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<document id="884DB6760110B0656FF8325348A8F19A" ID="10.11646/zootaxa.4243.2.5SLASH10497" ID-CLB-Dataset="36273" ID-DOI="10.11646/zootaxa.4243.2.5" ID-GBIF-Dataset="d919071f-81f4-4f70-8eae-f944fa975413" ID-ISSN="1175-5326" ID-Zenodo-Dep="399142" ID-ZooBank="F4484A86-0615-4396-B312-69ED7E7A59F6" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1489567041515" checkinUser="plazi" docAuthor="Hernando, Carles &amp; Ribera, Ignacio" docDate="2017" docId="E7618794532EFFC2B2DB2F109090FCFD" docLanguage="en" docName="zootaxa.4243.2.5SLASH10497.pdf" docOrigin="Zootaxa 4243 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Caccothryptus thai Hernando &amp; Ribera, 2017, sp. n." docType="treatment" docVersion="7" lastPageNumber="370" masterDocId="1B58FFEC532CFFC6B24C2909905DFE6A" masterDocTitle="Three new species of the genus Caccothryptus Sharp, 1902 from Asia (Coleoptera: Limnichidae)" masterLastPageNumber="370" masterPageNumber="366" pageNumber="368" updateTime="1698437990641" updateUser="ExternalLinkService">
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<mods:title id="9D77267E59961C99039CEC9B39C87DE5">Three new species of the genus Caccothryptus Sharp, 1902 from Asia (Coleoptera: Limnichidae)</mods:title>
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<mods:namePart id="C700A476980EC1A1B1875CBC2D210F42">Hernando, Carles</mods:namePart>
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<mods:namePart id="0F3A130124259DC1D4A0807360AC60CC">Ribera, Ignacio</mods:namePart>
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<mods:title id="AA06504BFA8CC1BDBBFDC43A9166EEF2">Zootaxa</mods:title>
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<mods:date id="881FC92D877E7031EFBCD770BFCFE64F">2017</mods:date>
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<mods:identifier id="04268E2353FF098DAE2686BCCDF69895" type="DOI">10.11646/zootaxa.4243.2.5</mods:identifier>
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<treatment id="E7618794532EFFC2B2DB2F109090FCFD" ID-DOI="http://doi.org/10.5281/zenodo.6002016" ID-GBIF-Taxon="127844210" ID-Zenodo-Dep="6002016" LSID="urn:lsid:plazi:treatment:E7618794532EFFC2B2DB2F109090FCFD" httpUri="http://treatment.plazi.org/id/E7618794532EFFC2B2DB2F109090FCFD" lastPageId="4" lastPageNumber="370" pageId="2" pageNumber="368">
<subSubSection id="27D26509532EFFC4B2DB2F1093DBF839" pageId="2" pageNumber="368" type="nomenclature">
<paragraph id="6F773682532EFFC4B2DB2F109262F859" blockId="2.[151,1436,1561,2035]" box="[151,575,1561,1587]" pageId="2" pageNumber="368">
<heading id="343F81EE532EFFC4B2DB2F109262F859" bold="true" box="[151,575,1561,1587]" fontSize="11" level="1" pageId="2" pageNumber="368" reason="1">
<emphasis id="5DBCEA90532EFFC4B2DB2F109262F859" bold="true" box="[151,575,1561,1587]" pageId="2" pageNumber="368">
<taxonomicName id="A8C84D01532EFFC4B2DB2F109126F859" box="[151,379,1561,1587]" class="Insecta" family="Limnichidae" genus="Caccothryptus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="368" phylum="Arthropoda" rank="species" species="thai" status="sp. nov.">
<emphasis id="5DBCEA90532EFFC4B2DB2F109126F859" bold="true" box="[151,379,1561,1587]" italics="true" pageId="2" pageNumber="368">Caccothryptus thai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="468F57EB532EFFC4B3CF2F109198F859" box="[387,453,1561,1587]" pageId="2" pageNumber="368" rank="species">sp. n.</taxonomicNameLabel>
(Figs 3,4)
</emphasis>
</heading>
</paragraph>
<paragraph id="6F773682532EFFC4B2DB2F3293DBF839" blockId="2.[151,1436,1561,2035]" box="[151,902,1595,1619]" pageId="2" pageNumber="368">
<treatmentCitationGroup id="4FD811AC532EFFC4B2DB2F3293DBF839" box="[151,902,1595,1619]" pageId="2" pageNumber="368">
<taxonomicName id="A8C84D01532EFFC4B2DB2F3291F6F839" ID-CoL="P5F6" box="[151,427,1595,1619]" class="Insecta" family="Limnichidae" genus="Caccothryptus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="368" phylum="Arthropoda" rank="species" species="compactus">
<emphasis id="5DBCEA90532EFFC4B2DB2F3291F6F839" box="[151,427,1595,1619]" italics="true" pageId="2" pageNumber="368">Caccothryptus compactus</emphasis>
</taxonomicName>
group sp2:
<treatmentCitation id="EE691093532EFFC4B0662F3593DEF839" author="Hernando" box="[554,899,1596,1619]" page="289" pageId="2" pageNumber="368" year="2014">
<bibRefCitation id="0B594B73532EFFC4B0662F3593DEF839" author="Hernando" box="[554,899,1596,1619]" pageId="2" pageNumber="368" refString="Hernando, C. &amp; Ribera, I. (2014) Taxonomic revision of the genus Caccothryptus Sharp (Coleoptera: Limnichidae). Koleopterologische Rundschau, 84, 281 - 304." type="journal article" year="2014">Hernando &amp; Ribera (2014: 289)</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="27D26509532EFFC2B2DB2F579118FFAD" lastPageId="4" lastPageNumber="370" pageId="2" pageNumber="368" type="materials_examined">
<paragraph id="6F773682532EFFC4B2DB2F579359F81F" blockId="2.[151,1436,1561,2035]" box="[151,772,1630,1653]" pageId="2" pageNumber="368">
<materialsCitation id="DFA03CDF532EFFC4B2DB2F579359F81F" ID-GBIF-Occurrence="1439686339" box="[151,772,1630,1653]" country="Thailand" location="Doi Inthanon" municipality="Bang Khun Klang" pageId="2" pageNumber="368" specimenCount="1" typeStatus="holotype">
<emphasis id="5DBCEA90532EFFC4B2DB2F57917BF81F" bold="true" box="[151,294,1630,1653]" pageId="2" pageNumber="368">
<typeStatus id="B0738820532EFFC4B2DB2F579092F81F" box="[151,207,1630,1653]" pageId="2" pageNumber="368">Type</typeStatus>
locality
</emphasis>
.
<location id="6A176059532EFFC4B3782F57919EF81F" LSID="urn:lsid:plazi:treatment:E7618794532EFFC2B2DB2F109090FCFD:6A176059532EFFC4B3782F57919EF81F" box="[308,451,1630,1653]" country="Thailand" municipality="Bang Khun Klang" name="Doi Inthanon" pageId="2" pageNumber="368">Doi Inthanon</location>
,
<collectingMunicipality id="8F13ACF8532EFFC4B3822F5792CEF81F" box="[462,659,1630,1653]" pageId="2" pageNumber="368">Bang Khun Klang</collectingMunicipality>
,
<collectingCountry id="17DF7612532EFFC4B0D22F57935DF81F" box="[670,768,1630,1653]" name="Thailand" pageId="2" pageNumber="368">Thailand</collectingCountry>
.
</materialsCitation>
</paragraph>
<paragraph id="6F773682532EFFC4B28B2F88923CF8B7" blockId="2.[151,1436,1561,2035]" pageId="2" pageNumber="368">
<emphasis id="5DBCEA90532EFFC4B28B2F88913AF8F2" bold="true" box="[199,359,1665,1688]" pageId="2" pageNumber="368">Type material</emphasis>
.
<emphasis id="5DBCEA90532EFFC4B33B2F889182F8F2" bold="true" box="[375,479,1665,1688]" pageId="2" pageNumber="368">Holotype</emphasis>
male (NMW): &quot;N-THAILAND, 3.-10.10. // 18°32'N 98°32'E // Doi Inthanon,
<quantity id="A8309B67532EFFC4B7762F8895DEF8F2" box="[1338,1411,1665,1688]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="2" pageNumber="368" unit="m" value="1200.0">1200m</quantity>
// Bang Khun Klang, Malicky // &amp; Chantaramongkol 1989&quot;; genitalia extracted and mounted on a transparent card pinned with the specimen; plus red holotype label.
</paragraph>
<paragraph id="6F773682532EFFC4B28B2FE0931EF92F" blockId="2.[151,1436,1561,2035]" pageId="2" pageNumber="368">
Paratypes.
<materialsCitation id="DFA03CDF532EFFC4B3722FE09460F96A" ID-GBIF-Occurrence="1439686334" box="[318,1085,1769,1792]" collectionCode="NMW" pageId="2" pageNumber="368" specimenCount="1" typeStatus="paratype">
4 exx (
<collectionCode id="09D9AE47532EFFC4B3C62FE09188F96A" box="[394,469,1769,1792]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="2" pageNumber="368">NMW</collectionCode>
): 2 exx. same data as holotype, with red
<typeStatus id="B0738820532EFFC4B1DB2FE093AEF96A" box="[919,1011,1769,1792]" pageId="2" pageNumber="368" type="paratype">paratype</typeStatus>
labels
</materialsCitation>
; 2 females &quot;NW-THAILAND // Doi Inthanon,
<quantity id="A8309B67532EFFC4B37E2E059127F949" box="[306,378,1804,1827]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="2" pageNumber="368" unit="m" value="1200.0">1200m</quantity>
// Bang Khun Klang / 98°32'E 18°32'N&quot;; &quot;1989 // Malicky &amp; // Chantaramongkol&quot;; &quot;
<taxonomicName id="A8C84D01532EFFC4B6B32E0595C1F949" box="[1279,1436,1804,1827]" class="Insecta" family="Limnichidae" genus="Caccothryptus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="368" phylum="Arthropoda" rank="genus" status="sp. nov.">Caccothryptus</taxonomicName>
<taxonomicNameLabel id="468F57EB532EFFC4B2DB2E279097F92F" box="[151,202,1838,1861]" pageId="2" pageNumber="368" rank="species">sp.n.</taxonomicNameLabel>
? // Hernando &amp; Ribera det 2002&quot;; plus red paratype labels.
</paragraph>
<paragraph id="6F773682532EFFC4B28B2E5891E9F9C7" blockId="2.[151,1436,1561,2035]" pageId="2" pageNumber="368">
Description.
<typeStatus id="B0738820532EFFC4B31D2E5891E5F902" box="[337,440,1873,1896]" pageId="2" pageNumber="368" type="holotype">Holotype</typeStatus>
: length
<quantity id="A8309B67532EFFC4B0412E58923FF902" box="[525,610,1873,1896]" metricMagnitude="-3" metricUnit="m" metricValue="4.9" pageId="2" pageNumber="368" unit="mm" value="4.9">4.9 mm</quantity>
, width
<quantity id="A8309B67532EFFC4B0FE2E58935AF902" box="[690,775,1873,1896]" metricMagnitude="-3" metricUnit="m" metricValue="2.6" pageId="2" pageNumber="368" unit="mm" value="2.6">2.6 mm</quantity>
;
<typeStatus id="B0738820532EFFC4B15F2E589323F902" box="[787,894,1873,1896]" pageId="2" pageNumber="368" type="paratype">paratypes</typeStatus>
: length
<quantity id="A8309B67532EFFC4B19E2E58940CF902" box="[978,1105,1873,1896]" metricMagnitude="-3" metricUnit="m" metricValue="4.699999999999999" metricValueMax="4.8" metricValueMin="4.6" pageId="2" pageNumber="368" unit="mm" value="4.699999999999999" valueMax="4.8" valueMin="4.6">4.6-4.8 mm</quantity>
, width
<quantity id="A8309B67532EFFC4B6EC2E58957DF902" box="[1184,1312,1873,1896]" metricMagnitude="-3" metricUnit="m" metricValue="2.55" metricValueMax="2.6" metricValueMin="2.5" pageId="2" pageNumber="368" unit="mm" value="2.55" valueMax="2.6" valueMin="2.5">2.5-2.6 mm</quantity>
. Body dark brown (most specimens are teneral, with a lighter colour); oval, elongated, parallel sided. Tarsi and antennae light brown, legs dark brown (as body).
</paragraph>
<paragraph id="6F773682532EFFC5B28B2EB095DDFEBB" blockId="2.[151,1436,1561,2035]" lastBlockId="3.[151,1436,152,556]" lastPageId="3" lastPageNumber="369" pageId="2" pageNumber="368">
Head: Eye margin strong, with a small denticle by insertion of antennae; with a very narrow supra-ocular sulcus. Eyes slightly prominent. Puncturation on head uniformly fine and very dense; space between punctures similar to their diameter. Two
<typeStatus id="B0738820532FFFC5B3722991912BFEC5" box="[318,374,152,175]" pageId="3" pageNumber="369">types</typeStatus>
of pubescence, one erect, castaneous limited to disk; a second golden, very dense, recumbent. Pubescence on frons orientated towards clypeum; pubescence on the lateral and basal areas orientated towards vertex.
</paragraph>
<paragraph id="6F773682532FFFC5B28B29D4937CFF36" blockId="3.[151,1436,152,556]" pageId="3" pageNumber="369">Pronotum: Transvere, same width than base of elytra, lateral margins regularly and slightly curved; strongly bordered, borders narrowing towards base. Puncturation similar to that on head, very uniform; spaces between punctures wider than diameter. Pubescence mostly formed by longer erect setae, castaneous; some shorter greyish setae on the lateral margins, absent from disk, which only has erect setae.</paragraph>
<paragraph id="6F773682532FFFC5B28B286193E3FC46" blockId="3.[151,1436,152,556]" pageId="3" pageNumber="369">Elytra: Disk with coarser puncturation than on pronotum and head; puncturation double, one very fine and dense, a second coarser, very irregular but with 3-4 poorly defined series on disc; some coarse punctures forming irregular 2-3 series on elytral margins. Elytral pubescence double; erect setae castaneous, recumbent setae generally golden, but in some areas darker (colour depends on angle of incidence of light). Recumbent seta with a zig-zag pattern, orientated towards apex. Elytra strongly bordered, border stronger in the humerus and less marked apically, from middle part to apex; almost indistinguishable at apex. Membranous wings well developed.</paragraph>
<caption id="3BB7660A532FFFC5B2DB2E3A94C4F920" box="[151,1177,1842,1866]" httpUri="https://zenodo.org/record/399457/files/figure.png" pageId="3" pageNumber="369" targetBox="[340,1247,610,1818]" targetPageId="3">
<paragraph id="6F773682532FFFC5B2DB2E3A94C4F920" blockId="3.[151,1177,1842,1866]" box="[151,1177,1842,1866]" pageId="3" pageNumber="369">
<emphasis id="5DBCEA90532FFFC5B2DB2E3A9144F920" bold="true" box="[151,281,1843,1866]" pageId="3" pageNumber="369">FIGURE 4.</emphasis>
Detail of the denticles in the main lobe of the aedeagus of
<taxonomicName id="A8C84D01532FFFC5B1C12E3B9404F920" box="[909,1113,1842,1866]" class="Insecta" family="Limnichidae" genus="Caccothryptus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="369" phylum="Arthropoda" rank="species" species="thai" status="sp. nov.">
<emphasis id="5DBCEA90532FFFC5B1C12E3B9404F920" box="[909,1113,1842,1866]" italics="true" pageId="3" pageNumber="369">Caccothryptus thai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="468F57EB532FFFC5B6132E3A94C4F920" box="[1119,1177,1843,1866]" pageId="3" pageNumber="369" rank="species">sp. n.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="6F773682532FFFC2B28B2E7191C5FEBE" blockId="3.[151,1435,1912,2004]" lastBlockId="4.[151,1436,152,663]" lastPageId="4" lastPageNumber="370" pageId="3" pageNumber="369">Venter: Uniformly dark brown. Fully pubescence except hypomeras, which are glabrous; prosternum strongly encased into mesosternum. Prosternum densely punctured, punctures deeper and larger than on dorsum; margins bordered; mesosternum with slightly shallower puncturation than on proternum; disk of metasternum with fine puncturation; lateral process of the metaventrite (metasternal wings) with very coarse puncturation. Glandular pores in 4th abdominal sternite only.</paragraph>
<paragraph id="6F7736825328FFC2B28B29E991B7FF56" blockId="4.[151,1436,152,663]" pageId="4" pageNumber="370">
Aedeagus (Figs 3,4): ventral margin of the median lobe with two series of 3-4 irregular, small dents (
<figureCitation id="F7F32A075328FFC2B6B329E9951CFE9D" box="[1279,1345,224,247]" captionStart="FIGURE 4" captionStartId="3.[151,255,1843,1866]" captionTargetBox="[340,1247,610,1818]" captionTargetId="figure@3.[340,1247,610,1818]" captionTargetPageId="3" captionText="FIGURE 4. Detail of the denticles in the main lobe of the aedeagus of Caccothryptus thai sp. n." httpUri="https://zenodo.org/record/399457/files/figure.png" pageId="4" pageNumber="370">Fig. 4</figureCitation>
). Dorsal part of base of parameres in lateral view with a small denticle; apex of parameres and median lobe narrow, fusiform; tips of parameres slightly divergent.
</paragraph>
<paragraph id="6F7736825328FFC2B28B28419118FFAD" blockId="4.[151,1436,152,663]" pageId="4" pageNumber="370">
<emphasis id="5DBCEA905328FFC2B28B284191C3FF35" bold="true" box="[199,414,328,351]" pageId="4" pageNumber="370">Comparative notes</emphasis>
. The species can be clearly separated by the presence of small dents in the median lobe, which is unique among the species of the
<taxonomicName id="A8C84D015328FFC2B06C286392EEFFEB" box="[544,691,362,385]" class="Insecta" family="Limnichidae" genus="Caccothryptus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="370" phylum="Arthropoda" rank="species" species="compactus">
<emphasis id="5DBCEA905328FFC2B06C286392EEFFEB" box="[544,691,362,385]" italics="true" pageId="4" pageNumber="370">C. compactus</emphasis>
</taxonomicName>
group. The shape of the aedeagus in ventral view (
<figureCitation id="F7F32A075328FFC2B6A32863956CFFEB" box="[1263,1329,362,385]" captionStart="FIGURES 1 – 3" captionStartId="2.[151,269,1388,1411]" captionTargetBox="[163,1424,311,1337]" captionTargetId="figure@2.[151,1436,298,1363]" captionTargetPageId="2" captionText="FIGURES 1 – 3. Male genitalia of: 1) Caccothryptus occidentalis sp. n., 2) Caccothryptus schillhammeri sp. n., 3) Caccothryptus thai sp. n. — aedeagus in A) lateral and B) ventral views; C) sternite XIX; D) sternite VIII. Scale bar: 0.5 mm." httpUri="https://zenodo.org/record/399456/files/figure.png" pageId="4" pageNumber="370">Fig. 3</figureCitation>
B) is also peculiar, with a very compact and fusiform apex, in contrast to all other known species of the group (
<figureCitation id="F7F32A075328FFC2B69F28849549FFCE" box="[1235,1300,397,420]" captionStart="FIGURES 1 – 3" captionStartId="2.[151,269,1388,1411]" captionTargetBox="[163,1424,311,1337]" captionTargetId="figure@2.[151,1436,298,1363]" captionTargetPageId="2" captionText="FIGURES 1 – 3. Male genitalia of: 1) Caccothryptus occidentalis sp. n., 2) Caccothryptus schillhammeri sp. n., 3) Caccothryptus thai sp. n. — aedeagus in A) lateral and B) ventral views; C) sternite XIX; D) sternite VIII. Scale bar: 0.5 mm." httpUri="https://zenodo.org/record/399456/files/figure.png" pageId="4" pageNumber="370">Fig. 2</figureCitation>
B;
<bibRefCitation id="0B594B735328FFC2B77E28849167FFAD" author="Hernando" pageId="4" pageNumber="370" refString="Hernando, C. &amp; Ribera, I. (2014) Taxonomic revision of the genus Caccothryptus Sharp (Coleoptera: Limnichidae). Koleopterologische Rundschau, 84, 281 - 304." type="journal article" year="2014">Hernando &amp; Ribera 2014</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="27D265095328FFC2B28B28DB94E4FF83" box="[199,1209,466,489]" pageId="4" pageNumber="370" type="etymology">
<paragraph id="6F7736825328FFC2B28B28DB94E4FF83" blockId="4.[151,1436,152,663]" box="[199,1209,466,489]" pageId="4" pageNumber="370">
<emphasis id="5DBCEA905328FFC2B28B28DB911DFF83" bold="true" box="[199,320,466,489]" pageId="4" pageNumber="370">Etymology</emphasis>
. Named in reference to their known distribution. The name is a noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="27D265095328FFC2B28B28FC934AFC66" box="[199,791,501,524]" pageId="4" pageNumber="370" type="distribution">
<paragraph id="6F7736825328FFC2B28B28FC934AFC66" blockId="4.[151,1436,152,663]" box="[199,791,501,524]" pageId="4" pageNumber="370">
<emphasis id="5DBCEA905328FFC2B28B28FC910FFC66" bold="true" box="[199,338,501,524]" pageId="4" pageNumber="370">Distribution</emphasis>
. So far only known from the
<typeStatus id="B07388205328FFC2B0C128FC92E6FC66" box="[653,699,501,524]" pageId="4" pageNumber="370">type</typeStatus>
locality.
</paragraph>
</subSubSection>
<subSubSection id="27D265095328FFC2B28B2B119090FCFD" pageId="4" pageNumber="370" type="discussion">
<paragraph id="6F7736825328FFC2B28B2B119090FCFD" blockId="4.[151,1436,152,663]" pageId="4" pageNumber="370">
<emphasis id="5DBCEA905328FFC2B28B2B119172FC45" bold="true" box="[199,303,536,559]" pageId="4" pageNumber="370">Remarks</emphasis>
. Two females of this species were recorded by
<bibRefCitation id="0B594B735328FFC2B17E2B119408FC45" author="Hernando" box="[818,1109,536,559]" pageId="4" pageNumber="370" refString="Hernando, C. &amp; Ribera, I. (2014) Taxonomic revision of the genus Caccothryptus Sharp (Coleoptera: Limnichidae). Koleopterologische Rundschau, 84, 281 - 304." type="journal article" year="2014">Hernando &amp; Ribera (2014)</bibRefCitation>
as &quot;
<taxonomicName id="A8C84D015328FFC2B6CA2B1E95C1FC45" box="[1158,1436,535,559]" class="Insecta" family="Limnichidae" genus="Caccothryptus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="370" phylum="Arthropoda" rank="species" species="compactus">
<emphasis id="5DBCEA905328FFC2B6CA2B1E95C1FC45" box="[1158,1436,535,559]" italics="true" pageId="4" pageNumber="370">Caccothryptus compactus</emphasis>
</taxonomicName>
group sp2&quot;, but not described due to the absence of males. The finding of males collected together with these females among the unsorted collections of the Naturhistorisches Museum Wien (NMW) allowed us to describe this new taxon here.
</paragraph>
</subSubSection>
</treatment>
</document>
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