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<document ID-DOI="http://dx.doi.org/10.11646/zootaxa.3826.1.1" ID-GBIF-Dataset="e7c519fb-b07e-452b-91a5-efdec51eb764" ID-ISSN="1175-5326" ID-Zenodo-Dep="286429" ID-ZooBank="B6414C18-599A-44CE-9FCA-F20C845DE79D" checkinTime="1460443804064" checkinUser="plazi" docAuthor="Bayer, Steffen" docDate="2014" docId="0383AE33C86CFFB1D2B4FEEDFC8CF43C" docLanguage="en" docName="zt03826p054.pdf" docOrigin="Zootaxa 3826 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Psechrus ancoralis" docType="treatment" docVersion="1" lastPageNumber="13" masterDocId="FFBAD64BC860FFBDD223FF82FFBAF113" masterDocTitle="Seven new species of Psechrus and additional taxonomic contributions to the knowledge of the spider family Psechridae (Araneae)" masterLastPageNumber="54" masterPageNumber="1" pageNumber="13" updateTime="1677763788444" updateUser="juliana">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Seven new species of Psechrus and additional taxonomic contributions to the knowledge of the spider family Psechridae (Araneae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Bayer, Steffen</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Zootaxa</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2014</mods:date>
<mods:detail type="pubDate">
<mods:number>2014-05-16</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>3826</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>1</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>54</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.11646/zootaxa.3826.1.1</mods:identifier>
<mods:identifier type="GBIF-Dataset">e7c519fb-b07e-452b-91a5-efdec51eb764</mods:identifier>
<mods:identifier type="ISSN">1175-5326</mods:identifier>
<mods:identifier type="Zenodo-Dep">286429</mods:identifier>
<mods:identifier type="ZooBank">B6414C18-599A-44CE-9FCA-F20C845DE79D</mods:identifier>
</mods:mods>
<treatment LSID="urn:lsid:plazi:treatment:0383AE33C86CFFB1D2B4FEEDFC8CF43C" httpUri="http://treatment.plazi.org/id/0383AE33C86CFFB1D2B4FEEDFC8CF43C" lastPageNumber="13" pageId="12" pageNumber="13">
<subSubSection box="[151,347,367,394]" pageId="12" pageNumber="13" type="nomenclature">
<paragraph blockId="12.[151,347,367,394]" box="[151,347,367,394]" pageId="12" pageNumber="13">
<heading bold="true" box="[151,347,367,394]" fontSize="11" level="1" pageId="12" pageNumber="13" reason="1">
<emphasis bold="true" box="[151,347,367,394]" pageId="12" pageNumber="13">
<taxonomicName ID-CoL="78B7S" ID-ENA="1304647" authority="Bayer &amp; Jäger, 2010" box="[151,273,367,393]" class="Arachnida" family="Psechridae" genus="Psechrus" higherTaxonomySource="CoL" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="ancoralis">
<emphasis bold="true" box="[151,273,367,393]" italics="true" pageId="12" pageNumber="13">ancoralis-</emphasis>
</taxonomicName>
group
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection pageId="12" pageNumber="13" type="diagnosis">
<paragraph blockId="12.[151,1437,438,1327]" pageId="12" pageNumber="13">
<emphasis bold="true" box="[151,266,438,463]" pageId="12" pageNumber="13">Diagnosis</emphasis>
(after
<bibRefCitation author="Bayer" box="[341,478,439,463]" firstAuthor="Bayer" pageId="12" pageNumber="13" refString="Bayer, S. (2012) The lace-sheet-weavers-a long story (Araneae: Psechridae: Psechrus). Zootaxa, 3379, 1 - 170." type="journal article" year="2012">
Bayer
<number box="[416,478,439,463]" pageId="12" pageNumber="13" value="2012.0">2012</number>
</bibRefCitation>
): Males with an elongated, filiform embolus arising retrolaterally on the tegulum and with a fleshy or membranous conductor (
<bibRefCitation author="Bayer" box="[611,746,475,499]" firstAuthor="Bayer" pageId="12" pageNumber="13" refString="Bayer, S. (2012) The lace-sheet-weavers-a long story (Araneae: Psechridae: Psechrus). Zootaxa, 3379, 1 - 170." type="journal article" year="2012">
Bayer
<number box="[687,746,475,499]" pageId="12" pageNumber="13" value="2012.0">2012</number>
</bibRefCitation>
, figs.
<number box="[813,841,474,499]" pageId="12" pageNumber="13" value="21.0">21</number>
b–c,
<number box="[895,923,474,499]" pageId="12" pageNumber="13" value="25.0">25</number>
b–c) and an unmodified palpal femur.
</paragraph>
<paragraph blockId="12.[151,1437,438,1327]" pageId="12" pageNumber="13">
Females are distinguished from those of all other species groups except the
<taxonomicName box="[1128,1271,510,534]" class="Arachnida" family="Psechridae" genus="Psechrus" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="himalayanus">
<emphasis box="[1128,1271,510,534]" italics="true" pageId="12" pageNumber="13">himalayanus</emphasis>
</taxonomicName>
-group by the following combination of characters: Epigyne without tegimentum (
<bibRefCitation author="Bayer" box="[959,1101,547,571]" firstAuthor="Bayer" pageId="12" pageNumber="13" refString="Bayer, S. (2012) The lace-sheet-weavers-a long story (Araneae: Psechridae: Psechrus). Zootaxa, 3379, 1 - 170." type="journal article" year="2012">
Bayer
<number box="[1042,1101,547,571]" pageId="12" pageNumber="13" value="2012.0">2012</number>
</bibRefCitation>
, figs
<number box="[1174,1204,546,571]" pageId="12" pageNumber="13" value="69.0">69</number>
a,
<quantity box="[1235,1286,547,571]" metricMagnitude="1" metricUnit="m" metricValue="8.9" pageId="12" pageNumber="13" unit="m" value="89.0">89m</quantity>
) or strongly sclerotised epigynal field (
<bibRefCitation author="Bayer" box="[446,580,583,607]" firstAuthor="Bayer" pageId="12" pageNumber="13" refString="Bayer, S. (2012) The lace-sheet-weavers-a long story (Araneae: Psechridae: Psechrus). Zootaxa, 3379, 1 - 170." type="journal article" year="2012">
Bayer
<number box="[521,580,583,607]" pageId="12" pageNumber="13" value="2012.0">2012</number>
</bibRefCitation>
, figs
<number box="[639,668,582,607]" pageId="12" pageNumber="13" value="87.0">87</number>
c–d), with small, slit-like copulatory openings (
<bibRefCitation author="Bayer" box="[1195,1428,582,607]" pageId="12" pageNumber="13" refString="Bayer, S. (2012) The lace-sheet-weavers-a long story (Araneae: Psechridae: Psechrus). Zootaxa, 3379, 1 - 170." type="journal article" year="2012" yearSuffix="d">
Bayer
<number box="[1270,1329,583,607]" pageId="12" pageNumber="13" value="2012.0">2012</number>
, fig.
<number box="[1385,1413,582,607]" pageId="12" pageNumber="13" value="37.0">37</number>
d
</bibRefCitation>
) and mostly converging anterior sections of epigynal slits (
<bibRefCitation author="Bayer" box="[835,974,619,643]" firstAuthor="Bayer" pageId="12" pageNumber="13" refString="Bayer, S. (2012) The lace-sheet-weavers-a long story (Araneae: Psechridae: Psechrus). Zootaxa, 3379, 1 - 170." type="journal article" year="2012">
Bayer
<number box="[915,974,619,643]" pageId="12" pageNumber="13" value="2012.0">2012</number>
</bibRefCitation>
, figs
<number box="[1043,1073,619,643]" pageId="12" pageNumber="13" value="27.0">27</number>
e,
<number box="[1102,1133,618,643]" pageId="12" pageNumber="13" value="36.0">36</number>
a). Median septum mostly broader than long; receptacula not distinctly round, copulatory ducts without distinctly rough surface (
<bibRefCitation author="Bayer" pageId="12" pageNumber="13" refString="Bayer, S. (2012) The lace-sheet-weavers-a long story (Araneae: Psechridae: Psechrus). Zootaxa, 3379, 1 - 170." type="journal article" year="2012" yearSuffix="b">
Bayer
<number box="[1373,1432,655,679]" pageId="12" pageNumber="13" value="2012.0">2012</number>
, fig.
<number box="[200,229,690,715]" pageId="12" pageNumber="13" value="80.0">80</number>
b
</bibRefCitation>
) and bulbous (
<bibRefCitation author="Bayer" box="[420,665,690,715]" pageId="12" pageNumber="13" refString="Bayer, S. (2012) The lace-sheet-weavers-a long story (Araneae: Psechridae: Psechrus). Zootaxa, 3379, 1 - 170." type="journal article" year="2012" yearSuffix="b">
Bayer
<number box="[499,558,691,715]" pageId="12" pageNumber="13" value="2012.0">2012</number>
, fig.
<number box="[622,651,690,715]" pageId="12" pageNumber="13" value="63.0">63</number>
b
</bibRefCitation>
) or twisted (
<bibRefCitation author="Bayer" box="[817,1062,690,715]" pageId="12" pageNumber="13" refString="Bayer, S. (2012) The lace-sheet-weavers-a long story (Araneae: Psechridae: Psechrus). Zootaxa, 3379, 1 - 170." type="journal article" year="2012" yearSuffix="b">
Bayer
<number box="[897,956,691,715]" pageId="12" pageNumber="13" value="2012.0">2012</number>
, fig.
<number box="[1019,1048,690,715]" pageId="12" pageNumber="13" value="59.0">59</number>
b
</bibRefCitation>
) sections and at most
<number box="[1330,1365,690,715]" pageId="12" pageNumber="13" value="1.5">1.5</number>
times broader than diameter of one receptaculum. Females are distinguished from those of the
<taxonomicName box="[1142,1285,726,750]" class="Arachnida" family="Psechridae" genus="Psechrus" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="himalayanus">
<emphasis box="[1142,1285,726,750]" italics="true" pageId="12" pageNumber="13">himalayanus</emphasis>
</taxonomicName>
-group by the spermathecal heads located upon the copulatory ducts (
<bibRefCitation author="Bayer" box="[803,943,763,787]" firstAuthor="Bayer" pageId="12" pageNumber="13" refString="Bayer, S. (2012) The lace-sheet-weavers-a long story (Araneae: Psechridae: Psechrus). Zootaxa, 3379, 1 - 170." type="journal article" year="2012">
Bayer
<number box="[884,943,763,787]" pageId="12" pageNumber="13" value="2012.0">2012</number>
</bibRefCitation>
, figs
<number box="[1012,1041,763,787]" pageId="12" pageNumber="13" value="21.0">21</number>
e,
<number box="[1072,1103,762,787]" pageId="12" pageNumber="13" value="29.0">29</number>
b); the latter longer than the diameter of receptaculum.
</paragraph>
<paragraph blockId="12.[151,1437,438,1327]" pageId="12" pageNumber="13">
The
<taxonomicName box="[250,356,835,859]" class="Arachnida" family="Psechridae" genus="Psechrus" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="ancoralis">
<emphasis box="[250,356,835,859]" italics="true" pageId="12" pageNumber="13">ancoralis</emphasis>
</taxonomicName>
-group includes
<number box="[540,554,834,859]" pageId="12" pageNumber="13" value="6.0">6</number>
species (
<taxonomicName authority="Bayer &amp; Jager, 2010" authorityName="Bayer &amp; Jager" authorityYear="2010" box="[660,1120,835,859]" class="Arachnida" family="Psechridae" genus="Psechrus" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="ancoralis">
<emphasis box="[660,876,835,859]" italics="true" pageId="12" pageNumber="13">Psechrus ancoralis</emphasis>
<bibRefCitation author="Bayer &amp; Jager" box="[884,1120,835,859]" firstAuthor="Bayer" pageId="12" pageNumber="13" refString="Bayer, S. &amp; Jager, P. (2010) Expected species richness in the genus Psechrus in Laos (Araneae: Psechridae). Revue suisse de Zoologie, 117, 57 - 75." type="journal article" year="2010">
Bayer &amp; Jäger,
<number box="[1061,1120,835,859]" pageId="12" pageNumber="13" value="2010.0">2010</number>
</bibRefCitation>
</taxonomicName>
,
<taxonomicName authority="Bayer &amp; Jager, 2010" authorityName="Bayer &amp; Jager" authorityYear="2010" class="Arachnida" family="Psechridae" genus="Psechrus" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="antraeus">
<emphasis box="[1132,1259,835,858]" italics="true" pageId="12" pageNumber="13">P. antraeus</emphasis>
<bibRefCitation author="Bayer &amp; Jager" firstAuthor="Bayer" pageId="12" pageNumber="13" refString="Bayer, S. &amp; Jager, P. (2010) Expected species richness in the genus Psechrus in Laos (Araneae: Psechridae). Revue suisse de Zoologie, 117, 57 - 75." type="journal article" year="2010">
Bayer &amp; Jäger,
<number box="[151,210,871,895]" pageId="12" pageNumber="13" value="2010.0">2010</number>
</bibRefCitation>
</taxonomicName>
,
<emphasis box="[222,294,870,894]" italics="true" pageId="12" pageNumber="13">P. l a os</emphasis>
<bibRefCitation author="Bayer" box="[306,447,871,895]" firstAuthor="Bayer" pageId="12" pageNumber="13" refString="Bayer, S. (2012) The lace-sheet-weavers-a long story (Araneae: Psechridae: Psechrus). Zootaxa, 3379, 1 - 170." type="journal article" year="2012">
Bayer,
<number box="[388,447,871,895]" pageId="12" pageNumber="13" value="2012.0">2012</number>
</bibRefCitation>
,
<emphasis box="[460,532,871,894]" italics="true" pageId="12" pageNumber="13">P. r a ni</emphasis>
<bibRefCitation author="Wang &amp; Yin" box="[544,762,870,895]" firstAuthor="Wang" pageId="12" pageNumber="13" refString="Wang, X. - P. &amp; Yin, C. - M. (2001) A review of the Chinese Psechridae (Araneae). Journal of Arachnology, 29, 330 - 344. http: // dx. doi. org / 10.1636 / 0161 - 8202 (2001) 029 [0330: arotcp] 2.0. co; 2" type="journal article" year="2001">
Wang &amp; Yin,
<number box="[703,762,871,895]" pageId="12" pageNumber="13" value="2001.0">2001</number>
</bibRefCitation>
,
<taxonomicName authority="Bayer &amp; Jager, 2010" authorityName="Bayer &amp; Jager" authorityYear="2010" box="[775,1135,871,895]" class="Arachnida" family="Psechridae" genus="Psechrus" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="steineri">
<emphasis box="[775,888,871,894]" italics="true" pageId="12" pageNumber="13">P. steineri</emphasis>
<bibRefCitation author="Bayer &amp; Jager" box="[897,1135,871,895]" firstAuthor="Bayer" pageId="12" pageNumber="13" refString="Bayer, S. &amp; Jager, P. (2010) Expected species richness in the genus Psechrus in Laos (Araneae: Psechridae). Revue suisse de Zoologie, 117, 57 - 75." type="journal article" year="2010">
Bayer &amp; Jäger,
<number box="[1076,1135,871,895]" pageId="12" pageNumber="13" value="2010.0">2010</number>
</bibRefCitation>
</taxonomicName>
, and
<taxonomicName authority="Jager, 2007" authorityName="Jager" authorityYear="2007" class="Arachnida" family="Psechridae" genus="Psechrus" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="khammouan">
<emphasis box="[1196,1363,870,894]" italics="true" pageId="12" pageNumber="13">P. khammouan</emphasis>
<bibRefCitation author="Jager" firstAuthor="Jager" pageId="12" pageNumber="13" refString="Jager, P. (2007) Spiders from Laos with descriptions of new species (Arachnida: Araneae). Acta Arachnologica, 56, 29 - 58. http: // dx. doi. org / 10.2476 / asjaa. 56.29" type="journal article" year="2007">
Jäger,
<number box="[151,209,907,931]" pageId="12" pageNumber="13" value="2007.0">2007</number>
</bibRefCitation>
</taxonomicName>
) and is distributed in
<collectingCountry box="[456,526,906,931]" name="China" pageId="12" pageNumber="13">China</collectingCountry>
,
<collectingCountry box="[537,635,906,931]" name="Vietnam" pageId="12" pageNumber="13">Vietnam</collectingCountry>
,
<collectingCountry box="[647,747,906,931]" name="Thailand" pageId="12" pageNumber="13">Thailand</collectingCountry>
and
<collectingCountry box="[802,860,907,930]" name="Laos" pageId="12" pageNumber="13">Laos</collectingCountry>
. It is not unlikely that representatives of this group also occur in
<collectingCountry box="[311,422,943,967]" name="Myanmar" pageId="12" pageNumber="13">Myanmar</collectingCountry>
. This species group is closely related to the
<taxonomicName box="[944,1087,942,966]" class="Arachnida" family="Psechridae" genus="Psechrus" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="himalayanus">
<emphasis box="[944,1087,942,966]" italics="true" pageId="12" pageNumber="13">himalayanus</emphasis>
</taxonomicName>
-group (currently including
<number box="[1409,1436,943,967]" pageId="12" pageNumber="13" value="11.0">11</number>
species, see below) (
<bibRefCitation author="Bayer &amp; Schonhofer" box="[402,723,978,1003]" firstAuthor="Bayer" pageId="12" pageNumber="13" refString="Bayer, S. &amp; Schonhofer, A. (2013) Phylogenetic relationships of the spider family Psechridae inferred from molecular data, with comments on the Lycosoidea (Arachnida: Araneae). Invertebrate Systematics, 27, 53 - 80. http: // dx. doi. org / 10.1071 / is 12017" type="journal article" year="2013">
Bayer &amp; Schönhofer
<number box="[664,723,979,1003]" pageId="12" pageNumber="13" value="2013.0">2013</number>
</bibRefCitation>
;
<bibRefCitation author="Bayer" box="[742,884,979,1003]" firstAuthor="Bayer" pageId="12" pageNumber="13" refString="Bayer, S. (2012) The lace-sheet-weavers-a long story (Araneae: Psechridae: Psechrus). Zootaxa, 3379, 1 - 170." type="journal article" year="2012">
Bayer
<number box="[823,884,979,1003]" pageId="12" pageNumber="13" value="2012.0">2012</number>
</bibRefCitation>
). The females show the basic patterns of the copulatory organs of the females of the
<taxonomicName box="[608,751,1014,1038]" class="Arachnida" family="Psechridae" genus="Psechrus" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="himalayanus">
<emphasis box="[608,751,1014,1038]" italics="true" pageId="12" pageNumber="13">himalayanus</emphasis>
</taxonomicName>
-group. However, several male and female copulatory organ characters that are typical of the representatives of the
<taxonomicName box="[764,907,1051,1075]" class="Arachnida" family="Psechridae" genus="Psechrus" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="himalayanus">
<emphasis box="[764,907,1051,1075]" italics="true" pageId="12" pageNumber="13">himalayanus</emphasis>
</taxonomicName>
-group strongly indicate that these species very likely had a common ancestor. The same argument can also be applied to the
<taxonomicName box="[1054,1159,1086,1110]" class="Arachnida" family="Psechridae" genus="Psechrus" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="ancoralis">
<emphasis box="[1054,1159,1086,1110]" italics="true" pageId="12" pageNumber="13">ancoralis</emphasis>
</taxonomicName>
-group. Nevertheless, as mentioned above, all the
<number box="[447,475,1123,1147]" pageId="12" pageNumber="13" value="17.0">17</number>
currently known representatives of both groups together represent a homogeneous group of species (clade), which is clearly delimited from the remaining
<taxonomicName box="[964,1067,1158,1182]" class="Arachnida" family="Psechridae" genus="Psechrus" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
<emphasis box="[964,1067,1158,1182]" italics="true" pageId="12" pageNumber="13">Psechrus</emphasis>
</taxonomicName>
species. In
<bibRefCitation author="Bayer &amp; Schonhofer" firstAuthor="Bayer" pageId="12" pageNumber="13" refString="Bayer, S. &amp; Schonhofer, A. (2013) Phylogenetic relationships of the spider family Psechridae inferred from molecular data, with comments on the Lycosoidea (Arachnida: Araneae). Invertebrate Systematics, 27, 53 - 80. http: // dx. doi. org / 10.1071 / is 12017" type="journal article" year="2013">
Bayer &amp; Schönhofer (
<number box="[159,218,1195,1219]" pageId="12" pageNumber="13" value="2013.0">2013</number>
)
</bibRefCitation>
the
<taxonomicName box="[276,419,1195,1219]" class="Arachnida" family="Psechridae" genus="Psechrus" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="himalayanus">
<emphasis box="[276,419,1195,1219]" italics="true" pageId="12" pageNumber="13">himalayanus</emphasis>
</taxonomicName>
-group was recovered paraphyletic including the representatives of the
<taxonomicName box="[1225,1330,1195,1219]" class="Arachnida" family="Psechridae" genus="Psechrus" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="ancoralis">
<emphasis box="[1225,1330,1195,1219]" italics="true" pageId="12" pageNumber="13">ancoralis</emphasis>
</taxonomicName>
-group in one of the inner branches of a subtree within a proposed phylogenetic tree. However, in only one of four analyses was this tree reliably recovered and it might be artificial. It remains to be seen how topology will change, if
<emphasis box="[1347,1436,1267,1291]" italics="true" pageId="12" pageNumber="13">de facto</emphasis>
all species of both groups are included in a molecular study.
</paragraph>
</subSubSection>
</treatment>
</document>
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