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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1175.107636" ID-Pensoft-Pub="1313-2970-1175-163" ID-Pensoft-UUID="CE03B1B394CF56B48E8F7E5026CD639C" ID-ZooBank="E88FF072E8C945AEA4944B2A5F7DD7E8" ModsDocID="1313-2970-1175-163" checkinTime="1692265381537" checkinUser="pensoft" docAuthor="Yin, Wenqi, Zhu, Pingzhou &amp; Shi, Hongliang" docDate="2023" docId="F17393A40072567C8060CCF5F507C7AB" docLanguage="en" docName="ZooKeys 1175: 163-185" docOrigin="ZooKeys 1175" docPubDate="2023-08-16" docSource="http://dx.doi.org/10.3897/zookeys.1175.107636" docTitle="Pterostichus (Orientostichus) jialini Yin &amp; Zhu &amp; Shi 2023, sp. nov." docType="treatment" docUuid="C0AC3F70-3137-4167-8308-44A1E4D1105E" docUuidSource="ZooBank" docVersion="1" id="CE03B1B394CF56B48E8F7E5026CD639C" lastPageNumber="163" masterDocId="CE03B1B394CF56B48E8F7E5026CD639C" masterDocTitle="Taxonomic study on the subgenus Orientostichus, the Pterostichus pulcher species group (Coleoptera, Carabidae, Pterostichus)" masterLastPageNumber="185" masterPageNumber="163" pageNumber="163" updateTime="1692265381537" updateUser="pensoft">
<mods:mods id="117A46E2CF6BF4F5F80C8EF878182B10" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="485AFF885C8F6F0E17D3E1AD8400518A">
<mods:title id="BAAC81A0828B2FF4FE33A4546A285529">Taxonomic study on the subgenus Orientostichus, the Pterostichus pulcher species group (Coleoptera, Carabidae, Pterostichus)</mods:title>
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<mods:name id="7CED99F04621ADF179A0963F6DC2338F" type="personal">
<mods:role id="89E0BF5289F6322CDCA951B28CAED8E2">
<mods:roleTerm id="32406A80BD7B520D3634F98CA5BE4457">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="1F2D319F25CD9C1F787A8A307C53E963">Yin, Wenqi</mods:namePart>
<mods:nameIdentifier id="33453617078C4CD86F61646315DAB723" type="ORCID">https://orcid.org/0000-0003-2903-1399</mods:nameIdentifier>
<mods:affiliation id="698EE5A01DDF3BF22DF4FF3627190FC8">College of Forestry, Beijing Forestry University, Beijing 100083, China</mods:affiliation>
</mods:name>
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<mods:role id="C386A268CCF4ECBC5415BEE8B07C87B3">
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<mods:namePart id="BAB2C97ED22B2459A61C8EBAC8D22EAA">Zhu, Pingzhou</mods:namePart>
<mods:nameIdentifier id="C121E328874E00C905C65974F51FE0D5" type="ORCID">https://orcid.org/0000-0003-2139-6764</mods:nameIdentifier>
<mods:affiliation id="27567A31156BD8BD885E5327F5133F34">Department of Entomology, College of Plant Protection, China Agricultural University, Beijing 100093, China</mods:affiliation>
</mods:name>
<mods:name id="6B6A46872E21987ADB0AC23546C2C69E" type="personal">
<mods:role id="78D778D20BECA5CE5D45158E83E750BB">
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<mods:namePart id="5C2E9AD469CB5CC55F8879302D0E5535">Shi, Hongliang</mods:namePart>
<mods:nameIdentifier id="7D22A04E28F860C207378CFB7BE7E9B9" type="ORCID">https://orcid.org/0000-0002-9989-5830</mods:nameIdentifier>
<mods:affiliation id="214EB8D0ADBD660CB693B057981B8D8C">College of Forestry, Beijing Forestry University, Beijing 100083, China</mods:affiliation>
<mods:nameIdentifier id="5B37C01EA76249E17D0CF2E14A5E8C06" type="email">shihl@bjfu.edu.cn</mods:nameIdentifier>
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<mods:typeOfResource id="847A676C33CF9EA3BA27429D4781ADCE">text</mods:typeOfResource>
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<mods:part id="0F6D82FD935471B8DFBE67282C36E56C">
<mods:date id="9BCDBE3D7DBE4AD3750078F4BB62C539">2023</mods:date>
<mods:detail id="FCD4FA11C77F8AF97C542FAEF0F2BAF1" type="pubDate">
<mods:number id="DA71696D3D3960D5DCEEB79058905FC6">2023-08-16</mods:number>
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<mods:detail id="31E8880E1A4CFC755C459465FD4B9B6A" type="volume">
<mods:number id="7EDDF002388B913FB81071741B6831B9">1175</mods:number>
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<mods:url id="0E120B73A9FD71E98EC61BB4E010FC5A">http://dx.doi.org/10.3897/zookeys.1175.107636</mods:url>
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<mods:classification id="6C8EC3910A7CFA7F5C1E42946A0348D2">journal article</mods:classification>
<mods:identifier id="6DE9AB50ABBF397B6C972A30ED4024E9" type="DOI">http://dx.doi.org/10.3897/zookeys.1175.107636</mods:identifier>
<mods:identifier id="4B16029310137DCE7C7DBB606CDA9606" type="Pensoft-Pub">1313-2970-1175-163</mods:identifier>
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<treatment id="F17393A40072567C8060CCF5F507C7AB" LSID="urn:lsid:zoobank.org:act:C0AC3F70-3137-4167-8308-44A1E4D1105E" httpUri="http://treatment.plazi.org/id/F17393A40072567C8060CCF5F507C7AB" lastPageNumber="163" pageId="0" pageNumber="163">
<subSubSection id="953D89F6F65B4A5CD5ECE924F47DBF89" pageId="0" pageNumber="163" type="nomenclature">
<paragraph id="48A77558965B8C9CCC60CDB4676D21A2" pageId="0" pageNumber="163">
<taxonomicName id="BB337F5F37B882545B951C898E696D48" LSID="https://zoobank.org/C0AC3F70-3137-4167-8308-44A1E4D1105E" authority="Yin &amp; Zhu &amp; Shi, 2023" authorityName="Yin &amp; Zhu &amp; Shi" authorityYear="2023" class="Insecta" family="Carabidae" genus="Pterostichus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pterostichus (Orientostichus) jialini" order="Coleoptera" pageId="0" pageNumber="163" phylum="Arthropoda" rank="species" species="jialini" status="sp. nov." subGenus="Orientostichus">Pterostichus (Orientostichus) jialini</taxonomicName>
<taxonomicNameLabel id="851638E8ED0F10BAD38BA1F91FDEF1CB" pageId="0" pageNumber="163">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="396C477B5DE817A477964C9EE06A8BA3" pageId="0" pageNumber="163" type="description">
<paragraph id="885F79A45ACBFA827D21EEF4293F8B93" pageId="0" pageNumber="163">
<figureCitation id="ACAA660AB15F41AE643F8EAD699A3EFB" captionStart="Figures 1–6" captionStartId="F1" captionText="Figures 1 - 6. Habitus and labels of Pterostichus pulcher species group 1 P. pemphis sp. nov., holotype 2 P. orbicollis sp. nov., holotype 3 P. leo sp. nov., holotype 4 P. liyuani sp. nov., holotype 5 P. condylus sp. nov., holotype 6 P. jialini sp. nov., holotype. Scale bar: 5 mm." figureDoi="10.3897/zookeys.1175.107636.figures1-6" httpUri="https://binary.pensoft.net/fig/893313" pageId="0" pageNumber="163">Figs 6</figureCitation>
<figureCitation id="ECD3DC11FE094CDCC051F1FCA52B9B1C" captionStart="Figures 7–12" captionStartId="F2" captionText="Figures 7 - 12. Pronota of Pterostichus pulcher species group 7 P. pemphis sp. nov., holotype 8 P. orbicollis sp. nov., holotype 9 P. leo sp. nov., holotype 10 P. liyuani sp. nov., holotype 11 P. condylus sp. nov., holotype 12 P. jialini sp. nov., holotype. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1175.107636.figures7-12" httpUri="https://binary.pensoft.net/fig/893314" pageId="0" pageNumber="163">, 12</figureCitation>
<figureCitation id="F7C00FE1DA3A58B9304E0D20AC49A936" captionStart="Figures 13–18" captionStartId="F3" captionText="Figures 13 - 18. Male genitalia of Pterostichus pulcher species group, left lateral view and dorsal view of median lobe of aedeagus 13 P. pemphis sp. nov., holotype 14 P. orbicollis sp. nov., holotype 15 P. leo sp. nov., holotype 16 P. liyuani sp. nov., holotype 17 P. condylus sp. nov., holotype 18 P. jialini sp. nov., holotype. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1175.107636.figures13-18" httpUri="https://binary.pensoft.net/fig/893315" pageId="0" pageNumber="163">, 18</figureCitation>
<figureCitation id="92016DC109913D21DD84065B53A7FDC0" captionStart="Figures 34–39" captionStartId="F7" captionText="Figures 34 - 39. Male sternum VII of Pterostichus pulcher species group spp. 34 P. pemphis sp. nov., holotype 35 P. orbicollis sp. nov., holotype 36 P. liyuani sp. nov., holotype 37 P. condylus sp. nov., holotype 38 P. jialini sp. nov., holotype 39 P. pulcher Sciaky &amp; Allegro, holotype. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1175.107636.figures34-39" httpUri="https://binary.pensoft.net/fig/893319" pageId="0" pageNumber="163">
, 38 Chinese vernacular name:
<normalizedToken id="EF862B3C8CAF3F9AF41AA2F83941EEFD" originalValue="家麟通缘步甲">家麟通缘步甲</normalizedToken>
</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="4AD20DA4D5DABA704C47829B5C4D2A65" pageId="0" pageNumber="163" type="type locality">
<paragraph id="E5212211264B33688DE5D36551AB95FB" pageId="0" pageNumber="163">Type locality.</paragraph>
<paragraph id="B0C02D1E0D477D9BA2160C8ED6D6206E" pageId="0" pageNumber="163">
China, Sichuan province: Liangshan Yi Autonomous Prefecture, Huidong county, Jiamashi pasture (
<geoCoordinate id="C8168C5CCBCD3F8D2F4E8ED605A9CA12" degrees="26.81" direction="north" orientation="latitude" precision="555" value="26.81">26.81N</geoCoordinate>
,
<geoCoordinate id="286F97ABA6DB0855AB51BF923D88BA05" degrees="102.68" direction="east" orientation="longitude" precision="555" value="102.68">102.68E</geoCoordinate>
, alt 3000 m).
</paragraph>
</subSubSection>
<subSubSection id="5AD32119F1FC8D642DCC4CDF0E633AA7" pageId="0" pageNumber="163" type="type material">
<paragraph id="FDF7B4846E7EDADE5A401E2DAEC744B8" pageId="0" pageNumber="163">Type material.</paragraph>
<paragraph id="E3B129B809ADF209389A2A5F8285F20D" pageId="0" pageNumber="163">
<emphasis id="041FB1B2375B1EF16F5084AFDA174253" bold="true" italics="true" pageId="0" pageNumber="163">Holotype</emphasis>
: ♂: &quot;China, Sichuan province, Liangshan autonomous prefecture, Huidong county, Jiamashi pasture, 3000 m, 2021.VII, pitfall trap&quot;, &quot;HOLOTYPE ♂
<taxonomicName id="C8303643B516397AF35A24D8DCDA99F5" authorityName="Yin &amp; Zhu &amp; Shi" authorityYear="2023" class="Insecta" family="Carabidae" genus="Pterostichus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pterostichus (Orientostichus) jialini" order="Coleoptera" pageId="0" pageNumber="163" phylum="Arthropoda" rank="species" species="jialini" subGenus="Orientostichus">Pterostichus (Orientostichus) jialini</taxonomicName>
sp. nov. det. Yin &amp; Shi, 2022&quot; [red label].
<emphasis id="4CB14E0F0F9185EA974B8FD28A0A588E" bold="true" italics="true" pageId="0" pageNumber="163">Paratype</emphasis>
: 1 ♂ (CYHL): the same data as holotype but labeled as paratype.
</paragraph>
</subSubSection>
<subSubSection id="F83D1C7ADEE10DD20EA12B15E5212B67" pageId="0" pageNumber="163" type="diagnosis">
<paragraph id="64F753C8C0536F7998CAC3347B4C27B2" pageId="0" pageNumber="163">Diagnosis.</paragraph>
<paragraph id="2A983A75CDB0B2FD8B0D0D72AF205868" pageId="0" pageNumber="163">
Elytra with several large foveate discal pores on intervals 3, 5, and 7, forming strong catenulate sculpturing. Antennomere 3 without accessory setae. Pronotum subcordate, strongly narrowed to base (PBW/PW = 0.65-0.67); posterior angles blunt and inconspicuously projected laterally; basal foveae depressed between inner and outer grooves which partly fused (Fig.
<figureCitation id="0583A8AB060F09D8662F0CF5E0F9BC10" captionStart="Figures 7–12" captionStartId="F2" captionText="Figures 7 - 12. Pronota of Pterostichus pulcher species group 7 P. pemphis sp. nov., holotype 8 P. orbicollis sp. nov., holotype 9 P. leo sp. nov., holotype 10 P. liyuani sp. nov., holotype 11 P. condylus sp. nov., holotype 12 P. jialini sp. nov., holotype. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1175.107636.figures7-12" httpUri="https://binary.pensoft.net/fig/893314" pageId="0" pageNumber="163">12</figureCitation>
). Sexual modification on sternite VII asymmetrical, with a large tubercle and two small protuberances on its right-basal side (Fig.
<figureCitation id="95748E8970CB79C09A8086D92018360D" captionStart="Figures 34–39" captionStartId="F7" captionText="Figures 34 - 39. Male sternum VII of Pterostichus pulcher species group spp. 34 P. pemphis sp. nov., holotype 35 P. orbicollis sp. nov., holotype 36 P. liyuani sp. nov., holotype 37 P. condylus sp. nov., holotype 38 P. jialini sp. nov., holotype 39 P. pulcher Sciaky &amp; Allegro, holotype. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1175.107636.figures34-39" httpUri="https://binary.pensoft.net/fig/893319" pageId="0" pageNumber="163">38</figureCitation>
). Apical lamella of male genitalia gradually narrowed to apex.
</paragraph>
</subSubSection>
<subSubSection id="703ECE9C590E5165A920E6EF9DF5F130" pageId="0" pageNumber="163" type="comparisons">
<paragraph id="1261FEB2A1B0AC57DE4A2AFAC1659C48" pageId="0" pageNumber="163">Comparison.</paragraph>
<paragraph id="899588EC22B6D7D1D959CA7D64617B80" pageId="0" pageNumber="163">
<taxonomicName id="97DA2542F4D10C4FE6A1D94F7854B499" lsidName="P. jialini" pageId="0" pageNumber="163" rank="species" species="jialini">
<emphasis id="50BB412A2ED62E5626AE8F4788A311A4" italics="true" pageId="0" pageNumber="163">P. jialini</emphasis>
</taxonomicName>
sp. nov. is most similar to
<taxonomicName id="24B7008945B9CE8E3D131D64F5DEE287" lsidName="P. pulcher" pageId="0" pageNumber="163" rank="species" species="pulcher">
<emphasis id="CDFE8C763A57C7748B68BB98CB339655" italics="true" pageId="0" pageNumber="163">P. pulcher</emphasis>
</taxonomicName>
as both species differ from other five species of the
<taxonomicName id="C45EB343D6CD265FFE5E0A57C3E46D19" lsidName="P. pulcher" pageId="0" pageNumber="163" rank="species" species="pulcher">
<emphasis id="755242C8459425EDE1F4BBD815369812" italics="true" pageId="0" pageNumber="163">P. pulcher</emphasis>
</taxonomicName>
species group in having large foveate discal pores on interval 7, forming catenulate sculpturing. Compared with
<taxonomicName id="A4DB58E7A2ECE6589318F22E1E334188" lsidName="P. pulcher" pageId="0" pageNumber="163" rank="species" species="pulcher">
<emphasis id="E39E65216D7A3FB815E8D163907547E9" italics="true" pageId="0" pageNumber="163">P. pulcher</emphasis>
</taxonomicName>
,
<taxonomicName id="CDADED35B0D42AD8380DD18BF0A6F2B7" lsidName="P. jialini" pageId="0" pageNumber="163" rank="species" species="jialini">
<emphasis id="F849B08A785AF122E378906F936AA658" italics="true" pageId="0" pageNumber="163">P. jialini</emphasis>
</taxonomicName>
sp. nov. is different in: (1) pronotum more strongly constricted to the base, with PBW/PW = 0.65-0.67 (vs PBW/PW = 0.75-0.81 in
<taxonomicName id="35A9DCD53413007B48CB0873CDC714C4" lsidName="P. pulcher" pageId="0" pageNumber="163" rank="species" species="pulcher">
<emphasis id="FFDF8474F99F4EFFC2BF8A8606A1869B" italics="true" pageId="0" pageNumber="163">P. pulcher</emphasis>
</taxonomicName>
); (2) pronotum posterior angles blunt and inconspicuously projected laterally, while
<taxonomicName id="15EC8EFBB3032B8D972A9C093A64C99F" lsidName="P. pulcher" pageId="0" pageNumber="163" rank="species" species="pulcher">
<emphasis id="7BBEA21BDA8CF9DD4007620C7F0AA301" italics="true" pageId="0" pageNumber="163">P. pulcher</emphasis>
</taxonomicName>
has posterior angles acute at apex and distinctly projected laterally; (3) pronotal basal foveae depressed between inner and outer grooves, but convex in
<taxonomicName id="F9BD680BB1C790774FD78158CE2EFE95" lsidName="P. pulcher" pageId="0" pageNumber="163" rank="species" species="pulcher">
<emphasis id="06BF44E8C7F4F2374116C6C18D57423C" italics="true" pageId="0" pageNumber="163">P. pulcher</emphasis>
</taxonomicName>
; (4) male sternite VII with distinct and asymmetrical secondary sexual characters, while in
<taxonomicName id="0B7CC5356242BD36EBE0710EA94DDAE8" lsidName="P. pulcher" pageId="0" pageNumber="163" rank="species" species="pulcher">
<emphasis id="D6E113D504F36DA2B775DB43C257EDB4" italics="true" pageId="0" pageNumber="163">P. pulcher</emphasis>
</taxonomicName>
male sternite VII shallowly and symmetrical tumid; (5) the apical lamella of male genitalia with its lateral margins convergent to apex, but in
<taxonomicName id="1E8C711DD91013CDAF38C441BEFFDF5C" lsidName="P. pulcher" pageId="0" pageNumber="163" rank="species" species="pulcher">
<emphasis id="5BF06BCA22AC2D083FBD838EE84EB8DB" italics="true" pageId="0" pageNumber="163">P. pulcher</emphasis>
</taxonomicName>
, the apical lamella with lateral margins subparallel to apex.
</paragraph>
</subSubSection>
<subSubSection id="7B3383061CFFD3D12DDD1C80F150997C" pageId="0" pageNumber="163" type="description">
<paragraph id="F639117B478515B13449DA33752F36CE" pageId="0" pageNumber="163">Description.</paragraph>
<paragraph id="9E984337E7B28B3B72E947B432CA7EEF" pageId="0" pageNumber="163">
BL = 15.4-15.5 mm, BW = 5.7-5.8 mm, dorsal surface and appendages black, elytra with very faint metallic luster. Antennomere 3 without accessory setae, only with primary setae forming apical ring. Pronotum subcordate, PW/PL = 1.13-1.15, widest near middle; anterior margin slightly wider than posterior margin; strongly narrowed to base (PBW/PW = 0.65-0.67); lateral margins evenly arched from anterior angles to middle, distinctly sinuate before posterior angles; posterior angles rather blunt, apex rounded-obtuse, inconspicuously projected laterally; lateral margins with 2-4 mid-lateral setae near maximum width; basal foveae impunctate, depressed between inner and outer grooves, making them seems partly fused together (Fig.
<figureCitation id="01F4D87D9657A1C34330B72AC96F7AFF" captionStart="Figures 7–12" captionStartId="F2" captionText="Figures 7 - 12. Pronota of Pterostichus pulcher species group 7 P. pemphis sp. nov., holotype 8 P. orbicollis sp. nov., holotype 9 P. leo sp. nov., holotype 10 P. liyuani sp. nov., holotype 11 P. condylus sp. nov., holotype 12 P. jialini sp. nov., holotype. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1175.107636.figures7-12" httpUri="https://binary.pensoft.net/fig/893314" pageId="0" pageNumber="163">12</figureCitation>
); disc with fine transverse wrinkles aside median line. Elytra oblong; parascutellar pore absent; intervals 3, 5, and 7 each with ≥ 3 large foveate discal pores, forming strong catenulate sculpturing. Male sternite VII with asymmetric modification, a large tubercle bending to right and two small protuberances on its right-basal side (Fig.
<figureCitation id="E815E57A5C42C01B86691C5A1A98510B" captionStart="Figures 34–39" captionStartId="F7" captionText="Figures 34 - 39. Male sternum VII of Pterostichus pulcher species group spp. 34 P. pemphis sp. nov., holotype 35 P. orbicollis sp. nov., holotype 36 P. liyuani sp. nov., holotype 37 P. condylus sp. nov., holotype 38 P. jialini sp. nov., holotype 39 P. pulcher Sciaky &amp; Allegro, holotype. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1175.107636.figures34-39" httpUri="https://binary.pensoft.net/fig/893319" pageId="0" pageNumber="163">38</figureCitation>
). Median lobe of aedeagus stout, strongly curved near basal 1/3; apical lamella gradually deflected ventrally; apical lamella slightly twisted longitudinally, (LL/LW = 1.75-1.9), subtriangular with lateral margins convergent to apex, apex rounded (Fig.
<figureCitation id="49A80DBABAA25043847787101910B7C2" captionStart="Figures 13–18" captionStartId="F3" captionText="Figures 13 - 18. Male genitalia of Pterostichus pulcher species group, left lateral view and dorsal view of median lobe of aedeagus 13 P. pemphis sp. nov., holotype 14 P. orbicollis sp. nov., holotype 15 P. leo sp. nov., holotype 16 P. liyuani sp. nov., holotype 17 P. condylus sp. nov., holotype 18 P. jialini sp. nov., holotype. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1175.107636.figures13-18" httpUri="https://binary.pensoft.net/fig/893315" pageId="0" pageNumber="163">18</figureCitation>
). Endophallus not studied due to restricted number of specimens.
</paragraph>
</subSubSection>
<subSubSection id="FEBDAA54E8A963A4FB728E6442C02584" pageId="0" pageNumber="163" type="distribution">
<paragraph id="5D1980B6F3A2CF8EB0EB8EF6CC1081A8" pageId="0" pageNumber="163">Distribution.</paragraph>
<paragraph id="19CDCA2F1C02E3E764E01C44C4E96700" pageId="0" pageNumber="163">
This species is only known from its type locality in Huidong county, Liangshan Yi Autonomous Prefecture (Fig.
<figureCitation id="04BB6E9FDF116A3C3DAF0BB2A8B816F8" captionStart="Figure 42" captionStartId="F9" captionText="Figure 42. Distribution of the taxa of Pterostichus pulcher species group in Sichuan province, China. Brown: P. pemphis sp. nov. Yellow: P. orbicollis sp. nov. Red: P. leo sp. nov. Purple: P. liyuani sp. nov. Blue: P. condylus sp. nov. Green: P. jialini sp. nov. Orange: P. pulcher Sciaky &amp; Allegro." figureDoi="10.3897/zookeys.1175.107636.figure42" httpUri="https://binary.pensoft.net/fig/893321" pageId="0" pageNumber="163">42</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="9FF37EE49A4C6FADA3830DFCDC0914D7" pageId="0" pageNumber="163" type="etymology">
<paragraph id="28122DF09A90785A0921E219939500FB" pageId="0" pageNumber="163">Etymology.</paragraph>
<paragraph id="5E08C1A8B6B41F514EF9CACB81DFB41D" pageId="0" pageNumber="163">The scientific name of the new species is dedicated to Mr. Jialin Tian, the collector of two type specimens.</paragraph>
</subSubSection>
</treatment>
</document>
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