283 lines
38 KiB
XML
283 lines
38 KiB
XML
<document id="C7CD0BA2A6709F38F716DD58F61AA6D6" ID-CLB-Dataset="31928" ID-DOI="10.11646/zootaxa.4299.4.8" ID-GBIF-Dataset="8eb6e6da-bfb9-40a0-b698-cbb1ef63d66e" ID-ISSN="1175-5326" ID-Zenodo-Dep="998520" ID-ZooBank="28Cb7701-A5E7-4912-Ae30-16Ce34132792" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1506584369341" checkinUser="plazi" docAuthor="Duarte, Marcelo" docDate="2017" docId="038F2526FFEA7D22899FFA08072DE0A4" docLanguage="en" docName="zootaxa.4299.4.8.pdf" docOrigin="Zootaxa 4299 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Limyra Duarte 2017, gen. nov." docType="treatment" docVersion="7" lastPageNumber="588" masterDocId="FFB65D5EFFEE7D258908FFF60501E67A" masterDocTitle="Description of Limyra, new genus with a new species and the redescription of Cibyra pluriargenteus (Viette), from southeastern Brazil (Lepidoptera: Hepialidae)" masterLastPageNumber="591" masterPageNumber="581" pageNumber="585" updateTime="1698469173363" updateUser="ExternalLinkService">
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<mods:title id="A712071B434C23A78FA82978893B4E6C">Description of Limyra, new genus with a new species and the redescription of Cibyra pluriargenteus (Viette), from southeastern Brazil (Lepidoptera: Hepialidae)</mods:title>
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<mods:namePart id="47433E5D670744BCEDEBCD753D24084F">Duarte, Marcelo</mods:namePart>
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<mods:title id="9CD1723C3FA01B9F0FD51701B270A885">Zootaxa</mods:title>
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<mods:part id="D948C9DA00E1BF7491088448E5A754F5">
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<mods:date id="F9F6FCD141DAF963488044B7D75E61B8">2017</mods:date>
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<mods:number id="A00AE7677DA8466EC97C4B542729714D">2017-07-31</mods:number>
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<mods:number id="63768FDA2D4988DCAAAD7853549E01D2">4299</mods:number>
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<mods:identifier id="6245A5F7A41EAEAB10D549A7FA937548" type="DOI">10.11646/zootaxa.4299.4.8</mods:identifier>
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<treatment id="038F2526FFEA7D22899FFA08072DE0A4" ID-DOI="http://doi.org/10.5281/zenodo.6024622" ID-GBIF-Taxon="134860846" ID-Zenodo-Dep="6024622" LSID="urn:lsid:plazi:treatment:038F2526FFEA7D22899FFA08072DE0A4" httpUri="http://treatment.plazi.org/id/038F2526FFEA7D22899FFA08072DE0A4" lastPageId="7" lastPageNumber="588" pageId="4" pageNumber="585">
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<subSubSection id="C33CC7BBFFEA7D21899FFA0804A5E041" pageId="4" pageNumber="585" type="nomenclature">
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<paragraph id="8B999430FFEA7D21899FFA08045AE06D" blockId="4.[151,420,1534,1595]" box="[151,347,1534,1559]" pageId="4" pageNumber="585">
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<heading id="D0D1235CFFEA7D21899FFA08045AE06D" bold="true" box="[151,347,1534,1559]" fontSize="10" level="2" pageId="4" pageNumber="585" reason="2">
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<emphasis id="B9524822FFEA7D21899FFA08045AE06D" bold="true" box="[151,347,1534,1559]" pageId="4" pageNumber="585">
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<taxonomicName id="4C26EFB3FFEA7D21899FFA0805EAE06C" authority="Duarte, 2017" authorityName="Duarte" authorityYear="2017" box="[151,235,1534,1558]" class="Insecta" family="Hepialidae" genus="Limyra" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="585" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="B9524822FFEA7D21899FFA0805EAE06C" bold="true" box="[151,235,1534,1558]" italics="true" pageId="4" pageNumber="585">Limyra</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A261F559FFEA7D2189FAFA08045AE06D" box="[242,347,1534,1559]" pageId="4" pageNumber="585" rank="genus">gen. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8B999430FFEA7D21899FF9D404A5E041" blockId="4.[151,420,1534,1595]" box="[151,420,1570,1595]" pageId="4" pageNumber="585">
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(
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<figureCitation id="131D88B5FFEA7D218997F9D4041EE041" box="[159,287,1570,1595]" captionStart="FIGURES 11 − 15" captionStartId="4.[151,264,1375,1398]" captionTargetBox="[151,1436,628,1354]" captionTargetId="figure@4.[151,1436,628,1354]" captionTargetPageId="4" captionText="FIGURES 11 − 15. Limyra silvai sp. nov. 11, male holotype, dorsal view; 12, male holotype, ventral view; 13, male paratype, dorsal view (MZSP 14.403); 14, female paratype, dorsal view (MZSP 14.404); 15, female paratype, ventral view (MZSP 14.404). Scale bar = 1 cm." httpUri="https://zenodo.org/record/998526/files/figure.png" pageId="4" pageNumber="585">Figs 11–15</figureCitation>
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,
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<figureCitation id="131D88B5FFEA7D218822F9D50472E041" box="[298,371,1571,1595]" captionStart-0="FIGURES 22 − 29" captionStart-1="FIGURE 30" captionStart-2="FIGURE 31" captionStartId-0="6.[151,264,1386,1409]" captionStartId-1="6.[151,250,2004,2026]" captionStartId-2="7.[151,250,664,686]" captionTargetBox-0="[152,1430,207,1365]" captionTargetId-0="figure@6.[151,1436,193,1365]" captionTargetPageId-0="6" captionText-0="FIGURES 22 − 29. Limyra silvai sp. nov. 22. Male abdomen (holotype), upper dorsal, lower ventral view. 23 − 25. Male genitalia (holotype): 23, ventral view; 24, lateral view; 25, anterior view (internal), fultura superior (arrow). 26 − 27. Female abdomen (MZSP 14.404). 26, upper dorsal, lower ventral view; 27, tergum and sternum VII and VIII (tergum VIII membranous), sternum VIII (arrow). 28 − 29. Female genitalia (MZSP 14.404). 28, lateral view; 29, posterior view. Scale bar = 1 mm, except Figs 22 and 26 = 1 cm." captionText-1="FIGURE 30. Limyra silvai sp. nov. Male genitalia, holotype." captionText-2="FIGURE 31. Limyra silvai sp. nov. Male (left) and female (right) venation." httpUri-0="https://zenodo.org/record/998530/files/figure.png" pageId="4" pageNumber="585">22–31</figureCitation>
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,
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<figureCitation id="131D88B5FFEA7D218876F9D5049AE041" box="[382,411,1571,1595]" captionStart="FIGURE 32" captionStartId="9.[151,250,730,752]" captionTargetBox="[340,1247,195,709]" captionTargetId="figure@9.[340,1247,195,709]" captionTargetPageId="9" captionText="FIGURE 32. Geographical distribution of Cibyra pluriargenteus (red circle) and Limyra silvai sp. nov. (blue star) in southeastern Brazil." httpUri="https://zenodo.org/record/998532/files/figure.png" pageId="4" pageNumber="585">32</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C33CC7BBFFEA7D21899FF99D0703E005" box="[151,514,1642,1664]" pageId="4" pageNumber="585" type="materials_examined">
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<paragraph id="8B999430FFEA7D21899FF99D0703E005" blockId="4.[151,514,1642,1664]" box="[151,514,1642,1664]" pageId="4" pageNumber="585">
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Type species:
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<taxonomicName id="4C26EFB3FFEA7D21882EF99D04ADE0FA" authority="Duarte, 2017" authorityName="Duarte" authorityYear="2017" box="[294,428,1642,1664]" class="Insecta" family="Hepialidae" genus="Limyra" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="585" phylum="Arthropoda" rank="species" species="silvai" status="sp. nov.">
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<emphasis id="B9524822FFEA7D21882EF99D04ADE0FA" box="[294,428,1642,1664]" italics="true" pageId="4" pageNumber="585">Limyra silvai</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A261F559FFEA7D2188BAF99C0703E005" box="[434,514,1642,1663]" pageId="4" pageNumber="585" rank="species">
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<emphasis id="B9524822FFEA7D2188BAF99C0703E005" bold="true" box="[434,514,1642,1663]" pageId="4" pageNumber="585">sp. nov.</emphasis>
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</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection id="C33CC7BBFFEA7D20899FF95A0730E13D" lastPageId="5" lastPageNumber="586" pageId="4" pageNumber="585" type="diagnosis">
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<paragraph id="8B999430FFEA7D21899FF95A0655E100" blockId="4.[151,1436,1708,1914]" pageId="4" pageNumber="585">
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<emphasis id="B9524822FFEA7D21899FF95A040BE0BF" bold="true" box="[151,266,1708,1733]" pageId="4" pageNumber="585">Diagnosis</emphasis>
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. Easily recognized by the distinctive male and female genitalia. In the male genitalia, saccus clearly in two fused parts, a dorsal and a ventral ones. Fultura superior present. In the female genitalia, the lamella antevaginalis is compound mainly by two tapered processes latero-anterior that extends and fuses posteriorly to a mesal process, forming a depression between both structures. Such structures are unique among all Neotropical genera. In addition, the new genus is diagnosed by the combination of the following characters in head micromorphology, wing venation, wing pattern, and abdomen:
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</paragraph>
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<paragraph id="8B999430FFEA7D21899FF85C06BAE1B8" blockId="4.[151,1038,1961,2022]" box="[151,955,1961,1986]" pageId="4" pageNumber="585">• male labial palpus with first and second palpomeres partially fused,</paragraph>
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<paragraph id="8B999430FFEA7D20899FF838077CE682" blockId="4.[151,1038,1961,2022]" lastBlockId="5.[151,1001,151,248]" lastPageId="5" lastPageNumber="586" pageId="4" pageNumber="585">
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• in the female the second palpomere twice the length of the first palpomere, • 'hepialine' venation (Rs2+Rs3 not stalked with Rs4) (
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<bibRefCitation id="EFB7E9C1FFEB7D208A12FF6106DCE6CA" author="Dumbleton, L. J." box="[794,989,151,176]" journalOrPublisher="New Zealand Journal of Science, 9" pageId="5" pageNumber="586" pagination="920 - 981" part="4" refString="Dumbleton, L. J. (1966) Genitalia, classification and zoogeography of the New Zealand Hepialidae (Lepidoptera). New Zealand Journal of Science, 9 (4), 920 - 981." title="Genitalia, classification and zoogeography of the New Zealand Hepialidae (Lepidoptera)" type="journal article" year="1966">Dumbleton 1966</bibRefCitation>
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), • absence of distinct banding or patches on the forewing, • sternum VIII of the male membranous.
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</paragraph>
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<caption id="DF59C4B8FFEB7D20899FFA4A05D8E055" httpUri="https://zenodo.org/record/998528/files/figure.png" pageId="5" pageNumber="586" targetBox="[160,1415,307,1447]" targetPageId="5">
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<paragraph id="8B999430FFEB7D20899FFA4A05D8E055" blockId="5.[151,1436,1468,1583]" pageId="5" pageNumber="586">
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<emphasis id="B9524822FFEB7D20899FFA4A0451E3A8" bold="true" box="[151,336,1468,1491]" pageId="5" pageNumber="586">FIGURES 16−21</emphasis>
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.
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<taxonomicName id="4C26EFB3FFEB7D208857FA4B0743E3A9" baseAuthorityName="Viette" baseAuthorityYear="1956" box="[351,578,1469,1491]" class="Insecta" family="Hepialidae" genus="Cibyra" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="586" phylum="Arthropoda" rank="species" species="pluriargenteus">
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<emphasis id="B9524822FFEB7D208857FA4B0743E3A9" box="[351,578,1469,1491]" italics="true" pageId="5" pageNumber="586">Cibyra pluriargenteus</emphasis>
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</taxonomicName>
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. 16−17. Male abdomen (MZSP 30.629): 16, upper dorsal, lower ventral view; 17, tergum and sternum VII and VIII; 18, Male genitalia, ventral view (MZSP 30.629); 19, Female tergum and sternum VII and VIII (MZSP 30.626); 20−21, Female genitalia: 20, lateral view (MZSP 30.625); 21, posterior view (MZSP 30.626). Scale bar = 1 mm.
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</paragraph>
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</caption>
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<paragraph id="8B999430FFEB7D2089CFF9A00730E13D" blockId="5.[151,1436,1622,2007]" pageId="5" pageNumber="586">
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The arched shape of the male genitalia resembles
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<taxonomicName id="4C26EFB3FFEB7D208BF6F9A006F2E015" authority="Viette, 1961" authorityName="Viette" authorityYear="1961" box="[766,1011,1622,1647]" class="Insecta" family="Hepialidae" genus="Phialuse" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="586" phylum="Arthropoda" rank="genus">
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<emphasis id="B9524822FFEB7D208BF6F9A00661E015" box="[766,864,1622,1647]" italics="true" pageId="5" pageNumber="586">Phialuse</emphasis>
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<bibRefCitation id="EFB7E9C1FFEB7D208A61F9A006F2E015" author="Viette, P." box="[873,1011,1622,1647]" journalOrPublisher="Opuscula Zoologica" pageId="5" pageNumber="586" pagination="1 - 7" part="55" refString="Viette, P. (1961) Beitrage zur Kenntnis der Insektenfauna Boliviens XV. Lepidoptera II. Contribution a l'etude des Hepialidae. 34 eme note. Especes sud-americaines nouvelles ou peu connues du Museum de Munich. Opuscula Zoologica, 55, 1 - 7." title="Beitrage zur Kenntnis der Insektenfauna Boliviens XV. Lepidoptera II. Contribution a l'etude des Hepialidae. 34 eme note. Especes sud-americaines nouvelles ou peu connues du Museum de Munich" type="journal article" year="1961">Viette, 1961</bibRefCitation>
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</taxonomicName>
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with respect to an extended structure attached to the ventral surface of the saccus, although no particular relationship can be inferred to present time, since other characters such as antennae and wing characteristics are very different. The membranous tergum VIII of the female suggests a close relation with
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<taxonomicName id="4C26EFB3FFEB7D208B84F93407DBE0A1" authorityName="Walker" authorityYear="1856" box="[652,730,1730,1755]" class="Insecta" family="Hepialidae" genus="Cibyra" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="586" phylum="Arthropoda" rank="genus">
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<emphasis id="B9524822FFEB7D208B84F93407DBE0A1" box="[652,730,1730,1755]" italics="true" pageId="5" pageNumber="586">Cibyra</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="4C26EFB3FFEB7D208A1BF9350132E0A1" authority="Herrich-Schaffer" authorityName="Herrich-Schaffer" box="[787,1075,1730,1755]" class="Insecta" family="Hepialidae" genus="Aepytus" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="586" phylum="Arthropoda" rank="genus">
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<emphasis id="B9524822FFEB7D208A1BF935066DE0A0" box="[787,876,1731,1754]" italics="true" pageId="5" pageNumber="586">Aepytus</emphasis>
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Herrich-Schäffer
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</taxonomicName>
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, [1856], but the presence of the fultura superior (trulleum) in the male genitalia contradicts its phylogenetic position from these genera. The shape of sternum II and its anterior lateral bar are also similar to other cibyrine genera, to
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<taxonomicName id="4C26EFB3FFEB7D208D4BF8FC006DE159" authority="Viette, 1951" authorityName="Viette" authorityYear="1951" box="[1091,1388,1802,1827]" class="Insecta" family="Hepialidae" genus="Hepialyxodes" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="586" phylum="Arthropoda" rank="genus">
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<emphasis id="B9524822FFEB7D208D4BF8FC01DDE159" box="[1091,1244,1802,1827]" italics="true" pageId="5" pageNumber="586">Hepialyxodes</emphasis>
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<bibRefCitation id="EFB7E9C1FFEB7D208DEBF8FC006DE159" author="Viette, P." box="[1251,1388,1802,1827]" journalOrPublisher="Bulletin mensuel de la Societe Linneenne de Lyon" pageId="5" pageNumber="586" pagination="95 - 96" part="20" publicationUrl="https://doi.org/10.3406/linly.1951.7396" refString="Viette, P. (1951) Contribution a l'etude des Hepialides (Lepid.) (24 note). Bulletin mensuel de la Societe Linneenne de Lyon, 20 (4), 95 - 96. https: // doi. org / 10.3406 / linly. 1951.7396" title="Contribution a l'etude des Hepialides (Lepid.) (24 note)" type="journal article" year="1951">Viette, 1951</bibRefCitation>
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</taxonomicName>
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and
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<taxonomicName id="4C26EFB3FFEB7D20899FF8D905F1E13C" box="[151,240,1839,1862]" class="Insecta" family="Hepialidae" genus="Aepytus" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="586" phylum="Arthropoda" rank="genus">
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<emphasis id="B9524822FFEB7D20899FF8D905F1E13C" box="[151,240,1839,1862]" italics="true" pageId="5" pageNumber="586">Aepytus</emphasis>
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</taxonomicName>
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in particular (
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<bibRefCitation id="EFB7E9C1FFEB7D208885F8D80724E13D" author="Grehan, J. R" box="[397,549,1838,1863]" journalOrPublisher="Bulletin of the Buffalo Society of Natural Sciences" pageId="5" pageNumber="586" pagination="1 - 30" part="42" refString="Grehan, J. R (2012) Morphological evidence for phylogenetic relationships within the Hepialidae (Lepidoptera: Exoporia). Bulletin of the Buffalo Society of Natural Sciences, 42, 1 - 30." title="Morphological evidence for phylogenetic relationships within the Hepialidae (Lepidoptera: Exoporia)" type="journal article" year="2012">Grehan 2012</bibRefCitation>
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).
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</paragraph>
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</subSubSection>
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||
<subSubSection id="C33CC7BBFFEB7D2289CFF8A40659E328" lastPageId="7" lastPageNumber="588" pageId="5" pageNumber="586" type="description">
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||
<paragraph id="8B999430FFEB7D2089CFF8A4009DE110" blockId="5.[151,1436,1622,2007]" box="[199,1436,1874,1899]" pageId="5" pageNumber="586">
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||
<emphasis id="B9524822FFEB7D2089CFF8A40450E111" bold="true" box="[199,337,1874,1899]" pageId="5" pageNumber="586">Description</emphasis>
|
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.
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<emphasis id="B9524822FFEB7D208857F8A40499E111" box="[351,408,1874,1899]" italics="true" pageId="5" pageNumber="586">Male</emphasis>
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(
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<figureCitation id="131D88B5FFEB7D2088AEF8A40727E111" box="[422,550,1874,1899]" captionStart="FIGURES 11 − 15" captionStartId="4.[151,264,1375,1398]" captionTargetBox="[151,1436,628,1354]" captionTargetId="figure@4.[151,1436,628,1354]" captionTargetPageId="4" captionText="FIGURES 11 − 15. Limyra silvai sp. nov. 11, male holotype, dorsal view; 12, male holotype, ventral view; 13, male paratype, dorsal view (MZSP 14.403); 14, female paratype, dorsal view (MZSP 14.404); 15, female paratype, ventral view (MZSP 14.404). Scale bar = 1 cm." httpUri="https://zenodo.org/record/998526/files/figure.png" pageId="5" pageNumber="586">Figs 11–13</figureCitation>
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,
|
||
<figureCitation id="131D88B5FFEB7D208B39F8A50750E111" box="[561,593,1875,1899]" captionStart="FIGURES 22 − 29" captionStartId="6.[151,264,1386,1409]" captionTargetBox="[152,1430,207,1365]" captionTargetId="figure@6.[151,1436,193,1365]" captionTargetPageId="6" captionText="FIGURES 22 − 29. Limyra silvai sp. nov. 22. Male abdomen (holotype), upper dorsal, lower ventral view. 23 − 25. Male genitalia (holotype): 23, ventral view; 24, lateral view; 25, anterior view (internal), fultura superior (arrow). 26 − 27. Female abdomen (MZSP 14.404). 26, upper dorsal, lower ventral view; 27, tergum and sternum VII and VIII (tergum VIII membranous), sternum VIII (arrow). 28 − 29. Female genitalia (MZSP 14.404). 28, lateral view; 29, posterior view. Scale bar = 1 mm, except Figs 22 and 26 = 1 cm." httpUri="https://zenodo.org/record/998530/files/figure.png" pageId="5" pageNumber="586">22</figureCitation>
|
||
). Medium sized moths with wingspan ranging wingspan from
|
||
<quantity id="4CDE39D5FFEB7D208C00F8A50099E110" box="[1288,1432,1875,1899]" metricMagnitude="-2" metricUnit="m" metricValue="3.85" metricValueMax="4.3" metricValueMin="3.4" pageId="5" pageNumber="586" unit="mm" value="38.5" valueMax="43.0" valueMin="34.0">34 to 43 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B999430FFEB7D2089CFF8800445E1AD" blockId="5.[151,1436,1622,2007]" pageId="5" pageNumber="586">
|
||
<emphasis id="B9524822FFEB7D2089CFF880040AE1F5" box="[199,267,1910,1935]" italics="true" pageId="5" pageNumber="586">Head.</emphasis>
|
||
Clypeus glabrous anteriorly and medially projected, and fairly well differentiated from the frons. Labial palpus stout, first and second palpomeres partially fused. Antenna bipectinate, sensilla trichodea and sensilla caetica present.
|
||
</paragraph>
|
||
<caption id="DF59C4B8FFE87D23899FFA9C0711E381" httpUri="https://zenodo.org/record/998530/files/figure.png" pageId="6" pageNumber="587" targetBox="[152,1430,207,1365]" targetPageId="6">
|
||
<paragraph id="8B999430FFE87D23899FFA9C0711E381" blockId="6.[151,1436,1386,1531]" pageId="6" pageNumber="587">
|
||
<emphasis id="B9524822FFE87D23899FFA9C0452E3FA" bold="true" box="[151,339,1386,1409]" pageId="6" pageNumber="587">FIGURES 22−29</emphasis>
|
||
.
|
||
<taxonomicName id="4C26EFB3FFE87D23886DFA9A04F1E3FB" authority="Duarte, 2017" authorityName="Duarte" authorityYear="2017" box="[357,496,1387,1409]" class="Insecta" family="Hepialidae" genus="Limyra" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="587" phylum="Arthropoda" rank="species" species="silvai" status="sp. nov.">
|
||
<emphasis id="B9524822FFE87D23886DFA9A04F1E3FB" box="[357,496,1387,1409]" italics="true" pageId="6" pageNumber="587">Limyra silvai</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A261F559FFE87D2388F4FA9D0750E3FA" box="[508,593,1387,1408]" pageId="6" pageNumber="587" rank="species">
|
||
<emphasis id="B9524822FFE87D2388F4FA9D0750E3FA" bold="true" box="[508,593,1387,1408]" pageId="6" pageNumber="587">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
22. Male abdomen (holotype), upper dorsal, lower ventral view. 23−25. Male genitalia (holotype): 23, ventral view; 24, lateral view; 25, anterior view (internal), fultura superior (arrow). 26−27. Female abdomen (MZSP 14.404). 26, upper dorsal, lower ventral view; 27, tergum and sternum VII and VIII (tergum VIII membranous), sternum VIII (arrow). 28−29. Female genitalia (MZSP 14.404). 28, lateral view; 29, posterior view. Scale bar = 1 mm, except Figs 22 and 26 = 1 cm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF59C4B8FFE87D23899FF8220606E190" box="[151,775,2004,2026]" pageId="6" pageNumber="587">
|
||
<paragraph id="8B999430FFE87D23899FF8220606E190" blockId="6.[151,775,2004,2026]" box="[151,775,2004,2026]" pageId="6" pageNumber="587">
|
||
<emphasis id="B9524822FFE87D23899FF8220419E193" bold="true" box="[151,280,2004,2026]" pageId="6" pageNumber="587">FIGURE 30</emphasis>
|
||
.
|
||
<taxonomicName id="4C26EFB3FFE87D23882DF82304ADE190" authority="Duarte, 2017" authorityName="Duarte" authorityYear="2017" box="[293,428,2004,2026]" class="Insecta" family="Hepialidae" genus="Limyra" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="587" phylum="Arthropoda" rank="species" species="silvai" status="sp. nov.">
|
||
<emphasis id="B9524822FFE87D23882DF82304ADE190" box="[293,428,2004,2026]" italics="true" pageId="6" pageNumber="587">Limyra silvai</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A261F559FFE87D2388BAF8230703E190" box="[434,514,2005,2026]" pageId="6" pageNumber="587" rank="species">
|
||
<emphasis id="B9524822FFE87D2388BAF8230703E190" bold="true" box="[434,514,2005,2026]" pageId="6" pageNumber="587">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
Male genitalia, holotype.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF59C4B8FFE97D22899FFD6E0699E4D4" box="[151,920,664,686]" pageId="7" pageNumber="588">
|
||
<paragraph id="8B999430FFE97D22899FFD6E0699E4D4" blockId="7.[151,920,664,686]" box="[151,920,664,686]" pageId="7" pageNumber="588">
|
||
<emphasis id="B9524822FFE97D22899FFD6E0419E4D7" bold="true" box="[151,280,664,686]" pageId="7" pageNumber="588">FIGURE 31</emphasis>
|
||
.
|
||
<taxonomicName id="4C26EFB3FFE97D22882DFD6F04ADE4D4" authority="Duarte, 2017" authorityName="Duarte" authorityYear="2017" box="[293,428,664,686]" class="Insecta" family="Hepialidae" genus="Limyra" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="588" phylum="Arthropoda" rank="species" species="silvai" status="sp. nov.">
|
||
<emphasis id="B9524822FFE97D22882DFD6F04ADE4D4" box="[293,428,664,686]" italics="true" pageId="7" pageNumber="588">Limyra silvai</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A261F559FFE97D2288BAFD6E0703E4D7" box="[434,514,664,685]" pageId="7" pageNumber="588" rank="species">
|
||
<emphasis id="B9524822FFE97D2288BAFD6E0703E4D7" bold="true" box="[434,514,664,685]" pageId="7" pageNumber="588">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
Male (left) and female (right) venation.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B999430FFE97D2289CFFD2306EAE56B" blockId="7.[151,1437,725,1758]" pageId="7" pageNumber="588">
|
||
<emphasis id="B9524822FFE97D2289CFFD230420E494" box="[199,289,725,750]" italics="true" pageId="7" pageNumber="588">Thorax:</emphasis>
|
||
Legs: distitarsus as in
|
||
<taxonomicName id="4C26EFB3FFE97D228B3AFD230781E494" authorityName="Walker" authorityYear="1856" box="[562,640,725,750]" class="Insecta" family="Hepialidae" genus="Cibyra" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="588" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9524822FFE97D228B3AFD230781E494" box="[562,640,725,750]" italics="true" pageId="7" pageNumber="588">Cibyra</emphasis>
|
||
</taxonomicName>
|
||
. Venation of fore- and hindwings as in
|
||
<taxonomicName id="4C26EFB3FFE97D228D5DFD2301A2E494" authorityName="Walker" authorityYear="1856" box="[1109,1187,725,750]" class="Insecta" family="Hepialidae" genus="Cibyra" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="588" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9524822FFE97D228D5DFD2301A2E494" box="[1109,1187,725,750]" italics="true" pageId="7" pageNumber="588">Cibyra</emphasis>
|
||
</taxonomicName>
|
||
, but in forewing Sc1 vestigial; in hindwing CuP vestigial, M2 complete and 2A present (
|
||
<figureCitation id="131D88B5FFE97D228A83FD0F06DFE56B" box="[907,990,761,786]" captionStart="FIGURE 31" captionStartId="7.[151,250,664,686]" captionText="FIGURE 31. Limyra silvai sp. nov. Male (left) and female (right) venation." pageId="7" pageNumber="588">Fig. 31</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B999430FFE97D2289CFFCEB0775E54C" blockId="7.[151,1437,725,1758]" box="[199,628,797,822]" pageId="7" pageNumber="588">
|
||
<emphasis id="B9524822FFE97D2289CFFCEB0430E54C" box="[199,305,797,822]" italics="true" pageId="7" pageNumber="588">Abdomen</emphasis>
|
||
: Sternum VIII membranous.
|
||
</paragraph>
|
||
<paragraph id="8B999430FFE97D2289CFFCB70656E5BC" blockId="7.[151,1437,725,1758]" pageId="7" pageNumber="588">
|
||
<emphasis id="B9524822FFE97D2289CFFCB7046AE520" box="[199,363,833,858]" italics="true" pageId="7" pageNumber="588">Male genitalia</emphasis>
|
||
(
|
||
<figureCitation id="131D88B5FFE97D228871FCB704FBE520" box="[377,506,833,858]" captionStart="FIGURES 22 − 29" captionStartId="6.[151,264,1386,1409]" captionTargetBox="[152,1430,207,1365]" captionTargetId="figure@6.[151,1436,193,1365]" captionTargetPageId="6" captionText="FIGURES 22 − 29. Limyra silvai sp. nov. 22. Male abdomen (holotype), upper dorsal, lower ventral view. 23 − 25. Male genitalia (holotype): 23, ventral view; 24, lateral view; 25, anterior view (internal), fultura superior (arrow). 26 − 27. Female abdomen (MZSP 14.404). 26, upper dorsal, lower ventral view; 27, tergum and sternum VII and VIII (tergum VIII membranous), sternum VIII (arrow). 28 − 29. Female genitalia (MZSP 14.404). 28, lateral view; 29, posterior view. Scale bar = 1 mm, except Figs 22 and 26 = 1 cm." httpUri="https://zenodo.org/record/998530/files/figure.png" pageId="7" pageNumber="588">Figs 23–25</figureCitation>
|
||
,
|
||
<figureCitation id="131D88B5FFE97D228B0EFCB70727E523" box="[518,550,833,857]" captionStart="FIGURE 30" captionStartId="6.[151,250,2004,2026]" captionText="FIGURE 30. Limyra silvai sp. nov. Male genitalia, holotype." pageId="7" pageNumber="588">30</figureCitation>
|
||
). Tegumen mostly fused to the pseudotegumen, ventral portion articulated with saccus. Saccus convex and in two fused parts. Tergal lobes softly sclerotized, resting on against the pseudotegumen. Pseudotegumen dorsally and ventrally unfused. Fultura superior present. Valvae slightly setose. Phallus membranous and when everted the vesica as a bladder.
|
||
</paragraph>
|
||
<paragraph id="8B999430FFE97D2289CFFC2706CDE590" blockId="7.[151,1437,725,1758]" box="[199,972,977,1002]" pageId="7" pageNumber="588">
|
||
<emphasis id="B9524822FFE97D2289CFFC27041DE590" box="[199,284,977,1002]" italics="true" pageId="7" pageNumber="588">Female</emphasis>
|
||
(
|
||
<figureCitation id="131D88B5FFE97D228821FC2704AAE590" box="[297,427,977,1002]" captionStart="FIGURES 11 − 15" captionStartId="4.[151,264,1375,1398]" captionTargetBox="[151,1436,628,1354]" captionTargetId="figure@4.[151,1436,628,1354]" captionTargetPageId="4" captionText="FIGURES 11 − 15. Limyra silvai sp. nov. 11, male holotype, dorsal view; 12, male holotype, ventral view; 13, male paratype, dorsal view (MZSP 14.403); 14, female paratype, dorsal view (MZSP 14.404); 15, female paratype, ventral view (MZSP 14.404). Scale bar = 1 cm." httpUri="https://zenodo.org/record/998526/files/figure.png" pageId="7" pageNumber="588">Figs 14, 15</figureCitation>
|
||
). Only differences from the male are mentioned.
|
||
</paragraph>
|
||
<paragraph id="8B999430FFE97D2289CFFC03014FE22F" blockId="7.[151,1437,725,1758]" pageId="7" pageNumber="588">
|
||
<emphasis id="B9524822FFE97D2289CFFC030405E274" box="[199,260,1013,1038]" italics="true" pageId="7" pageNumber="588">Head</emphasis>
|
||
: clypeus anteriorly and medially projected, glabrous and slightly differentiated from the frons. Labial palpus one-segmented. Antenna bipectinate with rami arising from a slight projection at the mid length of the antennomere, sensilla trichodea reduced in size and quantity, sensilla caetica present.
|
||
</paragraph>
|
||
<paragraph id="8B999430FFE97D2289CFFB97017CE203" blockId="7.[151,1437,725,1758]" box="[199,1149,1121,1146]" pageId="7" pageNumber="588">
|
||
<emphasis id="B9524822FFE97D2289CFFB97041EE200" box="[199,287,1121,1146]" italics="true" pageId="7" pageNumber="588">Thorax.</emphasis>
|
||
Venation of fore- and hindwings as in male, but hindwing with 3A (
|
||
<figureCitation id="131D88B5FFE97D228D15FB970171E203" box="[1053,1136,1121,1146]" captionStart="FIGURE 31" captionStartId="7.[151,250,664,686]" captionText="FIGURE 31. Limyra silvai sp. nov. Male (left) and female (right) venation." pageId="7" pageNumber="588">Fig. 31</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B999430FFE97D2289CFFB730695E2B8" blockId="7.[151,1437,725,1758]" pageId="7" pageNumber="588">
|
||
<emphasis id="B9524822FFE97D2289CFFB730430E2E4" box="[199,305,1157,1182]" italics="true" pageId="7" pageNumber="588">Abdomen</emphasis>
|
||
: tergum VIII as a thick membrane; sternum VIII compound as two lateral, slightly sclerotized, and semi-circular plates linked by a narrow sclerotized ridge posteriorly.
|
||
</paragraph>
|
||
<paragraph id="8B999430FFE97D2289CFFB3B0659E328" blockId="7.[151,1437,725,1758]" pageId="7" pageNumber="588">
|
||
<emphasis id="B9524822FFE97D2289CFFB3B0486E29C" box="[199,391,1229,1254]" italics="true" pageId="7" pageNumber="588">Female genitalia</emphasis>
|
||
(
|
||
<figureCitation id="131D88B5FFE97D22889EFB3B071AE29C" box="[406,539,1229,1254]" captionStart="FIGURES 22 − 29" captionStartId="6.[151,264,1386,1409]" captionTargetBox="[152,1430,207,1365]" captionTargetId="figure@6.[151,1436,193,1365]" captionTargetPageId="6" captionText="FIGURES 22 − 29. Limyra silvai sp. nov. 22. Male abdomen (holotype), upper dorsal, lower ventral view. 23 − 25. Male genitalia (holotype): 23, ventral view; 24, lateral view; 25, anterior view (internal), fultura superior (arrow). 26 − 27. Female abdomen (MZSP 14.404). 26, upper dorsal, lower ventral view; 27, tergum and sternum VII and VIII (tergum VIII membranous), sternum VIII (arrow). 28 − 29. Female genitalia (MZSP 14.404). 28, lateral view; 29, posterior view. Scale bar = 1 mm, except Figs 22 and 26 = 1 cm." httpUri="https://zenodo.org/record/998530/files/figure.png" pageId="7" pageNumber="588">Figs 28, 29</figureCitation>
|
||
). Anal papillae tapered, projected posteriorly. Lamella antevaginalis compound mainly by two processes separated meso-anteriorly that extends ventro-posteriorly fusing to a mesal process, thus forming a depression between both structures. Antrum situated anterior to the lamella. Subanal plate slightly sclerotized. Ductus bursae tube-like; corpus bursae cylindrical.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33CC7BBFFE97D2289CFFAAB0651E07C" pageId="7" pageNumber="588" type="etymology">
|
||
<paragraph id="8B999430FFE97D2289CFFAAB0651E07C" blockId="7.[151,1437,725,1758]" pageId="7" pageNumber="588">
|
||
<emphasis id="B9524822FFE97D2289CFFAAB0447E30C" bold="true" box="[199,326,1373,1398]" pageId="7" pageNumber="588">Etymology</emphasis>
|
||
. While forming the generic name
|
||
<taxonomicName id="4C26EFB3FFE97D228BC6FAA8061EE30F" authority="Duarte, 2017" authorityName="Duarte" authorityYear="2017" box="[718,799,1374,1397]" class="Insecta" family="Hepialidae" genus="Limyra" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="588" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B9524822FFE97D228BC6FAA8061EE30F" box="[718,799,1374,1397]" italics="true" pageId="7" pageNumber="588">Limyra</emphasis>
|
||
</taxonomicName>
|
||
we follow the tradition of
|
||
<bibRefCitation id="EFB7E9C1FFE97D228D5AFAAB01F6E30C" author="Walker, F." box="[1106,1271,1373,1398]" journalOrPublisher="Lepidoptera Heterocera. London (Trustees of BMNH)" pageId="7" pageNumber="588" pagination="1509" part="4" refString="Walker, F. (1856) List of the specimens of lepidopterous insects in the collection of the British Museum. Part VII. Lepidoptera Heterocera. London (Trustees of BMNH), [4] pp. + pp. [1509] - 1808." title="List of the specimens of lepidopterous insects in the collection of the British Museum. Part VII" type="journal article" year="1856">Walker (1856)</bibRefCitation>
|
||
, who gave the generic name
|
||
<taxonomicName id="4C26EFB3FFE97D22883EFA770485E3E0" authorityName="Walker" authorityYear="1856" box="[310,388,1409,1434]" class="Insecta" family="Hepialidae" genus="Cibyra" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="588" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9524822FFE97D22883EFA770485E3E0" box="[310,388,1409,1434]" italics="true" pageId="7" pageNumber="588">Cibyra</emphasis>
|
||
</taxonomicName>
|
||
for the genus of moths distributed in South America. We suppose that the name of the antic town Κιβυρα served as an inspiration. The name of the new genus
|
||
<taxonomicName id="4C26EFB3FFE97D228A8FFA5006D9E3C7" authority="Duarte, 2017" authorityName="Duarte" authorityYear="2017" box="[903,984,1446,1469]" class="Insecta" family="Hepialidae" genus="Limyra" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="588" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B9524822FFE97D228A8FFA5006D9E3C7" box="[903,984,1446,1469]" italics="true" pageId="7" pageNumber="588">Limyra</emphasis>
|
||
</taxonomicName>
|
||
also derives from the name of the small antic city (in Greek Λιµύρα) in Lycia on the southern coast of
|
||
<collectingCountry id="F331D4A0FFE97D228A6AFA3F06E7E398" box="[866,998,1481,1506]" name="Turkey" pageId="7" pageNumber="588">Asia Minor</collectingCountry>
|
||
, on the Limyrus River. Nowadays this place are ruins in
|
||
<collectingCountry id="F331D4A0FFE97D228856FA1B04B1E07C" box="[350,432,1517,1542]" name="Turkey" pageId="7" pageNumber="588">Turkey</collectingCountry>
|
||
. The gender of the name is feminine.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33CC7BBFFE97D2289CFF9E7072DE0A4" pageId="7" pageNumber="588" type="discussion">
|
||
<paragraph id="8B999430FFE97D2289CFF9E7072DE0A4" blockId="7.[151,1437,725,1758]" pageId="7" pageNumber="588">
|
||
<emphasis id="B9524822FFE97D2289CFF9E70435E050" bold="true" box="[199,308,1553,1578]" pageId="7" pageNumber="588">Remarks</emphasis>
|
||
. The presence of
|
||
<taxonomicName id="4C26EFB3FFE97D2288F6F9E4074EE053" authority="Duarte, 2017" authorityName="Duarte" authorityYear="2017" box="[510,591,1554,1577]" class="Insecta" family="Hepialidae" genus="Limyra" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="588" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B9524822FFE97D2288F6F9E4074EE053" box="[510,591,1554,1577]" italics="true" pageId="7" pageNumber="588">Limyra</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A261F559FFE97D228B5FF9E707C0E050" box="[599,705,1553,1578]" pageId="7" pageNumber="588" rank="genus">
|
||
<emphasis id="B9524822FFE97D228B5FF9E707C0E050" bold="true" box="[599,705,1553,1578]" pageId="7" pageNumber="588">gen. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
in southern
|
||
<collectingCountry id="F331D4A0FFE97D228A46F9E70693E050" box="[846,914,1553,1578]" name="Brazil" pageId="7" pageNumber="588">Brazil</collectingCountry>
|
||
is surprising since no other matching taxon is so far known. The presence of posterior lateral knob (not strongly developed), a broad intermediate connection between the tergosternal bar, the lateral ridge and a broken anterior margin at the tergosternal connection, along with a very close parallel position between the outer hindwing Sc and R veins support the inclusion of
|
||
<taxonomicName id="4C26EFB3FFE97D228C1BF9880065E0EF" authority="Duarte, 2017" authorityName="Duarte" authorityYear="2017" box="[1299,1380,1662,1685]" class="Insecta" family="Hepialidae" genus="Limyra" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="588" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B9524822FFE97D228C1BF9880065E0EF" box="[1299,1380,1662,1685]" italics="true" pageId="7" pageNumber="588">Limyra</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A261F559FFE97D228C63F98B05C9E0C0" pageId="7" pageNumber="588" rank="genus">
|
||
<emphasis id="B9524822FFE97D228C63F98B05C9E0C0" bold="true" pageId="7" pageNumber="588">gen. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
within the 'cibyrine' cluster of genera (
|
||
<bibRefCitation id="EFB7E9C1FFE97D228B91F9570634E0C3" author="Grehan, J. R" box="[665,821,1697,1722]" journalOrPublisher="Bulletin of the Buffalo Society of Natural Sciences" pageId="7" pageNumber="588" pagination="1 - 30" part="42" refString="Grehan, J. R (2012) Morphological evidence for phylogenetic relationships within the Hepialidae (Lepidoptera: Exoporia). Bulletin of the Buffalo Society of Natural Sciences, 42, 1 - 30." title="Morphological evidence for phylogenetic relationships within the Hepialidae (Lepidoptera: Exoporia)" type="journal article" year="2012">Grehan 2012</bibRefCitation>
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). Further investigation will be necessary for a better understanding of such relationships.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |