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<document id="EFEE2FDC6393B297E116D5FBDA053208" ID-DOI="10.5281/zenodo.195284" ID-GBIF-Dataset="e89fb81f-4b02-4f0c-a889-1e2072699196" ID-ISSN="1175-5326" ID-Zenodo-Dep="195284" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460225388371" checkinUser="plazi" docAuthor="Colloff, Matthew J." docDate="2010" docId="EC637008FFEE136FE38DDF37FCABFC54" docLanguage="en" docName="zt02475p038.pdf" docOrigin="Zootaxa 2475" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Scapheremaeus zephyrus Colloff, 2010, sp. nov." docType="treatment" docVersion="8" lastPageNumber="32" masterDocId="105A0870FFF31370E31ADB0AFF99FFA7" masterDocTitle="The hyperdiverse oribatid mite genus Scapheremaeus (Acari: Oribatida: Cymbaeremaeidae) in Australia, with descriptions of new species and consideration of biogeographical affinities" masterLastPageNumber="38" masterPageNumber="1" pageNumber="30" updateTime="1698238218813" updateUser="plazi">
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<mods:title id="B210077CFAB6CF028D778159F64FC9BD">The hyperdiverse oribatid mite genus Scapheremaeus (Acari: Oribatida: Cymbaeremaeidae) in Australia, with descriptions of new species and consideration of biogeographical affinities</mods:title>
</mods:titleInfo>
<mods:name id="E1F1E105503483BAB4A025DDCE2F25C6" type="personal">
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</mods:role>
<mods:namePart id="4E60D4AFF9870C40AA022BC1A41FF8E9">Colloff, Matthew J.</mods:namePart>
</mods:name>
<mods:typeOfResource id="0672E9EEB4575A3168D5271A37F862FB">text</mods:typeOfResource>
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<mods:title id="5422171B1AC753981DC072D348C1CE61">Zootaxa</mods:title>
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<mods:part id="A1EB12756A55F13254B05C51FC638921">
<mods:date id="8ECEE235DD2B7E3463EE3EE5AAE555B4">2010</mods:date>
<mods:detail id="95DBE4B578D2C804B0C6F526CC6F095B" type="volume">
<mods:number id="501ED3A56D86DE84B547594EBD84C9D9">2475</mods:number>
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<mods:classification id="B5A1E91A5F6FDC017BA5535A8F1EB049">journal article</mods:classification>
<mods:identifier id="B81BC34BA13756643ADABD2527F04CB5" type="DOI">10.5281/zenodo.195284</mods:identifier>
<mods:identifier id="D4F656227793E25B6F5F9365E52D7608" type="GBIF-Dataset">e89fb81f-4b02-4f0c-a889-1e2072699196</mods:identifier>
<mods:identifier id="DAE07BBA6101D7900086260D745FCA0D" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="481B62C110ED0275E394F0412A7819F1" type="Zenodo-Dep">195284</mods:identifier>
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<treatment id="EC637008FFEE136FE38DDF37FCABFC54" ID-DOI="http://doi.org/10.5281/zenodo.6199044" ID-GBIF-Taxon="119386164" ID-Zenodo-Dep="6199044" LSID="urn:lsid:plazi:treatment:EC637008FFEE136FE38DDF37FCABFC54" httpUri="http://treatment.plazi.org/id/EC637008FFEE136FE38DDF37FCABFC54" lastPageId="31" lastPageNumber="32" pageId="29" pageNumber="30">
<subSubSection id="2CD09295FFEE136DE38DDF37FF64FBDD" pageId="29" pageNumber="30" type="nomenclature">
<paragraph id="6475C11EFFEE136DE38DDF37FDB4FBF0" blockId="29.[151,557,1085,1146]" box="[151,557,1085,1111]" pageId="29" pageNumber="30">
<heading id="3F3D7672FFEE136DE38DDF37FDB4FBF0" bold="true" box="[151,557,1085,1111]" fontSize="11" level="1" pageId="29" pageNumber="30" reason="1">
<emphasis id="56BE1D0CFFEE136DE38DDF37FDB4FBF0" bold="true" box="[151,557,1085,1111]" pageId="29" pageNumber="30">
<taxonomicName id="A3CABA9DFFEE136DE38DDF37FE51FBF0" ID-CoL="8TBB6" box="[151,456,1085,1111]" class="Arachnida" family="Cymbaeremaeidae" genus="Scapheremaeus" kingdom="Animalia" order="Sarcoptiformes" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="zephyrus" status="sp. nov.">
<emphasis id="56BE1D0CFFEE136DE38DDF37FE51FBF0" bold="true" box="[151,456,1085,1111]" italics="true" pageId="29" pageNumber="30">Scapheremaeus zephyrus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4D8DA077FFEE136DE2D5DF37FDB4FBF0" box="[463,557,1085,1111]" pageId="29" pageNumber="30" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="6475C11EFFEE136DE38DDF6AFF64FBDD" blockId="29.[151,557,1085,1146]" box="[151,253,1120,1146]" pageId="29" pageNumber="30">
(
<figureCitation id="FCF1DD9BFFEE136DE385DF6AFF6DFBDD" box="[159,244,1120,1146]" captionStart="FIGURE 15" captionStartId="30.[151,255,1783,1807]" captionTargetBox="[191,1379,211,1743]" captionTargetId="figure@30.[151,1436,197,1777]" captionTargetPageId="30" captionText="FIGURE 15. Scapheremaeus zephyrus sp. nov. a) dorsal; b) ventral; c) lateral." httpUri="https://zenodo.org/record/195297/files/figure.png" pageId="29" pageNumber="30">Fig. 15</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="2CD09295FFEE136FE38DDFA0FE69FE67" lastPageId="31" lastPageNumber="32" pageId="29" pageNumber="30" type="description">
<paragraph id="6475C11EFFEE136DE38DDFA0FD65FB4D" blockId="29.[151,1437,1194,2004]" pageId="29" pageNumber="30">
<emphasis id="56BE1D0CFFEE136DE38DDFA0FEA8FB63" bold="true" box="[151,305,1194,1220]" pageId="29" pageNumber="30">Dimensions.</emphasis>
<typeStatus id="BB717FBCFFEE136DE223DFA0FE31FB63" box="[313,424,1194,1220]" pageId="29" pageNumber="30" type="holotype">Holotype</typeStatus>
female: length 384, breadth 202;
<typeStatus id="BB717FBCFFEE136DE025DFA0FC3CFB63" box="[831,933,1194,1220]" pageId="29" pageNumber="30" type="paratype">paratype</typeStatus>
female: length 391, breadth 211. Ratio of length of prodorsum to total length (
<typeStatus id="BB717FBCFFEE136DE125DFDAFD34FB4D" box="[575,685,1232,1258]" pageId="29" pageNumber="30" type="holotype">holotype</typeStatus>
): 0.28.
</paragraph>
<paragraph id="6475C11EFFEE136DE3DFDFFFFC21FA48" blockId="29.[151,1437,1194,2004]" pageId="29" pageNumber="30">
<emphasis id="56BE1D0CFFEE136DE3DFDFFFFEC2FAA8" bold="true" box="[197,347,1269,1295]" pageId="29" pageNumber="30">Prodorsum.</emphasis>
Rostrum broad, U-shaped, with transverse striae (
<figureCitation id="FCF1DD9BFFEE136DE0AEDFFFFB92FAA8" box="[948,1035,1269,1295]" captionStart="FIGURE 15" captionStartId="30.[151,255,1783,1807]" captionTargetBox="[191,1379,211,1743]" captionTargetId="figure@30.[151,1436,197,1777]" captionTargetPageId="30" captionText="FIGURE 15. Scapheremaeus zephyrus sp. nov. a) dorsal; b) ventral; c) lateral." httpUri="https://zenodo.org/record/195297/files/figure.png" pageId="29" pageNumber="30">Fig. 15</figureCitation>
a). Rostral seta (
<emphasis id="56BE1D0CFFEE136DE7D4DFFFFB7FFAA9" box="[1230,1254,1269,1294]" italics="true" pageId="29" pageNumber="30">ro</emphasis>
) smooth, short (9) straight, spiniform. Lateral carinae present as a pair of well-developed lateral ridges. Lamellar seta short (6), bacilliform, on short tubercles at apices of straight costulae. Intercostular area with slight, faint interlamellar ridge and mid-transcostular ridge; with large, polygonate plaques posterior of mid-intercostular ridge. Faint interbothridial ridge present. Interlamellar seta absent. Sensillus with large, club-shaped, darklypigmented head, sparingly ornamented with flat tubercles, ca. 24 long, 15 broad, emerging on short stalk from bothridium (diameter 11) positioned adjacent to dorsosejugal suture.
</paragraph>
<paragraph id="6475C11EFFEE136DE3DFDEF0FF64F873" blockId="29.[151,1437,1194,2004]" pageId="29" pageNumber="30">
<emphasis id="56BE1D0CFFEE136DE3DFDEF0FEC1F9B3" bold="true" box="[197,344,1530,1556]" pageId="29" pageNumber="30">Notogaster.</emphasis>
Length 272. Circumdorsal scissure entire, oval (
<figureCitation id="FCF1DD9BFFEE136DE0AADEF0FB90F9B3" box="[944,1033,1530,1556]" captionStart="FIGURE 15" captionStartId="30.[151,255,1783,1807]" captionTargetBox="[191,1379,211,1743]" captionTargetId="figure@30.[151,1436,197,1777]" captionTargetPageId="30" captionText="FIGURE 15. Scapheremaeus zephyrus sp. nov. a) dorsal; b) ventral; c) lateral." httpUri="https://zenodo.org/record/195297/files/figure.png" pageId="29" pageNumber="30">Fig. 15</figureCitation>
a), with well-developed diagonal humeral extensions. Lenticulus wedge-shaped, sub-rectangular, (27 long, 18 broad) with discrete margins and a ridge of cuticle immediately posterior of it. Without a shallow depression or tubercle either side of lenticulus. No humeral process evident in dorsal view, but lateral margin of notogaster concave immediately posterior of humeral region. Dorsal circumnotogastral plate sparingly plicate. Centrodorsal plate 211 long, 130 broad, oval, broadest at level of setae
<emphasis id="56BE1D0CFFEE136DE1B3DDBFFD58F969" box="[681,705,1717,1742]" italics="true" pageId="29" pageNumber="30">lp</emphasis>
. Centrodorsal ridge and posteriolateral concavities absent. Centrodorsal microsculpture consisting of large, sparsely-distributed, shallow alveoli. With nine pairs of short bacilliform notogastral setae, similar in size and shape, positioned on well-developed cylindrical or conical tubercles; two pairs of setae on centrodorsal plate, five pairs on dorsal circumnotogastral plate; two pairs of short, spiniform setae in
<emphasis id="56BE1D0CFFEE136DE2ADDC41FE5FF8C3" box="[439,454,1867,1892]" italics="true" pageId="29" pageNumber="30">p</emphasis>
series positioned on ventral circumnotogastral plate. Opening of
<emphasis id="56BE1D0CFFEE136DE7DADC41FB7CF8C3" box="[1216,1253,1867,1892]" italics="true" pageId="29" pageNumber="30">gla</emphasis>
about mid-way between setae
<emphasis id="56BE1D0CFFEE136DE258DC7AFEF9F82E" box="[322,352,1904,1929]" italics="true" pageId="29" pageNumber="30">lm</emphasis>
and
<emphasis id="56BE1D0CFFEE136DE283DC7AFE29F82E" box="[409,432,1904,1929]" italics="true" pageId="29" pageNumber="30">lp</emphasis>
. With three pairs of lyrifissures visible dorsally and one laterally. Posterior margin of dorsal circumnotogastral plate rounded, crenellated; circumnotogastral plate narrower in caudal region than laterally.
</paragraph>
<caption id="30B59196FFED136EE38DDDFDFC7AF8A9" box="[151,995,1783,1807]" httpUri="https://zenodo.org/record/195297/files/figure.png" pageId="30" pageNumber="31" targetBox="[191,1379,211,1743]" targetPageId="30">
<paragraph id="6475C11EFFED136EE38DDDFDFC7AF8A9" blockId="30.[151,995,1783,1807]" box="[151,995,1783,1807]" pageId="30" pageNumber="31">
<emphasis id="56BE1D0CFFED136EE38DDDFDFEBFF8A8" bold="true" box="[151,294,1783,1807]" pageId="30" pageNumber="31">FIGURE 15.</emphasis>
<taxonomicName id="A3CABA9DFFED136EE237DDFDFDA3F8A9" box="[301,570,1783,1806]" class="Arachnida" family="Cymbaeremaeidae" genus="Scapheremaeus" kingdom="Animalia" order="Sarcoptiformes" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="zephyrus" status="sp. nov.">
<emphasis id="56BE1D0CFFED136EE237DDFDFDA3F8A9" box="[301,570,1783,1806]" italics="true" pageId="30" pageNumber="31">Scapheremaeus zephyrus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4D8DA077FFED136EE15ADDFDFD0EF8A8" box="[576,663,1783,1807]" pageId="30" pageNumber="31" rank="species">
<emphasis id="56BE1D0CFFED136EE15ADDFDFD0EF8A8" bold="true" box="[576,663,1783,1807]" pageId="30" pageNumber="31">sp. nov.</emphasis>
</taxonomicNameLabel>
a) dorsal; b) ventral; c) lateral.
</paragraph>
</caption>
<paragraph id="6475C11EFFED136FE3DFDC35FD27FE8C" blockId="30.[151,1438,1855,2030]" lastBlockId="31.[151,1437,152,1011]" lastPageId="31" lastPageNumber="32" pageId="30" pageNumber="31">
<emphasis id="56BE1D0CFFED136EE3DFDC35FE0AF8FE" bold="true" box="[197,403,1855,1881]" pageId="30" pageNumber="31">Ventral region.</emphasis>
Epimeral setae setiform, thin, subequal in length, formula 3-1-2-2 (
<figureCitation id="FCF1DD9BFFED136EE600DC35FAE2F8FE" box="[1306,1403,1855,1881]" captionStart="FIGURE 15" captionStartId="30.[151,255,1783,1807]" captionTargetBox="[191,1379,211,1743]" captionTargetId="figure@30.[151,1436,197,1777]" captionTargetPageId="30" captionText="FIGURE 15. Scapheremaeus zephyrus sp. nov. a) dorsal; b) ventral; c) lateral." httpUri="https://zenodo.org/record/195297/files/figure.png" pageId="30" pageNumber="31">Fig. 15</figureCitation>
b). Microsculpture of ventral plate with transverse and longitudinal striae and alveoli in epimeral region. Subhexagonal genital plates as broad as long, striate, with six pairs of longitudinally-aligned genital setae positioned close to, but not on, medial edges. Genital setae sub-equal in length; same size and shape as epimeral setae. Anal setae setiform, 10, positioned on posterior half of anal plates, almost their length from median edge. With two pairs of subequal, spiniform adanal setae;
<emphasis id="56BE1D0CFFEC136FE0A1DB92FC40FF16" box="[955,985,152,177]" italics="true" pageId="31" pageNumber="32">ad</emphasis>
2 positioned at least their length from margin of anal plates; mutual distance between
<emphasis id="56BE1D0CFFEC136FE1DCDBCAFD7AFF7E" box="[710,739,192,217]" italics="true" pageId="31" pageNumber="32">ad</emphasis>
1 half that between
<emphasis id="56BE1D0CFFEC136FE0DEDBCAFC78FF7E" box="[964,993,192,217]" italics="true" pageId="31" pageNumber="32">ad</emphasis>
1 and
<emphasis id="56BE1D0CFFEC136FE738DBCAFBA6FF7E" box="[1058,1087,192,217]" italics="true" pageId="31" pageNumber="32">ad</emphasis>
2. Circumgastric scissure with two shallow V-shaped invaginations medial of setae
<emphasis id="56BE1D0CFFEC136FE017DBE3FC85FEA5" box="[781,796,233,258]" italics="true" pageId="31" pageNumber="32">p</emphasis>
1. Ventral circumnotogastral plate not plicate but with short, V-shaped, overlapping, scale-like ridges.
</paragraph>
<paragraph id="6475C11EFFEC136FE3DFDA3FFCFDFE3C" blockId="31.[151,1437,152,1011]" pageId="31" pageNumber="32">
<emphasis id="56BE1D0CFFEC136FE3DFDA3FFE1EFEE8" bold="true" box="[197,391,309,335]" pageId="31" pageNumber="32">Lateral aspect.</emphasis>
Mid-line of rostrum curved, acute (
<figureCitation id="FCF1DD9BFFEC136FE05EDA3CFC39FEF7" box="[836,928,310,336]" captionStart="FIGURE 15" captionStartId="30.[151,255,1783,1807]" captionTargetBox="[191,1379,211,1743]" captionTargetId="figure@30.[151,1436,197,1777]" captionTargetPageId="30" captionText="FIGURE 15. Scapheremaeus zephyrus sp. nov. a) dorsal; b) ventral; c) lateral." httpUri="https://zenodo.org/record/195297/files/figure.png" pageId="31" pageNumber="32">Fig. 15</figureCitation>
c). Hysterosoma markedly dorsoventrally flattened. Exobothridial seta absent. Humeral process consisting of broad, shallow, triangular region, bearing lyrifissure
<emphasis id="56BE1D0CFFEC136FE20FDA8BFEB5FE3D" box="[277,300,385,410]" italics="true" pageId="31" pageNumber="32">ia</emphasis>
ventrally. Centrodorsal region flat, ridge absent.
</paragraph>
<paragraph id="6475C11EFFEC136FE3DFDAAFFE69FE67" blockId="31.[151,1437,152,1011]" box="[197,496,421,448]" pageId="31" pageNumber="32">
<emphasis id="56BE1D0CFFEC136FE3DFDAAFFE9FFE18" bold="true" box="[197,262,421,447]" pageId="31" pageNumber="32">Legs.</emphasis>
Heterotridactylous.
</paragraph>
</subSubSection>
<subSubSection id="2CD09295FFEC136FE3DFDAC1FC04FD97" pageId="31" pageNumber="32" type="materials_examined">
<paragraph id="6475C11EFFEC136FE3DFDAC1FC04FD97" blockId="31.[151,1437,152,1011]" pageId="31" pageNumber="32">
<emphasis id="56BE1D0CFFEC136FE3DFDAC1FE5EFE42" bold="true" box="[197,455,459,485]" pageId="31" pageNumber="32">Material Examined.</emphasis>
<typeStatus id="BB717FBCFFEC136FE2D5DAC1FDA6FE42" box="[463,575,459,485]" pageId="31" pageNumber="32" type="holotype">Holotype</typeStatus>
female and
<typeStatus id="BB717FBCFFEC136FE1CCDAC1FCA7FE42" box="[726,830,459,485]" pageId="31" pageNumber="32" type="paratype">paratype</typeStatus>
female, ex. litter, coastal heathland, Little Target, Beecroft Peninsula (
<geoCoordinate id="01FEA7D9FFEC136FE290DAFBFE6CFDAC" box="[394,501,496,523]" direction="south" orientation="latitude" pageId="31" pageNumber="32" precision="925" value="-35.05">35°03´S</geoCoordinate>
,
<geoCoordinate id="01FEA7D9FFEC136FE119DAFBFDE7FDAC" box="[515,638,496,523]" direction="east" orientation="longitude" pageId="31" pageNumber="32" precision="925" value="150.8">150°48´E</geoCoordinate>
), New South
<collectingCountry id="1CDD818EFFEC136FE039DAFBFCF7FDAC" box="[803,878,497,523]" name="United Kingdom" pageId="31" pageNumber="32">Wales</collectingCountry>
, coll. D.J. Smith and P. Greenslade,
<date id="1074E7DEFFEC136FE63EDAFBFA01FDAC" box="[1316,1432,497,523]" pageId="31" pageNumber="32" value="1995-06-09">9.vi.1995</date>
. Non-type material: larva and protonymph (same sample as
<typeStatus id="BB717FBCFFEC136FE054D91CFC08FD97" box="[846,913,534,560]" pageId="31" pageNumber="32">types</typeStatus>
).
</paragraph>
</subSubSection>
<subSubSection id="2CD09295FFEC136FE3DFD931FF6CFD07" pageId="31" pageNumber="32" type="etymology">
<paragraph id="6475C11EFFEC136FE3DFD931FF6CFD07" blockId="31.[151,1437,152,1011]" pageId="31" pageNumber="32">
<emphasis id="56BE1D0CFFEC136FE3DFD931FECEFDF2" bold="true" box="[197,343,571,597]" pageId="31" pageNumber="32">Etymology.</emphasis>
<taxonomicName id="A3CABA9DFFEC136FE27AD931FD0DFDF3" box="[352,660,571,596]" class="Arachnida" family="Cymbaeremaeidae" genus="Scapheremaeus" kingdom="Animalia" order="Sarcoptiformes" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="zephyrus" status="sp. nov.">
<emphasis id="56BE1D0CFFEC136FE27AD931FD0DFDF3" box="[352,660,571,596]" italics="true" pageId="31" pageNumber="32">Scapheremaeus zephyrus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4D8DA077FFEC136FE186D931FD66FDF2" box="[668,767,571,597]" pageId="31" pageNumber="32" rank="species">
<emphasis id="56BE1D0CFFEC136FE186D931FD66FDF2" bold="true" box="[668,767,571,597]" pageId="31" pageNumber="32">sp. nov.</emphasis>
</taxonomicNameLabel>
is named from the Latin,
<emphasis id="56BE1D0CFFEC136FE759D931FB0AFDF3" box="[1091,1171,571,596]" italics="true" pageId="31" pageNumber="32">zephyr</emphasis>
, meaning ‘light [sea] breeze’, or ‘west wind’, an allusion to its lightly-sclerotised, lightly-ornamented cuticle and its coastal
<typeStatus id="BB717FBCFFEC136FE673D96BFA05FDDC" box="[1385,1436,609,635]" pageId="31" pageNumber="32">type</typeStatus>
locality.
</paragraph>
</subSubSection>
<subSubSection id="2CD09295FFEC136FE3DFD9A1FCABFC54" pageId="31" pageNumber="32" type="discussion">
<paragraph id="6475C11EFFEC136FE3DFD9A1FCA1FCFC" blockId="31.[151,1437,152,1011]" pageId="31" pageNumber="32">
<emphasis id="56BE1D0CFFEC136FE3DFD9A1FED9FD62" bold="true" box="[197,320,683,709]" pageId="31" pageNumber="32">Remarks.</emphasis>
<taxonomicName id="A3CABA9DFFEC136FE252D9A1FDECFD63" box="[328,629,683,708]" class="Arachnida" family="Cymbaeremaeidae" genus="Scapheremaeus" kingdom="Animalia" order="Sarcoptiformes" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="zephyrus" status="sp. nov.">
<emphasis id="56BE1D0CFFEC136FE252D9A1FDECFD63" box="[328,629,683,708]" italics="true" pageId="31" pageNumber="32">Scapheremaeus zephyrus</emphasis>
</taxonomicName>
<emphasis id="56BE1D0CFFEC136FE167D9A1FD44FD62" bold="true" box="[637,733,683,709]" pageId="31" pageNumber="32">
<taxonomicNameLabel id="4D8DA077FFEC136FE167D9A1FD44FD62" box="[637,733,683,709]" pageId="31" pageNumber="32" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
differs from other members of the genus by the following combination of characters: 1) the flat centrodorsal region, when viewed laterally; 2) the sparse, alveolate centrodorsal microsculpture; 3) the circumnotogastral plate much narrower caudally than laterally; 4) the scale-like ornamentation of the ventral circumnotogastral plates; 5) nine pairs of notogastral setae (missing setae
<emphasis id="56BE1D0CFFEC136FE3C3D84BFF71FCFD" box="[217,232,833,858]" italics="true" pageId="31" pageNumber="32">p</emphasis>
3) and two pairs of adanal setae (missing setae
<emphasis id="56BE1D0CFFEC136FE013D84BFCBFFCFD" box="[777,806,833,858]" italics="true" pageId="31" pageNumber="32">ad</emphasis>
3)
</paragraph>
<paragraph id="6475C11EFFEC136FE3DFD863FCABFC54" blockId="31.[151,1437,152,1011]" pageId="31" pageNumber="32">
<taxonomicName id="A3CABA9DFFEC136FE3DFD863FE77FC25" box="[197,494,873,898]" class="Arachnida" family="Cymbaeremaeidae" genus="Scapheremaeus" kingdom="Animalia" order="Sarcoptiformes" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="zephyrus" status="sp. nov.">
<emphasis id="56BE1D0CFFEC136FE3DFD863FE77FC25" box="[197,494,873,898]" italics="true" pageId="31" pageNumber="32">Scapheremaeus zephyrus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4D8DA077FFEC136FE2ECD862FDCCFC25" box="[502,597,872,898]" pageId="31" pageNumber="32" rank="species">
<emphasis id="56BE1D0CFFEC136FE2ECD862FDCCFC25" bold="true" box="[502,597,872,898]" pageId="31" pageNumber="32">sp. nov.</emphasis>
</taxonomicNameLabel>
is a member of the Carinatus species-group. It shares with
<taxonomicName id="A3CABA9DFFEC136FE600D863FA05FC25" box="[1306,1436,873,898]" class="Arachnida" family="Cymbaeremaeidae" genus="Scapheremaeus" kingdom="Animalia" order="Sarcoptiformes" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="alisonae" status="sp. nov.">
<emphasis id="56BE1D0CFFEC136FE600D863FA05FC25" box="[1306,1436,873,898]" italics="true" pageId="31" pageNumber="32">S. alisonae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4D8DA077FFEC136FE38DD884FF63FC0F" box="[151,250,910,936]" pageId="31" pageNumber="32" rank="species">
<emphasis id="56BE1D0CFFEC136FE38DD884FF63FC0F" bold="true" box="[151,250,910,936]" pageId="31" pageNumber="32">sp. nov.</emphasis>
</taxonomicNameLabel>
and
<taxonomicName id="A3CABA9DFFEC136FE222D884FE32FC00" box="[312,427,910,935]" class="Arachnida" family="Cymbaeremaeidae" genus="Scapheremaeus" kingdom="Animalia" order="Sarcoptiformes" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="nivalis" status="sp. nov.">
<emphasis id="56BE1D0CFFEC136FE222D884FE32FC00" box="[312,427,910,935]" italics="true" pageId="31" pageNumber="32">S. nivalis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4D8DA077FFEC136FE2A9D884FD8FFC0F" box="[435,534,910,936]" pageId="31" pageNumber="32" rank="species">
<emphasis id="56BE1D0CFFEC136FE2A9D884FD8FFC0F" bold="true" box="[435,534,910,936]" pageId="31" pageNumber="32">sp. nov.</emphasis>
</taxonomicNameLabel>
the character states of this group, as well as the reduction in notogastral setation (nine pairs) and adanal setation (two pairs); small, broad, triangular humeral processes, notched circumgastric scissure and the genital setae in alignment.
</paragraph>
</subSubSection>
</treatment>
</document>
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