234 lines
32 KiB
XML
234 lines
32 KiB
XML
<document id="291E0430DE0A5EF732EF83F2E2E2FF5A" ID-CLB-Dataset="28347" ID-DOI="10.11646/zootaxa.4567.3.1" ID-GBIF-Dataset="47813581-d217-4a19-8848-a3e9cf2ddaf4" ID-ISSN="1175-5326" ID-Zenodo-Dep="2598945" ID-ZooBank="0AEFAF80-B001-4A18-88AC-5B6A189F6DCD" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1553062614686" checkinUser="plazi" docAuthor="Staples, David A." docDate="2019" docId="03895C33290F4F27FF01FC74FDBEFC9F" docLanguage="en" docName="zootaxa.4567.3.1.pdf" docOrigin="Zootaxa 4567 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Hedgpethia filamentus Staples 2019, sp. nov." docType="treatment" docVersion="8" lastPageNumber="433" masterDocId="FFB0244B29114F07FF96FFB2FF80FFFD" masterDocTitle="Pycnogonids (Arthropoda, Pycnogonida) from the Southwest Indian Ridge" masterLastPageNumber="449" masterPageNumber="401" pageNumber="431" updateTime="1698701998649" updateUser="ExternalLinkService">
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<mods:titleInfo id="7137BACEA6015A89ED77F66FCB6F2292">
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<mods:title id="7FC2495C04CCCC48988747F1C208239B">Pycnogonids (Arthropoda, Pycnogonida) from the Southwest Indian Ridge</mods:title>
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<mods:name id="3C8EED8A4593585B816BA3F616D4F2EA" type="personal">
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<mods:roleTerm id="589016E281154DF2EF82F0DFE599D8E9">Author</mods:roleTerm>
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<mods:namePart id="B52F02532B8DBCAD88FC061E91A31868">Staples, David A.</mods:namePart>
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<mods:typeOfResource id="D90388DA8A5592CA399A90C2C4F6B9A2">text</mods:typeOfResource>
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<mods:title id="E840753649C15FDEEE9CE7A02C5B940A">Zootaxa</mods:title>
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<mods:part id="3E5ECF0F83AA306055C108E83B799B7E">
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<mods:date id="1235E637D827C3982888F50DA3C057E0">2019</mods:date>
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<mods:number id="C7015E768DAFFE3FF6DD4FB31C937041">2019-03-18</mods:number>
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<mods:number id="8F5684E2F969F6ABF56B9237A94B78C7">4567</mods:number>
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<mods:number id="C21B10F296EC7ACCD26B9BE8C4841957">3</mods:number>
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<mods:classification id="3D3E4F1E75D6B612422E2D6B9B5BEE38">journal article</mods:classification>
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<mods:identifier id="2ADAB247FAFA72FF13778A5468752F95" type="CLB-Dataset">28347</mods:identifier>
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<mods:identifier id="093BDF5797C0B56436613AD2DE2A8E66" type="DOI">10.11646/zootaxa.4567.3.1</mods:identifier>
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<mods:identifier id="B45D274606CB13593E97837FA464446D" type="GBIF-Dataset">47813581-d217-4a19-8848-a3e9cf2ddaf4</mods:identifier>
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<mods:identifier id="3C8B5D20F00C521971563B42FF4E117B" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="4C77C1B8BAEB682B0A4E9FD3E2C9B6C9" type="Zenodo-Dep">2598945</mods:identifier>
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<mods:identifier id="EED99438C8709C5C4097991F89B1C6D5" type="ZooBank">0AEFAF80-B001-4A18-88AC-5B6A189F6DCD</mods:identifier>
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</mods:mods>
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<treatment id="03895C33290F4F27FF01FC74FDBEFC9F" ID-DOI="http://doi.org/10.5281/zenodo.5944908" ID-GBIF-Taxon="154435135" ID-Zenodo-Dep="5944908" LSID="urn:lsid:plazi:treatment:03895C33290F4F27FF01FC74FDBEFC9F" httpUri="http://treatment.plazi.org/id/03895C33290F4F27FF01FC74FDBEFC9F" lastPageId="32" lastPageNumber="433" pageId="30" pageNumber="431">
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<subSubSection id="C33ABEAE290F4F19FF01FC74FD90FC1D" box="[151,528,966,992]" pageId="30" pageNumber="431" type="nomenclature">
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<paragraph id="8B9FED25290F4F19FF01FC74FD90FC1D" blockId="30.[151,528,966,1026]" box="[151,528,966,992]" pageId="30" pageNumber="431">
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<heading id="D0D75A49290F4F19FF01FC74FD90FC1D" bold="true" box="[151,528,966,992]" fontSize="11" level="1" pageId="30" pageNumber="431" reason="1">
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<emphasis id="B9543137290F4F19FF01FC74FD90FC1D" bold="true" box="[151,528,966,992]" pageId="30" pageNumber="431">
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<taxonomicName id="4C2096A6290F4F19FF01FC74FE2AFC1D" ID-CoL="1b0acde2-903f-4ac4-b61d-b717b4def07d" authority="Staples, 2019" authorityName="Staples" authorityYear="2019" box="[151,426,966,992]" class="Pycnogonida" family="Colossendeidae" genus="Hedgpethia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pantopoda" pageId="30" pageNumber="431" phylum="Arthropoda" rank="species" species="filamentus" status="sp. nov.">
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<emphasis id="B9543137290F4F19FF01FC74FE2AFC1D" bold="true" box="[151,426,966,992]" italics="true" pageId="30" pageNumber="431">Hedgpethia filamentus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A2678C4C290F4F19FE24FC74FD90FC1D" box="[434,528,966,992]" pageId="30" pageNumber="431" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C33ABEAE290F4F19FF01FC5BFE38FBFC" box="[151,440,1001,1026]" pageId="30" pageNumber="431" type="description">
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<paragraph id="8B9FED25290F4F19FF01FC5BFE38FBFC" blockId="30.[151,528,966,1026]" box="[151,440,1001,1026]" pageId="30" pageNumber="431">
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<figureCitation id="131BF1A0290F4F19FF01FC5BFE81FBFF" box="[151,257,1001,1026]" captionStart="FIGURE 15" captionStartId="31.[151,250,1931,1953]" captionTargetBox="[182,1424,162,1919]" captionTargetId="figure@31.[160,1427,153,1965]" captionTargetPageId="31" captionText="FIGURE 15. Hedgpethia filamentus sp. nov., holotype. A, trunk, dorsal view; B, trunk, lateral view; C, right oviger; D, right oviger claw; E, right palp; F, third leg; G, tarsus and propodus; H, abdomen, ventral view. Paratype, (JC066-4214). I, ocular tubercle." httpUri="https://zenodo.org/record/2598973/files/figure.png" pageId="30" pageNumber="431">Figure 15</figureCitation>
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A–I;
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<figureCitation id="131BF1A0290F4F19FEABFC5BFE38FBFC" box="[317,440,1001,1026]" captionStart="PLATE 5" captionStartId="44.[151,234,1740,1762]" captionTargetBox="[151,1436,193,1718]" captionTargetId="figure@44.[151,1436,193,1718]" captionTargetPageId="44" captionText="PLATE 5. Colossendeis sp. A, proboscis lateral; B, palp. Colossendeis melancholicus. C, specimen on rock wall. D, specimen with proboscis inserted into Isactinernus quadrilobatus. Hedgpethia filamentus sp. nov., holotype. E, trunk, lateral view; F, trunk, dorsal view. Hedgpethia shalei sp. nov., holotype. G, trunk, lateral view; H, trunk, dorsal view." httpUri="https://zenodo.org/record/2598997/files/figure.png" pageId="30" pageNumber="431">Plate 5E, F</figureCitation>
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C33ABEAE290F4F19FF01FB83FD95FADF" pageId="30" pageNumber="431" type="materials_examined">
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<paragraph id="8B9FED25290F4F19FF01FB83FE37FB6F" blockId="30.[151,1437,1073,2034]" pageId="30" pageNumber="431">
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<emphasis id="B9543137290F4F19FF01FB83FEFCFBB7" bold="true" box="[151,380,1073,1098]" pageId="30" pageNumber="431">Material examined</emphasis>
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.
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<materialsCitation id="3B48E778290F4F19FE1BFB83FE37FB6F" ID-GBIF-Occurrence="2013824389" collectingDate="2011-12-14" collectionCode="NHMUK" elevation="750" latitude="-32.714333" longLatPrecision="12" longitude="57.272335" pageId="30" pageNumber="431" specimenCode="NHMUK 2018.38" specimenCount="1" typeStatus="holotype">
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<typeStatus id="549B5387290F4F19FE1BFB83FE78FBB7" box="[397,504,1073,1098]" pageId="30" pageNumber="431" type="holotype">Holotype</typeStatus>
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, sex undetermined (
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<specimenCode id="DB86455E290F4F19FD74FB80FC37FBB7" box="[738,951,1074,1098]" collectionCode="NHMUK" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="30" pageNumber="431" type="Museum">NHMUK 2018.38</specimenCode>
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), Southwest Indian Ocean,
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<taxonomicName id="4C2096A6290F4F19FB61FB83FA18FBB7" authority="Bank" authorityName="Bank" box="[1271,1432,1073,1098]" class="Insecta" family="Curculionidae" genus="Atlantis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="431" phylum="Arthropoda" rank="genus">Atlantis Bank</taxonomicName>
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,
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<geoCoordinate id="EE148BE2290F4F19FF01FBE4FEA1FB93" box="[151,289,1109,1134]" direction="south" orientation="latitude" pageId="30" pageNumber="431" precision="9" value="-32.714333">32° 42.86´S</geoCoordinate>
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,
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<geoCoordinate id="EE148BE2290F4F19FEB8FBE7FE3AFB93" box="[302,442,1109,1134]" direction="east" orientation="longitude" pageId="30" pageNumber="431" precision="9" value="57.272335">57° 16.34´E</geoCoordinate>
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, ROV,
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<elevation id="000D0A16290F4F19FD98FBE7FDD9FB90" box="[526,601,1109,1134]" metricMagnitude="2" metricUnit="m" metricValue="7.5" pageId="30" pageNumber="431" unit="m" value="750.0">
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<quantity id="4CD840C0290F4F19FD98FBE7FDD9FB90" box="[526,601,1109,1134]" metricMagnitude="2" metricUnit="m" metricValue="7.5" pageId="30" pageNumber="431" unit="m" value="750.0">750 m</quantity>
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</elevation>
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, specimen JC066-4211C, stn 8.29, mooring site,
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<collectingMethod id="52619532290F4F19FB1CFBE4FB51FB90" box="[1162,1233,1110,1133]" pageId="30" pageNumber="431">on net</collectingMethod>
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containing whale bone,
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<date id="FF9ECBE5290F4F19FF4DFBC8FE32FB6F" box="[219,434,1145,1170]" pageId="30" pageNumber="431" value="2011-12-14">
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<collectingDate id="EFDA320D290F4F19FF4DFBC8FE32FB6F" box="[219,434,1145,1170]" pageId="30" pageNumber="431" value="2011-12-14">14 December 2011</collectingDate>
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</date>
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.
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</materialsCitation>
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</paragraph>
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<paragraph id="8B9FED25290F4F19FF51FB2FFD95FADF" blockId="30.[151,1437,1073,2034]" pageId="30" pageNumber="431">
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<materialsCitation id="3B48E778290F4F19FF51FB2FFD91FADF" ID-GBIF-Occurrence="2013824373" collectingDate="2011-12-14" collectionCode="NHMUK" elevation="750" latitude="-32.714333" longLatPrecision="12" longitude="57.272335" pageId="30" pageNumber="431" specimenCode="NHMUK 2018.39, NHMUK 2018.40" specimenCount="2" typeStatus="paratype">
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||
<emphasis id="B9543137290F4F19FF51FB2FFEC6FB4B" bold="true" box="[199,326,1181,1206]" pageId="30" pageNumber="431">
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||
<typeStatus id="549B5387290F4F19FF51FB2FFEC1FB4B" box="[199,321,1181,1206]" pageId="30" pageNumber="431" type="paratype">Paratypes</typeStatus>
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.
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</emphasis>
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Two specimens of undetermined sex and two smaller (subadult?) specimens (
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<specimenCode id="DB86455E290F4F19FB2AFB2CFA11FB4B" box="[1212,1425,1182,1206]" collectionCode="NHMUK" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="30" pageNumber="431" type="Museum">NHMUK 2018.39</specimenCode>
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), same collection details as
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<typeStatus id="549B5387290F4F19FE2FFB73FD9FFB27" box="[441,543,1217,1242]" pageId="30" pageNumber="431" type="holotype">holotype</typeStatus>
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||
. One damaged specimen (
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<specimenCode id="DB86455E290F4F19FCD7FB70FB92FB27" box="[833,1042,1218,1242]" collectionCode="NHMUK" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="30" pageNumber="431" type="Museum">NHMUK 2018.40</specimenCode>
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), Southwest Indian Ocean,
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<taxonomicName id="4C2096A6290F4F19FAD2FB73FF55FB03" authority="Bank" authorityName="Bank" class="Insecta" family="Curculionidae" genus="Atlantis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="431" phylum="Arthropoda" rank="genus">Atlantis Bank</taxonomicName>
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,
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<geoCoordinate id="EE148BE2290F4F19FF74FB54FEECFB03" box="[226,364,1253,1278]" direction="south" orientation="latitude" pageId="30" pageNumber="431" precision="9" value="-32.714333">32° 42.86´S</geoCoordinate>
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,
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<geoCoordinate id="EE148BE2290F4F19FEEFFB57FD84FB03" box="[377,516,1253,1278]" direction="east" orientation="longitude" pageId="30" pageNumber="431" precision="9" value="57.272335">57° 16.34´E</geoCoordinate>
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, ROV,
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<quantity id="4CD840C0290F4F19FDCFFB57FD24FB00" box="[601,676,1253,1278]" metricMagnitude="2" metricUnit="m" metricValue="7.5" pageId="30" pageNumber="431" unit="m" value="750.0">
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||
<elevation id="000D0A16290F4F19FDCFFB57FD24FB00" box="[601,676,1253,1278]" metricMagnitude="2" metricUnit="m" metricValue="7.5" pageId="30" pageNumber="431" unit="m" value="750.0">750 m</elevation>
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</quantity>
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, specimen JC066-4214C, stn 8.29, mooring site,
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<collectingMethod id="52619532290F4F19FB40FB54FA9DFB00" box="[1238,1309,1254,1277]" pageId="30" pageNumber="431">on net</collectingMethod>
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||
containing Mango wood,
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||
<date id="FF9ECBE5290F4F19FEAFFAB8FD91FADF" box="[313,529,1289,1314]" pageId="30" pageNumber="431" value="2011-12-14">
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||
<collectingDate id="EFDA320D290F4F19FEAFFAB8FD91FADF" box="[313,529,1289,1314]" pageId="30" pageNumber="431" value="2011-12-14">14 December 2011</collectingDate>
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||
</date>
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C33ABEAE290F4F27FF51FA9FFDB3FE73" lastPageId="32" lastPageNumber="433" pageId="30" pageNumber="431" type="description">
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<paragraph id="8B9FED25290F4F19FF51FA9FFB97FA4F" blockId="30.[151,1437,1073,2034]" pageId="30" pageNumber="431">
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||
<emphasis id="B9543137290F4F19FF51FA9FFED9FABB" bold="true" box="[199,345,1325,1350]" pageId="30" pageNumber="431">Description.</emphasis>
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<typeStatus id="549B5387290F4F19FEF5FA9FFE4FFABB" box="[355,463,1325,1350]" pageId="30" pageNumber="431" type="holotype">Holotype</typeStatus>
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||
. Leg span
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<quantity id="4CD840C0290F4F19FDDEFA9CFD1BFAB8" box="[584,667,1326,1350]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" pageId="30" pageNumber="431" unit="mm" value="12.0">12 mm</quantity>
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. A very fine, slender specimen. Trunk (
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<figureCitation id="131BF1A0290F4F19FBF8FA9FFA81FAB8" box="[1134,1281,1325,1350]" captionStart="FIGURE 15" captionStartId="31.[151,250,1931,1953]" captionTargetBox="[182,1424,162,1919]" captionTargetId="figure@31.[160,1427,153,1965]" captionTargetPageId="31" captionText="FIGURE 15. Hedgpethia filamentus sp. nov., holotype. A, trunk, dorsal view; B, trunk, lateral view; C, right oviger; D, right oviger claw; E, right palp; F, third leg; G, tarsus and propodus; H, abdomen, ventral view. Paratype, (JC066-4214). I, ocular tubercle." httpUri="https://zenodo.org/record/2598973/files/figure.png" pageId="30" pageNumber="431">Figs 15A, B</figureCitation>
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||
.
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||
<figureCitation id="131BF1A0290F4F19FA86FA9FFA14FABB" box="[1296,1428,1325,1350]" captionStart="PLATE 5" captionStartId="44.[151,234,1740,1762]" captionTargetBox="[151,1436,193,1718]" captionTargetId="figure@44.[151,1436,193,1718]" captionTargetPageId="44" captionText="PLATE 5. Colossendeis sp. A, proboscis lateral; B, palp. Colossendeis melancholicus. C, specimen on rock wall. D, specimen with proboscis inserted into Isactinernus quadrilobatus. Hedgpethia filamentus sp. nov., holotype. E, trunk, lateral view; F, trunk, dorsal view. Hedgpethia shalei sp. nov., holotype. G, trunk, lateral view; H, trunk, dorsal view." httpUri="https://zenodo.org/record/2598997/files/figure.png" pageId="30" pageNumber="431">Plate 5E, F</figureCitation>
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) segmented, smooth; lateral processes clearly separated by almost twice their basal width. Ocular tubercle placed on mid-region of cephalon, acuminate, height twice width at base, basal part broad, posterior surface inflated; four large eyes of equal size, posterior pair less pigmented compared to anterior pair.
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</paragraph>
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<paragraph id="8B9FED25290F4F19FF51FA0FFA93FA07" blockId="30.[151,1437,1073,2034]" pageId="30" pageNumber="431">Proboscis slightly longer than trunk, spindle-shaped, angled downwards from about one-third length, maximum width at about half length, tapering to rounded tip, narrow basal part about half width at tip.</paragraph>
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<paragraph id="8B9FED25290F4F19FF51F9B7FD33F9BF" blockId="30.[151,1437,1073,2034]" pageId="30" pageNumber="431">
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||
Abdomen (
|
||
<figureCitation id="131BF1A0290F4F19FED1F9B7FE2EF9E3" box="[327,430,1541,1566]" captionStart="FIGURE 15" captionStartId="31.[151,250,1931,1953]" captionTargetBox="[182,1424,162,1919]" captionTargetId="figure@31.[160,1427,153,1965]" captionTargetPageId="31" captionText="FIGURE 15. Hedgpethia filamentus sp. nov., holotype. A, trunk, dorsal view; B, trunk, lateral view; C, right oviger; D, right oviger claw; E, right palp; F, third leg; G, tarsus and propodus; H, abdomen, ventral view. Paratype, (JC066-4214). I, ocular tubercle." httpUri="https://zenodo.org/record/2598973/files/figure.png" pageId="30" pageNumber="431">Fig. 15H</figureCitation>
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) articulated at base, very short, base broad, implanted between fourth lateral processes, directed ventrally, barely evident in dorsal view.
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</paragraph>
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<paragraph id="8B9FED25290F4F19FF51F9FFFE12F99B" blockId="30.[151,1437,1073,2034]" box="[199,402,1613,1638]" pageId="30" pageNumber="431">Chelifores absent.</paragraph>
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<paragraph id="8B9FED25290F4F19FF51F9C3FE0CF953" blockId="30.[151,1437,1073,2034]" pageId="30" pageNumber="431">
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Palp ten-segmented (
|
||
<figureCitation id="131BF1A0290F4F19FE24F9C3FD97F977" box="[434,535,1649,1674]" captionStart="FIGURE 15" captionStartId="31.[151,250,1931,1953]" captionTargetBox="[182,1424,162,1919]" captionTargetId="figure@31.[160,1427,153,1965]" captionTargetPageId="31" captionText="FIGURE 15. Hedgpethia filamentus sp. nov., holotype. A, trunk, dorsal view; B, trunk, lateral view; C, right oviger; D, right oviger claw; E, right palp; F, third leg; G, tarsus and propodus; H, abdomen, ventral view. Paratype, (JC066-4214). I, ocular tubercle." httpUri="https://zenodo.org/record/2598973/files/figure.png" pageId="30" pageNumber="431">Fig. 15E</figureCitation>
|
||
), segment 3 about 1.9 times longer than segment 5, segments 7 to10 slender and of about equal length.
|
||
</paragraph>
|
||
<paragraph id="8B9FED25290F4F19FF51F90BFBD9F8E7" blockId="30.[151,1437,1073,2034]" pageId="30" pageNumber="431">
|
||
Oviger ten-segmented (
|
||
<figureCitation id="131BF1A0290F4F19FE46F90BFDB7F92F" box="[464,567,1721,1746]" captionStart="FIGURE 15" captionStartId="31.[151,250,1931,1953]" captionTargetBox="[182,1424,162,1919]" captionTargetId="figure@31.[160,1427,153,1965]" captionTargetPageId="31" captionText="FIGURE 15. Hedgpethia filamentus sp. nov., holotype. A, trunk, dorsal view; B, trunk, lateral view; C, right oviger; D, right oviger claw; E, right palp; F, third leg; G, tarsus and propodus; H, abdomen, ventral view. Paratype, (JC066-4214). I, ocular tubercle." httpUri="https://zenodo.org/record/2598973/files/figure.png" pageId="30" pageNumber="431">Fig. 15C</figureCitation>
|
||
), segment 6 longest, slightly longer than next longest segment 4, segments 7 to10 lined with four or five rows of spatulate spines, distal-most spines taller than proximal spines, claw short, broad, less than half length of segment 10 (
|
||
<figureCitation id="131BF1A0290F4F19FDECF8B3FD62F8E7" box="[634,738,1793,1818]" captionStart="FIGURE 15" captionStartId="31.[151,250,1931,1953]" captionTargetBox="[182,1424,162,1919]" captionTargetId="figure@31.[160,1427,153,1965]" captionTargetPageId="31" captionText="FIGURE 15. Hedgpethia filamentus sp. nov., holotype. A, trunk, dorsal view; B, trunk, lateral view; C, right oviger; D, right oviger claw; E, right palp; F, third leg; G, tarsus and propodus; H, abdomen, ventral view. Paratype, (JC066-4214). I, ocular tubercle." httpUri="https://zenodo.org/record/2598973/files/figure.png" pageId="30" pageNumber="431">Fig. 15D</figureCitation>
|
||
), cutting edge curved, blade-like.
|
||
</paragraph>
|
||
<paragraph id="8B9FED25290F4F19FF51F897FB72F80F" blockId="30.[151,1437,1073,2034]" pageId="30" pageNumber="431">
|
||
Third leg (
|
||
<figureCitation id="131BF1A0290F4F19FED6F897FE22F8C3" box="[320,418,1829,1854]" captionStart="FIGURE 15" captionStartId="31.[151,250,1931,1953]" captionTargetBox="[182,1424,162,1919]" captionTargetId="figure@31.[160,1427,153,1965]" captionTargetPageId="31" captionText="FIGURE 15. Hedgpethia filamentus sp. nov., holotype. A, trunk, dorsal view; B, trunk, lateral view; C, right oviger; D, right oviger claw; E, right palp; F, third leg; G, tarsus and propodus; H, abdomen, ventral view. Paratype, (JC066-4214). I, ocular tubercle." httpUri="https://zenodo.org/record/2598973/files/figure.png" pageId="30" pageNumber="431">Fig. 15F</figureCitation>
|
||
) slender, femur slightly inflated in mid-region, maximum width twice that of either tibia, few scattered spines and longer dorsodistal spine; tibiae with numerous scattered spinules, tibia 1 longest segment; tibia 2 only slightly longer than femur, tibia 2 with long dorsodistal spine about 3 times width of segment, tarsus more than half as long as propodus (
|
||
<figureCitation id="131BF1A0290F4F19FDA7F823FD19F857" box="[561,665,1937,1962]" captionStart="FIGURE 15" captionStartId="31.[151,250,1931,1953]" captionTargetBox="[182,1424,162,1919]" captionTargetId="figure@31.[160,1427,153,1965]" captionTargetPageId="31" captionText="FIGURE 15. Hedgpethia filamentus sp. nov., holotype. A, trunk, dorsal view; B, trunk, lateral view; C, right oviger; D, right oviger claw; E, right palp; F, third leg; G, tarsus and propodus; H, abdomen, ventral view. Paratype, (JC066-4214). I, ocular tubercle." httpUri="https://zenodo.org/record/2598973/files/figure.png" pageId="30" pageNumber="431">Fig. 15G</figureCitation>
|
||
), all spines fine, undifferentiated, 18 to 20 spines on ventral surface, propodus almost straight, heel without major heel spines,18 to 25 sole spines; terminal claw short, little less than half length of propodus, auxiliary claws absent. Cement glands not evident. Gonopores not evident.
|
||
</paragraph>
|
||
<caption id="DF5FBDAD290E4F18FF01F839FF70F823" httpUri="https://zenodo.org/record/2598973/files/figure.png" pageId="31" pageNumber="432" startId="31.[151,250,1931,1953]" targetBox="[182,1424,162,1919]" targetPageId="31">
|
||
<paragraph id="8B9FED25290E4F18FF01F839FF70F823" blockId="31.[151,1436,1931,2014]" pageId="31" pageNumber="432">
|
||
<emphasis id="B9543137290E4F18FF01F839FEA0F85C" bold="true" box="[151,288,1931,1953]" pageId="31" pageNumber="432">FIGURE 15.</emphasis>
|
||
<taxonomicName id="4C2096A6290E4F18FEB1F839FD8EF85C" authority="Staples, 2019" authorityName="Staples" authorityYear="2019" box="[295,526,1931,1953]" class="Pycnogonida" family="Colossendeidae" genus="Hedgpethia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pantopoda" pageId="31" pageNumber="432" phylum="Arthropoda" rank="species" species="filamentus" status="sp. nov.">
|
||
<emphasis id="B9543137290E4F18FEB1F839FD8EF85C" box="[295,526,1931,1953]" italics="true" pageId="31" pageNumber="432">Hedgpethia filamentus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9543137290E4F18FD83F839FDE7F85D" bold="true" box="[533,615,1931,1953]" pageId="31" pageNumber="432">
|
||
<taxonomicNameLabel id="A2678C4C290E4F18FD83F839FDE7F85D" box="[533,615,1931,1953]" pageId="31" pageNumber="432" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, holotype. A, trunk, dorsal view; B, trunk, lateral view; C, right oviger; D, right oviger claw; E, right palp; F, third leg; G, tarsus and propodus; H, abdomen, ventral view. Paratype, (JC066-4214). I, ocular tubercle.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B9FED2529314F27FF51FF29FDB3FE73" blockId="32.[151,1437,151,866]" pageId="32" pageNumber="433">
|
||
Measurements of
|
||
<typeStatus id="549B538729314F27FE01FF28FE79FF4E" box="[407,505,154,179]" pageId="32" pageNumber="433" type="holotype">holotype</typeStatus>
|
||
(mm). Trunk length (frontal margin of cephalon to tip of 4
|
||
<superScript id="7C55406D29314F27FB2EFF25FB44FF58" attach="left" box="[1208,1220,151,165]" fontSize="6" pageId="32" pageNumber="433">th</superScript>
|
||
lateral processes), 1.384; length of cephalon, 0.424; width across 2
|
||
<superScript id="7C55406D29314F27FD4AFF0FFD6CFF36" attach="left" box="[732,748,189,203]" fontSize="6" pageId="32" pageNumber="433">nd</superScript>
|
||
lateral processes, 0.600; proboscis length (lateral), 1.616; greatest diameter proboscis, 0.360. Third leg: coxa 1, 0.184; coxa 2, 0.136; coxa 3, 0.136; femur,1.416; tibia1,1.736; tibia 2, 1.400; tarsus, 0.256; propodus, 0.440; claw, 0.192. Palp: seg. 1, 0.112; seg. 2, 0.048; seg. 3, 1.024; seg. 4, 0.153; seg. 5, 0.536, seg. 6, 0.104; seg. 7, 0.168; seg. 8, 0.160; seg. 9, 0.160; seg. 10, 0.160. Oviger: seg.1, 0.080; seg. 2, 0.120; seg. 3, 0.216; seg. 4, 0.680; seg. 5, 0.168; seg. 6, 0.688; seg. 7, 0.264; seg. 8, 0.108; seg. 9, 0.200; seg. 10, 0.136; claw, 0.072.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33ABEAE29314F27FF51FE2BFBDAFE4F" box="[199,1114,409,434]" pageId="32" pageNumber="433" type="etymology">
|
||
<paragraph id="8B9FED2529314F27FF51FE2BFBDAFE4F" blockId="32.[151,1437,151,866]" box="[199,1114,409,434]" pageId="32" pageNumber="433">
|
||
<emphasis id="B954313729314F27FF51FE2BFECEFE4F" bold="true" box="[199,334,409,434]" pageId="32" pageNumber="433">Etymology.</emphasis>
|
||
The species name alludes to the filamentous habitus of the specimen.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33ABEAE29314F27FF51FE0FFBC0FD77" pageId="32" pageNumber="433" type="discussion">
|
||
<paragraph id="8B9FED2529314F27FF51FE0FFBC0FD77" blockId="32.[151,1437,151,866]" pageId="32" pageNumber="433">
|
||
<emphasis id="B954313729314F27FF51FE0FFEBBFE2B" bold="true" box="[199,315,445,470]" pageId="32" pageNumber="433">Remarks.</emphasis>
|
||
The
|
||
<typeStatus id="549B538729314F27FEECFE0FFE5CFE2B" box="[378,476,445,470]" pageId="32" pageNumber="433" type="holotype">holotype</typeStatus>
|
||
was chosen because it is the more complete of the two larger specimens. There is some variability in the shape of the ocular tubercle amongst these six specimens with the ocular tubercle of one
|
||
<typeStatus id="549B538729314F27FF01FDB4FF77FDE3" box="[151,247,518,542]" pageId="32" pageNumber="433" type="paratype">paratype</typeStatus>
|
||
(JC066-4214) being particularly acute (
|
||
<figureCitation id="131BF1A029314F27FD5BFDB7FCAEFDE3" box="[717,814,517,542]" captionStart="FIGURE 15" captionStartId="31.[151,250,1931,1953]" captionTargetBox="[182,1424,162,1919]" captionTargetId="figure@31.[160,1427,153,1965]" captionTargetPageId="31" captionText="FIGURE 15. Hedgpethia filamentus sp. nov., holotype. A, trunk, dorsal view; B, trunk, lateral view; C, right oviger; D, right oviger claw; E, right palp; F, third leg; G, tarsus and propodus; H, abdomen, ventral view. Paratype, (JC066-4214). I, ocular tubercle." httpUri="https://zenodo.org/record/2598973/files/figure.png" pageId="32" pageNumber="433">
|
||
Fig.
|
||
<date id="FF9ECBE529314F27FC93FDB7FCAEFDE3" box="[773,814,517,542]" pageId="32" pageNumber="433" value="2015-01">15I</date>
|
||
</figureCitation>
|
||
). Eyes are only evident in the larger specimens. The slightly smaller specimens are of a similar size to each other and have leg spans of
|
||
<quantity id="4CD840C029314F27FBCCFD98FB5BFDBC" box="[1114,1243,554,578]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" metricValueMax="10.0" metricValueMin="8.0" pageId="32" pageNumber="433" unit="mm" value="9.0" valueMax="10.0" valueMin="8.0">8 to10 mm</quantity>
|
||
. Apart from size other morphological difference are not evident and the size difference may indicate that they are males rather than an indication of their maturity. Gonopores are not apparent in any of the specimens.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33ABEAE29314F27FF51FD27FDBEFC9F" pageId="32" pageNumber="433" type="reference_group">
|
||
<paragraph id="8B9FED2529314F27FF51FD27FDBEFC9F" blockId="32.[151,1437,151,866]" pageId="32" pageNumber="433">
|
||
<taxonomicName id="4C2096A629314F27FF51FD27FE45FD53" ID-CoL="1b0acde2-903f-4ac4-b61d-b717b4def07d" authorityName="Staples" authorityYear="2019" box="[199,453,661,686]" class="Pycnogonida" family="Colossendeidae" genus="Hedgpethia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pantopoda" pageId="32" pageNumber="433" phylum="Arthropoda" rank="species" species="filamentus">
|
||
<emphasis id="B954313729314F27FF51FD27FE45FD53" box="[199,453,661,686]" italics="true" pageId="32" pageNumber="433">Hedgpethia filamentus</emphasis>
|
||
</taxonomicName>
|
||
shares a similarly shaped ocular tubercle with
|
||
<taxonomicName id="4C2096A629314F27FC5AFD25FB4BFD53" ID-CoL="3JZ9L" authority="Arango, 2009" authorityName="Arango" authorityYear="2009" box="[972,1227,661,686]" class="Pycnogonida" family="Colossendeidae" genus="Hedgpethia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pantopoda" pageId="32" pageNumber="433" phylum="Arthropoda" rank="species" species="calva">
|
||
<emphasis id="B954313729314F27FC5AFD25FBABFD53" box="[972,1067,661,686]" italics="true" pageId="32" pageNumber="433">H. calva</emphasis>
|
||
<bibRefCitation id="EFB190D429314F27FBA7FD24FB4BFD53" author="Arango, C. P." box="[1073,1227,662,686]" pageId="32" pageNumber="433" pagination="1 - 20" refId="ref26306" refString="Arango, C. P. (2009) New species and new records of sea spiders (Arthropoda: Pycnogonida) from deep waters in Western Australia. Zootaxa, 1977, 1 - 20." type="journal article" year="2009">Arango, 2009</bibRefCitation>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C2096A629314F27FB69FD25FEA1FD2F" ID-CoL="3JZ9R" authority="Child, 1998" authorityName="Child" authorityYear="1998" class="Pycnogonida" family="Colossendeidae" genus="Hedgpethia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pantopoda" pageId="32" pageNumber="433" phylum="Arthropoda" rank="species" species="eleommata">
|
||
<emphasis id="B954313729314F27FB69FD25FA1CFD53" box="[1279,1436,661,686]" italics="true" pageId="32" pageNumber="433">H. eleommata</emphasis>
|
||
<bibRefCitation id="EFB190D429314F27FF01FD0BFEA1FD2F" author="Child, C. A." box="[151,289,697,722]" pageId="32" pageNumber="433" pagination="1 - 71" refId="ref27031" refString="Child, C. A. (1998) The Marine Fauna of New Zealand: Pycnogonida (Sea Spiders). NIWA Biodiversity Memoir, 109, 1 - 71." type="journal article" year="1998">Child, 1998</bibRefCitation>
|
||
</taxonomicName>
|
||
but both those species have vastly longer propodal claws and in neither species are the lateral processes so widely spaced as in this species.
|
||
<taxonomicName id="4C2096A629314F27FD3BFD6FFBC2FD0B" ID-CoL="3JZ9G" authority="(Stock, 1970)" authorityName="Stock" authorityYear="1970" baseAuthorityName="Stock" baseAuthorityYear="1970" box="[685,1090,733,758]" class="Pycnogonida" family="Colossendeidae" genus="Hedgpethia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pantopoda" pageId="32" pageNumber="433" phylum="Arthropoda" rank="species" species="atlantica">
|
||
<emphasis id="B954313729314F27FD3BFD6FFC1EFD0B" box="[685,926,733,758]" italics="true" pageId="32" pageNumber="433">Hedgpethia atlantica</emphasis>
|
||
(
|
||
<bibRefCitation id="EFB190D429314F27FC39FD6FFBB9FD0B" author="Stock, J. H." box="[943,1081,733,758]" pageId="32" pageNumber="433" pagination="6 - 10" refId="ref29263" refString="Stock, J. H. (1970) The Pycnogonida collected off northwestern Africa during the cruise of the " Meteor ". Meteor Forschungsergebnisse (reihe D-Biol), 5, 6 - 10." type="journal article" year="1970">Stock, 1970</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
also possesses an acuminated ocular tubercle and shares a similar proboscis shape with
|
||
<taxonomicName id="4C2096A629314F27FCA7FCB0FC4BFCE7" ID-CoL="1b0acde2-903f-4ac4-b61d-b717b4def07d" authorityName="Staples" authorityYear="2019" box="[817,971,769,794]" class="Pycnogonida" family="Colossendeidae" genus="Hedgpethia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pantopoda" pageId="32" pageNumber="433" phylum="Arthropoda" rank="species" species="filamentus">
|
||
<emphasis id="B954313729314F27FCA7FCB0FC4BFCE7" box="[817,971,769,794]" italics="true" pageId="32" pageNumber="433">H. filamentus</emphasis>
|
||
</taxonomicName>
|
||
but differs in the arrangement of oviger spines on segments 7 to10 which are in one or two rows as opposed to four or five rows in the new species and in the much longer, pointed oviger claw.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |