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<document id="3B383D98CAC82541338CBB6639E4D157" ID-CLB-Dataset="30848" ID-DOI="10.11646/zootaxa.4377.4.10" ID-GBIF-Dataset="e01a3a46-9e8c-4866-b9f7-53f9ec24ba66" ID-ISSN="1175-5326" ID-Zenodo-Dep="1165782" ID-ZooBank="EF257818-5489-4D0D-A566-93A38568482F" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1517833634356" checkinUser="plazi" docAuthor="Sehnal, Richard" docDate="2018" docId="03E7057FFF84664D99EF7ABE386BFE91" docLanguage="en" docName="zootaxa.4377.4.10.pdf" docOrigin="Zootaxa 4377 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Oedanomerus lupanus Sehnal 2018, new species" docType="treatment" docVersion="7" lastPageNumber="598" masterDocId="FFDE7D07FF81664B99787B4C3A3EFFA8" masterDocTitle="Two new Oedanomerus species from Zimbabwe (Coleoptera: Scarabaeidae: Melolonthinae: Tanyproctini)" masterLastPageNumber="600" masterPageNumber="593" pageNumber="598" updateTime="1698518767090" updateUser="ExternalLinkService">
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<mods:title id="DA5C38F1DE788349C4A1647C29A1E619">Two new Oedanomerus species from Zimbabwe (Coleoptera: Scarabaeidae: Melolonthinae: Tanyproctini)</mods:title>
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<mods:namePart id="66F0BF9498D77771D19114CEDF1B8829">Sehnal, Richard</mods:namePart>
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<mods:title id="4BAAE8949375DC7CF5DE81D09F5EC7C8">Zootaxa</mods:title>
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<mods:part id="B3FA361ED04108533F67237786806051">
<mods:date id="D1C3B575FE8BF7EFE8C3F25A7A32A235">2018</mods:date>
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<mods:number id="76424D4C6C95EC23AE8B92D9AC34DF7C">2018-02-05</mods:number>
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<mods:number id="A034FE0CB8A93EF9D57B31741EE882F0">4377</mods:number>
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<treatment id="03E7057FFF84664D99EF7ABE386BFE91" ID-DOI="http://doi.org/10.5281/zenodo.5951256" ID-GBIF-Taxon="141125779" ID-Zenodo-Dep="5951256" LSID="urn:lsid:plazi:treatment:03E7057FFF84664D99EF7ABE386BFE91" httpUri="http://treatment.plazi.org/id/03E7057FFF84664D99EF7ABE386BFE91" lastPageId="6" lastPageNumber="598" pageId="5" pageNumber="598">
<subSubSection id="C354E7E2FF84664E99EF7ABE3E28FDA4" box="[151,1046,498,524]" pageId="5" pageNumber="598" type="nomenclature">
<paragraph id="8BF1B469FF84664E99EF7ABE3E28FDA4" blockId="5.[151,1436,498,2012]" box="[151,1046,498,524]" pageId="5" pageNumber="598">
<heading id="D0B90305FF84664E99EF7ABE3E28FDA4" bold="true" box="[151,1046,498,524]" fontSize="11" level="1" pageId="5" pageNumber="598" reason="1">
<emphasis id="B93A687BFF84664E99EF7ABE3E28FDA4" bold="true" box="[151,1046,498,524]" pageId="5" pageNumber="598">
<taxonomicName id="4C4ECFEAFF84664E99EF7ABE3839FDA4" ID-CoL="74JK6" authority="Sehnal" authorityName="Sehnal" authorityYear="2018" box="[151,519,498,524]" class="Insecta" family="Scarabaeidae" genus="Oedanomerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="598" phylum="Arthropoda" rank="species" species="lupanus" status="sp. nov.">
<emphasis id="B93A687BFF84664E99EF7ABE3B97FDA4" bold="true" box="[151,425,498,524]" italics="true" pageId="5" pageNumber="598">Oedanomerus lupanus</emphasis>
Sehnal
</taxonomicName>
,
<taxonomicNameLabel id="A209D500FF84664E9B6C7ABE389DFDA4" box="[532,675,498,524]" pageId="5" pageNumber="598" rank="species">new species</taxonomicNameLabel>
(
<figureCitation id="1375A8ECFF84664E9BCB7ABE3924FDA4" box="[691,794,498,524]" captionStart="FIGURES 1" captionStartId="1.[151,258,1690,1712]" captionTargetBox="[184,1405,218,1654]" captionTargetId="figure@1.[151,1435,191,1668]" captionTargetPageId="1" captionText="FIGURES 1AD. Habitus, dorsal and lateral views. A, Oedanomerus bidentatus Sehnal, new species, holotype male; B, Oedanomerus lupanus Sehnal, new species, holotype male; C, Oedanomerus squamosus Lacroix, 2005, holotype male; D, Oedanomerus uhligi Lacroix,2005,holotype male.Scalebar = 1 mm." httpUri="https://zenodo.org/record/1165784/files/figure.png" pageId="5" pageNumber="598">Figs. 1B</figureCitation>
,
<figureCitation id="1375A8ECFF84664E9A507ABE3972FDA4" box="[808,844,498,524]" captionStart="FIGURES 2" captionStartId="2.[151,258,1799,1821]" captionTargetBox="[167,1420,594,1757]" captionTargetId="figure@2.[151,1436,573,1778]" captionTargetPageId="2" captionText="FIGURES 2AL. Antenna (AD), clypeus (EH) and pronotal disc (IL). A, E, I, Oedanomerus bidentatus Sehnal, new species, holotype male; B, F, J, Oedanomerus lupanus Sehnal, new species, holotype male; C, G, K, Oedanomerus squamosus Lacroix, 2005, holotype male; D, H,L, Oedanomerusuhligi Lacroix,2005, holotype male.Scale bar = 1 mm." httpUri="https://zenodo.org/record/1165786/files/figure.png" pageId="5" pageNumber="598">2B</figureCitation>
,
<figureCitation id="1375A8ECFF84664E9A217ABE3943FDA4" box="[857,893,498,524]" captionStart="FIGURES 3" captionStartId="3.[151,258,1941,1963]" captionTargetBox="[168,1415,569,1876]" captionTargetId="figure@3.[150,1436,548,1898]" captionTargetPageId="3" captionText="FIGURES 3AD. Aedeagus, dorsal (left) and lateral (right) views. A, Oedanomerus bidentatus Sehnal, new species, holotype male; B, Oedanomerus lupanus Sehnal, new species, holotype male; C, Oedanomerus squamosus Lacroix, 2005, holotype male; D, Oedanomerus uhligi Lacroix,2005,holotype male.Scalebar = 1 mm." httpUri="https://zenodo.org/record/1165788/files/figure.png" pageId="5" pageNumber="598">3B</figureCitation>
,
<figureCitation id="1375A8ECFF84664E9AF27ABE39E4FDA4" box="[906,986,498,524]" captionStart="FIGURES4" captionStartId="4.[151,258,1795,1817]" captionTargetBox="[177,1409,204,1754]" captionTargetId="figure@4.[151,1436,191,1774]" captionTargetPageId="4" captionText="FIGURES4AH. Metafemur withmetatibia (AD)and protibia(EF).A, E, Oedanomerus bidentatusSehnal,new species, holotype male; B, F, Oedanomerus lupanus Sehnal, new species,holotype male; C, G, Oedanomerus squamosus Lacroix,2005,holotype male; D, H,Oedanomerus uhligi Lacroix,2005,holotype male.Scale bar = 1 mm." httpUri="https://zenodo.org/record/1165790/files/figure.png" pageId="5" pageNumber="598">4B, 4F</figureCitation>
,
<figureCitation id="1375A8ECFF84664E9A907ABE3E30FDA4" box="[1000,1038,498,524]" captionStart="FIGURE 5" captionStartId="6.[151,244,1524,1546]" captionTargetBox="[150,1436,367,1502]" captionTargetId="figure@6.[150,1436,367,1502]" captionTargetPageId="6" captionText="FIGURE 5. Map with currently known geographic distribution in southern Africa: A—Oedanomerus bicolor Evans, 1987; B— Oedanomerus capriviensis Evans, 1987; C—Oedanomerus hirsutus Waterhouse, 1875; D—Oedanomerus longicornis Arrow, 1987; E—Oedanomerus pilosus Frey, 1960; F—Oedanomerus snizeki Lacroix, 2005; G—Oedanomerus squamosus Lacroix, 2005; H— Oedanomerus uhligi Lacroix, 2005; J—Oedanomerus bidentatus Sehnal, new species; K—Oedanomerus lupanus Sehnal, new species.Reddots indicatenew species or new locality." httpUri="https://zenodo.org/record/1165792/files/figure.png" pageId="5" pageNumber="598">5K</figureCitation>
)
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C354E7E2FF84664E99EF79593B8FFD1F" pageId="5" pageNumber="598" type="materials_examined">
<paragraph id="8BF1B469FF84664E99EF795939F5FD84" blockId="5.[151,1436,498,2012]" box="[151,971,533,556]" pageId="5" pageNumber="598">
<materialsCitation id="3B26BE34FF84664E99EF795939F5FD84" ID-GBIF-Occurrence="1821772634" box="[151,971,533,556]" country="Zimbabwe" location="Hwange" pageId="5" pageNumber="598" specimenCount="1" stateProvince="Matabeleland North Province" typeStatus="holotype">
<emphasis id="B93A687BFF84664E99EF79593B10FD84" bold="true" box="[151,302,533,556]" pageId="5" pageNumber="598">
<typeStatus id="54F50ACBFF84664E99EF79593AF1FD84" box="[151,207,533,556]" pageId="5" pageNumber="598">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="F359F4F9FF84664E984C79593B97FD84" box="[308,425,533,556]" name="Zimbabwe" pageId="5" pageNumber="598">Zimbabwe</collectingCountry>
, [
<collectingRegion id="498A7A8BFF84664E98C1795938C2FD84" box="[441,764,533,556]" country="Zimbabwe" name="Matabeleland North" pageId="5" pageNumber="598">
Matabeleland
<collectingRegion id="498A7A8BFF84664E9B2B795938C2FD84" box="[595,764,533,556]" inferredBySuffix="true" pageId="5" pageNumber="598">North Province</collectingRegion>
</collectingRegion>
],
<location id="8E91E2B2FF84664E9A7579593956FD84" LSID="urn:lsid:plazi:treatment:03E7057FFF84664D99EF7ABE386BFE91:8E91E2B2FF84664E9A7579593956FD84" box="[781,872,533,556]" country="Zimbabwe" name="Hwange" pageId="5" pageNumber="598" stateProvince="Matabeleland North Province">Hwange</location>
, Lupane.
</materialsCitation>
</paragraph>
<paragraph id="8BF1B469FF84664E99BF79743802FDD9" blockId="5.[151,1436,498,2012]" pageId="5" pageNumber="598">
<emphasis id="B93A687BFF84664E99BF79743B52FDE7" bold="true" box="[199,364,568,591]" pageId="5" pageNumber="598">
<typeStatus id="54F50ACBFF84664E99BF79743AC1FDE7" box="[199,255,568,591]" pageId="5" pageNumber="598">Type</typeStatus>
material.
</emphasis>
<typeStatus id="54F50ACBFF84664E980A79743BEBFDE7" box="[370,469,568,591]" pageId="5" pageNumber="598" type="holotype">Holotype</typeStatus>
and two
<typeStatus id="54F50ACBFF84664E9B4F797438A3FDE7" box="[567,669,568,591]" pageId="5" pageNumber="598" type="paratype">paratypes</typeStatus>
(all males): “
<collectingCountry id="F359F4F9FF84664E9A5379743981FDE7" box="[811,959,568,591]" name="Zimbabwe" pageId="5" pageNumber="598">ZIMBABWE</collectingCountry>
W | SE Hwange |
<quantity id="4CB6198CFF84664E9DF879743EFDFDE7" box="[1152,1219,568,591]" metricMagnitude="4" metricUnit="m" metricValue="4.0" pageId="5" pageNumber="598" unit="km" value="40.0">40 km</quantity>
NW of Lupane |
<date id="FFF092A9FF84664E9C0279743ACDFDD9" pageId="5" pageNumber="598" value="2014-11-15">15. 11. 2014</date>
Snížek [white label, printed]”.
</paragraph>
<paragraph id="8BF1B469FF84664E99BF79313B8FFD1F" blockId="5.[151,1436,498,2012]" pageId="5" pageNumber="598">
<materialsCitation id="3B26BE34FF84664E99BF793139E1FD3C" ID-GBIF-Occurrence="1821772640" box="[199,991,637,660]" country="Czech Republic" location="Praha" pageId="5" pageNumber="598" specimenCount="2" stateProvince="Praha" typeStatus="holotype">
<emphasis id="B93A687BFF84664E99BF79313B41FD3C" bold="true" box="[199,383,637,660]" pageId="5" pageNumber="598">
<typeStatus id="54F50ACBFF84664E99BF79313AC1FD3C" box="[199,255,637,660]" pageId="5" pageNumber="598">Type</typeStatus>
depository
</emphasis>
.
<typeStatus id="54F50ACBFF84664E98F779313BCCFD3C" box="[399,498,637,660]" pageId="5" pageNumber="598" type="holotype">Holotype</typeStatus>
in Národní muzeum
<collectingRegion id="498A7A8BFF84664E9B9A7931391CFD3C" box="[738,802,637,660]" country="Czech Republic" name="Praha" pageId="5" pageNumber="598">Praha</collectingRegion>
,
<collectingCountry id="F359F4F9FF84664E9A57793139E1FD3C" box="[815,991,637,660]" name="Czech Republic" pageId="5" pageNumber="598">Czech Republic</collectingCountry>
</materialsCitation>
,
<materialsCitation id="3B26BE34FF84664E9A9479313B93FD1F" ID-GBIF-Occurrence="1821772637" country="Czech Republic" location="Richard Sehnal" municipality="Velenice" pageId="5" pageNumber="598" specimenCount="1" typeStatus="paratype">
<typeStatus id="54F50ACBFF84664E9A9479313E6CFD3C" box="[1004,1106,637,660]" pageId="5" pageNumber="598" type="paratype">paratypes</typeStatus>
in
<location id="8E91E2B2FF84664E9D0379313F1CFD3C" LSID="urn:lsid:plazi:treatment:03E7057FFF84664D99EF7ABE386BFE91:8E91E2B2FF84664E9D0379313F1CFD3C" box="[1147,1314,637,660]" country="Czech Republic" municipality="Velenice" name="Richard Sehnal" pageId="5" pageNumber="598">Richard Sehnal</location>
collection,
<collectingMunicipality id="6B952E13FF84664E99EF79EC3AC8FD1F" box="[151,246,672,695]" pageId="5" pageNumber="598">Velenice</collectingMunicipality>
,
<collectingCountry id="F359F4F9FF84664E987879EC3B93FD1F" box="[256,429,672,695]" name="Czech Republic" pageId="5" pageNumber="598">Czech Republic</collectingCountry>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C354E7E2FF84664E99BF798E3BA6F8F9" pageId="5" pageNumber="598" type="description">
<paragraph id="8BF1B469FF84664E99BF798E383DFCE9" blockId="5.[151,1436,498,2012]" pageId="5" pageNumber="598">
<emphasis id="B93A687BFF84664E99BF798E3815FD71" bold="true" box="[199,555,706,729]" pageId="5" pageNumber="598">
Description of
<typeStatus id="54F50ACBFF84664E980A798E3BEAFD71" box="[370,468,706,729]" pageId="5" pageNumber="598" type="holotype">holotype</typeStatus>
(male).
</emphasis>
Body length
<quantity id="4CB6198CFF84664E9BB9798E3926FD71" box="[705,792,706,729]" metricMagnitude="-3" metricUnit="m" metricValue="7.1" pageId="5" pageNumber="598" unit="mm" value="7.1">7.1 mm</quantity>
. Body elongate. Head, pronotum and scutellum black (
<figureCitation id="1375A8ECFF84664E9C08798E3A89FD54" captionStart="FIGURES 1" captionStartId="1.[151,258,1690,1712]" captionTargetBox="[184,1405,218,1654]" captionTargetId="figure@1.[151,1435,191,1668]" captionTargetPageId="1" captionText="FIGURES 1AD. Habitus, dorsal and lateral views. A, Oedanomerus bidentatus Sehnal, new species, holotype male; B, Oedanomerus lupanus Sehnal, new species, holotype male; C, Oedanomerus squamosus Lacroix, 2005, holotype male; D, Oedanomerus uhligi Lacroix,2005,holotype male.Scalebar = 1 mm." httpUri="https://zenodo.org/record/1165784/files/figure.png" pageId="5" pageNumber="598">Fig. 1B</figureCitation>
); antennae bicolorous, antennal club pale brown, antennal shaft black (
<figureCitation id="1375A8ECFF84664E9AB479A93E1CFD54" box="[972,1058,741,764]" captionStart="FIGURES 2" captionStartId="2.[151,258,1799,1821]" captionTargetBox="[167,1420,594,1757]" captionTargetId="figure@2.[151,1436,573,1778]" captionTargetPageId="2" captionText="FIGURES 2AL. Antenna (AD), clypeus (EH) and pronotal disc (IL). A, E, I, Oedanomerus bidentatus Sehnal, new species, holotype male; B, F, J, Oedanomerus lupanus Sehnal, new species, holotype male; C, G, K, Oedanomerus squamosus Lacroix, 2005, holotype male; D, H,L, Oedanomerusuhligi Lacroix,2005, holotype male.Scale bar = 1 mm." httpUri="https://zenodo.org/record/1165786/files/figure.png" pageId="5" pageNumber="598">Fig. 2B</figureCitation>
); elytra chesnut brown with black suture and humeral umbones (
<figureCitation id="1375A8ECFF84664E98A278443813FCB7" box="[474,557,776,799]" captionStart="FIGURES 1" captionStartId="1.[151,258,1690,1712]" captionTargetBox="[184,1405,218,1654]" captionTargetId="figure@1.[151,1435,191,1668]" captionTargetPageId="1" captionText="FIGURES 1AD. Habitus, dorsal and lateral views. A, Oedanomerus bidentatus Sehnal, new species, holotype male; B, Oedanomerus lupanus Sehnal, new species, holotype male; C, Oedanomerus squamosus Lacroix, 2005, holotype male; D, Oedanomerus uhligi Lacroix,2005,holotype male.Scalebar = 1 mm." httpUri="https://zenodo.org/record/1165784/files/figure.png" pageId="5" pageNumber="598">Fig. 1B</figureCitation>
). Pro-, meso- and metatibiae brown, protarsi testaceous, meso- and metatarsi dark brown. Abdomen brownish black.
</paragraph>
<paragraph id="8BF1B469FF84664E99BF78013B67FC64" blockId="5.[151,1436,498,2012]" pageId="5" pageNumber="598">
Head. Clypeus pointed; clypeal carina broad, prominent; clypeal plate vertical; shallowly punctate; each puncture bearing a long, erect, posteriorly inclined, yellow macrosetae (
<figureCitation id="1375A8ECFF84664E9A37783C3998FC2F" box="[847,934,880,903]" captionStart="FIGURES 2" captionStartId="2.[151,258,1799,1821]" captionTargetBox="[167,1420,594,1757]" captionTargetId="figure@2.[151,1436,573,1778]" captionTargetPageId="2" captionText="FIGURES 2AL. Antenna (AD), clypeus (EH) and pronotal disc (IL). A, E, I, Oedanomerus bidentatus Sehnal, new species, holotype male; B, F, J, Oedanomerus lupanus Sehnal, new species, holotype male; C, G, K, Oedanomerus squamosus Lacroix, 2005, holotype male; D, H,L, Oedanomerusuhligi Lacroix,2005, holotype male.Scale bar = 1 mm." httpUri="https://zenodo.org/record/1165786/files/figure.png" pageId="5" pageNumber="598">Fig. 2B</figureCitation>
). Labrum reduced, triangular; lobes rounded. Frons deeply rugose; edge of clypeus strongly punctate; each puncture bearing a long, semierect, posteriorly inclined, yellow macroseta.
</paragraph>
<paragraph id="8BF1B469FF84664E99BF78943F75FBD1" blockId="5.[151,1436,498,2012]" pageId="5" pageNumber="598">
Eyes large, exceeding genae externally in dorsal view. Genae rugopunctate, with group of long macrosetae. Antennae (
<figureCitation id="1375A8ECFF84664E987678B63B5FFBB9" box="[270,353,1018,1041]" captionStart="FIGURES 2" captionStartId="2.[151,258,1799,1821]" captionTargetBox="[167,1420,594,1757]" captionTargetId="figure@2.[151,1436,573,1778]" captionTargetPageId="2" captionText="FIGURES 2AL. Antenna (AD), clypeus (EH) and pronotal disc (IL). A, E, I, Oedanomerus bidentatus Sehnal, new species, holotype male; B, F, J, Oedanomerus lupanus Sehnal, new species, holotype male; C, G, K, Oedanomerus squamosus Lacroix, 2005, holotype male; D, H,L, Oedanomerusuhligi Lacroix,2005, holotype male.Scale bar = 1 mm." httpUri="https://zenodo.org/record/1165786/files/figure.png" pageId="5" pageNumber="598">Fig. 2B</figureCitation>
) with eight antennomeres; antennomeres 68 forming long, straight club at least 1.7x times longer than antennal shaft; antennomeres 15 with sparse, long macrosetae; antennal club completely densely punctate, without smooth areas. Pedicel bulbous and as long as antennomere 3. Terminal maxillary palpomere elongate, longer than palpomeres 2 and 3 combined; sub-basally with apically pointed, flat, oval alutaceous area tapering toward apex.
</paragraph>
<paragraph id="8BF1B469FF84664E99BF7FC93949FAAC" blockId="5.[151,1436,498,2012]" pageId="5" pageNumber="598">
Pronotum (
<figureCitation id="1375A8ECFF84664E98467FC93BAEFB34" box="[318,400,1157,1180]" captionStart="FIGURES 2" captionStartId="2.[151,258,1799,1821]" captionTargetBox="[167,1420,594,1757]" captionTargetId="figure@2.[151,1436,573,1778]" captionTargetPageId="2" captionText="FIGURES 2AL. Antenna (AD), clypeus (EH) and pronotal disc (IL). A, E, I, Oedanomerus bidentatus Sehnal, new species, holotype male; B, F, J, Oedanomerus lupanus Sehnal, new species, holotype male; C, G, K, Oedanomerus squamosus Lacroix, 2005, holotype male; D, H,L, Oedanomerusuhligi Lacroix,2005, holotype male.Scale bar = 1 mm." httpUri="https://zenodo.org/record/1165786/files/figure.png" pageId="5" pageNumber="598">Fig. 2B</figureCitation>
) weakly convex, approximately tetragonal, 1.28x times wider than long, widest approximately at middle; margin nearly completely bordered, anterior border broadly interrupted medially. Anterior and posterior angles broadly rounded, poorly defined. Surface with very long, yellow macrosetae, basal border with white macrosetae; disc shallowly punctate and wrinkled, posterior angles without punctures.
</paragraph>
<paragraph id="8BF1B469FF84664E99BF7E5C3B7DFAE1" blockId="5.[151,1436,498,2012]" pageId="5" pageNumber="598">Scutellum slightly longer than wide, matte, completly shallowly punctate, with long, fine macorseta, margins broadly arcuate.</paragraph>
<paragraph id="8BF1B469FF84664E99BF7E193E67FA19" blockId="5.[151,1436,498,2012]" pageId="5" pageNumber="598">
Elytra moderately convex, narrow. Disc wrinkled and punctate; punctures evenly distributed, separated by more than
<date id="FFF092A9FF84664E99EF7E343A8CFA27" box="[151,178,1400,1423]" pageId="5" pageNumber="598">2x</date>
the puncture diameter; macrosetae short and denser than on pronotum, inclined posteriorly, absent from distinct humeri. Sides punctate, with lateral margins and apices diffusely lighter-colored than disc.
</paragraph>
<paragraph id="8BF1B469FF84664E99BF7EF13E34F929" blockId="5.[151,1436,498,2012]" pageId="5" pageNumber="598">
<emphasis id="B93A687BFF84664E99BF7EF13B50FA7C" bold="true" box="[199,366,1469,1492]" pageId="5" pageNumber="598">Macropterous.</emphasis>
Legs. All femora shiny and irregularly, coarsely punctate; yellow macrosetae short and white macrosetae long. Protibia tridentate (
<figureCitation id="1375A8ECFF84664E9B3A7EAC38A2FA5F" box="[578,668,1504,1527]" captionStart="FIGURES4" captionStartId="4.[151,258,1795,1817]" captionTargetBox="[177,1409,204,1754]" captionTargetId="figure@4.[151,1436,191,1774]" captionTargetPageId="4" captionText="FIGURES4AH. Metafemur withmetatibia (AD)and protibia(EF).A, E, Oedanomerus bidentatusSehnal,new species, holotype male; B, F, Oedanomerus lupanus Sehnal, new species,holotype male; C, G, Oedanomerus squamosus Lacroix,2005,holotype male; D, H,Oedanomerus uhligi Lacroix,2005,holotype male.Scale bar = 1 mm." httpUri="https://zenodo.org/record/1165790/files/figure.png" pageId="5" pageNumber="598">Fig. 4F</figureCitation>
). Claws simple. Metafemora chesnut brown with intermixed long macrosetae and long macrosetae. Mesotibiae moderately expanded apically. Metatibiae black, strongly expanded apically, with one oblique carina externally; carina and apical edge with row of stout, yellow macrosetae; terminal calcars stout, long, lower calcar longer than upper calcar (
<figureCitation id="1375A8ECFF84664E9A757D04395DF9F7" box="[781,867,1608,1631]" captionStart="FIGURES4" captionStartId="4.[151,258,1795,1817]" captionTargetBox="[177,1409,204,1754]" captionTargetId="figure@4.[151,1436,191,1774]" captionTargetPageId="4" captionText="FIGURES4AH. Metafemur withmetatibia (AD)and protibia(EF).A, E, Oedanomerus bidentatusSehnal,new species, holotype male; B, F, Oedanomerus lupanus Sehnal, new species,holotype male; C, G, Oedanomerus squamosus Lacroix,2005,holotype male; D, H,Oedanomerus uhligi Lacroix,2005,holotype male.Scale bar = 1 mm." httpUri="https://zenodo.org/record/1165790/files/figure.png" pageId="5" pageNumber="598">Fig. 4B</figureCitation>
). Pro- and mesotarsomeres without patches of short, dense macrosetae; metatarsomeres ventrally covered with long, sparse macrosetae.
</paragraph>
<paragraph id="8BF1B469FF84664E99BF7DC13893F96F" blockId="5.[151,1436,498,2012]" pageId="5" pageNumber="598">Abdominal sternites brownish black, with yellowish-white, recumbent macrosetae. Pygidium chesnut brown, flat, finely punctate, with very long, white macrosetae.</paragraph>
<paragraph id="8BF1B469FF84664E99BF7D9E39E5F941" blockId="5.[151,1436,498,2012]" box="[199,987,1746,1769]" pageId="5" pageNumber="598">
Male genitalia (
<figureCitation id="1375A8ECFF84664E98177D9E3BFCF941" box="[367,450,1746,1769]" captionStart="FIGURES 3" captionStartId="3.[151,258,1941,1963]" captionTargetBox="[168,1415,569,1876]" captionTargetId="figure@3.[150,1436,548,1898]" captionTargetPageId="3" captionText="FIGURES 3AD. Aedeagus, dorsal (left) and lateral (right) views. A, Oedanomerus bidentatus Sehnal, new species, holotype male; B, Oedanomerus lupanus Sehnal, new species, holotype male; C, Oedanomerus squamosus Lacroix, 2005, holotype male; D, Oedanomerus uhligi Lacroix,2005,holotype male.Scalebar = 1 mm." httpUri="https://zenodo.org/record/1165788/files/figure.png" pageId="5" pageNumber="598">Fig. 3B</figureCitation>
). Aedeagus symmetrical, parameres slender, long.
</paragraph>
<paragraph id="8BF1B469FF84664E99BF7DB93E37F887" blockId="5.[151,1436,498,2012]" pageId="5" pageNumber="598">
<emphasis id="B93A687BFF84664E99BF7DB93B87F8A4" bold="true" box="[199,441,1781,1804]" pageId="5" pageNumber="598">Variability in males.</emphasis>
<typeStatus id="54F50ACBFF84664E98BE7DB93810F8A4" box="[454,558,1781,1804]" pageId="5" pageNumber="598" type="paratype">Paratypes</typeStatus>
somewhat variable in body length (
<quantity id="4CB6198CFF84664E9AAB7DB93E61F8A4" box="[979,1119,1781,1804]" metricMagnitude="-3" metricUnit="m" metricValue="7.35" metricValueMax="7.4" metricValueMin="7.3" pageId="5" pageNumber="598" unit="mm" value="7.35" valueMax="7.4" valueMin="7.3">7.37.4 mm</quantity>
), slightly variable in dorsal punctation density and length and distribution of macrosetae. Color as in
<typeStatus id="54F50ACBFF84664E9ADC7C543E3BF887" box="[932,1029,1816,1839]" pageId="5" pageNumber="598" type="holotype">holotype</typeStatus>
.
</paragraph>
<paragraph id="8BF1B469FF84664E99BF7C763BA6F8F9" blockId="5.[151,1436,498,2012]" box="[199,408,1850,1873]" pageId="5" pageNumber="598">
<emphasis id="B93A687BFF84664E99BF7C763B1FF8F9" bold="true" box="[199,289,1850,1873]" pageId="5" pageNumber="598">Female.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="C354E7E2FF84664D99BF7C11386BFE91" lastPageId="6" lastPageNumber="599" pageId="5" pageNumber="598" type="diagnosis">
<paragraph id="8BF1B469FF84664D99BF7C11386BFE91" blockId="5.[151,1436,498,2012]" lastBlockId="6.[151,1435,152,313]" lastPageId="6" lastPageNumber="599" pageId="5" pageNumber="598">
<emphasis id="B93A687BFF84664E99BF7C113B02F8DC" bold="true" box="[199,316,1885,1908]" pageId="5" pageNumber="598">Diagnosis.</emphasis>
Based on characters stated in
<bibRefCitation id="EFDFC998FF84664E9BF97C11392DF8DC" author="Evans" box="[641,787,1885,1908]" pageId="5" pageNumber="598" refString="Evans, A. V. (1987) Revision of the genus Oedanomerus Waterhouse (Coleoptera: Scarabaeidae: Melolonthinae). Journal of the Entomological Society of Southern Africa, 50, 357 - 362." type="journal article" year="1987">Evans (1987)</bibRefCitation>
, the new species differs from its congeners by the following combination of characters: smaller and narrow species; antennae with eight antennomeres; tridentate protibia; clypeus pointed apically; clypeal carina broad, prominent; posterior pronotal angles without punctures, basal border with very long, white macrosetae; head and pronotum black, elytra chestnut brown with black suture and humeral umbones; metafemur brown. Generally, the new species is similar to
<taxonomicName id="4C4ECFEAFF87664D9A527BD439A7FF07" authorityName="Frey" authorityYear="1960" box="[810,921,152,175]" class="Insecta" family="Scarabaeidae" genus="Oedanomerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="599" phylum="Arthropoda" rank="species" species="pilosus">
<emphasis id="B93A687BFF87664D9A527BD43903FF07" box="[810,829,152,175]" italics="true" pageId="6" pageNumber="599">O</emphasis>
.
<emphasis id="B93A687BFF87664D9A367BD439A7FF07" box="[846,921,152,175]" italics="true" pageId="6" pageNumber="599">pilosus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C4ECFEAFF87664D9AAC7BD43E59FF07" authority="Sehnal, 2018" authorityName="Sehnal" authorityYear="2018" box="[980,1127,152,175]" class="Insecta" family="Scarabaeidae" genus="Oedanomerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="599" phylum="Arthropoda" rank="species" species="bidentatus" status="sp. nov.">
<emphasis id="B93A687BFF87664D9AAC7BD439D9FF07" box="[980,999,152,175]" italics="true" pageId="6" pageNumber="599">O</emphasis>
.
<emphasis id="B93A687BFF87664D9A807BD43E59FF07" box="[1016,1127,152,175]" italics="true" pageId="6" pageNumber="599">bidentatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A209D500FF87664D9D0A7BD43EC7FF07" box="[1138,1273,152,175]" pageId="6" pageNumber="599" rank="species">new species</taxonomicNameLabel>
.
<taxonomicName id="4C4ECFEAFF87664D9C7E7BD43ADCFF79" authorityName="Frey" authorityYear="1960" class="Insecta" family="Scarabaeidae" genus="Oedanomerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="599" phylum="Arthropoda" rank="species" species="pilosus">
<emphasis id="B93A687BFF87664D9C7E7BD43ADCFF79" italics="true" pageId="6" pageNumber="599">Oedanomerus pilosus</emphasis>
</taxonomicName>
has head and pronotum piceous; anterior clypeal margin broadly rounded, clypeal subapical carina long; smaller body; and different shape of aedeagus (
<bibRefCitation id="EFDFC998FF87664D9B247B9138DAFF5C" author="Evans" box="[604,740,221,244]" pageId="6" pageNumber="599" refString="Evans, A. V. (1987) Revision of the genus Oedanomerus Waterhouse (Coleoptera: Scarabaeidae: Melolonthinae). Journal of the Entomological Society of Southern Africa, 50, 357 - 362." type="journal article" year="1987">Evans 1987</bibRefCitation>
).
<taxonomicName id="4C4ECFEAFF87664D9B857B913E30FF5C" authority="Sehnal, 2018" authorityName="Sehnal" authorityYear="2018" box="[765,1038,221,244]" class="Insecta" family="Scarabaeidae" genus="Oedanomerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="599" phylum="Arthropoda" rank="species" species="bidentatus" status="sp. nov.">
<emphasis id="B93A687BFF87664D9B857B913E30FF5C" box="[765,1038,221,244]" italics="true" pageId="6" pageNumber="599">Oedanomerus bidentatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A209D500FF87664D9D637B913E9FFF5C" box="[1051,1185,221,244]" pageId="6" pageNumber="599" rank="species">new species</taxonomicNameLabel>
has elytra black with brownish-black margins; narrower clypeus; bidentate protibia; smaller body; and differently shaped aedeagus and antennal club (
<figureCitation id="1375A8ECFF87664D984C7A6E3BACFE91" box="[308,402,290,313]" captionStart="FIGURES 1" captionStartId="1.[151,258,1690,1712]" captionTargetBox="[184,1405,218,1654]" captionTargetId="figure@1.[151,1435,191,1668]" captionTargetPageId="1" captionText="FIGURES 1AD. Habitus, dorsal and lateral views. A, Oedanomerus bidentatus Sehnal, new species, holotype male; B, Oedanomerus lupanus Sehnal, new species, holotype male; C, Oedanomerus squamosus Lacroix, 2005, holotype male; D, Oedanomerus uhligi Lacroix,2005,holotype male.Scalebar = 1 mm." httpUri="https://zenodo.org/record/1165784/files/figure.png" pageId="6" pageNumber="599">Figs. 1A</figureCitation>
,
<figureCitation id="1375A8ECFF87664D98E67A6E3BFFFE91" box="[414,449,290,313]" captionStart="FIGURES 2" captionStartId="2.[151,258,1799,1821]" captionTargetBox="[167,1420,594,1757]" captionTargetId="figure@2.[151,1436,573,1778]" captionTargetPageId="2" captionText="FIGURES 2AL. Antenna (AD), clypeus (EH) and pronotal disc (IL). A, E, I, Oedanomerus bidentatus Sehnal, new species, holotype male; B, F, J, Oedanomerus lupanus Sehnal, new species, holotype male; C, G, K, Oedanomerus squamosus Lacroix, 2005, holotype male; D, H,L, Oedanomerusuhligi Lacroix,2005, holotype male.Scale bar = 1 mm." httpUri="https://zenodo.org/record/1165786/files/figure.png" pageId="6" pageNumber="599">2A</figureCitation>
,
<figureCitation id="1375A8ECFF87664D98B47A6E3BD1FE91" box="[460,495,290,313]" captionStart="FIGURES 3" captionStartId="3.[151,258,1941,1963]" captionTargetBox="[168,1415,569,1876]" captionTargetId="figure@3.[150,1436,548,1898]" captionTargetPageId="3" captionText="FIGURES 3AD. Aedeagus, dorsal (left) and lateral (right) views. A, Oedanomerus bidentatus Sehnal, new species, holotype male; B, Oedanomerus lupanus Sehnal, new species, holotype male; C, Oedanomerus squamosus Lacroix, 2005, holotype male; D, Oedanomerus uhligi Lacroix,2005,holotype male.Scalebar = 1 mm." httpUri="https://zenodo.org/record/1165788/files/figure.png" pageId="6" pageNumber="599">3A</figureCitation>
,
<figureCitation id="1375A8ECFF87664D98827A6E3874FE91" box="[506,586,290,313]" captionStart="FIGURES4" captionStartId="4.[151,258,1795,1817]" captionTargetBox="[177,1409,204,1754]" captionTargetId="figure@4.[151,1436,191,1774]" captionTargetPageId="4" captionText="FIGURES4AH. Metafemur withmetatibia (AD)and protibia(EF).A, E, Oedanomerus bidentatusSehnal,new species, holotype male; B, F, Oedanomerus lupanus Sehnal, new species,holotype male; C, G, Oedanomerus squamosus Lacroix,2005,holotype male; D, H,Oedanomerus uhligi Lacroix,2005,holotype male.Scale bar = 1 mm." httpUri="https://zenodo.org/record/1165790/files/figure.png" pageId="6" pageNumber="599">4A, 4E</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>