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<document id="8F24F09B5B0A25FD00C8B39A5D40014A" ID-CLB-Dataset="30810" ID-DOI="10.11646/zootaxa.4378.1.10" ID-GBIF-Dataset="ab9b5bbb-4f34-41c7-8465-672e93791aae" ID-ISSN="1175-5326" ID-Zenodo-Dep="1168162" ID-ZooBank="5AC144DD-84CA-40C8-BEBC-7697E92BE0E9" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1517998502777" checkinUser="plazi" docAuthor="Alvarez, Fernando &amp; Villalobos, José Luis" docDate="2018" docId="0383CC7AFF9C936DCB86FADCFE2CBD7E" docLanguage="en" docName="zootaxa.4378.1.10.pdf.imf" docOrigin="Zootaxa 4378 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Rodriguezia adani Alvarez &amp; Villalobos 2018, n. sp." docType="treatment" docVersion="8" lastPageNumber="141" masterDocId="FFBAB402FF9D9369CB11FC2FFFADBC3E" masterDocTitle="A new species of stygobitic freshwater crab of the genus Rodriguezia Bott, 1969 (Crustacea: Decapoda: Trichodactylidae) from Tabasco, Mexico" masterLastPageNumber="143" masterPageNumber="137" pageNumber="138" updateTime="1702000180320" updateUser="plazi">
<mods:mods id="7B743C58BF4B5A8ED1AEBDE32E47902B" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="3616C1F7830A783EDCB4246BF3D15BEE">
<mods:title id="5E0B769E26939716B53ADA840B1349B7">A new species of stygobitic freshwater crab of the genus Rodriguezia Bott, 1969 (Crustacea: Decapoda: Trichodactylidae) from Tabasco, Mexico</mods:title>
</mods:titleInfo>
<mods:name id="BBACC136988904E972FC56C4FCA8A094" type="personal">
<mods:role id="FE63D8CE98543FEE30F5B6E9245B6B6F">
<mods:roleTerm id="331D87DA4DBB652D30A1751A0034F326">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="CEFA6351DEB3A7FE7AA77A933683B82B">Alvarez, Fernando</mods:namePart>
</mods:name>
<mods:name id="5EEBB0DB23BA0E181F4B0C05EDC61D55" type="personal">
<mods:role id="8A3DAB769A38D9F36A458454C496C7B8">
<mods:roleTerm id="1B50924D5BB5C38376CB1267931191B0">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="2EE2F1B72FF75D8D91F9B7372259D17D">Villalobos, José Luis</mods:namePart>
</mods:name>
<mods:typeOfResource id="7FC2E280C747CC13C2E3915C32209E16">text</mods:typeOfResource>
<mods:relatedItem id="01D09002C9673E9EA0F61BB4775FAF09" type="host">
<mods:titleInfo id="A48831ADAD0F77685A7164F0BAB76E9B">
<mods:title id="22A2C9023AEC06A56E296E21E006D77E">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="396361233675C76E24DCCE8F6C37C1C3">
<mods:date id="7FBB4172E61B9A1247B7736F116B54E9">2018</mods:date>
<mods:detail id="5A4C80F71BA24DCFEAE7D247E4E46F2C" type="pubDate">
<mods:number id="36EBAC6CECAA2A2C64532D212559F476">2018-02-07</mods:number>
</mods:detail>
<mods:detail id="4F47BD3B82B28E8CAFCD97C0E3AE2193" type="volume">
<mods:number id="AFAF73C5491EAC9B936745917321328B">4378</mods:number>
</mods:detail>
<mods:detail id="244643BA471378BACFB248034169E542" type="issue">
<mods:number id="FBFD6566A221899289104D759A6DA3C7">1</mods:number>
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<mods:start id="4CA8AEABED9771AA62F0A3A12507DBE9">137</mods:start>
<mods:end id="91C3A50C4DDB547C919DB7CDC1DC26B7">143</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="865AAECA38608C4F42D2201E648F1390">journal article</mods:classification>
<mods:identifier id="820382C0EB0EC42E0924A2D106EE9825" type="CLB-Dataset">30810</mods:identifier>
<mods:identifier id="4421693627D26FDE1C22DB70712C0DEC" type="DOI">10.11646/zootaxa.4378.1.10</mods:identifier>
<mods:identifier id="6A27577BD357F5FD78D90479363E7DC2" type="GBIF-Dataset">ab9b5bbb-4f34-41c7-8465-672e93791aae</mods:identifier>
<mods:identifier id="D489A10E09E29464113BAB316E2AFF4B" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="E906FF81D8A4304EEAAFAFDAB722B3AE" type="Zenodo-Dep">1168162</mods:identifier>
<mods:identifier id="FE56E68B890931FFFD882CEA7993758B" type="ZooBank">5AC144DD-84CA-40C8-BEBC-7697E92BE0E9</mods:identifier>
</mods:mods>
<treatment id="0383CC7AFF9C936DCB86FADCFE2CBD7E" ID-DOI="http://doi.org/10.5281/zenodo.5981646" ID-GBIF-Taxon="142183824" ID-Zenodo-Dep="5981646" LSID="urn:lsid:plazi:treatment:0383CC7AFF9C936DCB86FADCFE2CBD7E" httpUri="http://treatment.plazi.org/id/0383CC7AFF9C936DCB86FADCFE2CBD7E" lastPageId="4" lastPageNumber="141" pageId="1" pageNumber="138">
<subSubSection id="C3302EE7FF9C9368CB86FADCFEBFBB11" pageId="1" pageNumber="138" type="nomenclature">
<paragraph id="8B957D6CFF9C9368CB86FADCFE13BB33" blockId="1.[151,446,1779,1839]" box="[151,446,1779,1805]" pageId="1" pageNumber="138">
<heading id="D0DDCA00FF9C9368CB86FADCFE13BB33" bold="true" box="[151,446,1779,1805]" fontSize="11" level="1" pageId="1" pageNumber="138" reason="1">
<emphasis id="B95EA17EFF9C9368CB86FADCFE13BB33" bold="true" box="[151,446,1779,1805]" pageId="1" pageNumber="138">
<taxonomicName id="4C2A06EFFF9C9368CB86FADCFEDBBB33" ID-CoL="4T8VK" authority="Alvarez &amp; Villalobos, 2018" authorityName="Alvarez &amp; Villalobos" authorityYear="2018" box="[151,374,1779,1805]" class="Malacostraca" family="Trichodactylidae" genus="Rodriguezia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="138" phylum="Arthropoda" rank="species" species="adani" status="sp. nov.">
<emphasis id="B95EA17EFF9C9368CB86FADCFEDBBB33" bold="true" box="[151,374,1779,1805]" italics="true" pageId="1" pageNumber="138">Rodriguezia adani</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26D1C05FF9C9368CA6DFADCFE13BB33" box="[380,446,1779,1805]" pageId="1" pageNumber="138" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B957D6CFF9C9368CB86FB39FEBFBB11" blockId="1.[151,446,1779,1839]" box="[151,274,1814,1839]" pageId="1" pageNumber="138">
(
<figureCitation id="131161E9FF9C9368CB8EFB39FEA7BB11" box="[159,266,1814,1839]" captionStart-0="FIGURE 2" captionStart-1="FIGURE3" captionStart-2="FIGURE 4" captionStartId-0="3.[151,250,1993,2015]" captionStartId-1="4.[151,250,1978,2000]" captionStartId-2="5.[151,250,1315,1337]" captionTargetBox-0="[177,1419,333,1977]" captionTargetBox-1="[209,1398,386,1944]" captionTargetBox-2="[191,1392,246,1243]" captionTargetId-0="figure@3.[163,1427,293,1985]" captionTargetId-1="figure@4.[181,1405,368,1959]" captionTargetId-2="figure@5.[151,1436,210,1283]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionText-0="FIGURE 2. Rodriguezia adani n. sp. male holotype: A, B, dorsal view.Scale bar = 5 mm." captionText-1="FIGURE3. Rodrigueziaadani n. sp.: A,B, frontalview of carapace,male holotype; C, maleabdomen; D, femaleabdomen; E, antennule;F, antenna; G, third maxilliped; H, chelae.Scale bars: A, B, H = 5 mm; C, D = 3 mm; E, F = 1 mm; G = 2 mm." captionText-2="FIGURE 4. Rodriguezia adani n. sp. first gonopod: A, E, cephalothoracic view; B, F, lateral view; C, G, caudal view; D, H, mesial view." httpUri-0="https://zenodo.org/record/1168166/files/figure.png" httpUri-1="https://zenodo.org/record/1168168/files/figure.png" httpUri-2="https://zenodo.org/record/1168170/files/figure.png" pageId="1" pageNumber="138">Figs. 2–4</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3302EE7FF9C936BCB86FB71FCDEBCC6" lastPageId="2" lastPageNumber="139" pageId="1" pageNumber="138" type="materials_examined">
<paragraph id="8B957D6CFF9C9368CB86FB71FDF3BBDC" blockId="1.[151,1437,1886,2019]" pageId="1" pageNumber="138">
<emphasis id="B95EA17EFF9C9368CB86FB71FEE8BB49" bold="true" box="[151,325,1886,1911]" pageId="1" pageNumber="138">Type material.</emphasis>
<materialsCitation id="3B427731FF9C9368CA5AFB71FA30BBA4" ID-GBIF-Occurrence="1831402091" collectingDate="2016-07-16" collectionCode="CNCR" collectorName="A. Gómez &amp; Moreno" country="Mexico" elevation="133" latitude="17.621222" location="Grutas de Agua Blanca" longLatPrecision="1" longitude="-92.472664" municipality="Macuspana" pageId="1" pageNumber="138" specimenCode="CNCR 33912" specimenCount="1" stateProvince="Tabasco" typeStatus="holotype">
Male
<typeStatus id="5491C3CEFF9C9368CA9DFB71FE5CBB49" box="[396,497,1886,1911]" pageId="1" pageNumber="138" type="holotype">holotype</typeStatus>
, cl
<quantity id="4CD2D089FF9C9368C906FB71FDC2BB48" box="[535,623,1886,1911]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" pageId="1" pageNumber="138" unit="mm" value="9.5">9.5 mm</quantity>
, cb
<quantity id="4CD2D089FF9C9368C98AFB71FD59BB48" box="[667,756,1886,1911]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" pageId="1" pageNumber="138" unit="mm" value="9.5">9.5 mm</quantity>
;
<location id="8EF52BB7FF9C9368C811FB70FBA9BB49" LSID="urn:lsid:plazi:treatment:0383CC7AFF9C936DCB86FADCFE2CBD7E:8EF52BB7FF9C9368C811FB70FBA9BB49" box="[768,1028,1886,1911]" country="Mexico" latitude="17.621222" longLatPrecision="1" longitude="-92.472664" municipality="Macuspana" name="Grutas de Agua Blanca" pageId="1" pageNumber="138" stateProvince="Tabasco">Grutas de Agua Blanca</location>
(
<geoCoordinate id="EE1E1BABFF9C9368CF02FB71FB19BB49" box="[1043,1204,1886,1911]" direction="north" orientation="latitude" pageId="1" pageNumber="138" precision="1" value="17.621222">17°37’16.4”N</geoCoordinate>
,
<geoCoordinate id="EE1E1BABFF9C9368CFD1FB71FACBBB49" box="[1216,1382,1886,1911]" direction="west" orientation="longitude" pageId="1" pageNumber="138" precision="1" value="-92.472664">92°28’21.6”W</geoCoordinate>
;
<elevation id="00079A5FFF9C9368CE63FB70FF7BBBA5" metricMagnitude="2" metricUnit="m" metricValue="1.33" pageId="1" pageNumber="138" unit="m" value="133.0">
<quantity id="4CD2D089FF9C9368CE63FB70FF00BBA4" metricMagnitude="2" metricUnit="m" metricValue="1.33" pageId="1" pageNumber="138" unit="m" value="133.0">133 m</quantity>
asl
</elevation>
), municipality of
<collectingMunicipality id="6BF1E716FF9C9368CAB0FBACFD8EBBA4" box="[417,547,1923,1946]" pageId="1" pageNumber="138">Macuspana</collectingMunicipality>
,
<collectingRegion id="49EEB38EFF9C9368C921FBADFD23BBA5" box="[560,654,1922,1947]" country="Mexico" name="Tabasco" pageId="1" pageNumber="138">Tabasco</collectingRegion>
,
<collectingCountry id="F33D3DFCFF9C9368C98BFBADFD5EBBA5" box="[666,755,1922,1947]" name="Mexico" pageId="1" pageNumber="138">Mexico</collectingCountry>
,
<date id="FF945BACFF9C9368C9EEFBADFC39BBA5" box="[767,916,1922,1947]" pageId="1" pageNumber="138" value="2016-07-16">
<collectingDate id="EFD0A244FF9C9368C9EEFBADFC39BBA5" box="[767,916,1922,1947]" pageId="1" pageNumber="138" value="2016-07-16">16 July 2016</collectingDate>
</date>
, coll.
<collectorName id="26DF18BAFF9C9368C8C8FBADFBE0BBA4" box="[985,1101,1922,1946]" pageId="1" pageNumber="138">A. Gómez</collectorName>
&amp; E.
<collectorName id="26DF18BAFF9C9368CF82FBACFB42BBA4" box="[1171,1263,1923,1946]" pageId="1" pageNumber="138">Moreno</collectorName>
,
<specimenCode id="DB8CD517FF9C9368CFEAFBADFA35BBA4" box="[1275,1432,1922,1946]" collectionCode="CNCR" pageId="1" pageNumber="138">CNCR 33912</specimenCode>
.
</materialsCitation>
<materialsCitation id="3B427731FF9C9368CB86FB89FDF3BBDC" ID-GBIF-Occurrence="1831402183" collectingDate="2016-07-16" collectionCode="CNCR" collectorName="A. Gomez &amp; Moreno &amp; Female" country="Mexico" elevation="133" latitude="17.621222" location="Grutas de Agua Blanca" longLatPrecision="1" longitude="-92.472664" municipality="Macuspana" pageId="1" pageNumber="138" specimenCode="CNCR 33912" specimenCount="1" stateProvince="Tabasco" typeStatus="allotype">
<collectorName id="26DF18BAFF9C9368CB86FB89FF47BB81" box="[151,234,1958,1983]" pageId="1" pageNumber="138">Female</collectorName>
<typeStatus id="5491C3CEFF9C9368CBE1FB89FEE7BB81" box="[240,330,1958,1983]" pageId="1" pageNumber="138" type="allotype">allotype</typeStatus>
(carrying 4 crabs in abdomen, measuring cb
<quantity id="4CD2D089FF9C9368C854FB88FC31BB80" box="[837,924,1959,1983]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="1" pageNumber="138" unit="mm" value="1.7">1.7 mm</quantity>
), cl
<quantity id="4CD2D089FF9C9368C8C4FB88FB80BB80" box="[981,1069,1959,1983]" metricMagnitude="-3" metricUnit="m" metricValue="9.9" pageId="1" pageNumber="138" unit="mm" value="9.9">9.9 mm</quantity>
, cb
<quantity id="4CD2D089FF9C9368CF4BFB88FB6CBB80" box="[1114,1217,1959,1983]" metricMagnitude="-2" metricUnit="m" metricValue="1.04" pageId="1" pageNumber="138" unit="mm" value="10.4">10.4 mm</quantity>
; same locality, date and collector as
<typeStatus id="5491C3CEFF9C9368CA5FFBE5FE1EBBDD" box="[334,435,1994,2019]" pageId="1" pageNumber="138" type="holotype">holotype</typeStatus>
,
<specimenCode id="DB8CD517FF9C9368CAAFFBE5FDF6BBDC" box="[446,603,1994,2018]" collectionCode="CNCR" pageId="1" pageNumber="138">CNCR 33913</specimenCode>
.
</materialsCitation>
</paragraph>
<paragraph id="8B957D6CFF9F936BCBD6FCB7FCDEBCC6" blockId="2.[151,1437,151,2013]" pageId="2" pageNumber="139">
<materialsCitation id="3B427731FF9F936BCBD6FCB7FCADBC91" ID-GBIF-Occurrence="1831402184" box="[199,768,151,176]" pageId="2" pageNumber="139">
Paratypes. 2 males, cl 5.5,
<quantity id="4CD2D089FF9F936BCAFEFCB8FDEABC91" box="[495,583,151,176]" metricMagnitude="-3" metricUnit="m" metricValue="6.4" pageId="2" pageNumber="139" unit="mm" value="6.4">6.4 mm</quantity>
, cb 5.3,
<quantity id="4CD2D089FF9F936BC9B2FCB8FD51BC91" box="[675,764,151,176]" metricMagnitude="-3" metricUnit="m" metricValue="6.7" pageId="2" pageNumber="139" unit="mm" value="6.7">6.7 mm</quantity>
;
</materialsCitation>
<materialsCitation id="3B427731FF9F936BC816FCB7FCD8BCEA" ID-GBIF-Occurrence="1831402092" collectionCode="CNCR" pageId="2" pageNumber="139" specimenCode="CNCR 33914" typeStatus="holotype">
4 females, cl 4.6, 5.0, 6.2,
<quantity id="4CD2D089FF9F936BCF3AFCB7FB2EBC91" box="[1067,1155,152,176]" metricMagnitude="-3" metricUnit="m" metricValue="7.9" pageId="2" pageNumber="139" unit="mm" value="7.9">7.9 mm</quantity>
, cb 4.8, 5.2, 6.5,
<quantity id="4CD2D089FF9F936BCE51FCB8FA35BC91" box="[1344,1432,151,176]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" pageId="2" pageNumber="139" unit="mm" value="8.5">8.5 mm</quantity>
, same locality, date and collectors as for
<typeStatus id="5491C3CEFF9F936BC970FC93FD6BBCEB" box="[609,710,188,213]" pageId="2" pageNumber="139" type="holotype">holotype</typeStatus>
,
<specimenCode id="DB8CD517FF9F936BC9C2FC93FCDDBCEA" box="[723,880,188,212]" collectionCode="CNCR" pageId="2" pageNumber="139">CNCR 33914</specimenCode>
.
</materialsCitation>
<materialsCitation id="3B427731FF9F936BC86CFC93FCDEBCC6" ID-GBIF-Occurrence="1831402093" collectingDate="2015-07" collectionCode="CNCR" collectorName="A. Gomez &amp; C.N. Rico" pageId="2" pageNumber="139" specimenCode="CNCR 33915" specimenCount="1" specimenCount-male="1">
<specimenCount id="9D2CB6E5FF9F936BC86CFC93FC61BCEB" box="[893,972,188,213]" pageId="2" pageNumber="139" type="male">1 male</specimenCount>
, cl
<quantity id="4CD2D089FF9F936BC8E7FC93FBE2BCEA" box="[1014,1103,188,212]" metricMagnitude="-3" metricUnit="m" metricValue="8.7" pageId="2" pageNumber="139" unit="mm" value="8.7">8.7 mm</quantity>
, cb
<quantity id="4CD2D089FF9F936BCF91FC93FB74BCEA" box="[1152,1241,188,212]" metricMagnitude="-3" metricUnit="m" metricValue="9.1" pageId="2" pageNumber="139" unit="mm" value="9.1">9.1 mm</quantity>
, same locality as holotype;
<date id="FF945BACFF9F936BCA19FCF0FED4BCC6" box="[264,377,223,248]" pageId="2" pageNumber="139" value="2015-07">
<collectingDate id="EFD0A244FF9F936BCA19FCF0FED4BCC6" box="[264,377,223,248]" pageId="2" pageNumber="139" value="2015-07">July 2015</collectingDate>
</date>
; coll.
<collectorName id="26DF18BAFF9F936BCAADFCCFFD82BCC6" box="[444,559,224,248]" pageId="2" pageNumber="139">A. Gómez</collectorName>
&amp;
<collectorName id="26DF18BAFF9F936BC942FCCFFD6ABCC6" box="[595,711,223,248]" pageId="2" pageNumber="139">C.N. Rico</collectorName>
,
<specimenCode id="DB8CD517FF9F936BC9C3FCCFFCC2BCC6" box="[722,879,223,248]" collectionCode="CNCR" pageId="2" pageNumber="139">CNCR 33915</specimenCode>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3302EE7FF9F936BCBD6FD2BFDD6B866" pageId="2" pageNumber="139" type="description">
<paragraph id="8B957D6CFF9F936BCBD6FD2BFC6BBDCA" blockId="2.[151,1437,151,2013]" pageId="2" pageNumber="139">
<emphasis id="B95EA17EFF9F936BCBD6FD2BFEF4BD23" bold="true" box="[199,345,260,285]" pageId="2" pageNumber="139">Description.</emphasis>
Dorsal surface of carapace smooth; in dorsal view suborbicular, slightly wider than long; branchial, gastric, cardiac regions faintly marked; lateral margin complete, devoid of teeth; posterior margin slightly arched, almost straight; posterolateral and anterolateral angles rounded; anterior margin bilobed, central portion slightly depressed, laterally with small rounded projections that limit orbits (
<figureCitation id="131161E9FF9F936BCF4BFD40FB7ABDB6" box="[1114,1239,367,392]" captionStart="FIGURE 2" captionStartId="3.[151,250,1993,2015]" captionTargetBox="[177,1419,333,1977]" captionTargetId="figure@3.[163,1427,293,1985]" captionTargetPageId="3" captionText="FIGURE 2. Rodriguezia adani n. sp. male holotype: A, B, dorsal view.Scale bar = 5 mm." httpUri="https://zenodo.org/record/1168166/files/figure.png" pageId="2" pageNumber="139">Fig. 2A, B</figureCitation>
). In frontal view, superior border of carapace formed by minute tubercles; median portion elevated; laterally, portion between insertion of antennule and antenna slightly depressed, portion around orbits elevated. Internal orbital angle weakly marked, occlusive tooth rounded, buccal angle oval-shaped (
<figureCitation id="131161E9FF9F936BC82FFDF3FC15BDCA" box="[830,952,476,501]" captionStart="FIGURE3" captionStartId="4.[151,250,1978,2000]" captionTargetBox="[209,1398,386,1944]" captionTargetId="figure@4.[181,1405,368,1959]" captionTargetPageId="4" captionText="FIGURE3. Rodrigueziaadani n. sp.: A,B, frontalview of carapace,male holotype; C, maleabdomen; D, femaleabdomen; E, antennule;F, antenna; G, third maxilliped; H, chelae.Scale bars: A, B, H = 5 mm; C, D = 3 mm; E, F = 1 mm; G = 2 mm." httpUri="https://zenodo.org/record/1168168/files/figure.png" pageId="2" pageNumber="139">Fig. 3A, B</figureCitation>
).
</paragraph>
<paragraph id="8B957D6CFF9F936BCBD6FDD0FCA2BE5E" blockId="2.[151,1437,151,2013]" pageId="2" pageNumber="139">
Abdomen of male and female with all segments distinct. Male abdomen triangular, last segment with posterior margin rounded, twice as wide as long (
<figureCitation id="131161E9FF9F936BC976FE0BFD6FBE02" box="[615,706,548,573]" captionStart="FIGURE3" captionStartId="4.[151,250,1978,2000]" captionTargetBox="[209,1398,386,1944]" captionTargetId="figure@4.[181,1405,368,1959]" captionTargetPageId="4" captionText="FIGURE3. Rodrigueziaadani n. sp.: A,B, frontalview of carapace,male holotype; C, maleabdomen; D, femaleabdomen; E, antennule;F, antenna; G, third maxilliped; H, chelae.Scale bars: A, B, H = 5 mm; C, D = 3 mm; E, F = 1 mm; G = 2 mm." httpUri="https://zenodo.org/record/1168168/files/figure.png" pageId="2" pageNumber="139">Fig. 3C</figureCitation>
). Female abdomen broadly rounded, partially covering thoracic sternites, last segment 4 times as wide as long (
<figureCitation id="131161E9FF9F936BC9B9FE68FCAFBE5E" box="[680,770,583,608]" captionStart="FIGURE3" captionStartId="4.[151,250,1978,2000]" captionTargetBox="[209,1398,386,1944]" captionTargetId="figure@4.[181,1405,368,1959]" captionTargetPageId="4" captionText="FIGURE3. Rodrigueziaadani n. sp.: A,B, frontalview of carapace,male holotype; C, maleabdomen; D, femaleabdomen; E, antennule;F, antenna; G, third maxilliped; H, chelae.Scale bars: A, B, H = 5 mm; C, D = 3 mm; E, F = 1 mm; G = 2 mm." httpUri="https://zenodo.org/record/1168168/files/figure.png" pageId="2" pageNumber="139">Fig. 3D</figureCitation>
).
</paragraph>
<paragraph id="8B957D6CFF9F936BCBD6FE43FE30BECE" blockId="2.[151,1437,151,2013]" pageId="2" pageNumber="139">
Antennule and antenna, typical of genus (
<figureCitation id="131161E9FF9F936BC9A2FE43FC9CBEBA" box="[691,817,620,645]" captionStart="FIGURE3" captionStartId="4.[151,250,1978,2000]" captionTargetBox="[209,1398,386,1944]" captionTargetId="figure@4.[181,1405,368,1959]" captionTargetPageId="4" captionText="FIGURE3. Rodrigueziaadani n. sp.: A,B, frontalview of carapace,male holotype; C, maleabdomen; D, femaleabdomen; E, antennule;F, antenna; G, third maxilliped; H, chelae.Scale bars: A, B, H = 5 mm; C, D = 3 mm; E, F = 1 mm; G = 2 mm." httpUri="https://zenodo.org/record/1168168/files/figure.png" pageId="2" pageNumber="139">Fig. 3E, F</figureCitation>
). Eyes absent, ocular peduncle strongly reduced, not movable, appearing as blunt plug occupying most of orbit (
<figureCitation id="131161E9FF9F936BC86DFEA0FBAABE96" box="[892,1031,655,680]" captionStart="FIGURE3" captionStartId="4.[151,250,1978,2000]" captionTargetBox="[209,1398,386,1944]" captionTargetId="figure@4.[181,1405,368,1959]" captionTargetPageId="4" captionText="FIGURE3. Rodrigueziaadani n. sp.: A,B, frontalview of carapace,male holotype; C, maleabdomen; D, femaleabdomen; E, antennule;F, antenna; G, third maxilliped; H, chelae.Scale bars: A, B, H = 5 mm; C, D = 3 mm; E, F = 1 mm; G = 2 mm." httpUri="https://zenodo.org/record/1168168/files/figure.png" pageId="2" pageNumber="139">Fig. 3A, B</figureCitation>
). Third maxilliped approximately rectangular, ischium slightly wider than merus; exopod shorter than ischium and merus combined, flagellum with eight articles (
<figureCitation id="131161E9FF9F936BCA27FEF8FE3DBECE" box="[310,400,727,752]" captionStart="FIGURE3" captionStartId="4.[151,250,1978,2000]" captionTargetBox="[209,1398,386,1944]" captionTargetId="figure@4.[181,1405,368,1959]" captionTargetPageId="4" captionText="FIGURE3. Rodrigueziaadani n. sp.: A,B, frontalview of carapace,male holotype; C, maleabdomen; D, femaleabdomen; E, antennule;F, antenna; G, third maxilliped; H, chelae.Scale bars: A, B, H = 5 mm; C, D = 3 mm; E, F = 1 mm; G = 2 mm." httpUri="https://zenodo.org/record/1168168/files/figure.png" pageId="2" pageNumber="139">Fig. 3G</figureCitation>
).
</paragraph>
<paragraph id="8B957D6CFF9F936BCBD6FED3FB30BFBE" blockId="2.[151,1437,151,2013]" pageId="2" pageNumber="139">
Chelipeds subequal in size, carpi with tooth on internal dorsal margin, chelae with dorsal tooth on proximal third, fingers not gaping. Major chela with swollen palm, twice as long as high; fingers about one third length of chela, cutting edges with low irregular teeth, tips acute; in lateral view, movable finger approximately triangular (
<figureCitation id="131161E9FF9F936BCB8EFF48FF54BFBE" box="[159,249,871,896]" captionStart="FIGURE3" captionStartId="4.[151,250,1978,2000]" captionTargetBox="[209,1398,386,1944]" captionTargetId="figure@4.[181,1405,368,1959]" captionTargetPageId="4" captionText="FIGURE3. Rodrigueziaadani n. sp.: A,B, frontalview of carapace,male holotype; C, maleabdomen; D, femaleabdomen; E, antennule;F, antenna; G, third maxilliped; H, chelae.Scale bars: A, B, H = 5 mm; C, D = 3 mm; E, F = 1 mm; G = 2 mm." httpUri="https://zenodo.org/record/1168168/files/figure.png" pageId="2" pageNumber="139">Fig. 3H</figureCitation>
). Minor chela 2.4 times as long as high, fingers about half length of chela (
<figureCitation id="131161E9FF9F936BCF50FF48FB22BFBE" box="[1089,1167,871,896]" captionStart="FIGURE3" captionStartId="4.[151,250,1978,2000]" captionTargetBox="[209,1398,386,1944]" captionTargetId="figure@4.[181,1405,368,1959]" captionTargetPageId="4" captionText="FIGURE3. Rodrigueziaadani n. sp.: A,B, frontalview of carapace,male holotype; C, maleabdomen; D, femaleabdomen; E, antennule;F, antenna; G, third maxilliped; H, chelae.Scale bars: A, B, H = 5 mm; C, D = 3 mm; E, F = 1 mm; G = 2 mm." httpUri="https://zenodo.org/record/1168168/files/figure.png" pageId="2" pageNumber="139">
Fig.
<date id="FF945BACFF9F936BCF67FF47FB22BFBE" box="[1142,1167,872,896]" pageId="2" pageNumber="139">3I</date>
</figureCitation>
).
</paragraph>
<paragraph id="8B957D6CFF9F936BCBD6FFA3FD55BFF6" blockId="2.[151,1437,151,2013]" pageId="2" pageNumber="139">
Pereiopods 2–5, elongate, representing 1.84, 2.26, 2.48, 2.21 times carapace breadth (
<figureCitation id="131161E9FF9F936BCF8EFFA3FB51BF9A" box="[1183,1276,908,933]" captionStart="FIGURE 2" captionStartId="3.[151,250,1993,2015]" captionTargetBox="[177,1419,333,1977]" captionTargetId="figure@3.[163,1427,293,1985]" captionTargetPageId="3" captionText="FIGURE 2. Rodriguezia adani n. sp. male holotype: A, B, dorsal view.Scale bar = 5 mm." httpUri="https://zenodo.org/record/1168166/files/figure.png" pageId="2" pageNumber="139">Fig. 2A</figureCitation>
). Propodi and dactyli with fine pubescence, more dense on the latter.
</paragraph>
<paragraph id="8B957D6CFF9F936BCBD6FFFBFDD6B866" blockId="2.[151,1437,151,2013]" pageId="2" pageNumber="139">
First gonopod straight, slender, proximal third wider (
<figureCitation id="131161E9FF9F936BC80FFFFBFC3BBFD2" box="[798,918,980,1005]" captionStart="FIGURE 4" captionStartId="5.[151,250,1315,1337]" captionTargetBox="[191,1392,246,1243]" captionTargetId="figure@5.[151,1436,210,1283]" captionTargetPageId="5" captionText="FIGURE 4. Rodriguezia adani n. sp. first gonopod: A, E, cephalothoracic view; B, F, lateral view; C, G, caudal view; D, H, mesial view." httpUri="https://zenodo.org/record/1168170/files/figure.png" pageId="2" pageNumber="139">Fig. 4A, E</figureCitation>
). Middle constriction, typical of genus, faintly visible in lateral and mesial views (
<figureCitation id="131161E9FF9F936BC934FFD8FD30B82E" box="[549,669,1015,1040]" captionStart="FIGURE 4" captionStartId="5.[151,250,1315,1337]" captionTargetBox="[191,1392,246,1243]" captionTargetId="figure@5.[151,1436,210,1283]" captionTargetPageId="5" captionText="FIGURE 4. Rodriguezia adani n. sp. first gonopod: A, E, cephalothoracic view; B, F, lateral view; C, G, caudal view; D, H, mesial view." httpUri="https://zenodo.org/record/1168170/files/figure.png" pageId="2" pageNumber="139">Fig. 4C, E</figureCitation>
). Field of spines covering distal third except for lateral surface (
<figureCitation id="131161E9FF9F936BCE7EFFD8FF16B80A" captionStart="FIGURE 4" captionStartId="5.[151,250,1315,1337]" captionTargetBox="[191,1392,246,1243]" captionTargetId="figure@5.[151,1436,210,1283]" captionTargetPageId="5" captionText="FIGURE 4. Rodriguezia adani n. sp. first gonopod: A, E, cephalothoracic view; B, F, lateral view; C, G, caudal view; D, H, mesial view." httpUri="https://zenodo.org/record/1168170/files/figure.png" pageId="2" pageNumber="139">Fig. 4C</figureCitation>
); all spines similar in size and shape. Apical pore suboval, drop-shaped. Second pleopod less than half length of first, distal portion slender, acute (
<figureCitation id="131161E9FF9F936BC907F810FDC3B866" box="[534,622,1087,1112]" captionStart="FIGURE 4" captionStartId="5.[151,250,1315,1337]" captionTargetBox="[191,1392,246,1243]" captionTargetId="figure@5.[151,1436,210,1283]" captionTargetPageId="5" captionText="FIGURE 4. Rodriguezia adani n. sp. first gonopod: A, E, cephalothoracic view; B, F, lateral view; C, G, caudal view; D, H, mesial view." httpUri="https://zenodo.org/record/1168170/files/figure.png" pageId="2" pageNumber="139">Fig. 4B</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3302EE7FF9F936BCBD6F84BFADCB843" box="[199,1393,1124,1149]" pageId="2" pageNumber="139" type="distribution">
<paragraph id="8B957D6CFF9F936BCBD6F84BFADCB843" blockId="2.[151,1437,151,2013]" box="[199,1393,1124,1149]" pageId="2" pageNumber="139">
<emphasis id="B95EA17EFF9F936BCBD6F84BFECDB843" bold="true" box="[199,352,1124,1149]" pageId="2" pageNumber="139">Distribution.</emphasis>
The new species is only known from Grutas de Agua Blanca, Macuspana,
<collectingRegion id="49EEB38EFF9F936BCFBBF84BFAA4B843" box="[1194,1289,1124,1149]" country="Mexico" name="Tabasco" pageId="2" pageNumber="139">Tabasco</collectingRegion>
,
<collectingCountry id="F33D3DFCFF9F936BCE05F84BFAC0B843" box="[1300,1389,1124,1149]" name="Mexico" pageId="2" pageNumber="139">
<collectingRegion id="49EEB38EFF9F936BCE05F84BFAC0B843" box="[1300,1389,1124,1149]" country="Mexico" name="Mexico" pageId="2" pageNumber="139">Mexico</collectingRegion>
</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C3302EE7FF9F936BCBD6F8A8FC27B8FB" pageId="2" pageNumber="139" type="etymology">
<paragraph id="8B957D6CFF9F936BCBD6F8A8FC27B8FB" blockId="2.[151,1437,151,2013]" pageId="2" pageNumber="139">
<emphasis id="B95EA17EFF9F936BCBD6F8A8FEE3B89E" bold="true" box="[199,334,1159,1184]" pageId="2" pageNumber="139">Etymology.</emphasis>
We name the new species after Adán Gómez-González, explorer, biologist and friend, who found these crabs while exploring caves in
<collectingRegion id="49EEB38EFF9F936BC922F883FD23B8FB" box="[563,654,1196,1221]" country="Mexico" name="Tabasco" pageId="2" pageNumber="139">Tabasco</collectingRegion>
and
<collectingRegion id="49EEB38EFF9F936BC9D4F883FC8FB8FB" box="[709,802,1196,1221]" country="Mexico" name="Chiapas" pageId="2" pageNumber="139">Chiapas</collectingRegion>
,
<collectingCountry id="F33D3DFCFF9F936BC83CF883FC2BB8FB" box="[813,902,1196,1221]" name="Mexico" pageId="2" pageNumber="139">
<collectingRegion id="49EEB38EFF9F936BC83CF883FC2BB8FB" box="[813,902,1196,1221]" country="Mexico" name="Mexico" pageId="2" pageNumber="139">Mexico</collectingRegion>
</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C3302EE7FF9F936DCBD6F8E0FE2CBD7E" lastPageId="4" lastPageNumber="141" pageId="2" pageNumber="139" type="discussion">
<paragraph id="8B957D6CFF9F936BCBD6F8E0FD12B9DB" blockId="2.[151,1437,151,2013]" pageId="2" pageNumber="139">
<emphasis id="B95EA17EFF9F936BCBD6F8E0FE96B8D6" bold="true" box="[199,315,1231,1256]" pageId="2" pageNumber="139">Remarks.</emphasis>
<taxonomicName id="4C2A06EFFF9F936BCA57F8FFFDB6B8D6" authority="Alvarez &amp; Villalobos, 2018" authorityName="Alvarez &amp; Villalobos" authorityYear="2018" box="[326,539,1232,1256]" class="Malacostraca" family="Trichodactylidae" genus="Rodriguezia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="139" phylum="Arthropoda" rank="species" species="adani" status="n. sp. and R.">
<emphasis id="B95EA17EFF9F936BCA57F8FFFDB6B8D6" box="[326,539,1232,1256]" italics="true" pageId="2" pageNumber="139">Rodriguezia adani</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26D1C05FF9F936BC937F8E0FD6FB8D6" box="[550,706,1231,1256]" pageId="2" pageNumber="139" rank="subSpecies">
<emphasis id="B95EA17EFF9F936BC937F8E0FDC7B8D6" bold="true" box="[550,618,1231,1256]" pageId="2" pageNumber="139">n. sp.</emphasis>
and
<taxonomicName id="4C2A06EFFF9F936BC9BBF8FEFD6FB8D6" baseAuthorityName="Cottarelli &amp; Argano" baseAuthorityYear="1977" box="[682,706,1233,1256]" class="Malacostraca" family="Trichodactylidae" genus="Rodriguezia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="139" phylum="Arthropoda" rank="species" species="mensabak">
<emphasis id="B95EA17EFF9F936BC9BBF8FEFD6FB8D6" box="[682,706,1233,1256]" italics="true" pageId="2" pageNumber="139">R.</emphasis>
</taxonomicName>
</taxonomicNameLabel>
<emphasis id="B95EA17EFF9F936BC9DFF8FFFC93B8D6" box="[718,830,1232,1256]" italics="true" pageId="2" pageNumber="139">mensabak</emphasis>
are more similar to each other than either is to
<taxonomicName id="4C2A06EFFF9F936BCE95F8FEFEA0B932" baseAuthorityName="Rodriguez &amp; Manrique" baseAuthorityYear="1967" class="Malacostraca" family="Trichodactylidae" genus="Rodriguezia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="139" phylum="Arthropoda" rank="species" species="villalobosi">
<emphasis id="B95EA17EFF9F936BCE95F8FEFEA0B932" italics="true" pageId="2" pageNumber="139">R. villalobosi</emphasis>
</taxonomicName>
. While
<taxonomicName id="4C2A06EFFF9F936BCA62F8DAFDA4B932" baseAuthorityName="Cottarelli &amp; Argano" baseAuthorityYear="1977" box="[371,521,1268,1292]" class="Malacostraca" family="Trichodactylidae" genus="Rodriguezia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="139" phylum="Arthropoda" rank="species" species="mensabak">
<emphasis id="B95EA17EFF9F936BCA62F8DAFDA4B932" box="[371,521,1268,1292]" italics="true" pageId="2" pageNumber="139">R. mensabak</emphasis>
</taxonomicName>
and
<taxonomicName id="4C2A06EFFF9F936BC95DF8DAFD1FB932" authorityName="Alvarez &amp; Villalobos" authorityYear="2018" box="[588,690,1268,1292]" class="Malacostraca" family="Trichodactylidae" genus="Rodriguezia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="139" phylum="Arthropoda" rank="species" species="adani">
<emphasis id="B95EA17EFF9F936BC95DF8DAFD1FB932" box="[588,690,1268,1292]" italics="true" pageId="2" pageNumber="139">R. adani</emphasis>
</taxonomicName>
are stygobitic species with elongated legs, lacking eyes, with chelipeds subequal, and a male gonopod without an obvious middle constriction,
<taxonomicName id="4C2A06EFFF9F936BCF4AF936FB59B90E" baseAuthorityName="Rodriguez &amp; Manrique" baseAuthorityYear="1967" box="[1115,1268,1304,1328]" class="Malacostraca" family="Trichodactylidae" genus="Rodriguezia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="139" phylum="Arthropoda" rank="species" species="villalobosi">
<emphasis id="B95EA17EFF9F936BCF4AF936FB59B90E" box="[1115,1268,1304,1328]" italics="true" pageId="2" pageNumber="139">R. villalobosi</emphasis>
</taxonomicName>
is an epigean species with normal eyes, short pereiopods, stongly unequal chelipeds, and a male gonopod with a strong middle constriction.
<taxonomicName id="4C2A06EFFF9F936BCA3CF94FFDACB946" authorityName="Alvarez &amp; Villalobos" authorityYear="2018" box="[301,513,1376,1400]" class="Malacostraca" family="Trichodactylidae" genus="Rodriguezia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="139" phylum="Arthropoda" rank="species" species="adani">
<emphasis id="B95EA17EFF9F936BCA3CF94FFDACB946" box="[301,513,1376,1400]" italics="true" pageId="2" pageNumber="139">Rodriguezia adani</emphasis>
</taxonomicName>
differs from
<taxonomicName id="4C2A06EFFF9F936BC9B1F94EFC99B946" baseAuthorityName="Cottarelli &amp; Argano" baseAuthorityYear="1977" box="[672,820,1376,1400]" class="Malacostraca" family="Trichodactylidae" genus="Rodriguezia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="139" phylum="Arthropoda" rank="species" species="mensabak">
<emphasis id="B95EA17EFF9F936BC9B1F94EFC99B946" box="[672,820,1376,1400]" italics="true" pageId="2" pageNumber="139">R. mensabak</emphasis>
</taxonomicName>
in: the length of the pereiopods relative to carapace breadth, the new species has shorter legs; and in the extremely reduced ocular peduncle, which appears as a plug inside the orbit. Regarding other characters,
<taxonomicName id="4C2A06EFFF9F936BC994F986FD4EB9FE" authorityName="Alvarez &amp; Villalobos" authorityYear="2018" box="[645,739,1448,1472]" class="Malacostraca" family="Trichodactylidae" genus="Rodriguezia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="139" phylum="Arthropoda" rank="species" species="adani">
<emphasis id="B95EA17EFF9F936BC994F986FD4EB9FE" box="[645,739,1448,1472]" italics="true" pageId="2" pageNumber="139">R. adani</emphasis>
</taxonomicName>
and
<taxonomicName id="4C2A06EFFF9F936BC808F986FC0AB9FE" baseAuthorityName="Cottarelli &amp; Argano" baseAuthorityYear="1977" box="[793,935,1448,1472]" class="Malacostraca" family="Trichodactylidae" genus="Rodriguezia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="139" phylum="Arthropoda" rank="species" species="mensabak">
<emphasis id="B95EA17EFF9F936BC808F986FC0AB9FE" box="[793,935,1448,1472]" italics="true" pageId="2" pageNumber="139">R. mensabak</emphasis>
</taxonomicName>
are similar in the first gonopod and carapace morphology, and the shape of the male abdomen.
</paragraph>
<paragraph id="8B957D6CFF9F936BCBD6F9DEFC98BB3A" blockId="2.[151,1437,151,2013]" pageId="2" pageNumber="139">
Two interesting differences between
<taxonomicName id="4C2A06EFFF9F936BC97EF9DEFD7CBA36" authorityName="Alvarez &amp; Villalobos" authorityYear="2018" box="[623,721,1520,1544]" class="Malacostraca" family="Trichodactylidae" genus="Rodriguezia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="139" phylum="Arthropoda" rank="species" species="adani">
<emphasis id="B95EA17EFF9F936BC97EF9DEFD7CBA36" box="[623,721,1520,1544]" italics="true" pageId="2" pageNumber="139">R. adani</emphasis>
</taxonomicName>
. and
<taxonomicName id="4C2A06EFFF9F936BC80CF9DEFC02BA36" baseAuthorityName="Cottarelli &amp; Argano" baseAuthorityYear="1977" box="[797,943,1520,1544]" class="Malacostraca" family="Trichodactylidae" genus="Rodriguezia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="139" phylum="Arthropoda" rank="species" species="mensabak">
<emphasis id="B95EA17EFF9F936BC80CF9DEFC02BA36" box="[797,943,1520,1544]" italics="true" pageId="2" pageNumber="139">R. mensabak</emphasis>
</taxonomicName>
are the following. Grutas de Agua Blanca where the new species was collected is at an altitude of
<quantity id="4CD2D089FF9F936BC808FA3BFCCEBA12" box="[793,867,1556,1581]" metricMagnitude="2" metricUnit="m" metricValue="1.3599999999999999" pageId="2" pageNumber="139" unit="m" value="136.0">136 m</quantity>
, while Nicolás Bravo Cave near the town of Tila,
<collectingRegion id="49EEB38EFF9F936BCB86FA18FF59BA6E" box="[151,244,1591,1616]" country="Mexico" name="Chiapas" pageId="2" pageNumber="139">Chiapas</collectingRegion>
, the
<typeStatus id="5491C3CEFF9F936BCA3DFA16FEF1BA6F" box="[300,348,1593,1617]" pageId="2" pageNumber="139">type</typeStatus>
locality and only known occurrence of
<taxonomicName id="4C2A06EFFF9F936BC839FA16FC14BA6E" baseAuthorityName="Cottarelli &amp; Argano" baseAuthorityYear="1977" box="[808,953,1592,1616]" class="Malacostraca" family="Trichodactylidae" genus="Rodriguezia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="139" phylum="Arthropoda" rank="species" species="mensabak">
<emphasis id="B95EA17EFF9F936BC839FA16FC14BA6E" box="[808,953,1592,1616]" italics="true" pageId="2" pageNumber="139">R. mensabak</emphasis>
</taxonomicName>
, is at an altitude of
<quantity id="4CD2D089FF9F936BCFB2FA18FAACBA6E" box="[1187,1281,1591,1616]" metricMagnitude="3" metricUnit="m" metricValue="1.16" pageId="2" pageNumber="139" unit="m" value="1160.0">1,160 m</quantity>
(
<bibRefCitation id="EFBB009DFF9F936BCE03FA18FE80BA4A" pageId="2" pageNumber="139" refId="ref2501" refString="Cottarelli, V. &amp; Argano, R. (1977) Trichodactylus (Rodriguezia) mensabak n. sp. (Crustacea, Decapoda, Brachyura), granchio cieco delle acque sotterranee del Chiapas (Messico). Quaderni Problemi Attuali di Scienza e di Cultura, Accademia Nazionale dei Lincei, 171, 207 - 212." type="journal article">Cottarelli &amp; Argano 1977</bibRefCitation>
). Although the two caves are only
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apart, the former lies at the northern edge of the Sierra Norte de
<collectingRegion id="49EEB38EFF9F936BCBA9FA50FEB8BAA6" box="[184,277,1663,1688]" country="Mexico" name="Chiapas" pageId="2" pageNumber="139">Chiapas</collectingRegion>
, while the latter is well within the Altos de
<collectingRegion id="49EEB38EFF9F936BC9EBFA50FCF9BAA6" box="[762,852,1663,1688]" country="Mexico" name="Chiapas" pageId="2" pageNumber="139">Chiapas</collectingRegion>
region. The second difference is their body size, the largest
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<emphasis id="B95EA17EFF9F936BCBF8FA8AFEE5BA82" box="[233,328,1700,1724]" italics="true" pageId="2" pageNumber="139">R. adani</emphasis>
</taxonomicName>
, a female carrying four young crabs, has a carapace breadth of
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, whereas the
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of
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<emphasis id="B95EA17EFF9F936BCB86FAE6FE8BBADE" box="[151,294,1736,1760]" italics="true" pageId="2" pageNumber="139">R. mensabak</emphasis>
</taxonomicName>
has a carapace breadth of
<quantity id="4CD2D089FF9F936BC942FAE8FD15BADE" box="[595,696,1735,1760]" metricMagnitude="-2" metricUnit="m" metricValue="1.6199999999999999" pageId="2" pageNumber="139" unit="mm" value="16.2">16.2 mm</quantity>
(
<bibRefCitation id="EFBB009DFF9F936BC9D7FAE8FC43BADE" box="[710,1006,1735,1760]" pageId="2" pageNumber="139" refId="ref2501" refString="Cottarelli, V. &amp; Argano, R. (1977) Trichodactylus (Rodriguezia) mensabak n. sp. (Crustacea, Decapoda, Brachyura), granchio cieco delle acque sotterranee del Chiapas (Messico). Quaderni Problemi Attuali di Scienza e di Cultura, Accademia Nazionale dei Lincei, 171, 207 - 212." type="journal article">Cottarelli &amp; Argano 1977</bibRefCitation>
). The largest known adults of the new species are 2/3 the size of the largest adults of
<taxonomicName id="4C2A06EFFF9F936BC98EFAC2FC83BB3A" baseAuthorityName="Cottarelli &amp; Argano" baseAuthorityYear="1977" box="[671,814,1772,1796]" class="Malacostraca" family="Trichodactylidae" genus="Rodriguezia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="139" phylum="Arthropoda" rank="species" species="mensabak">
<emphasis id="B95EA17EFF9F936BC98EFAC2FC83BB3A" box="[671,814,1772,1796]" italics="true" pageId="2" pageNumber="139">R. mensabak</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8B957D6CFF9F936ACBD6FB20FCA7BCC6" blockId="2.[151,1437,151,2013]" lastBlockId="3.[151,1436,151,248]" lastPageId="3" lastPageNumber="140" pageId="2" pageNumber="139">
Regarding the habitat of
<taxonomicName id="4C2A06EFFF9F936BCAF5FB3FFD1BBB16" authority="Alvarez &amp; Villalobos, 2018" authorityName="Alvarez &amp; Villalobos" authorityYear="2018" box="[484,694,1808,1832]" class="Malacostraca" family="Trichodactylidae" genus="Rodriguezia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="139" phylum="Arthropoda" rank="species" species="adani" status="sp. nov.">
<emphasis id="B95EA17EFF9F936BCAF5FB3FFD1BBB16" box="[484,694,1808,1832]" italics="true" pageId="2" pageNumber="139">Rodriguezia adani</emphasis>
</taxonomicName>
<emphasis id="B95EA17EFF9F936BC9AEFB20FCADBB16" bold="true" box="[703,768,1807,1832]" pageId="2" pageNumber="139">
<taxonomicNameLabel id="A26D1C05FF9F936BC9AEFB20FCADBB16" box="[703,768,1807,1832]" pageId="2" pageNumber="139" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
in Grutas de Agua Blanca, the crabs were collected about
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from the cave entrance in complete darkness. There is a strong water flow near the entrance, which diminishes to an almost imperceptible flow in ponds that collect in the deeper part of this section that goes for
<specimenCount id="9D2CB6E5FF9F936BCE63FB77FF19BBAA" pageId="2" pageNumber="139" type="generic">
<quantity id="4CD2D089FF9F936BCE63FB77FF1DBBAA" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="2" pageNumber="139" unit="m" value="200.0">200 m</quantity>
.
</specimenCount>
The crabs were found along the edge of ponds under rocks in sandy substrate with clear water. Grutas de Agua Blanca is also the
<typeStatus id="5491C3CEFF9F936BCA79FB8EFE35BB87" box="[360,408,1953,1977]" pageId="2" pageNumber="139">type</typeStatus>
locality of the blind catfish
<taxonomicName id="4C2A06EFFF9F936BC9CDFB8FFABCBB86" authority="Weber &amp; Wilkens, 1998" authorityName="Weber &amp; Wilkens" authorityYear="1998" box="[732,1297,1952,1977]" class="Actinopterygii" family="Heptapteridae" genus="Rhamdia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="139" phylum="Chordata" rank="species" species="macuspanensis">
<emphasis id="B95EA17EFF9F936BC9CDFB8FFC59BB86" box="[732,1012,1952,1976]" italics="true" pageId="2" pageNumber="139">Rhamdia macuspanensis</emphasis>
<bibRefCitation id="EFBB009DFF9F936BC8EDFB8FFABCBB86" box="[1020,1297,1952,1977]" pageId="2" pageNumber="139" refId="ref2810" refString="Weber, A. &amp; Wilkens, H. (1998) Rhamdia macuspanensis: A new species of troglobitic pimelodid catfish (Siluriformes; Pimelodidae) from a cave in Tabasco, Mexico. Copeia, 1998 (4), 998 - 1004. https: // doi. org / 10.2307 / 1447347" type="journal article">Weber &amp; Wilkens, 1998</bibRefCitation>
</taxonomicName>
. Apparently the new species described herein was collected in the same section of the cave as
<taxonomicName id="4C2A06EFFF9F936ACF44FBEAFE93BC8E" authority="(Weber &amp; Wilkens 1998)" authorityName="Weber &amp; Wilkens" authorityYear="1998" baseAuthorityName="Weber &amp; Wilkens" baseAuthorityYear="1998" class="Actinopterygii" family="Heptapteridae" genus="Rhamdia" higherTaxonomySource="GBIF" kingdom="Animalia" lastPageId="3" lastPageNumber="140" order="Siluriformes" pageId="2" pageNumber="139" phylum="Chordata" rank="species" species="macuspanensis">
<emphasis id="B95EA17EFF9F936BCF44FBEAFA8DBBE2" box="[1109,1312,1989,2012]" italics="true" pageId="2" pageNumber="139">R. macuspanensis</emphasis>
(
<bibRefCitation id="EFBB009DFF9F936ACE23FBEBFE98BC8E" lastPageId="3" lastPageNumber="140" pageId="2" pageNumber="139" refId="ref2810" refString="Weber, A. &amp; Wilkens, H. (1998) Rhamdia macuspanensis: A new species of troglobitic pimelodid catfish (Siluriformes; Pimelodidae) from a cave in Tabasco, Mexico. Copeia, 1998 (4), 998 - 1004. https: // doi. org / 10.2307 / 1447347" type="journal article">Weber &amp; Wilkens 1998</bibRefCitation>
)
</taxonomicName>
. Other organisms collected in the cave are the pseudothelphusid crab
<taxonomicName id="4C2A06EFFF9E936ACF52FCB7FE7EBCEA" authority="Alvarez &amp; Villalobos, 1998" authorityName="Alvarez &amp; Villalobos" authorityYear="1998" class="Malacostraca" family="Pseudothelphusidae" genus="Odontothelphusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="140" phylum="Arthropoda" rank="species" species="palenquensis">
<emphasis id="B95EA17EFF9E936ACF52FCB7FA31BC8E" box="[1091,1436,152,176]" italics="true" pageId="3" pageNumber="140">Odontothelphusa palenquensis</emphasis>
Alvarez &amp; Villalobos, 1998
</taxonomicName>
, an undescribed species of freshwater prawn of the genus
<taxonomicName id="4C2A06EFFF9E936ACF78FC93FAB3BCEA" box="[1129,1310,188,212]" class="Malacostraca" family="Palaemonidae" genus="Macrobrachium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="140" phylum="Arthropoda" rank="genus">
<emphasis id="B95EA17EFF9E936ACF78FC93FAB3BCEA" box="[1129,1310,188,212]" italics="true" pageId="3" pageNumber="140">Macrobrachium</emphasis>
</taxonomicName>
, a juvenile crayfish of the genus
<taxonomicName id="4C2A06EFFF9E936ACA99FCCFFDB2BCC6" authorityName="Ortmann" authorityYear="1905" box="[392,543,224,248]" class="Malacostraca" family="Cambaridae" genus="Procambarus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="140" phylum="Arthropoda" rank="genus">
<emphasis id="B95EA17EFF9E936ACA99FCCFFDB2BCC6" box="[392,543,224,248]" italics="true" pageId="3" pageNumber="140">Procambarus</emphasis>
</taxonomicName>
, and asellid isopods.
</paragraph>
<caption id="DF552DE4FF9E936ACB86FBE6FB83BBE1" box="[151,1070,1993,2015]" httpUri="https://zenodo.org/record/1168166/files/figure.png" pageId="3" pageNumber="140" startId="3.[151,250,1993,2015]" targetBox="[177,1419,333,1977]" targetPageId="3">
<paragraph id="8B957D6CFF9E936ACB86FBE6FB83BBE1" blockId="3.[151,1070,1993,2015]" box="[151,1070,1993,2015]" pageId="3" pageNumber="140">
<emphasis id="B95EA17EFF9E936ACB86FBE6FEBFBBE0" bold="true" box="[151,274,1993,2015]" pageId="3" pageNumber="140">FIGURE 2.</emphasis>
<taxonomicName id="4C2A06EFFF9E936ACA08FBE6FE78BBE1" authority="Alvarez &amp; Villalobos, 2018" authorityName="Alvarez &amp; Villalobos" authorityYear="2018" box="[281,469,1993,2015]" class="Malacostraca" family="Trichodactylidae" genus="Rodriguezia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="140" phylum="Arthropoda" rank="species" species="adani" status="sp. nov.">
<emphasis id="B95EA17EFF9E936ACA08FBE6FE78BBE1" box="[281,469,1993,2015]" italics="true" pageId="3" pageNumber="140">Rodriguezia adani</emphasis>
</taxonomicName>
<emphasis id="B95EA17EFF9E936ACACAFBE6FDB9BBE0" bold="true" box="[475,532,1993,2014]" pageId="3" pageNumber="140">
<taxonomicNameLabel id="A26D1C05FF9E936ACACAFBE6FDB9BBE0" box="[475,532,1993,2014]" pageId="3" pageNumber="140" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
male holotype: A, B, dorsal view. Scale bar = 5 mm.
</paragraph>
</caption>
<paragraph id="8B957D6CFF99936DCBD6FCB7FE2CBD7E" blockId="4.[151,1437,151,320]" pageId="4" pageNumber="141">
It remains a relevant question for future studies to determine if the two stygobitic species originated from the epigean
<taxonomicName id="4C2A06EFFF99936DCBE4FC92FE27BCEA" baseAuthorityName="Rodriguez &amp; Manrique" baseAuthorityYear="1967" box="[245,394,188,212]" class="Malacostraca" family="Trichodactylidae" genus="Rodriguezia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="141" phylum="Arthropoda" rank="species" species="villalobosi">
<emphasis id="B95EA17EFF99936DCBE4FC92FE27BCEA" box="[245,394,188,212]" italics="true" pageId="4" pageNumber="141">R. villalobosi</emphasis>
</taxonomicName>
, or if they represent two separate lineages. In case they do, a complex geological setting will have to be invoked since
<taxonomicName id="4C2A06EFFF99936DCAD0FCCEFDF4BCC6" baseAuthorityName="Rodriguez &amp; Manrique" baseAuthorityYear="1967" box="[449,601,224,248]" class="Malacostraca" family="Trichodactylidae" genus="Rodriguezia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="141" phylum="Arthropoda" rank="species" species="villalobosi">
<emphasis id="B95EA17EFF99936DCAD0FCCEFDF4BCC6" box="[449,601,224,248]" italics="true" pageId="4" pageNumber="141">R. villalobosi</emphasis>
</taxonomicName>
is the species occurring farther inland, and would have had to reach its present geographic position very early in the formation of this part of southern
<collectingCountry id="F33D3DFCFF99936DCF36FD2BFBD2BD23" box="[1063,1151,260,285]" name="Mexico" pageId="4" pageNumber="141">
<collectingRegion id="49EEB38EFF99936DCF36FD2BFBD2BD23" box="[1063,1151,260,285]" country="Mexico" name="Mexico" pageId="4" pageNumber="141">Mexico</collectingRegion>
</collectingCountry>
, during early Cretaceous times (
<bibRefCitation id="EFBB009DFF99936DCBF3FD07FED8BD7E" box="[226,373,296,320]" pageId="4" pageNumber="141" refId="ref2561" refString="Ecosur (2001) Geological map of Chiapas, Mexico (1: 2,000,000). El Colegio de la Frontera Sur - INEGI. Available from: www. ecosur. mx / sitios / images / stories / ecosur / Servicios / LAIGE / mapas _ PEOT / glmp _ geologia. jpg (accessed 8 August 2018)" type="journal article">Ecosur 2001</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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