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<document id="1E5531D2B8D33EFD3326607B5821CF45" ID-DOI="10.11646/zootaxa.3964.3.1" ID-GBIF-Dataset="d80458ea-4fb4-4999-92c2-b8445c9dd19a" ID-ISSN="1175-5326" ID-Zenodo-Dep="231957" ID-ZooBank="88702846-1180-400A-B7DB-4154ACD4EC49" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461074488649" checkinUser="plazi" docAuthor="Wesener, Thomas" docDate="2015" docId="EC3687B45819FF89FF3DFD1DFC230A88" docLanguage="en" docName="zt03964p320.pdf" docOrigin="Zootaxa 3964 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Zephronia nepalensis Wesener, 2015, new species" docType="treatment" docVersion="9" lastPageNumber="308" masterDocId="100FFFCC581DFF8EFFAAF841FFEB0C68" masterDocTitle="The Giant Pill-Millipedes of Nepal (Diplopoda, Sphaerotheriida, Zephroniidae) *" masterLastPageNumber="320" masterPageNumber="301" pageNumber="305" updateTime="1698586053603" updateUser="plazi">
<mods:mods id="F8D02E91F94AB32912B4411B43A06B0F" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="A89EEE9E8D95C753D147A5B53E3CBB69">The Giant Pill-Millipedes of Nepal (Diplopoda, Sphaerotheriida, Zephroniidae) *</mods:title>
</mods:titleInfo>
<mods:name id="9881FBF3B61A18A33BE5BB5159BB0E98" type="personal">
<mods:role id="7D1FCB19C89E1D91BCD88AEC9462F64D">
<mods:roleTerm id="6EDD60045ECF07E3BE00FAD8517CE5C0">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="35DBF98D16045C50C0CE2B2599A1B130">Wesener, Thomas</mods:namePart>
</mods:name>
<mods:typeOfResource id="CC8B0AC7DFD048707E29277CEE465BB2">text</mods:typeOfResource>
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<mods:title id="497BD656CF47A0EA13E299F924070C73">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="91FD662642B30C9F046A5AD147C1827E">
<mods:date id="8D342E9D327DB5513B051777A95E81BF">2015</mods:date>
<mods:detail id="4D50FD17FB20A753C601E2118D7A9260" type="volume">
<mods:number id="AA1FF2AC4DDF4B3CAC72B06DBFEC1C5B">3964</mods:number>
</mods:detail>
<mods:detail id="361BB0F714ED63A7A173B41F8A1FC118" type="issue">
<mods:number id="FD81EAB6AC92108C445E036A338B0034">3</mods:number>
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<mods:start id="99B9D5D285DBC3BCCC4F4ACD06A3A8AD">301</mods:start>
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<mods:classification id="C286F2D42C3BA5EC7A0143DB68F28D4C">journal article</mods:classification>
<mods:identifier id="99D515003064AEF9723CEC9FBC78AABB" type="DOI">10.11646/zootaxa.3964.3.1</mods:identifier>
<mods:identifier id="37B8A881B217B4E72FFB477729278AA6" type="GBIF-Dataset">d80458ea-4fb4-4999-92c2-b8445c9dd19a</mods:identifier>
<mods:identifier id="9BA0D7AC511A17F04FE8813976C84A5B" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="D355EE3050FDB62A54E14583FE4AA89A" type="Zenodo-Dep">231957</mods:identifier>
<mods:identifier id="63574F163DAA2F1537F2CAD86DB42F47" type="ZooBank">88702846-1180-400A-B7DB-4154ACD4EC49</mods:identifier>
</mods:mods>
<treatment id="EC3687B45819FF89FF3DFD1DFC230A88" ID-DOI="http://doi.org/10.5281/zenodo.6100359" ID-GBIF-Taxon="119617613" ID-Zenodo-Dep="6100359" LSID="urn:lsid:plazi:treatment:EC3687B45819FF89FF3DFD1DFC230A88" httpUri="http://treatment.plazi.org/id/EC3687B45819FF89FF3DFD1DFC230A88" lastPageId="7" lastPageNumber="308" pageId="4" pageNumber="305">
<subSubSection id="2C8565295819FF8AFF3DFD1DFE7309F0" pageId="4" pageNumber="305" type="nomenclature">
<paragraph id="642036A25819FF8AFF3DFD1DFDD8091E" blockId="4.[151,563,1372,1432]" box="[151,563,1372,1398]" pageId="4" pageNumber="305">
<heading id="3F6881CE5819FF8AFF3DFD1DFDD8091E" bold="true" box="[151,563,1372,1398]" fontSize="11" level="1" pageId="4" pageNumber="305" reason="1">
<emphasis id="56EBEAB05819FF8AFF3DFD1DFDD8091E" bold="true" box="[151,563,1372,1398]" pageId="4" pageNumber="305">
<taxonomicName id="A39F4D215819FF8AFF3DFD1DFE77091E" box="[151,412,1372,1398]" class="Diplopoda" family="Zephronidae" genus="Zephronia" kingdom="Animalia" order="Sphaerotheriida" pageId="4" pageNumber="305" phylum="Arthropoda" rank="species" species="nepalensis" status="sp. nov.">
<emphasis id="56EBEAB05819FF8AFF3DFD1DFE77091E" bold="true" box="[151,412,1372,1398]" italics="true" pageId="4" pageNumber="305">Zephronia nepalensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4DD857CB5819FF8AFE0EFD1DFDD8091E" box="[420,563,1372,1398]" pageId="4" pageNumber="305" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="642036A25819FF8AFF3DFD3EFE7309F0" blockId="4.[151,563,1372,1432]" box="[151,408,1407,1432]" pageId="4" pageNumber="305">Figures: 2B, 3A, 5A–F</paragraph>
</subSubSection>
<subSubSection id="2C8565295819FF8AFF3DFD83FBFE09B3" box="[151,1045,1474,1499]" pageId="4" pageNumber="305" type="etymology">
<paragraph id="642036A25819FF8AFF3DFD83FBFE09B3" blockId="4.[151,1436,1474,2031]" box="[151,1045,1474,1499]" pageId="4" pageNumber="305">
<emphasis id="56EBEAB05819FF8AFF3DFD83FEFD09B3" bold="true" box="[151,278,1474,1499]" pageId="4" pageNumber="305">Etymology</emphasis>
.
<taxonomicName id="A39F4D215819FF8AFE8EFD83FE7109B3" box="[292,410,1474,1499]" class="Diplopoda" family="Zephronidae" genus="Zephronia" kingdom="Animalia" order="Sphaerotheriida" pageId="4" pageNumber="305" phylum="Arthropoda" rank="species" species="nepalensis">
<emphasis id="56EBEAB05819FF8AFE8EFD83FE7109B3" box="[292,410,1474,1499]" italics="true" pageId="4" pageNumber="305">nepalensis</emphasis>
</taxonomicName>
, adjective, after the origin of the
<typeStatus id="BB2488005819FF8AFCBAFD82FCAB09B3" box="[784,832,1475,1499]" pageId="4" pageNumber="305">type</typeStatus>
specimens,
<collectingCountry id="1C8876325819FF8AFC60FD83FBFA09B3" box="[970,1041,1474,1499]" name="Nepal" pageId="4" pageNumber="305">Nepal</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="2C8565295819FF8AFF6DFDA4FE660A9E" pageId="4" pageNumber="305" type="materials_examined">
<paragraph id="642036A25819FF8AFF6DFDA4FAEC0A4A" blockId="4.[151,1436,1474,2031]" pageId="4" pageNumber="305">
<emphasis id="56EBEAB05819FF8AFF6DFDA4FE400996" bold="true" box="[199,427,1509,1534]" pageId="4" pageNumber="305">Material examined</emphasis>
.
<typeStatus id="BB2488005819FF8AFE16FDA4FDC20996" box="[444,553,1509,1534]" pageId="4" pageNumber="305" type="holotype">Holotype</typeStatus>
: 1 M (
<emphasis id="56EBEAB05819FF8AFDD2FDA4FC950996" bold="true" box="[632,894,1509,1534]" pageId="4" pageNumber="305">
deposited at the
<collectionCode id="028EAE675819FF8AFCE9FDA4FC950996" LSID="urn:lsid:biocol.org:col:34838" box="[835,894,1509,1534]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="4" pageNumber="305">SMF</collectionCode>
</emphasis>
),
<emphasis id="56EBEAB05819FF8AFC32FDA4FC510996" bold="true" box="[920,954,1509,1534]" pageId="4" pageNumber="305">N1</emphasis>
field label
<emphasis id="56EBEAB05819FF8AFBEAFDA4FB9A0996" bold="true" box="[1088,1137,1509,1534]" pageId="4" pageNumber="305">234,</emphasis>
<collectingCountry id="1C8876325819FF8AFBD0FDA4FB2B0996" box="[1146,1216,1509,1534]" name="Nepal" pageId="4" pageNumber="305">Nepal</collectingCountry>
, Ilam District, Mai Pokhari [
<geoCoordinate id="01AB50655819FF8AFF56FE4BFE4A0A4A" box="[252,417,1545,1570]" direction="north" orientation="latitude" pageId="4" pageNumber="305" precision="1" value="27.048512">27° 2'54.64&quot;N</geoCoordinate>
,
<geoCoordinate id="01AB50655819FF8AFE06FE48FDAC0A4A" box="[428,583,1545,1570]" direction="east" orientation="longitude" pageId="4" pageNumber="305" precision="1" value="87.90027">87°54'0.97&quot;E</geoCoordinate>
],
<quantity id="A3679B475819FF8AFDF3FE48FD1D0A49" box="[601,758,1545,1570]" metricMagnitude="3" metricUnit="m" metricValue="2.2" metricValueMax="2.25" metricValueMin="2.15" pageId="4" pageNumber="305" unit="m" value="2200.0" valueMax="2250.0" valueMin="2150.0">2150–2250 m</quantity>
, leg. J. Martens &amp; B. Daams,
<date id="102110625819FF8AFBEDFE48FAE80A4A" box="[1095,1283,1545,1570]" pageId="4" pageNumber="305" value="1983-08-23" valueMax="1983-08-25" valueMin="1983-08-23">23.–25.viii.1983</date>
.
</paragraph>
<paragraph id="642036A25819FF8AFF6DFE6FFB990A2E" blockId="4.[151,1436,1474,2031]" box="[199,1138,1581,1606]" pageId="4" pageNumber="305">
<typeStatus id="BB2488005819FF8AFF6DFE6FFED30A2E" box="[199,312,1582,1606]" pageId="4" pageNumber="305" type="paratype">Paratypes</typeStatus>
: 1 M (
<emphasis id="56EBEAB05819FF8AFE2BFE6CFDA30A2E" bold="true" box="[385,584,1581,1606]" pageId="4" pageNumber="305">
<collectionCode id="028EAE675819FF8AFE2BFE6CFE3E0A2E" LSID="urn:lsid:biocol.org:col:34613" box="[385,469,1581,1606]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="4" pageNumber="305">ZFMK</collectionCode>
Myr4173
</emphasis>
), 1 F (
<emphasis id="56EBEAB05819FF8AFD3EFE6CFC8D0A2E" bold="true" box="[660,870,1581,1606]" pageId="4" pageNumber="305">
<collectionCode id="028EAE675819FF8AFD3EFE6CFD030A2E" LSID="urn:lsid:biocol.org:col:34613" box="[660,744,1581,1606]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="4" pageNumber="305">ZFMK</collectionCode>
Myr4172)
</emphasis>
, same data as
<typeStatus id="BB2488005819FF8AFBA2FE6CFB850A2E" box="[1032,1134,1581,1606]" pageId="4" pageNumber="305" type="holotype">holotype</typeStatus>
.
</paragraph>
<paragraph id="642036A25819FF8AFF6DFE11FE660A9E" blockId="4.[151,1436,1474,2031]" pageId="4" pageNumber="305">
Other material examined: 2 M (
<emphasis id="56EBEAB05819FF8AFD96FE11FCE30A01" bold="true" box="[572,776,1616,1641]" pageId="4" pageNumber="305">
<collectionCode id="028EAE675819FF8AFD96FE11FD7B0A01" LSID="urn:lsid:biocol.org:col:34613" box="[572,656,1616,1641]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="4" pageNumber="305">ZFMK</collectionCode>
Myr4175
</emphasis>
),
<emphasis id="56EBEAB05819FF8AFC89FE11FCAE0A01" bold="true" box="[803,837,1616,1641]" pageId="4" pageNumber="305">N4</emphasis>
field label
<emphasis id="56EBEAB05819FF8AFC64FE11FC130A01" bold="true" box="[974,1016,1616,1641]" pageId="4" pageNumber="305">258</emphasis>
,
<collectingCountry id="1C8876325819FF8AFBA0FE11FBBB0A01" box="[1034,1104,1616,1641]" name="Nepal" pageId="4" pageNumber="305">Nepal</collectingCountry>
, Panchthar District, between Deorali, Puspati and Sheldoti [
<geoCoordinate id="01AB50655819FF8AFE47FE35FD6F0AE4" box="[493,644,1652,1676]" direction="north" orientation="latitude" pageId="4" pageNumber="305" precision="1" value="27.116667">27° 7'0.00&quot;N</geoCoordinate>
,
<geoCoordinate id="01AB50655819FF8AFD25FE35FCC10AE5" box="[655,810,1652,1677]" direction="east" orientation="longitude" pageId="4" pageNumber="305" precision="1" value="87.88333">87°53'0.00&quot;E</geoCoordinate>
], 2600–
<quantity id="A3679B475819FF8AFC29FE35FC310AE4" box="[899,986,1652,1677]" metricMagnitude="3" metricUnit="m" metricValue="2.5" pageId="4" pageNumber="305" unit="m" value="2500.0">2500 m</quantity>
,
<taxonomicName id="A39F4D215819FF8AFC4FFE34FB6E0AE5" box="[997,1157,1652,1677]" class="Pinopsida" family="Pinaceae" genus="Tsuga" kingdom="Plantae" order="Pinales" pageId="4" pageNumber="305" phylum="Tracheophyta" rank="species" species="dumosa">
<emphasis id="56EBEAB05819FF8AFC4FFE34FB6E0AE5" box="[997,1157,1652,1677]" italics="true" pageId="4" pageNumber="305">Tsuga dumosa</emphasis>
</taxonomicName>
forest, leg. J. Martens &amp; B. Daams,
<date id="102110625819FF8AFEB1FED9FE5A0AD8" box="[283,433,1688,1712]" pageId="4" pageNumber="305" value="1983-08-28">28.VIII.1983</date>
; 1 F (
<emphasis id="56EBEAB05819FF8AFE57FED6FD230AD8" bold="true" box="[509,712,1687,1712]" pageId="4" pageNumber="305">
<collectionCode id="028EAE675819FF8AFE57FED6FDBA0AD8" LSID="urn:lsid:biocol.org:col:34613" box="[509,593,1687,1712]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="4" pageNumber="305">ZFMK</collectionCode>
Myr4174
</emphasis>
),
<emphasis id="56EBEAB05819FF8AFD4EFED6FCEC0AD8" bold="true" box="[740,775,1687,1712]" pageId="4" pageNumber="305">N5</emphasis>
field label
<emphasis id="56EBEAB05819FF8AFC3BFED6FC500AD8" bold="true" box="[913,955,1687,1712]" pageId="4" pageNumber="305">270</emphasis>
,
<collectingCountry id="1C8876325819FF8AFC66FED6FBF90AD8" box="[972,1042,1687,1712]" name="Nepal" pageId="4" pageNumber="305">Nepal</collectingCountry>
, Taplejung District, Kabeli Khola valley, above Yamputhin [
<geoCoordinate id="01AB50655819FF8AFE65FEFBFD900ABB" box="[463,635,1722,1747]" direction="north" orientation="latitude" pageId="4" pageNumber="305" precision="1" value="27.433332">27°25'60.00&quot;N</geoCoordinate>
,
<geoCoordinate id="01AB50655819FF8AFD27FEFBFCC30ABB" box="[653,808,1722,1747]" direction="east" orientation="longitude" pageId="4" pageNumber="305" precision="1" value="87.9">87°54'0.00&quot;E</geoCoordinate>
], 2000-
<quantity id="A3679B475819FF8AFC29FEFAFC0B0ABA" box="[899,992,1723,1747]" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="4" pageNumber="305" unit="m" value="1700.0">1700 m</quantity>
, forest in ravine, leg.
<date id="102110625819FF8AFB5CFEFAFA930ABB" box="[1270,1400,1723,1747]" pageId="4" pageNumber="305" value="1983-09-03">03.IX.1983</date>
, J. Martens &amp; B. Daams.
</paragraph>
</subSubSection>
<subSubSection id="2C8565295819FF8AFF6DFF40FAC30B87" pageId="4" pageNumber="305" type="diagnosis">
<paragraph id="642036A25819FF8AFF6DFF40FAC30B87" blockId="4.[151,1436,1474,2031]" pageId="4" pageNumber="305">
<emphasis id="56EBEAB05819FF8AFF6DFF40FED10B72" bold="true" box="[199,314,1793,1818]" pageId="4" pageNumber="305">Diagnosis</emphasis>
. Descriptions of the apparently widespread
<taxonomicName id="A39F4D215819FF8AFC84FF40FB670B72" authority="Butler, 1882" authorityName="Butler" authorityYear="1882" box="[814,1164,1793,1818]" class="Diplopoda" family="Zephronidae" genus="Zephronia" kingdom="Animalia" order="Sphaerotheriida" pageId="4" pageNumber="305" phylum="Arthropoda" rank="species" species="tumida">
<emphasis id="56EBEAB05819FF8AFC84FF40FC130B72" box="[814,1016,1793,1818]" italics="true" pageId="4" pageNumber="305">Zephronia tumida</emphasis>
<bibRefCitation id="000E4B535819FF8AFBAAFF40FB670B72" author="Butler" box="[1024,1164,1793,1818]" pageId="4" pageNumber="305" refString="Butler, A. G. (1882) Descriptions of new species of Myriopoda of the genus Zephronia from lndia and Sumatra. Annals and Magazine of Natural History, Series 5, 9, 196 - 198." type="journal article" year="1882">Butler, 1882</bibRefCitation>
</taxonomicName>
seem to resemble those of
<taxonomicName id="A39F4D215819FF8AFF1EFF67FEA30B56" box="[180,328,1829,1854]" class="Diplopoda" family="Zephronidae" genus="Zephronia" kingdom="Animalia" order="Sphaerotheriida" pageId="4" pageNumber="305" phylum="Arthropoda" rank="species" species="nepalensis">
<emphasis id="56EBEAB05819FF8AFF1EFF67FEA30B56" box="[180,328,1829,1854]" italics="true" pageId="4" pageNumber="305">Z. nepalensis</emphasis>
</taxonomicName>
. However, a comparison of the
<typeStatus id="BB2488005819FF8AFD01FF67FD300B56" box="[683,731,1830,1854]" pageId="4" pageNumber="305">type</typeStatus>
of
<taxonomicName id="A39F4D215819FF8AFCAAFF67FC800B56" box="[768,875,1829,1854]" class="Diplopoda" family="Zephronidae" genus="Zephronia" kingdom="Animalia" order="Sphaerotheriida" pageId="4" pageNumber="305" phylum="Arthropoda" rank="species" species="tumida">
<emphasis id="56EBEAB05819FF8AFCAAFF67FC800B56" box="[768,875,1829,1854]" italics="true" pageId="4" pageNumber="305">Z. tumida</emphasis>
</taxonomicName>
with
<taxonomicName id="A39F4D215819FF8AFC03FF67FBD70B56" box="[937,1084,1829,1854]" class="Diplopoda" family="Zephronidae" genus="Zephronia" kingdom="Animalia" order="Sphaerotheriida" pageId="4" pageNumber="305" phylum="Arthropoda" rank="species" species="nepalensis">
<emphasis id="56EBEAB05819FF8AFC03FF67FBD70B56" box="[937,1084,1829,1854]" italics="true" pageId="4" pageNumber="305">Z. nepalensis</emphasis>
</taxonomicName>
clearly shows that both species differ vastly in their locking carinae, size, legs and numerous other characters.
<taxonomicName id="A39F4D215819FF8AFB91FF0BFB380B09" box="[1083,1235,1864,1889]" class="Diplopoda" family="Zephronidae" genus="Zephronia" kingdom="Animalia" order="Sphaerotheriida" pageId="4" pageNumber="305" phylum="Arthropoda" rank="species" species="nepalensis">
<emphasis id="56EBEAB05819FF8AFB91FF0BFB380B09" box="[1083,1235,1864,1889]" italics="true" pageId="4" pageNumber="305">Z. nepalensis</emphasis>
</taxonomicName>
has a differently shaped vulval operculum from all other
<taxonomicName id="A39F4D215819FF8AFDD4FF2DFD190BED" box="[638,754,1900,1925]" class="Diplopoda" family="Zephronidae" genus="Zephronia" kingdom="Animalia" order="Sphaerotheriida" pageId="4" pageNumber="305" phylum="Arthropoda" rank="genus">
<emphasis id="56EBEAB05819FF8AFDD4FF2DFD190BED" box="[638,754,1900,1925]" italics="true" pageId="4" pageNumber="305">Zephronia</emphasis>
</taxonomicName>
species with two locking carinae described from nearby Darjeeling, such as
<taxonomicName id="A39F4D215819FF8AFED6FFD1FD770BC0" authority="Attems, 1936" authorityName="Attems" authorityYear="1936" box="[380,668,1935,1960]" class="Diplopoda" family="Zephronidae" genus="Zephronia" kingdom="Animalia" order="Sphaerotheriida" pageId="4" pageNumber="305" phylum="Arthropoda" rank="species" species="debilis">
<emphasis id="56EBEAB05819FF8AFED6FFD1FE0C0BC0" box="[380,487,1935,1960]" italics="true" pageId="4" pageNumber="305">Z. debilis</emphasis>
(
<bibRefCitation id="000E4B535819FF8AFE52FFD1FD7F0BC0" author="Attems" box="[504,660,1935,1960]" pageId="4" pageNumber="305" refString="Attems, C. (1936) Diplopoda of India. Memoirs of the Indian Museum, 11, 133 - 323." type="journal article" year="1936">Attems, 1936</bibRefCitation>
)
</taxonomicName>
and
<taxonomicName id="A39F4D215819FF8AFD72FFD1FC100BC0" authority="Butler, 1872" authorityName="Butler" authorityYear="1872" box="[728,1019,1935,1960]" class="Diplopoda" family="Zephronidae" genus="Zephronia" kingdom="Animalia" order="Sphaerotheriida" pageId="4" pageNumber="305" phylum="Arthropoda" rank="species" species="nigrinota">
<emphasis id="56EBEAB05819FF8AFD72FFD1FC8B0BCF" box="[728,864,1936,1959]" italics="true" pageId="4" pageNumber="305">Z. nigrinota</emphasis>
Butler, 1872
</taxonomicName>
. The species
<taxonomicName id="A39F4D215819FF8AFB32FFD1FAC50BC0" box="[1176,1326,1935,1960]" class="Diplopoda" family="Zephronidae" genus="Zephronia" kingdom="Animalia" order="Sphaerotheriida" pageId="4" pageNumber="305" phylum="Arthropoda" rank="species" species="nepalensis">
<emphasis id="56EBEAB05819FF8AFB32FFD1FAC50BC0" box="[1176,1326,1935,1960]" italics="true" pageId="4" pageNumber="305">Z. nepalensis</emphasis>
</taxonomicName>
seems to resemble most closely is
<taxonomicName id="A39F4D215819FF8AFE17FFF5FD300BA4" authority="Attems, 1936" authorityName="Attems" authorityYear="1936" box="[445,731,1971,1996]" class="Diplopoda" family="Zephronidae" genus="Zephronia" kingdom="Animalia" order="Sphaerotheriida" pageId="4" pageNumber="305" phylum="Arthropoda" rank="species" species="alticola">
<emphasis id="56EBEAB05819FF8AFE17FFF5FDDF0BA4" box="[445,564,1971,1996]" italics="true" pageId="4" pageNumber="305">Z. alticola</emphasis>
<bibRefCitation id="000E4B535819FF8AFD94FFF2FD300BA4" author="Attems" box="[574,731,1971,1996]" pageId="4" pageNumber="305" refString="Attems, C. (1936) Diplopoda of India. Memoirs of the Indian Museum, 11, 133 - 323." type="journal article" year="1936">Attems, 1936</bibRefCitation>
</taxonomicName>
, which has a centrally elongated and 'flapped' operculum.
<taxonomicName id="A39F4D215819FF8AFA2FFFF5FEE60B87" class="Diplopoda" family="Zephronidae" genus="Zephronia" kingdom="Animalia" order="Sphaerotheriida" pageId="4" pageNumber="305" phylum="Arthropoda" rank="species" species="nepalensis">
<emphasis id="56EBEAB05819FF8AFA2FFFF5FEE60B87" italics="true" pageId="4" pageNumber="305">Z. nepalensis</emphasis>
</taxonomicName>
is distinguishable from the three aforementioned species as an apical spine on leg 3 is absent.
</paragraph>
</subSubSection>
<caption id="30E0662A5818FF8BFF3DFF25FC8A0B9D" httpUri="https://zenodo.org/record/231961/files/figure.png" pageId="5" pageNumber="306" targetBox="[163,1423,195,1868]" targetPageId="5">
<paragraph id="642036A25818FF8BFF3DFF25FC8A0B9D" blockId="5.[151,1436,1892,2037]" pageId="5" pageNumber="306">
<emphasis id="56EBEAB05818FF8BFF3DFF25FEE60B12" bold="true" box="[151,269,1892,1914]" pageId="5" pageNumber="306">FIGURE 4</emphasis>
.
<taxonomicName id="A39F4D215818FF8BFEB0FF25FDBC0B12" authority="Butler, 1882" authorityName="Butler" authorityYear="1882" box="[282,599,1892,1914]" class="Diplopoda" family="Zephronidae" genus="Zephronia" kingdom="Animalia" order="Sphaerotheriida" pageId="5" pageNumber="306" phylum="Arthropoda" rank="species" species="tumida">
<emphasis id="56EBEAB05818FF8BFEB0FF25FE240B12" box="[282,463,1892,1914]" italics="true" pageId="5" pageNumber="306">Zephronia tumida</emphasis>
Butler, 1882
</taxonomicName>
, holotype male,
<emphasis id="56EBEAB05818FF8BFD55FF25FC250B12" bold="true" box="[767,974,1892,1914]" pageId="5" pageNumber="306">NHML BM1882.11</emphasis>
.
<emphasis id="56EBEAB05818FF8BFC71FF25FC060B12" bold="true" box="[987,1005,1892,1914]" pageId="5" pageNumber="306">A</emphasis>
: left leg 9, posterior view;
<emphasis id="56EBEAB05818FF8BFAA9FF25FAFF0B12" bold="true" box="[1283,1300,1892,1914]" pageId="5" pageNumber="306">B</emphasis>
: right half of anterior telopod, anterior view;
<emphasis id="56EBEAB05818FF8BFE7DFFC2FE020BF1" bold="true" box="[471,489,1923,1945]" pageId="5" pageNumber="306">C</emphasis>
: tip of right anterior telopod, posterior view;
<emphasis id="56EBEAB05818FF8BFC19FFC2FC2E0BF1" bold="true" box="[947,965,1923,1945]" pageId="5" pageNumber="306">D</emphasis>
: right anterior telopod, podomeres 2–4, lateral view;
<emphasis id="56EBEAB05818FF8BFF7FFFE3FF0D0BD0" bold="true" box="[213,230,1954,1976]" pageId="5" pageNumber="306">E</emphasis>
: right posterior telopod, podomeres 2–4, posterior view;
<emphasis id="56EBEAB05818FF8BFC82FFE3FCDC0BD0" bold="true" box="[808,823,1954,1976]" pageId="5" pageNumber="306">F</emphasis>
: right posterior telopod, anterior view.
<emphasis id="56EBEAB05818FF8BFB6EFFE3FAB60BD0" bold="true" box="[1220,1373,1954,1976]" pageId="5" pageNumber="306">Abbreviations</emphasis>
: Co = coxa; cr-T = crenulated teeth; Fe = femur; ML = membranous ledge or lobe; Po = postfemur; Pre = prefemur; Syn = syncoxite; Ta = tarsus; Ti = tibia; numbers refer to podomere. Scale bars = 1 mm.
</paragraph>
</caption>
<caption id="30E0662A581BFF88FF3DFF7EFC7E0BB8" httpUri="https://zenodo.org/record/231962/files/figure.png" pageId="6" pageNumber="307" targetBox="[166,1417,193,1833]" targetPageId="6">
<paragraph id="642036A2581BFF88FF3DFF7EFC7E0BB8" blockId="6.[151,1436,1855,2000]" pageId="6" pageNumber="307">
<emphasis id="56EBEAB0581BFF88FF3DFF7EFEE60B3D" bold="true" box="[151,269,1855,1877]" pageId="6" pageNumber="307">FIGURE 5</emphasis>
.
<taxonomicName id="A39F4D21581BFF88FEB0FF7EFE1F0B3D" box="[282,500,1855,1877]" class="Diplopoda" family="Zephronidae" genus="Zephronia" kingdom="Animalia" order="Sphaerotheriida" pageId="6" pageNumber="307" phylum="Arthropoda" rank="species" species="nepalensis" status="sp. nov.">
<emphasis id="56EBEAB0581BFF88FEB0FF7EFE1F0B3D" box="[282,500,1855,1877]" italics="true" pageId="6" pageNumber="307">Zephronia nepalensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4DD857CB581BFF88FE50FF7EFDDF0B3D" box="[506,564,1855,1877]" pageId="6" pageNumber="307" rank="species">
<emphasis id="56EBEAB0581BFF88FE50FF7EFDDF0B3D" bold="true" box="[506,564,1855,1877]" pageId="6" pageNumber="307">n. sp.</emphasis>
</taxonomicNameLabel>
, B = female paratype,
<emphasis id="56EBEAB0581BFF88FCB7FF7EFC340B3D" bold="true" box="[797,991,1855,1877]" pageId="6" pageNumber="307">ZFMK MYR4172</emphasis>
, A, C–F male holotype,
<emphasis id="56EBEAB0581BFF88FB71FF7EFAE50B3D" bold="true" box="[1243,1294,1855,1877]" pageId="6" pageNumber="307">SMF</emphasis>
.
<emphasis id="56EBEAB0581BFF88FAB1FF7EFAC60B3D" bold="true" box="[1307,1325,1855,1877]" pageId="6" pageNumber="307">A</emphasis>
: left leg 9, posterior view;
<emphasis id="56EBEAB0581BFF88FE9FFF1FFEAD0B1C" bold="true" box="[309,326,1886,1908]" pageId="6" pageNumber="307">B</emphasis>
: right coxa and prefemur 2, with vulva;
<emphasis id="56EBEAB0581BFF88FD74FF1FFD1B0B1C" bold="true" box="[734,752,1886,1908]" pageId="6" pageNumber="307">C</emphasis>
: left half of anterior telopod, anterior view;
<emphasis id="56EBEAB0581BFF88FB04FF1FFB2B0B1C" bold="true" box="[1198,1216,1886,1908]" pageId="6" pageNumber="307">D</emphasis>
: left anterior telopod, podomeres 2–4, lateral view;
<emphasis id="56EBEAB0581BFF88FE68FF3DFE380BFA" bold="true" box="[450,467,1916,1938]" pageId="6" pageNumber="307">E</emphasis>
: right posterior telopod, anterior view;
<emphasis id="56EBEAB0581BFF88FCC9FF3DFC990BFA" bold="true" box="[867,882,1916,1938]" pageId="6" pageNumber="307">F</emphasis>
: left posterior telopod, posterior view.
<emphasis id="56EBEAB0581BFF88FB56FF3DFA7E0BFA" bold="true" box="[1276,1429,1916,1938]" pageId="6" pageNumber="307">Abbreviations</emphasis>
: Bu = bursa; Co = coxa; cr-T = crenulated teeth; IH = position of inner horns; ML = membranous lobe; O = operculum; Pre = prefemur; Syn = syncoxite; numbers refer to podomere. Scale bars = 1 mm.
</paragraph>
</caption>
<subSubSection id="2C856529581AFF89FF6DF8D6FCF90AD4" pageId="7" pageNumber="308" type="description">
<paragraph id="642036A2581AFF89FF6DF8D6FC970CBD" blockId="7.[151,1437,151,1760]" pageId="7" pageNumber="308">
<emphasis id="56EBEAB0581AFF89FF6DF8D6FEBA0CD8" bold="true" box="[199,337,151,176]" pageId="7" pageNumber="308">Description</emphasis>
. Measurements: male
<typeStatus id="BB248800581AFF89FDFBF8D6FD530CD8" box="[593,696,151,176]" pageId="7" pageNumber="308" type="holotype">holotype</typeStatus>
:
<quantity id="A3679B47581AFF89FD69F8D6FCC30CC7" box="[707,808,151,176]" metricMagnitude="-2" metricUnit="m" metricValue="2.35" pageId="7" pageNumber="308" unit="mm" value="23.5">23.5 mm</quantity>
long,
<quantity id="A3679B47581AFF89FCDAF8D9FC3E0CC7" box="[880,981,152,176]" metricMagnitude="-2" metricUnit="m" metricValue="1.04" pageId="7" pageNumber="308" unit="mm" value="10.4">10.4 mm</quantity>
(2nd) wide,
<quantity id="A3679B47581AFF89FBCCF8D6FB560CC7" box="[1126,1213,151,176]" metricMagnitude="-3" metricUnit="m" metricValue="6.3" pageId="7" pageNumber="308" unit="mm" value="6.3">6.3 mm</quantity>
(2nd) high. Female
<typeStatus id="BB248800581AFF89FF3DF8FCFF170CBD" box="[151,252,189,213]" pageId="7" pageNumber="308" type="paratype">paratype</typeStatus>
:
<quantity id="A3679B47581AFF89FEACF8FDFE810CBC" box="[262,362,188,213]" metricMagnitude="-2" metricUnit="m" metricValue="2.57" pageId="7" pageNumber="308" unit="mm" value="25.7">25.7 mm</quantity>
long,
<quantity id="A3679B47581AFF89FE1AF8FDFDF80CBC" box="[432,531,188,212]" metricMagnitude="-2" metricUnit="m" metricValue="1.1199999999999999" pageId="7" pageNumber="308" unit="mm" value="11.2">11.2 mm</quantity>
(2nd) wide,
<quantity id="A3679B47581AFF89FD08F8FDFD130CBC" box="[674,760,188,212]" metricMagnitude="-3" metricUnit="m" metricValue="7.1" pageId="7" pageNumber="308" unit="mm" value="7.1">7.1 mm</quantity>
(2nd) high.
</paragraph>
<paragraph id="642036A2581AFF89FF6DF89EFB6B0D75" blockId="7.[151,1437,151,1760]" pageId="7" pageNumber="308">Colouration faded because of long preservation in ethanol. Tergites greenish-brown, head and collum darker, of chestnut colour. Underside, legs and antennae light brown, tip of appendages greenish.</paragraph>
<paragraph id="642036A2581AFF89FF3DF966FB020DC5" blockId="7.[151,1437,151,1760]" pageId="7" pageNumber="308">
Head: Eyes with ca. 60 ocelli. Antennae short, protruding back to coxa 4. Antennomeres 1 and 2 with few longer setae, 3–6 more densely pubescent. Antennomere 6 laterally flattened, 2 times longer than antennomere 5. Size relations of antennomeres 1=2=3=4=5&lt;&lt;6. Disc with 45/43 apical cones (male
<typeStatus id="BB248800581AFF89FBA4F92EFB9F0DE0" box="[1038,1140,367,392]" pageId="7" pageNumber="308" type="holotype">holotype</typeStatus>
), female with 22/36 apical cones. Mouthparts not dissected. Palpi of gnathochilarium with sensory cones arranged in clusters.
</paragraph>
<paragraph id="642036A2581AFF89FF6DF9F6FD400DA7" blockId="7.[151,1437,151,1760]" box="[199,683,439,464]" pageId="7" pageNumber="308">Collum glabrous except for marginal setae.</paragraph>
<paragraph id="642036A2581AFF89FF6DF99DFE0C0E55" blockId="7.[151,1437,151,1760]" pageId="7" pageNumber="308">Thoracic shield surface like those of following tergites. Grooves deep, without any ledges. Tergites 3–12 anteriorly covered in longer hair, posteriorly smooth, glabrous, shiny. Paratergite tips of midbody tergites only weakly projecting posteriorly.</paragraph>
<paragraph id="642036A2581AFF89FF6DFA06FAE30EED" blockId="7.[151,1437,151,1760]" pageId="7" pageNumber="308">
Anal shield massive, well-rounded (
<figureCitation id="FCA42A27581AFF89FDC7FA06FD580E08" box="[621,691,583,608]" captionStart="FIGURE 3" captionStartId="4.[151,250,614,636]" captionTargetBox="[150,1435,193,592]" captionTargetId="figure@4.[150,1436,193,593]" captionTargetPageId="4" captionText="FIGURE 3. Multi-layer photographs, anal shield and posterior tergites, lateral view. All specimens were males. A: Zephronia nepalensis n. sp., holotype, SMF; B: Kophosphaera politissima Attems, 1935, ZFMK MYR 4176; C: K. shivapuri n. sp., holotype, SMF; D: K. martensi n. sp., holotype, SMF. Abbreviations: 12 T = tergite 12; AS = anal shield. Not to scale." httpUri="https://zenodo.org/record/231960/files/figure.png" pageId="7" pageNumber="308">Fig. 3</figureCitation>
A), shiny and glabrous. Underside carrying two well-developed locking carinae, posterior carina 2–2.5 times longer than anterior one. Carina located close to margin.
</paragraph>
<paragraph id="642036A2581AFF89FF6DFACEFC4C0E98" blockId="7.[151,1437,151,1760]" pageId="7" pageNumber="308">
Endotergum inner section lacking spines, only with few very short setae (
<figureCitation id="FCA42A27581AFF89FBB9FACEFBB10EC0" box="[1043,1114,655,680]" captionStart="FIGURE 2" captionStartId="3.[151,250,1883,1905]" captionTargetBox="[170,1425,769,1857]" captionTargetId="figure@3.[170,1427,769,1858]" captionTargetPageId="3" captionText="FIGURE 2. SEM of tergites or endoterga of different Zephronia and Kophosphaera species. A: Zephronia montana (Karsch, 1881), ZMB 55, tergite surface; B: Zephronia nepalensis n. sp., holotype, SMF, endotergum; C: Kophosphaera politissima Attems, 1935, ZFMK MYR 4176, endotergum; D: K. shivapuri n. sp., holotype, SMF, endotergum; E: K. martensi n. sp., holotype, SMF, endotergum." httpUri="https://zenodo.org/record/231959/files/figure.png" pageId="7" pageNumber="308">Fig. 2</figureCitation>
B). Between ridge and inner section, single row of large, elliptical cuticular impressions. Externally, two rows of marginal bristles (
<figureCitation id="FCA42A27581AFF89FA9EFAF5FA900EA4" box="[1332,1403,692,717]" captionStart="FIGURE 2" captionStartId="3.[151,250,1883,1905]" captionTargetBox="[170,1425,769,1857]" captionTargetId="figure@3.[170,1427,769,1858]" captionTargetPageId="3" captionText="FIGURE 2. SEM of tergites or endoterga of different Zephronia and Kophosphaera species. A: Zephronia montana (Karsch, 1881), ZMB 55, tergite surface; B: Zephronia nepalensis n. sp., holotype, SMF, endotergum; C: Kophosphaera politissima Attems, 1935, ZFMK MYR 4176, endotergum; D: K. shivapuri n. sp., holotype, SMF, endotergum; E: K. martensi n. sp., holotype, SMF, endotergum." httpUri="https://zenodo.org/record/231959/files/figure.png" pageId="7" pageNumber="308">Fig. 2</figureCitation>
B). Bristles very long, strongly protruding above tergite margin (
<figureCitation id="FCA42A27581AFF89FCE8FA96FC6D0E98" box="[834,902,727,752]" captionStart="FIGURE 2" captionStartId="3.[151,250,1883,1905]" captionTargetBox="[170,1425,769,1857]" captionTargetId="figure@3.[170,1427,769,1858]" captionTargetPageId="3" captionText="FIGURE 2. SEM of tergites or endoterga of different Zephronia and Kophosphaera species. A: Zephronia montana (Karsch, 1881), ZMB 55, tergite surface; B: Zephronia nepalensis n. sp., holotype, SMF, endotergum; C: Kophosphaera politissima Attems, 1935, ZFMK MYR 4176, endotergum; D: K. shivapuri n. sp., holotype, SMF, endotergum; E: K. martensi n. sp., holotype, SMF, endotergum." httpUri="https://zenodo.org/record/231959/files/figure.png" pageId="7" pageNumber="308">Fig. 2</figureCitation>
B).
</paragraph>
<paragraph id="642036A2581AFF89FF6DFABDFC7E0F7C" blockId="7.[151,1437,151,1760]" box="[199,917,764,789]" pageId="7" pageNumber="308">First stigma-carrying plate with a well-rounded projecting apex.</paragraph>
<paragraph id="642036A2581AFF89FF6DFB5EFDF00F34" blockId="7.[151,1437,151,1760]" pageId="7" pageNumber="308">Pleurite 1 laterally elongated into a slender, well-rounded process. Pleurite 2 also weakly elongated into a broader and well-rounded process.</paragraph>
<paragraph id="642036A2581AFF89FF6DFB29FD610F85" blockId="7.[151,1437,151,1760]" pageId="7" pageNumber="308">
Leg 1 with 3 or 4, 2 with 4 or 5, 3 with 5 or 6 ventral spines. Leg pairs 4–21 with 6–8 ventral spines, many of them broken off. Legs 4–21 with single apical spine, which is absent in leg pairs 1–3. Coxa process weakly developed and well-rounded (
<figureCitation id="FCA42A27581AFF89FE4CFBEEFDC20FA0" box="[486,553,943,968]" captionStart="FIGURE 5" captionStartId="6.[151,250,1855,1877]" captionTargetBox="[166,1417,193,1833]" captionTargetId="figure@6.[166,1420,193,1834]" captionTargetPageId="6" captionText="FIGURE 5. Zephronia nepalensis n. sp., B = female paratype, ZFMK MYR 4172, A, C – F male holotype, SMF. A: left leg 9, posterior view; B: right coxa and prefemur 2, with vulva; C: left half of anterior telopod, anterior view; D: left anterior telopod, podomeres 2 – 4, lateral view; E: right posterior telopod, anterior view; F: left posterior telopod, posterior view. Abbreviations: Bu = bursa; Co = coxa; cr-T = crenulated teeth; IH = position of inner horns; ML = membranous lobe; O = operculum; Pre = prefemur; Syn = syncoxite; numbers refer to podomere. Scale bars = 1 mm." httpUri="https://zenodo.org/record/231962/files/figure.png" pageId="7" pageNumber="308">Fig. 5</figureCitation>
A), absent from leg pair 1 and 2. Femur 1.5, tarsus 3.6 times longer than wide. Femur with toothed ridge of medium length.
</paragraph>
<paragraph id="642036A2581AFF89FF6DFBB6FD87085C" blockId="7.[151,1437,151,1760]" pageId="7" pageNumber="308">Male gonopore simple pit located centrally at the mesal margin of coxa 2, gonopore unusually large, spanning almost half of the total height of the coxa.</paragraph>
<paragraph id="642036A2581AFF89FF6DFC7EFBB80964" blockId="7.[151,1437,151,1760]" pageId="7" pageNumber="308">
Anterior telopod (
<figureCitation id="FCA42A27581AFF89FE3CFC7EFE0B0830" box="[406,480,1087,1112]" captionStart="FIGURE 5" captionStartId="6.[151,250,1855,1877]" captionTargetBox="[166,1417,193,1833]" captionTargetId="figure@6.[166,1420,193,1834]" captionTargetPageId="6" captionText="FIGURE 5. Zephronia nepalensis n. sp., B = female paratype, ZFMK MYR 4172, A, C – F male holotype, SMF. A: left leg 9, posterior view; B: right coxa and prefemur 2, with vulva; C: left half of anterior telopod, anterior view; D: left anterior telopod, podomeres 2 – 4, lateral view; E: right posterior telopod, anterior view; F: left posterior telopod, posterior view. Abbreviations: Bu = bursa; Co = coxa; cr-T = crenulated teeth; IH = position of inner horns; ML = membranous lobe; O = operculum; Pre = prefemur; Syn = syncoxite; numbers refer to podomere. Scale bars = 1 mm." httpUri="https://zenodo.org/record/231962/files/figure.png" pageId="7" pageNumber="308">Figs 5</figureCitation>
C, D): syncoxite wide, with few small triangular spines. Podomere 1 only slightly larger than podomere 2 or 3, rectangular. Podomere 2 large, posterior process well visible in anterior view, apical part of process slender, strongly curved towards podomere 3 and 4. Process with few sclerotized spots. Podomere 3 as long as but much slenderer than podomere 2. Posterior side of podomere 3 carrying nine large crenulated teeth. Podomere 4 long, reaching 1/3 the size of podomere 3, towards process of podomere 2 with a membranous area carrying two long sclerotized spines. All podomeres densely covered with long setae.
</paragraph>
<paragraph id="642036A2581AFF89FF6DFD56FAF5098D" blockId="7.[151,1437,151,1760]" pageId="7" pageNumber="308">
Posterior telopod (
<figureCitation id="FCA42A27581AFF89FE3FFD56FE300958" box="[405,475,1303,1328]" captionStart="FIGURE 5" captionStartId="6.[151,250,1855,1877]" captionTargetBox="[166,1417,193,1833]" captionTargetId="figure@6.[166,1420,193,1834]" captionTargetPageId="6" captionText="FIGURE 5. Zephronia nepalensis n. sp., B = female paratype, ZFMK MYR 4172, A, C – F male holotype, SMF. A: left leg 9, posterior view; B: right coxa and prefemur 2, with vulva; C: left half of anterior telopod, anterior view; D: left anterior telopod, podomeres 2 – 4, lateral view; E: right posterior telopod, anterior view; F: left posterior telopod, posterior view. Abbreviations: Bu = bursa; Co = coxa; cr-T = crenulated teeth; IH = position of inner horns; ML = membranous lobe; O = operculum; Pre = prefemur; Syn = syncoxite; numbers refer to podomere. Scale bars = 1 mm." httpUri="https://zenodo.org/record/231962/files/figure.png" pageId="7" pageNumber="308">Figs 5</figureCitation>
E, F): typical for
<taxonomicName id="A39F4D21581AFF89FD30FD56FCE50958" box="[666,782,1303,1328]" class="Diplopoda" family="Zephronidae" genus="Zephronia" kingdom="Animalia" order="Sphaerotheriida" pageId="7" pageNumber="308" phylum="Arthropoda" rank="genus">
<emphasis id="56EBEAB0581AFF89FD30FD56FCE50958" box="[666,782,1303,1328]" italics="true" pageId="7" pageNumber="308">Zephronia</emphasis>
</taxonomicName>
, with four podomeres, two membranous ledges and a twotipped membranous ledge at the basis of the immovable finger. Podomere 3 straight, 2.5 times longer than wide. Hollowed-out inner margin with membranous ledge, no visible spines, and 15 or 16 crenulated teeth. Podomere 4 well-developed, lacking teeth, but with a short membranous ledge and two sclerotized spines. Immovable finger wide, running parallel to podomere 3 and 4. Podomere 1 and 2 completely covered with setae, only process of podomere
<quantity id="A3679B47581AFF89FEA4FD8DFED2098D" box="[270,313,1484,1509]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="7" pageNumber="308" unit="in" value="2.0">2 in</quantity>
posterior view glabrous. Podomere 3 and 4 with few marginal setae, otherwise glabrous.
</paragraph>
<paragraph id="642036A2581AFF89FF6DFDAEFC140A38" blockId="7.[151,1437,151,1760]" pageId="7" pageNumber="308">
Female with a large subanal plate that is not subdivided, but recessed at the central margin. Female vulva with black margins towards the opening. Operculum large, as large as basal plates (
<figureCitation id="FCA42A27581AFF89FBEEFE55FB670A45" box="[1092,1164,1556,1581]" captionStart="FIGURE 5" captionStartId="6.[151,250,1855,1877]" captionTargetBox="[166,1417,193,1833]" captionTargetId="figure@6.[166,1420,193,1834]" captionTargetPageId="6" captionText="FIGURE 5. Zephronia nepalensis n. sp., B = female paratype, ZFMK MYR 4172, A, C – F male holotype, SMF. A: left leg 9, posterior view; B: right coxa and prefemur 2, with vulva; C: left half of anterior telopod, anterior view; D: left anterior telopod, podomeres 2 – 4, lateral view; E: right posterior telopod, anterior view; F: left posterior telopod, posterior view. Abbreviations: Bu = bursa; Co = coxa; cr-T = crenulated teeth; IH = position of inner horns; ML = membranous lobe; O = operculum; Pre = prefemur; Syn = syncoxite; numbers refer to podomere. Scale bars = 1 mm." httpUri="https://zenodo.org/record/231962/files/figure.png" pageId="7" pageNumber="308">Fig. 5</figureCitation>
B). Operculum mesally projecting into a wide, triangular tip, protruding above basal half of prefemur.
</paragraph>
<paragraph id="642036A2581AFF89FF6DFE1DFCF90AD4" blockId="7.[151,1437,151,1760]" pageId="7" pageNumber="308">
<emphasis id="56EBEAB0581AFF89FF6DFE1DFE3F0A1D" bold="true" box="[199,468,1628,1653]" pageId="7" pageNumber="308">Intraspecific variation</emphasis>
. The males from the Parchthar District are smaller than the
<typeStatus id="BB248800581AFF89FB25FE1DFB1E0A1D" box="[1167,1269,1628,1653]" pageId="7" pageNumber="308" type="holotype">holotype</typeStatus>
; it is therefore unclear if the observed morphological differences (fewer teeth at the anterior and posterior telopods, fewer ocelli and apical cones on the antennae) are size-related or not.
</paragraph>
</subSubSection>
<subSubSection id="2C856529581AFF89FF6DFE86FC230A88" box="[199,968,1735,1760]" pageId="7" pageNumber="308" type="distribution">
<paragraph id="642036A2581AFF89FF6DFE86FC230A88" blockId="7.[151,1437,151,1760]" box="[199,968,1735,1760]" pageId="7" pageNumber="308">
<emphasis id="56EBEAB0581AFF89FF6DFE86FEB20A88" bold="true" box="[199,345,1735,1760]" pageId="7" pageNumber="308">Distribution</emphasis>
. Apparently widespread species in eastern-most
<collectingCountry id="1C887632581AFF89FCD7FE86FC2F0A88" box="[893,964,1735,1760]" name="Nepal" pageId="7" pageNumber="308">Nepal</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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