569]" collectionCode="RGM" pageId="11" pageNumber="162" specimenCode="RGM 1310206" specimenCount="36">
|
||
<specimenCount id="9D10FD3CC84AFFBCFE22FDC0FD85A74F" box="[392,544,542,568]" pageId="11" pageNumber="162" type="generic">36 specimens</specimenCount>
|
||
(
|
||
<specimenCode id="DBB09ECEC84AFFBCFD84FDC0FD7BA74E" box="[558,734,542,569]" collectionCode="RGM" country="Netherlands" httpUri="http://grbio.org/cool/47nz-wbm0" name="National Museum of Natural History, Naturalis" pageId="11" pageNumber="162" type="Museum">RGM 1310206</specimenCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30C653EC84AFFBCFF6DFD9CFEB1A7F7" pageId="11" pageNumber="162" type="etymology">
|
||
<paragraph id="8BA936B5C84AFFBCFF6DFD9CFEB1A7F7" blockId="11.[151,1437,361,712]" pageId="11" pageNumber="162">
|
||
<emphasis id="B962EAA7C84AFFBCFF6DFD9CFEEBA72B" bold="true" box="[199,334,578,604]" pageId="11" pageNumber="162">Etymology.</emphasis>
|
||
Referring to the supposed position as predecessor to modern
|
||
<taxonomicName id="4C164D36C84AFFBCFBA4FD9CFB2AA72B" box="[1038,1167,578,604]" class="Kinetofragminophora" family="Corynophryidae" genus="Andrusovia" kingdom="Protozoa" order="Suctorida" pageId="11" pageNumber="162" phylum="Ciliophora" rank="genus">
|
||
<emphasis id="B962EAA7C84AFFBCFBA4FD9CFB2AA72B" box="[1038,1167,578,604]" italics="true" pageId="11" pageNumber="162">Andrusovia</emphasis>
|
||
</taxonomicName>
|
||
. The name is a noun in apposition.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30C653EC84AFFBCFF6DFD57FBE5A7BF" pageId="11" pageNumber="162" type="diagnosis">
|
||
<paragraph id="8BA936B5C84AFFBCFF6DFD57FBE5A7BF" blockId="11.[151,1437,361,712]" pageId="11" pageNumber="162">
|
||
<emphasis id="B962EAA7C84AFFBCFF6DFD57FEE7A7D4" bold="true" box="[199,322,649,675]" pageId="11" pageNumber="162">Diagnosis.</emphasis>
|
||
Low trochiform, conical, comparably large
|
||
<taxonomicName id="4C164D36C84AFFBCFC9FFD54FC13A7D3" box="[821,950,650,676]" class="Kinetofragminophora" family="Corynophryidae" genus="Andrusovia" kingdom="Protozoa" order="Suctorida" pageId="11" pageNumber="162" phylum="Ciliophora" rank="genus">
|
||
<emphasis id="B962EAA7C84AFFBCFC9FFD54FC13A7D3" box="[821,950,650,676]" italics="true" pageId="11" pageNumber="162">Andrusovia</emphasis>
|
||
</taxonomicName>
|
||
with up to four well rounded whorls, wide umbilicus, broadly ovoid aperture, weakly sinuate and weakly thickened peristome.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF69663DC84AFFBCFF3DF852FEC4A29B" ID-DOI="http://doi.org/10.5281/zenodo.4550158" ID-Zenodo-Dep="4550158" httpUri="https://zenodo.org/record/4550158/files/figure.png" pageId="11" pageNumber="162" startId="11.[151,250,1932,1956]" targetBox="[151,1436,736,1909]" targetPageId="11">
|
||
<paragraph id="8BA936B5C84AFFBCFF3DF852FEC4A29B" blockId="11.[151,1437,1932,2029]" pageId="11" pageNumber="162">
|
||
<emphasis id="B962EAA7C84AFFBCFF3DF852FEB6A2D3" bold="true" box="[151,275,1932,1956]" pageId="11" pageNumber="162">FIGURE 4.</emphasis>
|
||
<taxonomicName id="4C164D36C84AFFBCFEB3F852FE5AA2D3" authority="Anistratenko & Neubauer & Anistratenko & Kijashko & Wesselingh, 2021" authorityName="Anistratenko & Neubauer & Anistratenko & Kijashko & Wesselingh" authorityYear="2021" box="[281,511,1932,1956]" class="Kinetofragminophora" family="Corynophryidae" genus="Andrusovia" kingdom="Protozoa" order="Suctorida" pageId="11" pageNumber="162" phylum="Ciliophora" rank="species" species="antecessor" status="sp. nov.">
|
||
<emphasis id="B962EAA7C84AFFBCFEB3F852FE5AA2D3" box="[281,511,1932,1956]" italics="true" pageId="11" pageNumber="162">Andrusovia antecessor</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B962EAA7C84AFFBCFDACF852FDF3A2D3" bold="true" box="[518,598,1932,1956]" pageId="11" pageNumber="162">
|
||
<taxonomicNameLabel id="A25157DCC84AFFBCFDACF852FDF3A2D3" box="[518,598,1932,1956]" pageId="11" pageNumber="162" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
from the early Late Pleistocene of Selitrennoye, Astrakhan region, Russia (locality 1). (a, d, g) Paratype (RGM 1309840). (b, c, h) Holotype (RGM 1309839). (e, f, i) Paratype (LV 201509). Scale bars = 500 μm (a–f), 100 μm (g–i).
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C30C653EC84DFFBBFF6DFF48FF75A483" pageId="12" pageNumber="163" type="description">
|
||
<paragraph id="8BA936B5C84DFFBBFF6DFF48FF75A483" blockId="12.[151,1437,150,1041]" pageId="12" pageNumber="163">
|
||
<emphasis id="B962EAA7C84DFFBBFF6DFF48FC81A5C7" bold="true" box="[199,804,150,177]" pageId="12" pageNumber="163">
|
||
Description (emended from
|
||
<bibRefCitation id="EF874B44C84DFFBBFDB1FF48FCB3A5C7" author="Neubauer, T. A. & van de Velde, S. & Yanina, T. A. & Wesselingh, F. P." box="[539,790,150,177]" pageId="12" pageNumber="163" pagination="43 - 103" refId="ref32549" refString="Neubauer, T. A., van de Velde, S., Yanina, T. A. & Wesselingh, F. P. (2018) A late Pleistocene gastropod fauna from the northern Caspian Sea with implications for Pontocaspian gastropod taxonomy. ZooKeys, 770, 43 - 103. https: // doi. org / 10.3897 / zookeys. 770.25365" type="journal article" year="2018">
|
||
Neubauer
|
||
<emphasis id="B962EAA7C84DFFBBFD33FF49FD69A5C6" bold="true" box="[665,716,150,177]" italics="true" pageId="12" pageNumber="163">et al</emphasis>
|
||
. 2018
|
||
</bibRefCitation>
|
||
).
|
||
</emphasis>
|
||
Shell broad trochiform, about as high as wide, with up to 4 whorls. Rarely, specimens with slightly elevated spire occur. Protoconch high domical, about semi-circular in profile; initial part immersed; consists of 1.1 whorls, measuring 300 μm in diameter; nucleus about 90 μm wide; protoconch surface finely but strongly malleate near lower suture, rest appears eroded; P/T boundary sharp, marked by massive growth constrictions near lower suture. Teleoconch whorls highly convex, with maximum convexity in adapical half, producing slightly stepped spire. Last whorl attains 74–81% of shell height. Aperture broadly ovoid, slightly inclined, with faint concavity at contact to penultimate whorl. Peristome slightly thickened and expanded at columella and base; sinuate in lateral view, with weakly protruding central part and weak adapical indentation. Umbilicus wide, deep. Fine prosocyrt growth lines cover shell. On one specimen, traces of spiral threads occur on base.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30C653EC84DFFBBFF6DFE20FAA6A6D3" pageId="12" pageNumber="163" type="discussion">
|
||
<paragraph id="8BA936B5C84DFFBBFF6DFE20FB4DA7BA" blockId="12.[151,1437,150,1041]" pageId="12" pageNumber="163">
|
||
<emphasis id="B962EAA7C84DFFBBFF6DFE20FE9EA76F" bold="true" box="[199,315,510,536]" pageId="12" pageNumber="163">Remarks.</emphasis>
|
||
<bibRefCitation id="EF874B44C84DFFBBFEE9FE20FDE1A76F" author="Neubauer, T. A. & van de Velde, S. & Yanina, T. A. & Wesselingh, F. P." box="[323,580,510,537]" pageId="12" pageNumber="163" pagination="43 - 103" refId="ref32549" refString="Neubauer, T. A., van de Velde, S., Yanina, T. A. & Wesselingh, F. P. (2018) A late Pleistocene gastropod fauna from the northern Caspian Sea with implications for Pontocaspian gastropod taxonomy. ZooKeys, 770, 43 - 103. https: // doi. org / 10.3897 / zookeys. 770.25365" type="journal article" year="2018">
|
||
Neubauer
|
||
<emphasis id="B962EAA7C84DFFBBFE12FE21FE4FA76F" box="[440,490,510,536]" italics="true" pageId="12" pageNumber="163">et al</emphasis>
|
||
. (2018)
|
||
</bibRefCitation>
|
||
wrongfully attributed this species to the extant
|
||
<taxonomicName id="4C164D36C84DFFBBFBC8FE20FAEDA76F" authorityName="Starobogatov" authorityYear="2000" box="[1122,1352,510,536]" class="Kinetofragminophora" family="Corynophryidae" genus="Andrusovia" kingdom="Protozoa" order="Suctorida" pageId="12" pageNumber="163" phylum="Ciliophora" rank="species" species="brusinai">
|
||
<emphasis id="B962EAA7C84DFFBBFBC8FE20FAEDA76F" box="[1122,1352,510,536]" italics="true" pageId="12" pageNumber="163">Andrusovia brusinai</emphasis>
|
||
</taxonomicName>
|
||
, which is here considered a junior synonym of
|
||
<taxonomicName id="4C164D36C84DFFBBFDF2FDFCFCF4A74B" authorityName="Brusina" authorityYear="1902" box="[600,849,546,572]" class="Kinetofragminophora" family="Corynophryidae" genus="Andrusovia" kingdom="Protozoa" order="Suctorida" pageId="12" pageNumber="163" phylum="Ciliophora" rank="species" species="dybowskii">
|
||
<emphasis id="B962EAA7C84DFFBBFDF2FDFCFCF4A74B" box="[600,849,546,572]" italics="true" pageId="12" pageNumber="163">Andrusovia dybowskii</emphasis>
|
||
</taxonomicName>
|
||
(see above).
|
||
<taxonomicName id="4C164D36C84DFFBBFC43FDFCFB4FA74B" authority="Anistratenko & Neubauer & Anistratenko & Kijashko & Wesselingh, 2021" authorityName="Anistratenko & Neubauer & Anistratenko & Kijashko & Wesselingh" authorityYear="2021" box="[1001,1258,546,572]" class="Kinetofragminophora" family="Corynophryidae" genus="Andrusovia" kingdom="Protozoa" order="Suctorida" pageId="12" pageNumber="163" phylum="Ciliophora" rank="species" species="antecessor" status="sp. nov.">
|
||
<emphasis id="B962EAA7C84DFFBBFC43FDFCFB4FA74B" box="[1001,1258,546,572]" italics="true" pageId="12" pageNumber="163">Andrusovia antecessor</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B962EAA7C84DFFBBFB5BFDFCFAE9A74B" bold="true" box="[1265,1356,546,572]" pageId="12" pageNumber="163">
|
||
<taxonomicNameLabel id="A25157DCC84DFFBBFB5BFDFCFAE9A74B" box="[1265,1356,546,572]" pageId="12" pageNumber="163" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
differs from that species in the larger size, the more whorls while having an equally small (compared to subfossil shells) or smaller (relative to recent shells) protoconch, the more regular coiling resulting in a perfectly conical shape, and the rounded whorls. The overall similarity (and the lack of other known species) suggests that both species are closely related; given the age,
|
||
<taxonomicName id="4C164D36C84DFFBBFE3FFD6DFD88A7BB" authority="Anistratenko & Neubauer & Anistratenko & Kijashko & Wesselingh, 2021" authorityName="Anistratenko & Neubauer & Anistratenko & Kijashko & Wesselingh" authorityYear="2021" box="[405,557,691,717]" class="Kinetofragminophora" family="Corynophryidae" genus="Andrusovia" kingdom="Protozoa" order="Suctorida" pageId="12" pageNumber="163" phylum="Ciliophora" rank="species" species="antecessor" status="sp. nov.">
|
||
<emphasis id="B962EAA7C84DFFBBFE3FFD6DFD88A7BB" box="[405,557,691,717]" italics="true" pageId="12" pageNumber="163">A. antecessor</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B962EAA7C84DFFBBFD99FD6CFD28A7BB" bold="true" box="[563,653,690,716]" pageId="12" pageNumber="163">
|
||
<taxonomicNameLabel id="A25157DCC84DFFBBFD99FD6CFD28A7BB" box="[563,653,690,716]" pageId="12" pageNumber="163" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
might be a phylogenetic predecessor of
|
||
<taxonomicName id="4C164D36C84DFFBBFBF8FD6DFB44A7BB" authorityName="Brusina" authorityYear="1902" box="[1106,1249,690,717]" class="Kinetofragminophora" family="Corynophryidae" genus="Andrusovia" kingdom="Protozoa" order="Suctorida" pageId="12" pageNumber="163" phylum="Ciliophora" rank="species" species="dybowskii">
|
||
<emphasis id="B962EAA7C84DFFBBFBF8FD6DFB44A7BB" box="[1106,1249,690,717]" italics="true" pageId="12" pageNumber="163">A. dybowskii</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BA936B5C84DFFBBFF6DFD08FAA6A6D3" blockId="12.[151,1437,150,1041]" pageId="12" pageNumber="163">
|
||
The smaller protoconch (and thus smaller hatching size) combined with a larger adult size of
|
||
<taxonomicName id="4C164D36C84DFFBBFB71FD08FF7AA663" authority="Anistratenko & Neubauer & Anistratenko & Kijashko & Wesselingh, 2021" authorityName="Anistratenko & Neubauer & Anistratenko & Kijashko & Wesselingh" authorityYear="2021" class="Kinetofragminophora" family="Corynophryidae" genus="Andrusovia" kingdom="Protozoa" order="Suctorida" pageId="12" pageNumber="163" phylum="Ciliophora" rank="species" species="antecessor" status="sp. nov.">
|
||
<emphasis id="B962EAA7C84DFFBBFB71FD08FF7AA663" italics="true" pageId="12" pageNumber="163">Andrusovia antecessor</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B962EAA7C84DFFBBFF4DFD24FEE4A663" bold="true" box="[231,321,762,788]" pageId="12" pageNumber="163">
|
||
<taxonomicNameLabel id="A25157DCC84DFFBBFF4DFD24FEE4A663" box="[231,321,762,788]" pageId="12" pageNumber="163" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
compared to modern
|
||
<taxonomicName id="4C164D36C84DFFBBFD90FD25FD6FA663" authorityName="Brusina" authorityYear="1902" box="[570,714,762,789]" class="Kinetofragminophora" family="Corynophryidae" genus="Andrusovia" kingdom="Protozoa" order="Suctorida" pageId="12" pageNumber="163" phylum="Ciliophora" rank="species" species="dybowskii">
|
||
<emphasis id="B962EAA7C84DFFBBFD90FD25FD6FA663" box="[570,714,762,789]" italics="true" pageId="12" pageNumber="163">A. dybowskii</emphasis>
|
||
</taxonomicName>
|
||
suggests a faster growth rate or, alternatively, a longer lifetime. Growth rates in gastropods typically correlate positively with temperature (e.g.
|
||
<bibRefCitation id="EF874B44C84DFFBBFBB6FCC0FAE7A64E" author="Kevrekidis, T. & Wilke, T." box="[1052,1346,798,825]" pageId="12" pageNumber="163" pagination="375 - 382" refId="ref31352" refString="Kevrekidis, T. & Wilke, T. (2005) Life cycle, population dynamics and productivity of Ventrosia maritima in the Evros Delta (northern Aegean Sea). Journal of the Marine Biological Association of the United Kingdom, 85, 375 - 382. https: // doi. org / 10.1017 / S 0025315405011306 h" type="journal article" year="2005">Kevrekidis & Wilke 2005</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EF874B44C84DFFBBFAE4FCC0FEE1A62A" author="Shanahan, T. M. & Pigati, J. S. & Dettman, D. L. & Quade, J." pageId="12" pageNumber="163" pagination="3949 - 3966" refId="ref33310" refString="Shanahan, T. M., Pigati, J. S., Dettman, D. L. & Quade, J. (2005) Isotopic variability in the aragonite shells of freshwater gastropods living in springs with nearly constant temperature and isotopic composition. Geochimica et Cosmochimica Acta, 69 (16), 3949 - 3966. https: // doi. org / 10.1016 / j. gca. 2005.03.049" type="journal article" year="2005">
|
||
Shanahan
|
||
<emphasis id="B962EAA7C84DFFBBFF62FC9DFF5FA62B" box="[200,250,834,860]" italics="true" pageId="12" pageNumber="163">et al</emphasis>
|
||
. 2005
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EF874B44C84DFFBBFEFAFC9CFDECA62B" author="McCreesh, N. & Arinaitwe, M. & Arineitwe, W. & Tukahebwa, E. M. & Booth, M." box="[336,585,834,861]" pageId="12" pageNumber="163" pagination="503" refId="ref32321" refString="McCreesh, N., Arinaitwe, M., Arineitwe, W., Tukahebwa, E. M. & Booth, M. (2014) Effect of water temperature and population density on the population dynamics of Schistosoma mansoni intermediate host snails. Parasites & Vectors, 7, 503. https: // doi. org / 10.1186 / s 13071 - 014 - 0503 - 9" type="journal article" year="2014">
|
||
McCreesh
|
||
<emphasis id="B962EAA7C84DFFBBFE61FC9DFE5BA62B" box="[459,510,834,860]" italics="true" pageId="12" pageNumber="163">et al</emphasis>
|
||
. 2014
|
||
</bibRefCitation>
|
||
). Similarly, the time of hatching can occur earlier under warmer conditions (
|
||
<bibRefCitation id="EF874B44C84DFFBBFF35FCB8FED8A6F7" author="Cancino, J. M. & Gallardo, J. A. & Torres, F. A." box="[159,381,870,897]" pageId="12" pageNumber="163" pagination="133 - 139" refId="ref29851" refString="Cancino, J. M., Gallardo, J. A. & Torres, F. A. (2003) Combined effects of dissolved oxygen concentration and water temperature on embryonic development and larval shell secretion in the marine snail Chorus giganteus (Gastropoda: Muricidae). Marine Biology, 142, 133 - 139. https: // doi. org / 10.1007 / s 00227 - 002 - 0925 - 3" type="journal article" year="2003">
|
||
Cancino
|
||
<emphasis id="B962EAA7C84DFFBBFEAEFCB9FE96A6F7" box="[260,307,870,896]" italics="true" pageId="12" pageNumber="163">et al</emphasis>
|
||
. 2003
|
||
</bibRefCitation>
|
||
). The scenario of an increased growth rate in
|
||
<taxonomicName id="4C164D36C84DFFBBFCC4FCB9FBA1A6F7" authority="Anistratenko & Neubauer & Anistratenko & Kijashko & Wesselingh, 2021" authorityName="Anistratenko & Neubauer & Anistratenko & Kijashko & Wesselingh" authorityYear="2021" box="[878,1028,871,897]" class="Kinetofragminophora" family="Corynophryidae" genus="Andrusovia" kingdom="Protozoa" order="Suctorida" pageId="12" pageNumber="163" phylum="Ciliophora" rank="species" species="antecessor" status="sp. nov.">
|
||
<emphasis id="B962EAA7C84DFFBBFCC4FCB9FBA1A6F7" box="[878,1028,871,897]" italics="true" pageId="12" pageNumber="163">A. antecessor</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B962EAA7C84DFFBBFBA3FCB8FBC7A6F7" bold="true" box="[1033,1122,870,896]" pageId="12" pageNumber="163">
|
||
<taxonomicNameLabel id="A25157DCC84DFFBBFBA3FCB8FBC7A6F7" box="[1033,1122,870,896]" pageId="12" pageNumber="163" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
as a function of temperature fits well to the supposed warmer climate during the Hyrcanian (
|
||
<bibRefCitation id="EF874B44C84DFFBBFCCBFC54FC51A6D3" author="Yanina, T. A." box="[865,1012,906,932]" pageId="12" pageNumber="163" pagination="88 - 99" refId="ref34506" refString="Yanina, T. A. (2014) The Ponto-Caspian region: Environmental consequences of climate change during the Late Pleistocene. Quaternary International, 345, 88 - 99. https: // doi. org / 10.1016 / j. quaint. 2014.01.045" type="journal article" year="2014">Yanina 2014</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EF874B44C84DFFBBFC54FC54FB5DA6D3" author="Krijgsman, W. & Tesakov, A. & Yanina, T. & Lazarev, S. & Danukalova, G. & Van Baak, C. G. C. & Agusti, J. & Alcicek, M. C. & Aliyeva, E. & Bista, D. & Bruch, A. & Bukhsianidze, M. & Flecker, R. & Frolov, P. & Hoyle, T. M. & Jorissen, E. L. & Kirscher, U. & Koriche, S. A. & Kroonenberg, S. B. & Lordkipanidze, D. & Oms, O. & Rausch, L. & Singarayer, J. & Stoica, M. & van de Velde, S. & Titov, V. V. & Wesselingh, F. P." box="[1022,1272,906,933]" pageId="12" pageNumber="163" pagination="1 - 40" refId="ref31554" refString="Krijgsman, W., Tesakov, A., Yanina, T., Lazarev, S., Danukalova, G., Van Baak, C. G. C., Agusti, J., Alcicek, M. C., Aliyeva, E., Bista, D., Bruch, A., B ʾ y ʾ kmeric, Y., Bukhsianidze, M., Flecker, R., Frolov, P., Hoyle, T. M., Jorissen, E. L., Kirscher, U., Koriche, S. A., Kroonenberg, S. B., Lordkipanidze, D., Oms, O., Rausch, L., Singarayer, J., Stoica, M., van de Velde, S., Titov, V. V. & Wesselingh, F. P. (2019) Rise and fall of the Paratethys Sea during the Messinian Salinity Crisis. Earth-Science Reviews, 188, 1 - 40. https: // doi. org / 10.1016 / j. earscirev. 2018.10.013" type="journal article" year="2019">
|
||
Krijgsman
|
||
<emphasis id="B962EAA7C84DFFBBFBD6FC55FB08A6D3" box="[1148,1197,906,932]" italics="true" pageId="12" pageNumber="163">et al</emphasis>
|
||
. 2019
|
||
</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30C653EC84DFFBBFF6DFC70FE5BA167" pageId="12" pageNumber="163" type="distribution">
|
||
<paragraph id="8BA936B5C84DFFBBFF6DFC70FE5BA167" blockId="12.[151,1437,150,1041]" pageId="12" pageNumber="163">
|
||
<emphasis id="B962EAA7C84DFFBBFF6DFC70FEC4A6BF" bold="true" box="[199,353,942,968]" pageId="12" pageNumber="163">Distribution.</emphasis>
|
||
Endemic to the Caspian Sea, known only from the Late Pleistocene strata of the
|
||
<typeStatus id="54AD8817C84DFFBBFAA0FC71FA9FA6BE" box="[1290,1338,943,969]" pageId="12" pageNumber="163">type</typeStatus>
|
||
locality. Reconstructions of the paleoecology suggest a shallow (
|
||
<quantity id="4CEE9B50C84DFFBBFCA7FC0CFCD0A69A" box="[781,885,978,1005]" metricMagnitude="1" metricUnit="m" metricValue="2.0" metricValueMax="2.5" metricValueMin="1.5" pageId="12" pageNumber="163" unit="m" value="20.0" valueMax="25.0" valueMin="15.0">15–25 m</quantity>
|
||
), lower mesohaline (5–8 psu) open lake environment (
|
||
<bibRefCitation id="EF874B44C84DFFBBFF74FC28FE56A167" author="van de Velde, S. & Yanina, T. A. & Neubauer, T. A. & Wesselingh, F. P." box="[222,499,1014,1041]" pageId="12" pageNumber="163" pagination="1227 - 1239" refId="ref34097" refString="van de Velde, S., Yanina, T. A., Neubauer, T. A. & Wesselingh, F. P. (2020) The Late Pleistocene mollusk fauna of Selitrennoye (Astrakhan province, Russia): A natural baseline for endemic Caspian Sea faunas. Journal of Great Lakes Research, 46 (5), 1227 - 1239. https: // doi. org / 10.1016 / j. jglr. 2019.04.001" type="journal article" year="2020">
|
||
van de Velde
|
||
<emphasis id="B962EAA7C84DFFBBFEDDFC29FE0DA167" box="[375,424,1014,1040]" italics="true" pageId="12" pageNumber="163">et al</emphasis>
|
||
. 2020
|
||
</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |