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<document id="EE6E05DD527207A71BC10ADA725FD13A" ID-CLB-Dataset="7428" ID-DOI="10.11646/zootaxa.4949.1.2" ID-GBIF-Dataset="617a0e17-b4c7-4ac5-b371-73293b34d738" ID-ISSN="1175-5326" ID-Zenodo-Dep="4677338" ID-ZooBank="A4B4DDB2-381A-4C48-A3E6-9268D7E17601" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1616660462658" checkinUser="plazi" docAuthor="Quah, Evan S. H., Lim, Kelvin K. P. &amp; Grismer, L. Lee" docDate="2021" docId="DA028789FE16FF87FF4A5142FD0BF8C3" docLanguage="en" docName="zootaxa.4949.1.2.pdf" docOrigin="Zootaxa 4949 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Asthenodipsas vertebralis" docType="treatment" docVersion="8" lastPageNumber="33" masterDocId="263BFFF1FE1FFF8EFFDD502CFFADFF8E" masterDocTitle="On the taxonomic status of Asthenodipsas vertebralis (Boulenger, 1900) (Squamata Pareidae) in Borneo with the description of a new species" masterLastPageNumber="44" masterPageNumber="24" pageNumber="33" updateTime="1698934911108" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="4120B4C08BA03D205D4C8C2E3DD8160B" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="AA6510D2D3F534C72D542415E2157E18">
<mods:title id="6A83C02DC38FB6AA27CFA25D89487924">On the taxonomic status of Asthenodipsas vertebralis (Boulenger, 1900) (Squamata Pareidae) in Borneo with the description of a new species</mods:title>
</mods:titleInfo>
<mods:name id="B9798AC8D26DF6EB58700D34F44323F2" type="personal">
<mods:role id="6BE6D244F19FFA2538E86A9115855D30">
<mods:roleTerm id="B8D72AE5A73C143D45A7CCA48E8350D9">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="A886FDEBDD4058C1E7C88BA3E6D0703D">Quah, Evan S. H.</mods:namePart>
<mods:nameIdentifier id="EB5223DEBCA7E40FDE1CC19BD233A116" type="ORCID">0000-0002-5357-1953</mods:nameIdentifier>
<mods:affiliation id="B77EF381FF64BCF1E26C4F2BBFD48A70">Institute of Tropical Biodiversity and Sustainable Development, Universiti Malaysia Terengganu, 21030 Kuala Nerus, Terengganu, &amp; Lee Kong Chian Natural History Museum, National University of Singapore, 2 Conservatory Drive, 117377 Singapore. kelvinlim @ nus. edu. sg; https: // orcid. org / 0000 - 0002 - 0638 - 9198</mods:affiliation>
<mods:nameIdentifier id="D38058741D07AE640374EA423A074A44" type="email">kelvinlim@nus.edu.sg</mods:nameIdentifier>
</mods:name>
<mods:name id="21E422CA9739E1B30F2BFA4553648B00" type="personal">
<mods:role id="DB17E70F7453313074484F487B2DE472">
<mods:roleTerm id="209A7482A7C12415353F142E9841AFBA">Author</mods:roleTerm>
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<mods:namePart id="BC721D87C0F9446CE1FFB5D83703FA46">Lim, Kelvin K. P.</mods:namePart>
<mods:nameIdentifier id="AED1CC92BB2D0805113DDBCFEC051002" type="ORCID">0000-0002-0638-9198</mods:nameIdentifier>
<mods:affiliation id="A351260A310BD5D97D21F8F1CA642925">Lee Kong Chian Natural History Museum, National University of Singapore, 2 Conservatory Drive, 117377 Singapore. kelvinlim @ nus. edu. sg; https: // orcid. org / 0000 - 0002 - 0638 - 9198</mods:affiliation>
<mods:nameIdentifier id="8641C5EC8920C95DBA826D27F2112463" type="email">kelvinlim@nus.edu.sg</mods:nameIdentifier>
</mods:name>
<mods:name id="39CDFE28E8C7B59E7D23AB22C787DA5D" type="personal">
<mods:role id="DA185906DCB2923D7E93579A5B9C8396">
<mods:roleTerm id="167D8904AF68EC96689FE1ABE775DE54">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="E5995E0057582FA19777A62405CD3E1A">Grismer, L. Lee</mods:namePart>
<mods:nameIdentifier id="40D0CDA6BF55956E23044A05800E72BF" type="ORCID">0000-0001-8422-3698</mods:nameIdentifier>
<mods:affiliation id="A36D1B67ED975D3B61FB9A701AF18143">Herpetology Laboratory, Department of Biology, La Sierra University, 4500 Riverwalk Parkway, Riverside, California 92515, USA. lgrismer @ lasierra. edu; https: // orcid. org / 0000 - 0001 - 8422 - 3698</mods:affiliation>
<mods:nameIdentifier id="C099B277369D718404B8EAC9CBA77CD8" type="email">lgrismer@lasierra.edu</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="42B2529F3A8A0198F8F09BD323BD6AD9">text</mods:typeOfResource>
<mods:relatedItem id="6DD8C6D7E4CDB28982B330FC8F390097" type="host">
<mods:titleInfo id="2EB07AF40743F3835A611B2B35A93633">
<mods:title id="75A9EBE47313B1E72A852AC2EC231DF5">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="BE4F1C49A2F821AC7BB36CA3AB602BD8">
<mods:date id="4974ED5A23BB8EECF7DB10726A851D2E">2021</mods:date>
<mods:detail id="E0397A6FE90B3597F1C0D859143DFCA6" type="pubDate">
<mods:number id="ED465B99A5FEECB8486E7D10892A5309">2021-03-24</mods:number>
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<mods:detail id="58B1022011E10AC811E762EA47E6717C" type="volume">
<mods:number id="DA6937D7BB2548F1F36BF7296009E623">4949</mods:number>
</mods:detail>
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<mods:number id="EB25436CA9FDA4384952AB142256BE3F">1</mods:number>
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<mods:start id="04BF0346BDA67565013E77998D2D269A">24</mods:start>
<mods:end id="2E9879C5291155EB7107E9986DDD9436">44</mods:end>
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<mods:classification id="74A4C8834CAF208458B2ADF78FCDEC41">journal article</mods:classification>
<mods:identifier id="93F86F18C52D9A42B7A0D07234B646EA" type="CLB-Dataset">7428</mods:identifier>
<mods:identifier id="35CA4F2C05896A98EAC72654BA67016E" type="DOI">10.11646/zootaxa.4949.1.2</mods:identifier>
<mods:identifier id="CB130A3C9AD37BF06856A9F8E9CC4045" type="GBIF-Dataset">617a0e17-b4c7-4ac5-b371-73293b34d738</mods:identifier>
<mods:identifier id="934EC4147D50A32403DC898C4081EEE8" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="E1EB97BED6A606693D1111063C246CEE" type="Zenodo-Dep">4677338</mods:identifier>
<mods:identifier id="DD3442DF3DE27C7176B8F1679BF19C65" type="ZooBank">A4B4DDB2-381A-4C48-A3E6-9268D7E17601</mods:identifier>
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<treatment id="DA028789FE16FF87FF4A5142FD0BF8C3" ID-DOI="http://doi.org/10.5281/zenodo.4677405" ID-GBIF-Taxon="180888385" ID-Zenodo-Dep="4677405" LSID="urn:lsid:plazi:treatment:DA028789FE16FF87FF4A5142FD0BF8C3" httpUri="http://treatment.plazi.org/id/DA028789FE16FF87FF4A5142FD0BF8C3" lastPageNumber="33" pageId="9" pageNumber="33">
<subSubSection id="1AB16514FE16FF87FF4A5142FD03FE06" box="[151,686,366,393]" pageId="9" pageNumber="33" type="nomenclature">
<paragraph id="5214369FFE16FF87FF4A5142FD03FE06" blockId="9.[151,686,366,429]" box="[151,686,366,393]" pageId="9" pageNumber="33">
<heading id="095C81F3FE16FF87FF4A5142FD03FE06" bold="true" box="[151,686,366,393]" fontSize="11" level="1" pageId="9" pageNumber="33" reason="1">
<taxonomicName id="95AB4D1CFE16FF87FF4A5142FD03FE06" authority="(Boulenger, 1900)" baseAuthorityName="Boulenger" baseAuthorityYear="1900" box="[151,686,366,393]" class="Reptilia" family="Pareatidae" genus="Asthenodipsas" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="33" phylum="Chordata" rank="species" species="vertebralis">
<emphasis id="60DFEA8DFE16FF87FF4A5142FD03FE06" bold="true" box="[151,686,366,393]" pageId="9" pageNumber="33">
<emphasis id="60DFEA8DFE16FF87FF4A5142FE61FE07" bold="true" box="[151,460,366,393]" italics="true" pageId="9" pageNumber="33">Asthenodipsas vertebralis</emphasis>
(Boulenger, 1900)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="1AB16514FE16FF87FF4A51BEFD08FB4B" pageId="9" pageNumber="33" type="description">
<paragraph id="5214369FFE16FF87FF4A51BEFE7BFE22" blockId="9.[151,686,366,429]" box="[151,470,402,429]" pageId="9" pageNumber="33">
(
<figureCitation id="CA902A1AFE16FF87FF7C51BEFF43FE22" box="[161,238,402,428]" captionStart="FIGURE 3" captionStartId="7.[151,250,1374,1398]" captionTargetBox="[157,1430,531,1343]" captionTargetId="figure-224@7.[151,1437,525,1350]" captionTargetPageId="7" captionText="FIGURE 3. A &amp; B. Dorsum and venter of Asthenodipsas vertebralis (ZRC 2.6619) from Gunung Berumput, Sarawak. Figure not to scale." figureDoi="http://doi.org/10.5281/zenodo.4677344" httpUri="https://zenodo.org/record/4677344/files/figure.png" pageId="9" pageNumber="33">Figs. 3</figureCitation>
,
<figureCitation id="CA902A1AFE16FF87FF2651BEFEA4FE22" box="[251,265,402,428]" captionStart="FIGURE 4" captionStartId="8.[151,250,1550,1574]" captionTargetBox="[151,1436,181,1527]" captionTargetId="figure-16@8.[151,1436,181,1527]" captionTargetPageId="8" captionText="FIGURE 4. Arrangement of supralabials in: A. Holotype of Asthenodipsas ingeri sp. nov. (MCZ R43592) from Lumu Lumu, Mount Kinabalu, Sabah. B. Adult Asthenodipsas cf. ingeri sp. nov. (LSUDPC 12633) from Payeh Maga Highlands, Sarawak. C &amp; D. Adults of A. ingeri sp. nov. from Kinabalu National Park, Sabah (LSUDPC 12634 &amp; 12636 respectively). E. Adult A. vertebralis (LSUHC 11709) from Bukit Larut, Perak. F. Asthenodipsas vertebralis (ZRC 2.6619) from Gunung Berumput, Sarawak. Figures not to scale. (Photographs by Evan Quah [A &amp; E], Chien C. Lee [B], Jannico Kelk [C], Björn Lardner [D], &amp; Kelvin K.P. Lim [F])." figureDoi="http://doi.org/10.5281/zenodo.4677348" httpUri="https://zenodo.org/record/4677348/files/figure.png" pageId="9" pageNumber="33">4</figureCitation>
&amp;
<figureCitation id="CA902A1AFE16FF87FEF051BEFE90FE23" box="[301,317,402,429]" captionStart="FIGURE 5" captionStartId="10.[151,250,1862,1886]" captionTargetBox="[161,1426,181,1839]" captionTargetId="figure-16@10.[161,1426,181,1839]" captionTargetPageId="10" captionText="FIGURE 5. Arrangement of scales on the underside of head in: A. Holotype of Asthenodipsas ingeri sp. nov. (MCZ R43592) from Lumu Lumu, Mount Kinabalu, Sabah. B. Adult Asthenodipsas ingeri sp. nov. (LSUDPC 12636) from Kinabalu National Park, Sabah. C. Asthenodipsas vertebralis (ZRC 2.6619) from Gunung Berumput, Sarawak. D. Adult A. vertebralis (LSUHC 12758) from Genting Highlands, Pahang. Figures not to scale. IL = infralabials, M = mental, SaIM = single anterior inframaxillary, pIM = pairs of posterior inframaxillaries. (Photographs by Evan Quah [A &amp; D], Björn Lardner [B], &amp; Kelvin K.P. Lim [C])." figureDoi="http://doi.org/10.5281/zenodo.4677352" httpUri="https://zenodo.org/record/4677352/files/figure.png" pageId="9" pageNumber="33">5</figureCitation>
;
<tableCitation id="1F290324FE16FF87FE9451BEFE31FE22" box="[329,412,402,428]" captionStart="TABLE 2" captionStartId="5.[151,239,260,284]" captionTargetPageId="5" captionText="TABLE 2. Scale counts and selected measurements (mm) of Asthenodipsas ingeri sp. nov. and A. vertebralis (ZRC 2.6619) from Borneo." httpUri="http://table.plazi.org/id/06D46617FE1AFF8BFF4A5128FE2CFECE" pageId="9" pageNumber="33" tableUuid="06D46617FE1AFF8BFF4A5128FE2CFECE">Table 2</tableCitation>
&amp;
<tableCitation id="1F290324FE16FF87FE6251BEFE60FE22" box="[447,461,402,428]" captionStart="TABLE 3" captionStartId="12.[151,239,462,488]" captionTargetBox="[165,2011,538,1687]" captionText="TABLE 3. Measurements and scale counts of Asthenodipsas ingeri sp. nov., A. lasgalenensis, A. tropidonotus, and A. vertebralis from Peninsular Malaysia and Borneo (ZRC 2.6619). (*measurements from Grossmann &amp; Tillack, 2003; NA = measurement not available). Bold character states are those that discretely separate A. ingeri sp. nov. from the others." httpUri="http://table.plazi.org/id/06D46617FE13FF82FF4A51E2F81BFD82" pageId="9" pageNumber="33" tableUuid="06D46617FE13FF82FF4A51E2F81BFD82">3</tableCitation>
)
</paragraph>
<paragraph id="5214369FFE16FF87FF4A51F6FE90FD96" blockId="9.[151,1437,474,1869]" pageId="9" pageNumber="33">
<emphasis id="60DFEA8DFE16FF87FF4A51F6FED2FE7A" bold="true" box="[151,383,474,500]" pageId="9" pageNumber="33">Referred specimen.</emphasis>
<materialsCitation id="E2C33CC2FE16FF87FE5A51F6FA35FE7A" ID-GBIF-Occurrence="3069689301" box="[391,1432,474,501]" collectingDate="2002-05-12" collectionCode="ZRC" collectorName="Indraneil Das" country="Malaysia" location="Gunung Berumput" pageId="9" pageNumber="33" specimenCode="ZRC 2.6619" specimenCount="1">
<specimenCode id="020D9EE4FE16FF87FE5A51F6FDBBFE7A" box="[391,534,474,500]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="9" pageNumber="33">ZRC 2.6619</specimenCode>
, collected by
<collectorName id="FF5E5349FE16FF87FD6F51F6FCE6FE7B" box="[690,843,474,501]" pageId="9" pageNumber="33">Indraneil Das</collectorName>
on
<date id="2615105FFE16FF87FCAA51F6FBA3FE7A" box="[887,1038,474,501]" pageId="9" pageNumber="33" value="2002-05-12">
<collectingDate id="3651E9B7FE16FF87FCAA51F6FBA3FE7A" box="[887,1038,474,501]" pageId="9" pageNumber="33" value="2002-05-12">12 May 2002</collectingDate>
</date>
from
<location id="57746044FE16FF87FB8951F6FA84FE7B" LSID="urn:lsid:plazi:treatment:DA028789FE16FF87FF4A5142FD0BF8C3:57746044FE16FF87FB8951F6FA84FE7B" box="[1108,1321,474,501]" country="Malaysia" name="Gunung Berumput" pageId="9" pageNumber="33">Gunung Berumput</location>
,
<collectingCountry id="2ABC760FFE16FF87FAE851F6FA35FE7A" box="[1333,1432,474,500]" name="Malaysia" pageId="9" pageNumber="33">Sarawak</collectingCountry>
</materialsCitation>
, East
<collectingCountry id="2ABC760FFE16FF87FF1351D2FE95FD96" box="[206,312,510,536]" name="Malaysia" pageId="9" pageNumber="33">Malaysia</collectingCountry>
.
</paragraph>
<paragraph id="5214369FFE16FF87FF1A520EFEC6FC46" blockId="9.[151,1437,474,1869]" pageId="9" pageNumber="33">
<emphasis id="60DFEA8DFE16FF87FF1A520EFDD0FDB2" bold="true" box="[199,637,546,572]" pageId="9" pageNumber="33">Description of specimen ZRC 2.6619</emphasis>
. Adult male SVL
<quantity id="95539B7AFE16FF87FC94520EFC0AFDB3" box="[841,935,546,573]" metricMagnitude="-1" metricUnit="m" metricValue="5.02" pageId="9" pageNumber="33" unit="mm" value="502.0">502 mm</quantity>
and TaL
<quantity id="95539B7AFE16FF87FBCD520EFBC2FDB3" box="[1040,1135,546,573]" metricMagnitude="-1" metricUnit="m" metricValue="1.14" pageId="9" pageNumber="33" unit="mm" value="114.0">114 mm</quantity>
; rostral slightly wider than tall; head bulbous, longer than wide (HL/HW 1.97); nasals undivided; internasals shorter than prefrontals; posterior margin of prefrontals contact eye; frontal hexagonal, longer than wide; loreals present, 1/1, longer than tall; supraoculars subpentagonal, approximately 2/3 the length and half the width of frontal; preoculars absent; upper and lower postoculars present on both left and right side, lower postocular extending to below orbit; suboculars absent; supralabials 7/7 with 3
<superScript id="A5DE9BD7FE16FF87FE3352FAFE51FD6A" attach="left" box="[494,508,726,740]" fontSize="6" pageId="9" pageNumber="33">rd</superScript>
&amp; 4
<superScript id="A5DE9BD7FE16FF87FDF352FAFD96FD6A" attach="left" box="[558,571,726,740]" fontSize="6" pageId="9" pageNumber="33">th</superScript>
contacting orbit and 7th elongate; temporals 2+2+2/2+2+2 (
<figureCitation id="CA902A1AFE16FF87FB3B52FAFA95FD7F" box="[1254,1336,726,753]" captionStart="FIGURE 4" captionStartId="8.[151,250,1550,1574]" captionTargetBox="[151,1436,181,1527]" captionTargetId="figure-16@8.[151,1436,181,1527]" captionTargetPageId="8" captionText="FIGURE 4. Arrangement of supralabials in: A. Holotype of Asthenodipsas ingeri sp. nov. (MCZ R43592) from Lumu Lumu, Mount Kinabalu, Sabah. B. Adult Asthenodipsas cf. ingeri sp. nov. (LSUDPC 12633) from Payeh Maga Highlands, Sarawak. C &amp; D. Adults of A. ingeri sp. nov. from Kinabalu National Park, Sabah (LSUDPC 12634 &amp; 12636 respectively). E. Adult A. vertebralis (LSUHC 11709) from Bukit Larut, Perak. F. Asthenodipsas vertebralis (ZRC 2.6619) from Gunung Berumput, Sarawak. Figures not to scale. (Photographs by Evan Quah [A &amp; E], Chien C. Lee [B], Jannico Kelk [C], Björn Lardner [D], &amp; Kelvin K.P. Lim [F])." figureDoi="http://doi.org/10.5281/zenodo.4677348" httpUri="https://zenodo.org/record/4677348/files/figure.png" pageId="9" pageNumber="33">Fig. 4F</figureCitation>
); mental triangular, wider than long; anterior inframaxillary pentagonal, in contact with 1
<superScript id="A5DE9BD7FE16FF87FBC052D6FB85FC86" attach="left" box="[1053,1064,762,776]" fontSize="6" pageId="9" pageNumber="33">st</superScript>
and 2
<superScript id="A5DE9BD7FE16FF87FBB352D6FBD3FC86" attach="left" box="[1134,1150,762,776]" fontSize="6" pageId="9" pageNumber="33">nd</superScript>
pair of infralabials; three pairs of posterior inframaxillaries following anterior inframaxillary, first pair of posterior inframaxillaries slightly oval, rhomboid-shaped, second and third pair square to hexagonal; infralabials 6/7 with 1
<superScript id="A5DE9BD7FE16FF87FB4C536EFB31FCDE" attach="left" box="[1169,1180,834,848]" fontSize="6" pageId="9" pageNumber="33">st</superScript>
pair in medial contact (
<figureCitation id="CA902A1AFE16FF87FF7C534AFF55FC0F" box="[161,248,870,897]" captionStart="FIGURE 5" captionStartId="10.[151,250,1862,1886]" captionTargetBox="[161,1426,181,1839]" captionTargetId="figure-16@10.[161,1426,181,1839]" captionTargetPageId="10" captionText="FIGURE 5. Arrangement of scales on the underside of head in: A. Holotype of Asthenodipsas ingeri sp. nov. (MCZ R43592) from Lumu Lumu, Mount Kinabalu, Sabah. B. Adult Asthenodipsas ingeri sp. nov. (LSUDPC 12636) from Kinabalu National Park, Sabah. C. Asthenodipsas vertebralis (ZRC 2.6619) from Gunung Berumput, Sarawak. D. Adult A. vertebralis (LSUHC 12758) from Genting Highlands, Pahang. Figures not to scale. IL = infralabials, M = mental, SaIM = single anterior inframaxillary, pIM = pairs of posterior inframaxillaries. (Photographs by Evan Quah [A &amp; D], Björn Lardner [B], &amp; Kelvin K.P. Lim [C])." figureDoi="http://doi.org/10.5281/zenodo.4677352" httpUri="https://zenodo.org/record/4677352/files/figure.png" pageId="9" pageNumber="33">Fig. 5D</figureCitation>
). Body long, slender, laterally compressed, bearing a prominent keel-shaped vertebral region; dorsal scales smooth, in 15/15/15 rows, vertebrals greatly enlarged; 190 ventrals; 68 divided subcaudals; cloacal scute entire; tail tapering to a point.
</paragraph>
<paragraph id="5214369FFE16FF87FF1A53FEFD08FB4B" blockId="9.[151,1437,474,1869]" pageId="9" pageNumber="33">
<emphasis id="60DFEA8DFE16FF87FF1A53FEFDC3FC62" bold="true" box="[199,622,978,1004]" pageId="9" pageNumber="33">
Colouration in preservation (
<figureCitation id="CA902A1AFE16FF87FDFC53FEFDC8FC62" box="[545,613,978,1004]" captionStart="FIGURE 3" captionStartId="7.[151,250,1374,1398]" captionTargetBox="[157,1430,531,1343]" captionTargetId="figure-224@7.[151,1437,525,1350]" captionTargetPageId="7" captionText="FIGURE 3. A &amp; B. Dorsum and venter of Asthenodipsas vertebralis (ZRC 2.6619) from Gunung Berumput, Sarawak. Figure not to scale." figureDoi="http://doi.org/10.5281/zenodo.4677344" httpUri="https://zenodo.org/record/4677344/files/figure.png" pageId="9" pageNumber="33">Fig. 3</figureCitation>
)
</emphasis>
. The ground colour of the head, body and tail is dark-brown with a series of approximately 40 black, poorly defined, rhomboidal bands ranging from 1–3 dorsal scales in length running along the dorsum from the nape to the base of the tail. The tail is irregularly mottled with dark spots. The supralabials, chin and throat are light-yellow with some fine speckling along the edges of the supralabials, mental, infralabials and inframaxillaries. The venter is yellowish cream with dark speckling along the lateral edges. A series of dark lateral spots are spaced along the belly, and are formed from the dorsal bands meeting the ventral scales. A thin, broken, light-yellow vertebral stripe is present.
</paragraph>
</subSubSection>
<subSubSection id="1AB16514FE16FF87FF1A54E2FD0BF8C3" pageId="9" pageNumber="33" type="distribution">
<paragraph id="5214369FFE16FF87FF1A54E2FF77FA82" blockId="9.[151,1437,474,1869]" pageId="9" pageNumber="33">
<emphasis id="60DFEA8DFE16FF87FF1A54E2FE73FB66" bold="true" box="[199,478,1230,1256]" pageId="9" pageNumber="33">Distribution in Borneo</emphasis>
.
<taxonomicName id="95AB4D1CFE16FF87FE2C54E2FCBAFB66" baseAuthorityName="Boulenger" baseAuthorityYear="1900" box="[497,791,1230,1256]" class="Reptilia" family="Pareatidae" genus="Asthenodipsas" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="33" phylum="Chordata" rank="species" species="vertebralis">
<emphasis id="60DFEA8DFE16FF87FE2C54E2FCBAFB66" box="[497,791,1230,1256]" italics="true" pageId="9" pageNumber="33">Asthenodipsas vertebralis</emphasis>
</taxonomicName>
is currently only known from Gunung Berumput,
<collectingCountry id="2ABC760FFE16FF87FAB354E2FF7BFA82" name="Malaysia" pageId="9" pageNumber="33">Sarawak</collectingCountry>
.
</paragraph>
<paragraph id="5214369FFE16FF87FF1A553AFC7EF987" blockId="9.[151,1437,474,1869]" pageId="9" pageNumber="33">
<emphasis id="60DFEA8DFE16FF87FF1A553AFD02FABE" bold="true" box="[199,687,1302,1328]" pageId="9" pageNumber="33">Comparison and comments on taxonomy</emphasis>
.
<taxonomicName id="95AB4D1CFE16FF87FD61553AFB8CFABE" authority=", ZRC" authorityName="ZRC" box="[700,1057,1302,1329]" class="Reptilia" family="Pareatidae" genus="Asthenodipsas" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="33" phylum="Chordata" rank="species" species="vertebralis">
<emphasis id="60DFEA8DFE16FF87FD61553AFC70FABE" box="[700,989,1302,1328]" italics="true" pageId="9" pageNumber="33">Asthenodipsas vertebralis</emphasis>
, ZRC
</taxonomicName>
2.6619 can be differentiated from
<taxonomicName id="95AB4D1CFE16FF87FF4A5517FF54FADA" authorityName="Dumeril, Bibron &amp; Dumeril" authorityYear="1854" box="[151,249,1338,1365]" class="Reptilia" family="Pareatidae" genus="Pareas" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="33" phylum="Chordata" rank="species" species="laevis">
<emphasis id="60DFEA8DFE16FF87FF4A5517FF54FADA" box="[151,249,1338,1365]" italics="true" pageId="9" pageNumber="33">A. laevis</emphasis>
</taxonomicName>
,
<taxonomicName id="95AB4D1CFE16FF87FED75517FE09FADA" authorityName="Quah, Grismer, Lim, Shahrul &amp; Chan" authorityYear="2020" box="[266,420,1338,1365]" class="Reptilia" family="Pareatidae" genus="Asthenodipsas" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="33" phylum="Chordata" rank="species" species="borneensis">
<emphasis id="60DFEA8DFE16FF87FED75517FE09FADA" box="[266,420,1338,1365]" italics="true" pageId="9" pageNumber="33">A. borneensis</emphasis>
</taxonomicName>
,
<taxonomicName id="95AB4D1CFE16FF87FE685517FDFCFADA" authorityName="Peters" authorityYear="1864" box="[437,593,1338,1365]" class="Reptilia" family="Pareatidae" genus="Asthenodipsas" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="33" phylum="Chordata" rank="species" species="malaccana">
<emphasis id="60DFEA8DFE16FF87FE685517FDFCFADA" box="[437,593,1338,1365]" italics="true" pageId="9" pageNumber="33">A. malaccana</emphasis>
</taxonomicName>
,
<taxonomicName id="95AB4D1CFE16FF87FDBC5517FD57FADA" authorityName="Quah, Grismer, Lim, Shahrul &amp; Imbun" authorityYear="2019" box="[609,762,1338,1365]" class="Reptilia" family="Pareatidae" genus="Asthenodipsas" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="33" phylum="Chordata" rank="species" species="jamilinaisi">
<emphasis id="60DFEA8DFE16FF87FDBC5517FD57FADA" box="[609,762,1338,1365]" italics="true" pageId="9" pageNumber="33">A. jamilinaisi</emphasis>
</taxonomicName>
and
<taxonomicName id="95AB4D1CFE16FF87FCE85517FC10FADA" authorityName="Quah, Grismer, Lim, Shahrul &amp; Imbun" authorityYear="2019" box="[821,957,1338,1365]" class="Reptilia" family="Pareatidae" genus="Asthenodipsas" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="33" phylum="Chordata" rank="species" species="stuebingi">
<emphasis id="60DFEA8DFE16FF87FCE85517FC10FADA" box="[821,957,1338,1365]" italics="true" pageId="9" pageNumber="33">A. stuebingi</emphasis>
</taxonomicName>
by its higher number of ventrals (190 vs. 148–179), higher number of subcaudals (68 vs. 26–53), and the pairs of infralabials in contact (1
<superScript id="A5DE9BD7FE16FF87FB045572FB49FAE2" attach="left" box="[1241,1252,1374,1388]" fontSize="6" pageId="9" pageNumber="33">st</superScript>
vs. 2
<superScript id="A5DE9BD7FE16FF87FAFC5572FA9CFAE2" attach="left" box="[1313,1329,1374,1388]" fontSize="6" pageId="9" pageNumber="33">nd</superScript>
or 3
<superScript id="A5DE9BD7FE16FF87FABB5572FAD9FAE2" attach="none" box="[1382,1396,1374,1388]" fontSize="6" pageId="9" pageNumber="33">rd</superScript>
). It can be further separated from
<taxonomicName id="95AB4D1CFE16FF87FE2C55AFFDFEFA12" authorityName="Dumeril, Bibron &amp; Dumeril" authorityYear="1854" box="[497,595,1410,1437]" class="Reptilia" family="Pareatidae" genus="Pareas" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="33" phylum="Chordata" rank="species" species="laevis">
<emphasis id="60DFEA8DFE16FF87FE2C55AFFDFEFA12" box="[497,595,1410,1437]" italics="true" pageId="9" pageNumber="33">A. laevis</emphasis>
</taxonomicName>
by its number of dorsal scale rows (15/15/15 vs. 15/15/13) and the sharp vertebral keel (present vs. absent) (
<bibRefCitation id="363A4B6EFE16FF87FDF9558AFD4BFA4E" author="Quah, E. S. H. &amp; Grismer, L. L. &amp; Lim, K. K. P. &amp; Shahrul, A. M. S. &amp; Imbun, P. Y." box="[548,742,1446,1473]" pageId="9" pageNumber="33" pagination="501 - 526" refId="ref13775" refString="Quah, E. S. H., Grismer, L. L., Lim, K. K. P., Shahrul, A. M. S. &amp; Imbun, P. Y. (2019) A taxonomic reappraisal of the Smooth Slug Snake (Asthenodipsas laevis) (Boie 1827) (Squamata: Pareidae) in Borneo with the description of two new species. Zootaxa, 4646 (3), 501 - 526. https: // doi. org / 10.11646 / zootaxa. 4646.3.4" type="journal article" year="2019">
Quah
<emphasis id="60DFEA8DFE16FF87FDB7558BFD31FA4E" box="[618,668,1446,1472]" italics="true" pageId="9" pageNumber="33">et al</emphasis>
. 2019
</bibRefCitation>
,
<bibRefCitation id="363A4B6EFE16FF87FD2F558AFC82FA4E" author="Quah, E. S. H. &amp; Grismer, L. L. &amp; Lim, K. K. P. &amp; Shahrul, A. M. S. &amp; Chan, K. O." box="[754,815,1446,1472]" pageId="9" pageNumber="33" pagination="1 - 24" refId="ref13872" refString="Quah, E. S. H., Grismer, L. L., Lim, K. K. P., Shahrul, A. M. S. &amp; Chan, K. O. (2020) A taxonomic revision of Asthenodipsas malaccana Peters, 1864 (Squamata: Pareidae) with a description of a new species from Borneo. Zootaxa, 4729 (1), 1 - 24. https: // doi. org / 10.11646 / zootaxa. 4729.1.1" type="journal article" year="2020">2020</bibRefCitation>
). ZRC 2.6619 can also be differentiated from
<taxonomicName id="95AB4D1CFE16FF87FAE4558BFA31FA4E" box="[1337,1436,1447,1473]" class="Reptilia" family="Pareatidae" genus="Asthenodipsas" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="33" phylum="Chordata" rank="species" species="ingeri" status="sp. nov.">
<emphasis id="60DFEA8DFE16FF87FAE4558BFA31FA4E" box="[1337,1436,1447,1473]" italics="true" pageId="9" pageNumber="33">A. ingeri</emphasis>
</taxonomicName>
<emphasis id="60DFEA8DFE16FF87FF4A55E6FF5CFA6A" bold="true" box="[151,241,1482,1508]" pageId="9" pageNumber="33">
<taxonomicNameLabel id="7BEC57F6FE16FF87FF4A55E6FF5CFA6A" box="[151,241,1482,1508]" pageId="9" pageNumber="33" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
by the pairs of infralabials in contact (1
<superScript id="A5DE9BD7FE16FF87FD6E55E6FD13FA56" attach="left" box="[691,702,1482,1496]" fontSize="6" pageId="9" pageNumber="33">st</superScript>
vs. 1
<superScript id="A5DE9BD7FE16FF87FD2655E6FCABFA56" attach="left" box="[763,774,1482,1496]" fontSize="6" pageId="9" pageNumber="33">st</superScript>
&amp; 3
<superScript id="A5DE9BD7FE16FF87FCE555E6FCEBFA56" attach="none" box="[824,838,1482,1496]" fontSize="6" pageId="9" pageNumber="33">rd</superScript>
), the number of supralabials in contact with the orbit (3 &amp; 4 vs 3–5) and more pairs of posterior inframaxillaries (three vs two).
</paragraph>
<paragraph id="5214369FFE16FF87FF1A563EFD0BF8C3" blockId="9.[151,1437,474,1869]" pageId="9" pageNumber="33">
ZRC 2.6619 is identified as
<taxonomicName id="95AB4D1CFE16FF87FE22563FFD3AF9A2" box="[511,663,1554,1581]" class="Reptilia" family="Pareatidae" genus="Asthenodipsas" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="33" phylum="Chordata" rank="species" species="vertebralis">
<emphasis id="60DFEA8DFE16FF87FE22563FFD3AF9A2" box="[511,663,1554,1581]" italics="true" pageId="9" pageNumber="33">A. vertebralis</emphasis>
</taxonomicName>
by its possession of three pairs of posterior inframaxillaries, first pair of infralabials in medial contact, yellowish labials and ventrals, and a banded dorsal pattern on the body with a thin, light vertebral stripe (
<bibRefCitation id="363A4B6EFE16FF87FE5B5676FDF7F9FA" author="Loredo, A. I. &amp; Wood, Jr., P. L. &amp; Quah, E. S. H. &amp; Anuar, S. &amp; Greer, L. F. &amp; Ahmad, N. &amp; Grismer, L. L." box="[390,602,1626,1653]" pageId="9" pageNumber="33" pagination="505 - 524" refId="ref13028" refString="Loredo, A. I., Wood, Jr., P. L., Quah, E. S. H., Anuar, S., Greer, L. F., Ahmad, N. &amp; Grismer, L. L. (2013) Cryptic speciation within Asthenodipsas vertebralis (Boulenger, 1900) (Squamata: Pareatidae), the description of a new species from Peninsular Malaysia, and the resurrection of A. tropidonotus (Lidth de Jude, 1923) from Sumatra: an integrative taxonomic analysis. Zootaxa, 3664 (4), 505 - 524. https: // doi. org / 10.11646 / zootaxa. 3664.4.5" type="journal article" year="2013">
Loredo
<emphasis id="60DFEA8DFE16FF87FE025677FDBDF9FA" box="[479,528,1626,1652]" italics="true" pageId="9" pageNumber="33">et al</emphasis>
. 2013
</bibRefCitation>
). Its colour pattern rules it out as being
<taxonomicName id="95AB4D1CFE16FF87FBD05677FB6EF9FA" box="[1037,1219,1626,1653]" class="Reptilia" family="Pareatidae" genus="Asthenodipsas" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="33" phylum="Chordata" rank="species" species="lasgalenensis">
<emphasis id="60DFEA8DFE16FF87FBD05677FB6EF9FA" box="[1037,1219,1626,1653]" italics="true" pageId="9" pageNumber="33">A. lasgalenensis</emphasis>
</taxonomicName>
whose adult colour pattern consists of a solid, dark-brown to black dorsum with white labials and venter. It is further distinguished from
<taxonomicName id="95AB4D1CFE16FF87FF08568FFE2FF932" baseAuthorityName="Lidth de Jeude" baseAuthorityYear="1923" box="[213,386,1698,1725]" class="Reptilia" family="Pareatidae" genus="Asthenodipsas" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="33" phylum="Chordata" rank="species" species="tropidonotus">
<emphasis id="60DFEA8DFE16FF87FF08568FFE2FF932" box="[213,386,1698,1725]" italics="true" pageId="9" pageNumber="33">A. tropidonotus</emphasis>
</taxonomicName>
by its lower number of subcaudals (68 vs. 73–76) and higher number of postoculars (2 vs. 1) and falls within the range of other scale and meristic characters reported for
<taxonomicName id="95AB4D1CFE16FF87FC2E56EBFB21F96E" box="[1011,1164,1734,1761]" class="Reptilia" family="Pareatidae" genus="Asthenodipsas" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="33" phylum="Chordata" rank="species" species="vertebralis">
<emphasis id="60DFEA8DFE16FF87FC2E56EBFB21F96E" box="[1011,1164,1734,1761]" italics="true" pageId="9" pageNumber="33">A. vertebralis</emphasis>
</taxonomicName>
(
<tableCitation id="1F290324FE16FF87FB4656EAFB5FF96E" box="[1179,1266,1734,1760]" captionStart="TABLE 3" captionStartId="12.[151,239,462,488]" captionTargetBox="[165,2011,538,1687]" captionText="TABLE 3. Measurements and scale counts of Asthenodipsas ingeri sp. nov., A. lasgalenensis, A. tropidonotus, and A. vertebralis from Peninsular Malaysia and Borneo (ZRC 2.6619). (*measurements from Grossmann &amp; Tillack, 2003; NA = measurement not available). Bold character states are those that discretely separate A. ingeri sp. nov. from the others." httpUri="http://table.plazi.org/id/06D46617FE13FF82FF4A51E2F81BFD82" pageId="9" pageNumber="33" tableUuid="06D46617FE13FF82FF4A51E2F81BFD82">Table 3</tableCitation>
;
<bibRefCitation id="363A4B6EFE16FF87FB2256EAFE82F88A" author="Grossmann, W. &amp; Tillack, F." pageId="9" pageNumber="33" pagination="175 - 90" refId="ref12606" refString="Grossmann, W. &amp; Tillack, F. (2003) On the Taxonomic Status of Asthenodipsas tropidonotus (Van Lidth de Jeude, 1923) and Pareas vertebralis (Boulenger, 1900) (Serpentes: Colubridae: Pareatinae). Russian Journal of Herpetology, 10 (3), 175 - 90." type="journal article" year="2003">Grossmann &amp; Tillack, 2003</bibRefCitation>
;
<bibRefCitation id="363A4B6EFE16FF87FEE656C6FDB5F88A" author="Loredo, A. I. &amp; Wood, Jr., P. L. &amp; Quah, E. S. H. &amp; Anuar, S. &amp; Greer, L. F. &amp; Ahmad, N. &amp; Grismer, L. L." box="[315,536,1770,1797]" pageId="9" pageNumber="33" pagination="505 - 524" refId="ref13028" refString="Loredo, A. I., Wood, Jr., P. L., Quah, E. S. H., Anuar, S., Greer, L. F., Ahmad, N. &amp; Grismer, L. L. (2013) Cryptic speciation within Asthenodipsas vertebralis (Boulenger, 1900) (Squamata: Pareatidae), the description of a new species from Peninsular Malaysia, and the resurrection of A. tropidonotus (Lidth de Jude, 1923) from Sumatra: an integrative taxonomic analysis. Zootaxa, 3664 (4), 505 - 524. https: // doi. org / 10.11646 / zootaxa. 3664.4.5" type="journal article" year="2013">
Loredo
<emphasis id="60DFEA8DFE16FF87FE4956C7FE6BF88A" box="[404,454,1770,1796]" italics="true" pageId="9" pageNumber="33">et al</emphasis>
., 2013
</bibRefCitation>
). However, in light of the resurrection of
<taxonomicName id="95AB4D1CFE16FF87FC3156C7FB34F88A" baseAuthorityName="Lidth de Jeude" baseAuthorityYear="1923" box="[1004,1177,1770,1797]" class="Reptilia" family="Pareatidae" genus="Asthenodipsas" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="33" phylum="Chordata" rank="species" species="tropidonotus">
<emphasis id="60DFEA8DFE16FF87FC3156C7FB34F88A" box="[1004,1177,1770,1797]" italics="true" pageId="9" pageNumber="33">A. tropidonotus</emphasis>
</taxonomicName>
from the synonymy of
<taxonomicName id="95AB4D1CFE16FF87FF4A5723FE9DF8A6" box="[151,304,1806,1833]" class="Reptilia" family="Pareatidae" genus="Asthenodipsas" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="33" phylum="Chordata" rank="species" species="vertebralis">
<emphasis id="60DFEA8DFE16FF87FF4A5723FE9DF8A6" box="[151,304,1806,1833]" italics="true" pageId="9" pageNumber="33">A. vertebralis</emphasis>
</taxonomicName>
with the inclusion of genetic data, we only tentatively assign the specimen as
<taxonomicName id="95AB4D1CFE16FF87FB7F5723FA97F8A6" box="[1186,1338,1806,1833]" class="Reptilia" family="Pareatidae" genus="Asthenodipsas" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="33" phylum="Chordata" rank="species" species="vertebralis">
<emphasis id="60DFEA8DFE16FF87FB7F5723FA97F8A6" box="[1186,1338,1806,1833]" italics="true" pageId="9" pageNumber="33">A. vertebralis</emphasis>
</taxonomicName>
pending molecular analyses of specimens from Borneo.
</paragraph>
</subSubSection>
</treatment>
</document>
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