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<document id="DB88E815F00C1552D0490DBBC4A1EC2D" ID-CLB-Dataset="37404" ID-DOI="10.11646/zootaxa.4205.4.1" ID-GBIF-Dataset="eb5dfbad-e024-4318-b58e-6d9b9e19eac3" ID-ISSN="1175-5326" ID-Zenodo-Dep="208241" ID-ZooBank="BAC8F107-21D6-49FE-BAC7-BF4EE6C3E6A4" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1482105213028" checkinUser="plazi" docAuthor="Kolesik, Peter &amp; Gagné, Raymond J." docDate="2016" docId="D759878B0E6EEE415BF6FF4D2BE5FBE6" docLanguage="en" docName="Zootaxa.4205.4.301-338" docOrigin="Zootaxa 4205 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Divellepidosis indubitata Skuse 1888, new combination" docType="treatment" docVersion="12" lastPageNumber="318" masterDocId="2B60FFF30E7FEE505B61FFDA2A35FFDF" masterDocTitle="Revision of early taxa of Australian gall midges (Diptera: Cecidomyiidae)" masterLastPageNumber="338" masterPageNumber="301" pageNumber="318" updateTime="1698434883341" updateUser="ExternalLinkService">
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<mods:title id="7497DE1ECBB79C07C429E74D5206EFC9">Revision of early taxa of Australian gall midges (Diptera: Cecidomyiidae)</mods:title>
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<mods:namePart id="A3C156C3976B54176CB5AC6F5FA7C609">Kolesik, Peter</mods:namePart>
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<mods:namePart id="976E5A45D99303DA8B6FC359AFA8A7EE">Gagné, Raymond J.</mods:namePart>
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<mods:date id="F3D3D6E42F09A8BD13BF4FFCC7486D66">2016</mods:date>
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<treatment id="D759878B0E6EEE415BF6FF4D2BE5FBE6" ID-DOI="http://doi.org/10.5281/zenodo.4752523" ID-GBIF-Taxon="125140076" ID-Zenodo-Dep="4752523" LSID="urn:lsid:plazi:treatment:D759878B0E6EEE415BF6FF4D2BE5FBE6" httpUri="http://treatment.plazi.org/id/D759878B0E6EEE415BF6FF4D2BE5FBE6" lastPageNumber="318" pageId="17" pageNumber="318">
<subSubSection id="17EA65160E6EEE415BF6FF4D2B26FF0C" pageId="17" pageNumber="318" type="nomenclature">
<paragraph id="5F4F369D0E6EEE415BF6FF4D2966FF6D" blockId="17.[151,851,151,211]" box="[151,851,151,178]" pageId="17" pageNumber="318">
<heading id="040781F10E6EEE415BF6FF4D2966FF6D" bold="true" box="[151,851,151,178]" fontSize="11" level="1" pageId="17" pageNumber="318" reason="1">
<emphasis id="6D84EA8F0E6EEE415BF6FF4D2966FF6D" bold="true" box="[151,851,151,178]" pageId="17" pageNumber="318">
<taxonomicName id="98F04D1E0E6EEE415BF6FF4D285AFF6D" ID-CoL="92R6H" authority="Skuse, 1888" authorityName="Skuse" authorityYear="1888" box="[151,623,151,178]" class="Insecta" family="Cecidomyiidae" genus="Divellepidosis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="318" phylum="Arthropoda" rank="species" species="indubitata" status="comb. nov.">
<emphasis id="6D84EA8F0E6EEE415BF6FF4D2BF4FF6E" bold="true" box="[151,449,151,177]" italics="true" pageId="17" pageNumber="318">Divellepidosis indubitata</emphasis>
(
<bibRefCitation id="3B614B6C0E6EEE415AB1FF42285DFF6D" author="Skuse" box="[464,616,152,178]" pageId="17" pageNumber="318" refString="Skuse, F. A. A. (1888) Diptera of Australia. Part I. Proceedings of the Linnean Society of New South Wales, 3, 7 - 144. [pls. 2 - 3.]" type="journal article" year="1888">Skuse, 1888</bibRefCitation>
)
</taxonomicName>
,
<taxonomicNameLabel id="76B757F40E6EEE41591CFF422966FF6D" box="[637,851,152,178]" pageId="17" pageNumber="318">new combination</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="5F4F369D0E6EEE415BF6FF602B26FF0C" blockId="17.[151,851,151,211]" box="[151,275,186,211]" pageId="17" pageNumber="318">
[
<figureCitation id="C7CB2A180E6EEE415BFCFF602AD0FF0C" box="[157,229,186,211]" captionStart="FIGURE 9" captionStartId="16.[151,250,1831,1853]" captionTargetBox="[167,1422,635,1796]" captionTargetId="figure@16.[151,1436,609,1821]" captionTargetPageId="16" captionText="FIGURE 9. Divellepidosis pallidina (Skuse) (a h), Divellepidosis indubitata (Skuse) (i n). (a f, i n) male, (g h) female. (a) terminalia in dorsal view (left cercus lobe in lateral view), (b) wing, (c) sixth flagellomere in ventral view, (d) sixth flagellomere in dorsal view, (e) first tarsomere of hindleg, (f) last tarsomere of hindleg with claw and empodium, (g) sixth flagellomere in ventral view, (h) sixth flagellomere in dorsal view, (i) terminalia in dorsal view, (j) sixth flagellomere in ventral view, (k) sixth flagellomere in dorsal view, (l) wing, (m) first tarsomere of midleg, (n) palpus with palpiger." httpUri="https://zenodo.org/record/208250/files/figure.png" pageId="17" pageNumber="318">
Figs
<date id="2B4E105D0E6EEE415BB6FF602AD0FF0C" box="[215,229,186,211]" pageId="17" pageNumber="318">9</date>
</figureCitation>
in]
</paragraph>
</subSubSection>
<subSubSection id="17EA65160E6EEE415BF6FED82991FEE8" pageId="17" pageNumber="318" type="reference_group">
<paragraph id="5F4F369D0E6EEE415BF6FED8289AFEC7" blockId="17.[151,932,258,311]" box="[151,687,258,280]" pageId="17" pageNumber="318">
<treatmentCitationGroup id="7FE011B30E6EEE415BF6FED8289AFEC7" box="[151,687,258,280]" pageId="17" pageNumber="318">
<taxonomicName id="98F04D1E0E6EEE415BF6FED8289EFEC7" ID-CoL="8BCDK" authority="Skuse, 1888: 114" authorityName="Skuse" authorityPageNumber="114" authorityYear="1888" box="[151,683,258,280]" class="Insecta" family="Cecidomyiidae" genus="Cecidomyia" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="318" phylum="Arthropoda" rank="species" species="indubitata" subGenus="Colpodia">
<emphasis id="6D84EA8F0E6EEE415BF6FED82B25FEC7" box="[151,272,258,280]" italics="true" pageId="17" pageNumber="318">Cecidomyia</emphasis>
(
<emphasis id="6D84EA8F0E6EEE415A7FFED82B49FEC7" box="[286,380,258,280]" italics="true" pageId="17" pageNumber="318">Colpodia</emphasis>
)
<emphasis id="6D84EA8F0E6EEE415AEDFED82BC1FEC7" box="[396,500,258,280]" italics="true" pageId="17" pageNumber="318">indubitata</emphasis>
<treatmentCitation id="DE51108C0E6EEE415A9BFED8289EFEC7" author="Skuse" box="[506,683,258,280]" page="114" pageId="17" pageNumber="318" year="1888">
<bibRefCitation id="3B614B6C0E6EEE415A9BFED8284CFEC7" author="Skuse" box="[506,633,258,280]" pageId="17" pageNumber="318" refString="Skuse, F. A. A. (1888) Diptera of Australia. Part I. Proceedings of the Linnean Society of New South Wales, 3, 7 - 144. [pls. 2 - 3.]" type="journal article" year="1888">Skuse, 1888</bibRefCitation>
: 114
</treatmentCitation>
</taxonomicName>
.
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</paragraph>
<paragraph id="5F4F369D0E6EEE415BF6FEFB2991FEE8" blockId="17.[151,932,258,311]" box="[151,932,289,311]" pageId="17" pageNumber="318">
<treatmentCitationGroup id="7FE011B30E6EEE415BF6FEFB2991FEE8" box="[151,932,289,311]" pageId="17" pageNumber="318">
<taxonomicName id="98F04D1E0E6EEE415BF6FEFB2BC7FEE8" authority="Skuse 1888" authorityName="Skuse" authorityYear="1888" box="[151,498,289,311]" class="Insecta" family="Cecidomyiidae" genus="Colpodia" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="318" phylum="Arthropoda" rank="species" species="indubitata" status="comb. nov.">
<emphasis id="6D84EA8F0E6EEE415BF6FEFB2B56FEE8" box="[151,355,289,311]" italics="true" pageId="17" pageNumber="318">Colpodia indubitata</emphasis>
(
<bibRefCitation id="3B614B6C0E6EEE415A10FEFB2BDEFEE8" author="Skuse" box="[369,491,289,311]" pageId="17" pageNumber="318" refString="Skuse, F. A. A. (1888) Diptera of Australia. Part I. Proceedings of the Linnean Society of New South Wales, 3, 7 - 144. [pls. 2 - 3.]" type="journal article" year="1888">Skuse 1888</bibRefCitation>
)
</taxonomicName>
,
<taxonomicNameLabel id="76B757F40E6EEE415A9CFEF82899FEE8" box="[509,684,289,311]" pageId="17" pageNumber="318">new combination</taxonomicNameLabel>
by
<treatmentCitation id="DE51108C0E6EEE4159B3FEFB2995FEE8" author="Gagne" box="[722,928,289,311]" page="155" pageId="17" pageNumber="318" year="1989">
<bibRefCitation id="3B614B6C0E6EEE4159B3FEFB2995FEE8" author="Gagne" box="[722,928,289,311]" pageId="17" pageNumber="318" refString="Gagne, R. J. (1989 b) Family Cecidomyiidae. In: Evenhuis, N. L. (Ed.), &quot; Catalog of the Diptera of the Australasian and Oceanian Regions &quot; (Bishop Museum Press and E. J. Brill, Honolulu), pp. 152 - 163." type="book chapter" year="1989" yearSuffix="b">Gagné (1989b: 155)</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="17EA65160E6EEE415BF6FEBE2B30FE1A" pageId="17" pageNumber="318" type="materials_examined">
<paragraph id="5F4F369D0E6EEE415BF6FEBE2B30FE1A" blockId="17.[151,1437,356,1081]" pageId="17" pageNumber="318">
<emphasis id="6D84EA8F0E6EEE415BF6FEBE2BEFFEA2" bold="true" box="[151,474,356,381]" pageId="17" pageNumber="318">Material studied. Holotype</emphasis>
male (ANIC 29-38489), “Middle Harbour (Skuse). September”. The type retains the wings, head with partially shrivelled flagellomeres of one complete antenna, thorax with one hindleg, and the abdomen.
</paragraph>
</subSubSection>
<subSubSection id="17EA65160E6EEE415BA6FE152878FC86" pageId="17" pageNumber="318" type="description">
<paragraph id="5F4F369D0E6EEE415BA6FE152878FC86" blockId="17.[151,1437,356,1081]" pageId="17" pageNumber="318">
<emphasis id="6D84EA8F0E6EEE415BA6FE152B6CFE37" bold="true" box="[199,345,463,488]" pageId="17" pageNumber="318">Description.</emphasis>
Wing
<quantity id="98089B780E6EEE415AC5FE0A2BCEFE38" box="[420,507,464,488]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="17" pageNumber="318" unit="mm" value="1.8">1.8 mm</quantity>
long,
<quantity id="98089B780E6EEE415923FE1528ACFE38" box="[578,665,463,488]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="17" pageNumber="318" unit="mm" value="0.5">0.5 mm</quantity>
wide; R4+5 strongly curved at apical third, joining C at wing apex, C broken at juncture with R4+5, Rs fully developed, halfway between arculus and end of R1; Cu forked, Cu2 stronger than Cu1; wingfold weak. Head: palpus 4-segmented, palpiger present; flagellomeres 14, with single node, bearing single horizontal circumfilar band, neck 1.5x longer than node. First tarsomere with small blunt apicoventral projection, tarsal claws simple, empodium shrivelled on the remaining leg of
<typeStatus id="804B883F0E6EEE415F79FDB22E4BFD5E" box="[1048,1150,616,641]" pageId="17" pageNumber="318" type="holotype">holotype</typeStatus>
. Terminalia: gonocoxites short, without apical lobe dorsally; gonocoxal apodemes separate, ending anteriad of gonocoxites; gonostylus wide basally, with inner edge straight, outer edge curved, abruptly tapering towards apical comb-like tooth, entirely microtrichose and setose; cerci large, bluntly triangular, shallowly divided, with several apical setae; hypoproct concave, slightly shorter than cerci, with single seta on either lobe; tegmen stout, slightly concave apically, longer than cerci, bearing no papillae, ejaculatory apodeme starting between base of gonocoxal apodemes and base of aedeagus and reaching tip of aedeagus.
</paragraph>
</subSubSection>
<subSubSection id="17EA65160E6EEE415BA6FCBE2BE5FBE6" pageId="17" pageNumber="318" type="discussion">
<paragraph id="5F4F369D0E6EEE415BA6FCBE2BE5FBE6" blockId="17.[151,1437,356,1081]" pageId="17" pageNumber="318">
<emphasis id="6D84EA8F0E6EEE415BA6FCBE2B0EFCA2" bold="true" box="[199,315,868,893]" pageId="17" pageNumber="318">Remarks.</emphasis>
This species and
<taxonomicName id="98F04D1E0E6EEE41596DFCBF28A0FCA3" box="[524,661,868,892]" class="Insecta" family="Cecidomyiidae" genus="Divellepidosis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="318" phylum="Arthropoda" rank="species" species="pallidina">
<emphasis id="6D84EA8F0E6EEE41596DFCBF28A0FCA3" box="[524,661,868,892]" italics="true" pageId="17" pageNumber="318">D. pallidina</emphasis>
</taxonomicName>
represent the first records of
<taxonomicName id="98F04D1E0E6EEE41588FFCBE2FA9FCA2" authority="Fedotova &amp; Sidorenko" authorityName="Fedotova &amp; Sidorenko" box="[1006,1436,868,893]" class="Insecta" family="Cecidomyiidae" genus="Divellepidosis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="318" phylum="Arthropoda" rank="genus">
<emphasis id="6D84EA8F0E6EEE41588FFCBE2EBAFCA3" box="[1006,1167,868,892]" italics="true" pageId="17" pageNumber="318">Divellepidosis</emphasis>
Fedotova &amp; Sidorenko
</taxonomicName>
(
<taxonomicName id="98F04D1E0E6EEE415BFFFC522B62FC7E" box="[158,343,904,929]" class="Insecta" family="Cecidomyiidae" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="318" phylum="Arthropoda" rank="subFamily" subFamily="Porricondylinae">Porricondylinae</taxonomicName>
:
<taxonomicName id="98F04D1E0E6EEE415A05FC522832FC7E" box="[356,519,904,929]" class="Insecta" family="Cecidomyiidae" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="318" phylum="Arthropoda" rank="tribe" tribe="Porricondylini">Porricondylini</taxonomicName>
) in
<collectingCountry id="27E7760D0E6EEE415957FC522895FC7E" box="[566,672,904,929]" name="Australia" pageId="17" pageNumber="318">Australia</collectingCountry>
. The two species differ in the following characters: in
<taxonomicName id="98F04D1E0E6EEE415E72FC532FA9FC7F" box="[1299,1436,904,928]" class="Insecta" family="Cecidomyiidae" genus="Divellepidosis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="318" phylum="Arthropoda" rank="species" species="pallidina">
<emphasis id="6D84EA8F0E6EEE415E72FC532FA9FC7F" box="[1299,1436,904,928]" italics="true" pageId="17" pageNumber="318">D. pallidina</emphasis>
</taxonomicName>
the wing vein Rs is closer to the distal end of R1 than to the arculus, the gonocoxite is long with an acute dorsoapical lobe and the gonoxocal apodemes reach the base of gonocoxites; in
<taxonomicName id="98F04D1E0E6EEE415F2BFC0E2ED6FC34" box="[1098,1251,979,1003]" class="Insecta" family="Cecidomyiidae" genus="Colpodia" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="318" phylum="Arthropoda" rank="species" species="indubitata">
<emphasis id="6D84EA8F0E6EEE415F2BFC0E2ED6FC34" box="[1098,1251,979,1003]" italics="true" pageId="17" pageNumber="318">D. indubitata</emphasis>
</taxonomicName>
R S lies halfway between R1 and the arculus, the gonocoxite has no apical lobe, and the gonoxocal apodemes reach far anteriad of the base of the gonocoxites.
</paragraph>
</subSubSection>
</treatment>
</document>