285 lines
33 KiB
XML
285 lines
33 KiB
XML
<document id="484B42156601DC933083B8067AD09637" ID-DOI="10.5281/zenodo.199558" ID-GBIF-Dataset="7b7b9345-9632-469e-a881-e8d6403e3f4e" ID-ISSN="1175-5326" ID-Zenodo-Dep="199558" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460339995128" checkinUser="plazi" docAuthor="Campos, Ernesto & Hernández-Ávila, Iván" docDate="2010" docId="D51D87C0FFC3C7624B9C5739FC6EDCBF" docLanguage="en" docName="zt02691p052.pdf" docOrigin="Zootaxa 2691" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Calyptraeotheres pepeluisi Campos & Hernández-Ávila, 2010, new species" docType="treatment" docVersion="9" lastPageNumber="50" masterDocId="2924FFB8FFC7C76B4B0B503BFFD8D973" masterDocTitle="Phylogeny of Calyptraeotheres Campos, 1990 (Crustacea, Decapoda, Brachyura, Pinnotheridae) with the description of C. pepeluisi new species from the tropical Mexican Pacific" masterLastPageNumber="52" masterPageNumber="41" pageNumber="45" updateTime="1698260868845" updateUser="plazi">
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<mods:title id="87D247E93A2959889F28F88E29E9742B">Phylogeny of Calyptraeotheres Campos, 1990 (Crustacea, Decapoda, Brachyura, Pinnotheridae) with the description of C. pepeluisi new species from the tropical Mexican Pacific</mods:title>
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<mods:namePart id="0A4F310BC5AE3CABFD8338A653641169">Campos, Ernesto</mods:namePart>
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<mods:roleTerm id="2EC0812843E7EDDDA27275DECFC80843">Author</mods:roleTerm>
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<mods:namePart id="0F81F9CFFB24F1CBF7CD5B55242B7945">Hernández-Ávila, Iván</mods:namePart>
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<mods:typeOfResource id="E1FE094B37B2D02B75EC970189DF8611">text</mods:typeOfResource>
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<mods:title id="FD3221553F6EC383EA0162CD45E329EB">Zootaxa</mods:title>
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<mods:part id="06F45F4242A4C88DFC5E63C4045897BF">
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<mods:date id="ED24BA0CD9F1AB21F446E20C4DF4E501">2010</mods:date>
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<mods:detail id="9BCC6A8CC36F661712CC0A0F96E2DA59" type="volume">
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<mods:number id="9EBC06FDA64CF6D5E86BA6FEB870A66A">2691</mods:number>
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<mods:identifier id="446E6E9587570436F762124D77A68853" type="DOI">10.5281/zenodo.199558</mods:identifier>
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<mods:identifier id="014A169C1E1B7217A2C4206F4FE15155" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="799B765DC377AD37DD94BFA1591A3B18" type="Zenodo-Dep">199558</mods:identifier>
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<treatment id="D51D87C0FFC3C7624B9C5739FC6EDCBF" ID-DOI="http://doi.org/10.5281/zenodo.6208225" ID-GBIF-Taxon="119406312" ID-Zenodo-Dep="6208225" LSID="urn:lsid:plazi:treatment:D51D87C0FFC3C7624B9C5739FC6EDCBF" httpUri="http://treatment.plazi.org/id/D51D87C0FFC3C7624B9C5739FC6EDCBF" lastPageId="9" lastPageNumber="50" pageId="4" pageNumber="45">
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<subSubSection id="15AE655DFFC3C76F4B9C5739FEC0DE4C" pageId="4" pageNumber="45" type="nomenclature">
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<paragraph id="5D0B36D6FFC3C76F4B9C5739FDB5DE6F" blockId="4.[151,621,1794,1855]" box="[151,621,1794,1820]" pageId="4" pageNumber="45">
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<heading id="064381BAFFC3C76F4B9C5739FDB5DE6F" bold="true" box="[151,621,1794,1820]" fontSize="11" level="1" pageId="4" pageNumber="45" reason="1">
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<emphasis id="6FC0EAC4FFC3C76F4B9C5739FDB5DE6F" bold="true" box="[151,621,1794,1820]" pageId="4" pageNumber="45">
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<taxonomicName id="9AB44D55FFC3C76F4B9C5739FE0EDE6F" ID-CoL="Q77X" box="[151,470,1794,1820]" class="Malacostraca" family="Pinnotheridae" genus="Calyptraeotheres" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="45" phylum="Arthropoda" rank="species" species="pepeluisi" status="sp. nov.">
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<emphasis id="6FC0EAC4FFC3C76F4B9C5739FE0EDE6F" bold="true" box="[151,470,1794,1820]" italics="true" pageId="4" pageNumber="45">Calyptraeotheres pepeluisi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="74F357BFFFC3C76F4AD55739FDB5DE6F" box="[478,621,1794,1820]" pageId="4" pageNumber="45" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="5D0B36D6FFC3C76F4B9C571EFEC0DE4C" blockId="4.[151,621,1794,1855]" box="[151,280,1829,1855]" pageId="4" pageNumber="45">
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(
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<figureCitation id="C58F2A53FFC3C76F4B94571EFF2ADE4C" box="[159,242,1829,1855]" captionStart="FIGURE 3" captionStartId="7.[151,257,1740,1764]" captionTargetBox="[155,1423,181,1739]" captionTargetId="figure@7.[151,1436,175,1761]" captionTargetPageId="7" captionText="FIGURE 3. Calyptraeotheres pepeluisi new species, off Lázaro Cárdenas port, Michoacán, Mexico (CNCR 26183). Adult female: A, dorsal view; B, partial frontal view." httpUri="https://zenodo.org/record/199561/files/figure.png" pageId="4" pageNumber="45">Figs. 3</figureCitation>
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,
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<figureCitation id="C58F2A53FFC3C76F4BF4571EFED7DE4C" box="[255,271,1829,1855]" captionStart="FIGURE 4" captionStartId="8.[151,257,1927,1951]" captionTargetBox="[264,1308,199,1893]" captionTargetId="figure@8.[255,1323,194,1898]" captionTargetPageId="8" captionText="FIGURE 4. Calyptraeotheres pepeluisi new species, off Lázaro Cárdenas port, Michoacán, Mexico (CNCR 26183). Adult female: A, abdomen, somites 3 – 6 and telson; B – C, MXP 3, inner and outer views respectively; D, cheliped; E, tip of cheliped." httpUri="https://zenodo.org/record/199562/files/figure.png" pageId="4" pageNumber="45">4</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="15AE655DFFC3C76F4B9C5751FE52DE87" pageId="4" pageNumber="45" type="materials_examined">
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<paragraph id="5D0B36D6FFC3C76F4B9C5751FABFDEBC" blockId="4.[151,1435,1898,2036]" pageId="4" pageNumber="45">
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<emphasis id="6FC0EAC4FFC3C76F4B9C5751FED3DEF7" bold="true" box="[151,267,1898,1924]" pageId="4" pageNumber="45">Material.</emphasis>
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<typeStatus id="820F8874FFC3C76F4A185751FE58DEF7" box="[275,384,1898,1924]" pageId="4" pageNumber="45" type="holotype">Holotype</typeStatus>
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(
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<collectionCode id="3BA5AE13FFC3C76F4A9B5751FE3CDEF7" box="[400,484,1898,1924]" pageId="4" pageNumber="45">CNCR</collectionCode>
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26183), ovigerous female,
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<date id="290A1016FFC3C76F482B5751FC64DEF7" box="[800,956,1898,1924]" pageId="4" pageNumber="45" value="1983-07-10">10 July 1983</date>
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, collected by trawl, off Lázaro Cardenas port, Michoacán, Pacific coast of
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<collectingCountry id="25A37646FFC3C76F492B57ABFDA4DED9" box="[544,636,1936,1962]" name="Mexico" pageId="4" pageNumber="45">Mexico</collectingCountry>
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, oceanographic expedition Atlas IV on board of the R/V El Puma of the
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<collectionCode id="3BA5AE13FFC3C76F4BC9578EFEC7DEBC" box="[194,287,1973,1999]" httpUri="http://grbio.org/cool/10ei-x9nj" name="Universidad Nacional Autonoma de Mexico" pageId="4" pageNumber="45">UNAM</collectionCode>
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,
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<date id="290A1016FFC3C76F4A21578EFE7CDEBC" box="[298,420,1973,1999]" pageId="4" pageNumber="45" value="1983-06">June 1983</date>
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, locality A-IV, C4-13,
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<geoCoordinate id="38805011FFC3C76F49BB578EFD14DEBC" box="[688,716,1973,1999]" direction="north" orientation="latitude" pageId="4" pageNumber="45" precision="15" value="18.07889">18</geoCoordinate>
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º 04’44” N
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<geoCoordinate id="38805011FFC3C76F4858578EFC21DEBC" box="[851,1017,1973,1999]" direction="north" orientation="latitude" pageId="4" pageNumber="45" precision="15" value="18.083889">18º 05’ 02” N</geoCoordinate>
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,
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<geoCoordinate id="38805011FFC3C76F4F0F578EFB65DEBC" box="[1028,1213,1973,1999]" direction="west" orientation="longitude" pageId="4" pageNumber="45" precision="15" value="-102.555">
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102º 33’ 18”
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<geoCoordinate id="38805011FFC3C76F4FAA578EFB65DEBC" box="[1185,1213,1973,1999]" direction="west" orientation="longitude" pageId="4" pageNumber="45" precision="15" value="-102.55112">W</geoCoordinate>
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</geoCoordinate>
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102 º 33’ 04”.
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</paragraph>
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<paragraph id="5D0B36D6FFC3C76F4BCE57E1FE52DE87" blockId="4.[151,1435,1898,2036]" box="[197,394,2010,2036]" pageId="4" pageNumber="45">
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<emphasis id="6FC0EAC4FFC3C76F4BCE57E1FEDFDE87" bold="true" box="[197,263,2010,2036]" pageId="4" pageNumber="45">Host.</emphasis>
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Unknown.
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</paragraph>
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</subSubSection>
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<caption id="09CB665EFFC2C76E4B9C5779FF05DEAA" httpUri="https://zenodo.org/record/199559/files/figure.png" pageId="5" pageNumber="46" targetBox="[180,1406,198,1827]" targetPageId="5">
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<paragraph id="5D0B36D6FFC2C76E4B9C5779FF05DEAA" blockId="5.[151,1436,1858,2009]" pageId="5" pageNumber="46">
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<emphasis id="6FC0EAC4FFC2C76E4B9C5779FEC1DE29" bold="true" box="[151,281,1858,1882]" pageId="5" pageNumber="46">FIGURE 1.</emphasis>
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Strict consensus cladogram of 19 most parsimonious trees (length=27 CI = 0.667; RI = 0.625; RC = 0.417) obtained from heuristic parsimony analyses of unweighted and unordered characters. Subfamily name as proposed by Campos (2009) is indicated. Pinnotherid I–V and Pinnixid I–II are explained in Table 1.
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<taxonomicName id="9AB44D55FFC2C76E4F5B57B9FAB3DEEA" box="[1104,1387,1922,1945]" class="Malacostraca" family="Varunidae" genus="Asthenognathus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="46" phylum="Arthropoda" rank="species" species="atlanticus">
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<emphasis id="6FC0EAC4FFC2C76E4F5B57B9FAB3DEEA" box="[1104,1387,1922,1945]" italics="true" pageId="5" pageNumber="46">Asthenognathus atlanticus</emphasis>
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</taxonomicName>
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was used as the outgroup. Numbers above branches are the percentage of 1,000 bootstrap replicates that support the relevant nodes.
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</paragraph>
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</caption>
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<caption id="09CB665EFFC1C76D4B9C570EFB48DEBF" httpUri="https://zenodo.org/record/199560/files/figure.png" pageId="6" pageNumber="47" targetBox="[189,1377,226,1810]" targetPageId="6">
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<paragraph id="5D0B36D6FFC1C76D4B9C570EFB48DEBF" blockId="6.[151,1436,1845,1996]" pageId="6" pageNumber="47">
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<emphasis id="6FC0EAC4FFC1C76D4B9C570EFEC2DE3E" bold="true" box="[151,282,1845,1869]" pageId="6" pageNumber="47">FIGURE 2.</emphasis>
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Strict consensus cladogram of 13.78 most parsimonious trees (length=13.45 CI = 0.865; RI = 0.896; RC = 0.7751) obtained from heuristic parsimony analyses of unordered characters. The data were
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<emphasis id="6FC0EAC4FFC1C76D4F93576EFAC2DE1F" box="[1176,1306,1877,1900]" italics="true" pageId="6" pageNumber="47">a posteriori</emphasis>
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reweighted using the rescaled consistency index (RC). Subfamily name as proposed by Campos (2009) is indicated. Pinnotherid I–V and Pinnixid I–II are explained in Table 1.
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<taxonomicName id="9AB44D55FFC1C76D496757AEFBCADEDF" box="[620,1042,1941,1964]" class="Malacostraca" family="Varunidae" genus="Asthenognathus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="47" phylum="Arthropoda" rank="species" species="atlanticus">
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<emphasis id="6FC0EAC4FFC1C76D496757AEFC53DEDF" box="[620,907,1941,1964]" italics="true" pageId="6" pageNumber="47">Asthenognathus atlanticus</emphasis>
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(Varunidae)
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</taxonomicName>
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was used as the outgroup. Numbers above branches are the percentage of 1,000 bootstrap replicates that support the relevant nodes.
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</paragraph>
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</caption>
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<caption id="09CB665EFFC0C76C4B9C56F7FD14DE77" httpUri="https://zenodo.org/record/199561/files/figure.png" pageId="7" pageNumber="48" targetBox="[155,1423,181,1739]" targetPageId="7">
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<paragraph id="5D0B36D6FFC0C76C4B9C56F7FD14DE77" blockId="7.[151,1437,1740,1796]" pageId="7" pageNumber="48">
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<emphasis id="6FC0EAC4FFC0C76C4B9C56F7FEC5DF97" bold="true" box="[151,285,1740,1764]" pageId="7" pageNumber="48">FIGURE 3.</emphasis>
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<taxonomicName id="9AB44D55FFC0C76C4A2E56F6FD95DF97" box="[293,589,1741,1764]" class="Malacostraca" family="Pinnotheridae" genus="Calyptraeotheres" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="48" phylum="Arthropoda" rank="species" species="pepeluisi" status="sp. nov.">
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<emphasis id="6FC0EAC4FFC0C76C4A2E56F6FD95DF97" box="[293,589,1741,1764]" italics="true" pageId="7" pageNumber="48">Calyptraeotheres pepeluisi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="74F357BFFFC0C76C495D56F7FD06DF97" box="[598,734,1740,1764]" pageId="7" pageNumber="48" rank="species">
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<emphasis id="6FC0EAC4FFC0C76C495D56F7FD06DF97" bold="true" box="[598,734,1740,1764]" pageId="7" pageNumber="48">new species</emphasis>
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</taxonomicNameLabel>
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, off Lázaro Cárdenas port, Michoacán, Mexico (CNCR26183). Adult female: A, dorsal view; B, partial frontal view.
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</paragraph>
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||
</caption>
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<subSubSection id="15AE655DFFC0C7624BCE5709FF3CDB71" lastPageId="9" lastPageNumber="50" pageId="7" pageNumber="48" type="description">
|
||
<paragraph id="5D0B36D6FFC0C7624BCE5709FF3CDB71" blockId="7.[151,1437,1842,2018]" lastBlockId="9.[151,1438,152,1484]" lastPageId="9" lastPageNumber="50" pageId="7" pageNumber="48">
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||
<emphasis id="6FC0EAC4FFC0C76C4BCE5709FDBBDE3F" bold="true" box="[197,611,1842,1868]" pageId="7" pageNumber="48">
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||
Description of
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<typeStatus id="820F8874FFC0C76C4A815709FE23DE3F" box="[394,507,1842,1868]" pageId="7" pageNumber="48" type="holotype">holotype</typeStatus>
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female.
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</emphasis>
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Carapace suborbicular, with pilose sub-rectangular front, slightly projected, with medial shallow sulcus; anterolateral margin pilose, arcuate, cristate; regions poorly defined; posterior margin slightly curved; 2 cervical depressions from orbits to gastric region, converging posteriorly. Eyes visible in dorsal view. MXP3 placed obliquely, exopod with unsegmented flagellum; endopod with ischium and merus fused, widening distally, palp 2-segmented, articulating distally on inner margin; carpus sub-trapezoidal, larger than sub-conical propodus; dactylus absent. Chelipeds stout, dorsodistal margin of carpus with small rounded projection; chela longer than combined merus-carpus length, palm widening distally, dorsal margin unarmed, ventral margin of propodus slightly curved with fringe of minute setae, dactylus curved, cutting margin with proximal tooth inserted in notch of propodus, fingers without gape when closed, tips crossed, both fingers with additional minute teeth; outer surface of chela slightly convex, inner surface convex dorso-proximally, slightly concave distally, with ventro-proximal depression. Relative length of WL in decreasing order 3 =2>1>4, margins unarmed, WL 1–3 of similar shape, WL 4 relatively slender. Dactyl falcate, relative length 4> 3> 2> 1; dactyli shorter than propodi in WL 1–3, dactylus of WL 4 slightly larger than propodus. Abdomen with 6-somites, telson distinctly separated, covering sternum, reaching buccal cavity.
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</paragraph>
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||
</subSubSection>
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||
<caption id="09CB665EFFCFC7634B9C57BCFECDDE96" httpUri="https://zenodo.org/record/199562/files/figure.png" pageId="8" pageNumber="49" targetBox="[264,1308,199,1893]" targetPageId="8">
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||
<paragraph id="5D0B36D6FFCFC7634B9C57BCFECDDE96" blockId="8.[151,1437,1927,2021]" pageId="8" pageNumber="49">
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||
<emphasis id="6FC0EAC4FFCFC7634B9C57BCFEC5DEEC" bold="true" box="[151,285,1927,1951]" pageId="8" pageNumber="49">FIGURE 4.</emphasis>
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||
<taxonomicName id="9AB44D55FFCFC7634A2E57B3FD95DEEC" box="[293,589,1928,1951]" class="Malacostraca" family="Pinnotheridae" genus="Calyptraeotheres" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="49" phylum="Arthropoda" rank="species" species="pepeluisi" status="sp. nov.">
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||
<emphasis id="6FC0EAC4FFCFC7634A2E57B3FD95DEEC" box="[293,589,1928,1951]" italics="true" pageId="8" pageNumber="49">Calyptraeotheres pepeluisi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="6FC0EAC4FFCFC763495D57BCFD06DEEC" bold="true" box="[598,734,1927,1951]" pageId="8" pageNumber="49">
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||
<taxonomicNameLabel id="74F357BFFFCFC763495D57BCFD06DEEC" box="[598,734,1927,1951]" pageId="8" pageNumber="49" rank="species">new species</taxonomicNameLabel>
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||
</emphasis>
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||
, off Lázaro Cárdenas port, Michoacán, Mexico (CNCR26183). Adult female: A, abdomen, somites 3–6 and telson; B–C, MXP3, inner and outer views respectively; D, cheliped; E, tip of cheliped.
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||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="15AE655DFFCEC7624BCE5236FB19DB01" pageId="9" pageNumber="50" type="etymology">
|
||
<paragraph id="5D0B36D6FFCEC7624BCE5236FB19DB01" blockId="9.[151,1438,152,1484]" pageId="9" pageNumber="50">
|
||
<emphasis id="6FC0EAC4FFCEC7624BCE5236FE8ADB54" bold="true" box="[197,338,525,551]" pageId="9" pageNumber="50">Etymology.</emphasis>
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This species is named in honor of our colleague and friend Pepe-Luis, the nickname of José Luis Villalobos-Hiriart (CNCR, Instituto de Biología, UNAM) for his numerous contributions to systematics and phylogeny of
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<taxonomicName id="9AB44D55FFCEC7624A605263FE38DB01" box="[363,480,600,626]" class="Malacostraca" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="50" phylum="Arthropoda" rank="order">Decapoda</taxonomicName>
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and his invaluable support to our studies on pinnotherid crabs.
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</paragraph>
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</subSubSection>
|
||
<subSubSection id="15AE655DFFCEC7624BCE5246FC6EDCBF" pageId="9" pageNumber="50" type="materials_examined">
|
||
<paragraph id="5D0B36D6FFCEC7624BCE5246FD9DDBCF" blockId="9.[151,1438,152,1484]" pageId="9" pageNumber="50">
|
||
<emphasis id="6FC0EAC4FFCEC7624BCE5246FD82DBE4" bold="true" box="[197,602,637,663]" pageId="9" pageNumber="50">
|
||
<typeStatus id="820F8874FFCEC7624BCE5246FEDEDBE4" box="[197,262,637,663]" pageId="9" pageNumber="50">Type</typeStatus>
|
||
locality and distribution.
|
||
</emphasis>
|
||
Known only from the
|
||
<typeStatus id="820F8874FFCEC76248765246FC6BDBE4" box="[893,947,637,663]" pageId="9" pageNumber="50">type</typeStatus>
|
||
locality, off the Lázaro Cárdenas port, Michoacán, Pacific coast of
|
||
<collectingCountry id="25A37646FFCEC7624AE85299FD99DBCF" box="[483,577,674,700]" name="Mexico" pageId="9" pageNumber="50">Mexico</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="5D0B36D6FFCEC7624BCE52FCFC6EDCBF" blockId="9.[151,1438,152,1484]" pageId="9" pageNumber="50">
|
||
<emphasis id="6FC0EAC4FFCEC7624BCE52FCFE1DDB92" bold="true" box="[197,453,711,737]" pageId="9" pageNumber="50">Taxonomic remarks.</emphasis>
|
||
<taxonomicName id="9AB44D55FFCEC7624AC752F3FCDFDB92" box="[460,775,712,737]" class="Malacostraca" family="Pinnotheridae" genus="Calyptraeotheres" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="50" phylum="Arthropoda" rank="species" species="pepeluisi" status="sp. nov.">
|
||
<emphasis id="6FC0EAC4FFCEC7624AC752F3FCDFDB92" box="[460,775,712,737]" italics="true" pageId="9" pageNumber="50">Calyptraeotheres pepeluisi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="74F357BFFFCEC762480552FCFC45DB92" box="[782,925,711,737]" pageId="9" pageNumber="50" rank="species">
|
||
<emphasis id="6FC0EAC4FFCEC762480552FCFC45DB92" bold="true" box="[782,925,711,737]" pageId="9" pageNumber="50">new species</emphasis>
|
||
</taxonomicNameLabel>
|
||
is the third species of the genus recorded in the Pacific Ocean. It differs from the two austral (southern hemisphere) species,
|
||
<taxonomicName id="9AB44D55FFCEC7624F4B52D6FB72DA75" box="[1088,1194,749,774]" class="Malacostraca" family="Varunidae" genus="Neoxenophtalmus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="50" phylum="Arthropoda" rank="species" species="garthi">
|
||
<emphasis id="6FC0EAC4FFCEC7624F4B52D6FB72DA75" box="[1088,1194,749,774]" italics="true" pageId="9" pageNumber="50">C. garthi</emphasis>
|
||
</taxonomicName>
|
||
(Atlantic:
|
||
<collectingCountry id="25A37646FFCEC7624E2C52D6FA44DA74" box="[1319,1436,749,775]" name="Argentina" pageId="9" pageNumber="50">Argentina</collectingCountry>
|
||
and
|
||
<collectingCountry id="25A37646FFCEC7624BC35329FEC9DA5F" box="[200,273,786,812]" name="Brazil" pageId="9" pageNumber="50">Brazil</collectingCountry>
|
||
) and
|
||
<taxonomicName id="9AB44D55FFCEC7624A5A5329FE1BDA58" box="[337,451,786,811]" class="Malacostraca" family="Pinnotheridae" genus="Caytraeotheres" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="50" phylum="Arthropoda" rank="species" species="politus">
|
||
<emphasis id="6FC0EAC4FFCEC7624A5A5329FE1BDA58" box="[337,451,786,811]" italics="true" pageId="9" pageNumber="50">C. politus</emphasis>
|
||
</taxonomicName>
|
||
(Pacific:
|
||
<collectingCountry id="25A37646FFCEC76249385329FDB1DA5F" box="[563,617,786,812]" name="Peru" pageId="9" pageNumber="50">Peru</collectingCountry>
|
||
and
|
||
<collectingCountry id="25A37646FFCEC76249AA5329FD3ADA5F" box="[673,738,786,812]" name="Chile" pageId="9" pageNumber="50">Chile</collectingCountry>
|
||
) by its 2-segmented MXP3 palp (
|
||
<figureCitation id="C58F2A53FFCEC7624F605329FB69DA5F" box="[1131,1201,786,812]" captionStart="FIGURE 4" captionStartId="8.[151,257,1927,1951]" captionTargetBox="[264,1308,199,1893]" captionTargetId="figure@8.[255,1323,194,1898]" captionTargetPageId="8" captionText="FIGURE 4. Calyptraeotheres pepeluisi new species, off Lázaro Cárdenas port, Michoacán, Mexico (CNCR 26183). Adult female: A, abdomen, somites 3 – 6 and telson; B – C, MXP 3, inner and outer views respectively; D, cheliped; E, tip of cheliped." httpUri="https://zenodo.org/record/199562/files/figure.png" pageId="9" pageNumber="50">Fig. 4</figureCitation>
|
||
B–C) instead of a 3- segmented palp (see
|
||
<bibRefCitation id="39254B27FFCEC7624A935303FD9CDA21" author="Campos" box="[408,580,824,850]" pageId="9" pageNumber="50" refString="Campos, E. (1999) Inclusion of the austral species Pinnotheres politus (Smith, 1869) and Pinnotheres garthi Fenucci, 1975 within the genus Calyptraeotheres Campos, 1990 (Crustacea: Brachyura: Pinnotheridae). Proceedings of the Biological Society of Washington, 112, 536 - 540." type="journal article" year="1999">Campos 1999</bibRefCitation>
|
||
,
|
||
<figureCitation id="C58F2A53FFCEC76249595303FD42DA21" box="[594,666,824,850]" captionStart="FIGURE 4" captionStartId="8.[151,257,1927,1951]" captionTargetBox="[264,1308,199,1893]" captionTargetId="figure@8.[255,1323,194,1898]" captionTargetPageId="8" captionText="FIGURE 4. Calyptraeotheres pepeluisi new species, off Lázaro Cárdenas port, Michoacán, Mexico (CNCR 26183). Adult female: A, abdomen, somites 3 – 6 and telson; B – C, MXP 3, inner and outer views respectively; D, cheliped; E, tip of cheliped." httpUri="https://zenodo.org/record/199562/files/figure.png" pageId="9" pageNumber="50">Fig. 4</figureCitation>
|
||
A–B). Both
|
||
<taxonomicName id="9AB44D55FFCEC76248315303FC3EDA22" box="[826,998,824,849]" class="Malacostraca" family="Pinnotheridae" genus="Calyptraeotheres" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="50" phylum="Arthropoda" rank="species" species="hernandezi">
|
||
<emphasis id="6FC0EAC4FFCEC76248315303FC3EDA22" box="[826,998,824,849]" italics="true" pageId="9" pageNumber="50">C. hernandezi</emphasis>
|
||
</taxonomicName>
|
||
(Caribbean Sea:
|
||
<collectingCountry id="25A37646FFCEC7624FB75303FAE3DA21" box="[1212,1339,824,850]" name="Venezuela" pageId="9" pageNumber="50">Venezuela</collectingCountry>
|
||
) and
|
||
<taxonomicName id="9AB44D55FFCEC7624E8B5303FF3DDA05" class="Malacostraca" family="Pinnotheridae" genus="Calyptraeotheres" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="50" phylum="Arthropoda" rank="species" species="granti">
|
||
<emphasis id="6FC0EAC4FFCEC7624E8B5303FA4BDA22" box="[1408,1427,824,849]" italics="true" pageId="9" pageNumber="50">C</emphasis>
|
||
.
|
||
<emphasis id="6FC0EAC4FFCEC7624B9C5366FF3DDA05" box="[151,229,861,886]" italics="true" pageId="9" pageNumber="50">granti</emphasis>
|
||
</taxonomicName>
|
||
(Mexican Pacific) share a 2-segmented MXP3 palp with
|
||
<taxonomicName id="9AB44D55FFCEC76248CF5366FB85DA05" box="[964,1117,861,886]" class="Malacostraca" family="Pinnotheridae" genus="Calyptraeotheres" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="50" phylum="Arthropoda" rank="species" species="pepeluisi">
|
||
<emphasis id="6FC0EAC4FFCEC76248CF5366FB85DA05" box="[964,1117,861,886]" italics="true" pageId="9" pageNumber="50">C. pepeluisi</emphasis>
|
||
</taxonomicName>
|
||
, but this latter species is morphologically closer to the former species.
|
||
<taxonomicName id="9AB44D55FFCEC76249E153B9FB86DAE8" box="[746,1118,898,923]" class="Malacostraca" family="Pinnotheridae" genus="Calyptraeotheres" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="50" phylum="Arthropoda" rank="species" species="hernandezi">
|
||
<emphasis id="6FC0EAC4FFCEC76249E153B9FB86DAE8" box="[746,1118,898,923]" italics="true" pageId="9" pageNumber="50">Calyptraeotheres hernandezi</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="9AB44D55FFCEC7624F9553B9FAE1DAE8" box="[1182,1337,898,923]" class="Malacostraca" family="Pinnotheridae" genus="Calyptraeotheres" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="50" phylum="Arthropoda" rank="species" species="pepeluisi">
|
||
<emphasis id="6FC0EAC4FFCEC7624F9553B9FAE1DAE8" box="[1182,1337,898,923]" italics="true" pageId="9" pageNumber="50">C. pepeluisi</emphasis>
|
||
</taxonomicName>
|
||
share a subarcuate carapace, with a fringe of setae on its anterolateral and frontal margins (
|
||
<figureCitation id="C58F2A53FFCEC7624F625393FB76DAB1" box="[1129,1198,936,962]" captionStart="FIGURE 3" captionStartId="7.[151,257,1740,1764]" captionTargetBox="[155,1423,181,1739]" captionTargetId="figure@7.[151,1436,175,1761]" captionTargetPageId="7" captionText="FIGURE 3. Calyptraeotheres pepeluisi new species, off Lázaro Cárdenas port, Michoacán, Mexico (CNCR 26183). Adult female: A, dorsal view; B, partial frontal view." httpUri="https://zenodo.org/record/199561/files/figure.png" pageId="9" pageNumber="50">Fig. 3</figureCitation>
|
||
A), a conical MXP3 propodus, and dorsally visible eyes. This resemblance provides evidence that
|
||
<taxonomicName id="9AB44D55FFCEC7624F5553F6FB2ADA95" box="[1118,1266,973,998]" class="Malacostraca" family="Pinnotheridae" genus="Calyptraeotheres" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="50" phylum="Arthropoda" rank="species" species="pepeluisi">
|
||
<emphasis id="6FC0EAC4FFCEC7624F5553F6FB2ADA95" box="[1118,1266,973,998]" italics="true" pageId="9" pageNumber="50">C. pepeluisi</emphasis>
|
||
</taxonomicName>
|
||
is the Pacific counterpart of
|
||
<taxonomicName id="9AB44D55FFCEC7624A4853C9FE3FDD78" box="[323,487,1010,1035]" class="Malacostraca" family="Pinnotheridae" genus="Calyptraeotheres" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="50" phylum="Arthropoda" rank="species" species="hernandezi">
|
||
<emphasis id="6FC0EAC4FFCEC7624A4853C9FE3FDD78" box="[323,487,1010,1035]" italics="true" pageId="9" pageNumber="50">C. hernandezi</emphasis>
|
||
</taxonomicName>
|
||
and that both species probably evolved after the complete closure of the Isthmus of
|
||
<collectingCountry id="25A37646FFCEC7624BB35423FECDDD41" box="[184,277,1048,1074]" name="Panama" pageId="9" pageNumber="50">Panama</collectingCountry>
|
||
3.1–3.5 MYA (
|
||
<bibRefCitation id="39254B27FFCEC7624ADB5423FD5EDD41" author="Keigwin" box="[464,646,1048,1074]" pageId="9" pageNumber="50" refString="Keigwin, L. D. (1982) Isotopic paleoceanography of the Caribbean and east Pacific: role of Panama uplift in late Neogene time. Science, 217, 350 - 352." type="journal article" year="1982">Keigwin, 1982</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="39254B27FFCEC76249985423FC60DD41" author="Coates" box="[659,952,1048,1074]" pageId="9" pageNumber="50" refString="Coates A. G. & Obando, J. A. (1996) The geological evolution of the Central American Isthmus. In: J. B. C. Jackson, A. F. Budd & A. G. Coates (Eds.), Evolution and Environment in Tropical America. University of Chicago Press. pp. 21 - 56." type="book chapter" year="1996">Coates & Obando, 1996</bibRefCitation>
|
||
). In spite of the fact that
|
||
<taxonomicName id="9AB44D55FFCEC7624FE85423FAAADD42" box="[1251,1394,1048,1073]" class="Malacostraca" family="Pinnotheridae" genus="Calyptraeotheres" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="50" phylum="Arthropoda" rank="species" species="pepeluisi">
|
||
<emphasis id="6FC0EAC4FFCEC7624FE85423FAAADD42" box="[1251,1394,1048,1073]" italics="true" pageId="9" pageNumber="50">C. pepeluisi</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="9AB44D55FFCEC7624E8A5423FEC2DD25" class="Malacostraca" family="Pinnotheridae" genus="Calyptraeotheres" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="50" phylum="Arthropoda" rank="species" species="hernandezi">
|
||
<emphasis id="6FC0EAC4FFCEC7624E8A5423FEC2DD25" italics="true" pageId="9" pageNumber="50">C. hernandezi</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="9AB44D55FFCEC7624A5F5406FE18DD25" box="[340,448,1085,1110]" class="Malacostraca" family="Pinnotheridae" genus="Calyptraeotheres" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="50" phylum="Arthropoda" rank="species" species="granti">
|
||
<emphasis id="6FC0EAC4FFCEC7624A5F5406FE18DD25" box="[340,448,1085,1110]" italics="true" pageId="9" pageNumber="50">C. granti</emphasis>
|
||
</taxonomicName>
|
||
have a 2-segmented MXP3 palp, it is remarkable that the shape of the propodus of this appendage in the former two species (conical) is quite different from that observed in the latter species (subtrapezoidal). Furthermore, in
|
||
<taxonomicName id="9AB44D55FFCEC762492C54B3FD4ADDD2" box="[551,658,1160,1185]" class="Malacostraca" family="Pinnotheridae" genus="Calyptraeotheres" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="50" phylum="Arthropoda" rank="species" species="granti">
|
||
<emphasis id="6FC0EAC4FFCEC762492C54B3FD4ADDD2" box="[551,658,1160,1185]" italics="true" pageId="9" pageNumber="50">C. granti</emphasis>
|
||
</taxonomicName>
|
||
the eyes are not visible in dorsal view and it has a subpentagonal carapace with a glabrous margin instead of a subarcuate carapace, with the margin setose and eyes visible in dorsal view.
|
||
<taxonomicName id="9AB44D55FFCEC7624A2654E8FDB3DD9F" box="[301,619,1235,1260]" class="Malacostraca" family="Pinnotheridae" genus="Calyptraeotheres" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="50" phylum="Arthropoda" rank="species" species="pepeluisi">
|
||
<emphasis id="6FC0EAC4FFCEC7624A2654E8FDB3DD9F" box="[301,619,1235,1260]" italics="true" pageId="9" pageNumber="50">Calyptraeotheres pepeluisi</emphasis>
|
||
</taxonomicName>
|
||
can be separated from
|
||
<taxonomicName id="9AB44D55FFCEC762488B54E8FBFEDD9F" box="[896,1062,1235,1260]" class="Malacostraca" family="Pinnotheridae" genus="Calyptraeotheres" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="50" phylum="Arthropoda" rank="species" species="hernandezi">
|
||
<emphasis id="6FC0EAC4FFCEC762488B54E8FBFEDD9F" box="[896,1062,1235,1260]" italics="true" pageId="9" pageNumber="50">C. hernandezi</emphasis>
|
||
</taxonomicName>
|
||
as follows: eyes proportionally smaller; posterior margin of carapace less concave; carapace without a shallow transverse T-shaped depression connecting both longitudinal sulci; and margins of ischium-merus and external margin of carpus with pinnate setae (simple setae in
|
||
<taxonomicName id="9AB44D55FFCEC76249415578FD2FDC2F" box="[586,759,1347,1372]" class="Malacostraca" family="Pinnotheridae" genus="Calyptraeotheres" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="50" phylum="Arthropoda" rank="species" species="hernandezi">
|
||
<emphasis id="6FC0EAC4FFCEC76249415578FD2FDC2F" box="[586,759,1347,1372]" italics="true" pageId="9" pageNumber="50">C. hernandezi</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
<taxonomicName id="9AB44D55FFCEC76248195578FB85DC2F" box="[786,1117,1347,1372]" class="Malacostraca" family="Pinnotheridae" genus="Calyptraeotheres" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="50" phylum="Arthropoda" rank="species" species="pepeluisi">
|
||
<emphasis id="6FC0EAC4FFCEC76248195578FB85DC2F" box="[786,1117,1347,1372]" italics="true" pageId="9" pageNumber="50">Calyptraeotheres pepeluisi</emphasis>
|
||
</taxonomicName>
|
||
can be separated from its Pacific congener,
|
||
<taxonomicName id="9AB44D55FFCEC7624A675553FE00DCF2" box="[364,472,1384,1409]" class="Malacostraca" family="Pinnotheridae" genus="Calyptraeotheres" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="50" phylum="Arthropoda" rank="species" species="granti">
|
||
<emphasis id="6FC0EAC4FFCEC7624A675553FE00DCF2" box="[364,472,1384,1409]" italics="true" pageId="9" pageNumber="50">C. granti</emphasis>
|
||
</taxonomicName>
|
||
, as follows: carapace with the lateral margin arcuate instead of subparallel; eyes visible in dorsal view (not visible in dorsal view in
|
||
<taxonomicName id="9AB44D55FFCEC762480155B6FCAFDCD5" box="[778,887,1421,1446]" class="Malacostraca" family="Pinnotheridae" genus="Calyptraeotheres" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="50" phylum="Arthropoda" rank="species" species="granti">
|
||
<emphasis id="6FC0EAC4FFCEC762480155B6FCAFDCD5" box="[778,887,1421,1446]" italics="true" pageId="9" pageNumber="50">C. granti</emphasis>
|
||
</taxonomicName>
|
||
), and MXP3 propodus conical, not obliquely truncated as in
|
||
<taxonomicName id="9AB44D55FFCEC7624A425588FE6ADCBF" box="[329,434,1459,1484]" class="Malacostraca" family="Pinnotheridae" genus="Calyptraeotheres" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="50" phylum="Arthropoda" rank="species" species="granti">
|
||
<emphasis id="6FC0EAC4FFCEC7624A425588FE6ADCBF" box="[329,434,1459,1484]" italics="true" pageId="9" pageNumber="50">C. granti</emphasis>
|
||
</taxonomicName>
|
||
. The host for the new species is not known.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |