251 lines
20 KiB
XML
251 lines
20 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/jhr.89.80163" ID-GBIF-Dataset="0177c1fd-eb14-43e2-b8f7-4b45ef4316cc" ID-Pensoft-Pub="1314-2607-89-19" ID-Pensoft-UUID="E7C0AB37B92C5EAF9DE5F0CED4F15BF3" ID-Zenodo-Dep="6336552" ID-ZooBank="AACD3DB78F694230B12F5E811CA88A15" ModsDocID="1314-2607-89-19" checkinTime="1646131449304" checkinUser="pensoft" docAuthor="Viertler, Alexandra, Klopfstein, Seraina, Jouault, Corentin & Spasojevic, Tamara" docDate="2022" docId="CC581986AB8D5A6D9B87D7A36FB64040" docLanguage="en" docName="JourHymenoptRes 89: 19-45" docOrigin="Journal of Hymenoptera Research 89" docPubDate="2022-02-28" docSource="http://dx.doi.org/10.3897/jhr.89.80163" docTitle="Madma oisella Viertler & Klopfstein & Jouault & Spasojevic 2022, sp. nov." docType="treatment" docUuid="83D47279-D75A-4BEB-80E9-9F3BA89F0A30" docUuidSource="ZooBank" docVersion="4" id="E7C0AB37B92C5EAF9DE5F0CED4F15BF3" lastPageNumber="19" masterDocId="E7C0AB37B92C5EAF9DE5F0CED4F15BF3" masterDocTitle="Darwin wasps (Hymenoptera, Ichneumonidae) in Lower Eocene amber from the Paris basin" masterLastPageNumber="45" masterPageNumber="19" pageNumber="19" updateTime="1678757445716" updateUser="pensoft">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Darwin wasps (Hymenoptera, Ichneumonidae) in Lower Eocene amber from the Paris basin</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Viertler, Alexandra</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-9729-5439</mods:nameIdentifier>
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<mods:affiliation>Naturhistorisches Museum Basel, Augustinergasse 2, CH- 4001 Basel, Switzerland & Institute of Ecology and Evolution, University of Bern, Baltzerstrasse 6, CH- 3012 Bern, Switzerland</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Klopfstein, Seraina</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-4025-975X</mods:nameIdentifier>
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<mods:affiliation>Institute of Ecology and Evolution, University of Bern, Baltzerstrasse 6, CH- 3012 Bern, Switzerland & Naturhistorisches Museum Basel, Augustinergasse 2, CH- 4001 Basel, Switzerland</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Jouault, Corentin</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-3680-5172</mods:nameIdentifier>
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<mods:affiliation>Institut de Syste ́ matique, E ́ volution, Biodiversite ́ (ISYEB), Muse ́ um national d'Histoire naturelle, CNRS, Sorbonne Universite ́, EPHE, Universite ́ des Antilles, CP 50, 57 rue Cuvier, 75005 Paris, France & Institut des Sciences de l'E ́ volution de Montpellier, CNRS, Place Euge ̀ ne Bataillon, UMR 5554, 34095 Montpellier, France & Univ. Rennes, CNRS, Ge ́ osciences Rennes, UMR 6118, 35000 Rennes, France</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Spasojevic, Tamara</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-5301-5722</mods:nameIdentifier>
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<mods:affiliation>Naturhistorisches Museum Basel, Augustinergasse 2, CH- 4001 Basel, Switzerland & Institute of Ecology and Evolution, University of Bern, Baltzerstrasse 6, CH- 3012 Bern, Switzerland</mods:affiliation>
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<mods:nameIdentifier type="email">spasojevic.ta@gmail.com</mods:nameIdentifier>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>Journal of Hymenoptera Research</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2022</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2022-02-28</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>89</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>19</mods:start>
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<mods:end>45</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/jhr.89.80163</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/jhr.89.80163</mods:identifier>
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||
<mods:identifier type="Pensoft-Pub">1314-2607-89-19</mods:identifier>
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||
<mods:identifier type="ZooBank">AACD3DB78F694230B12F5E811CA88A15</mods:identifier>
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<mods:identifier type="Pensoft-UUID">E7C0AB37B92C5EAF9DE5F0CED4F15BF3</mods:identifier>
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||
<mods:identifier type="Zenodo-Dep">6336552</mods:identifier>
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||
</mods:mods>
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<subSection pageId="0" pageNumber="19" type="systematic palaeontology">
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<treatment ID-GBIF-Taxon="193849817" LSID="urn:lsid:zoobank.org:act:83D47279-D75A-4BEB-80E9-9F3BA89F0A30" httpUri="http://treatment.plazi.org/id/CC581986AB8D5A6D9B87D7A36FB64040" lastPageNumber="19" pageId="0" pageNumber="19">
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<subSubSection pageId="0" pageNumber="19" type="nomenclature">
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<paragraph pageId="0" pageNumber="19">
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<taxonomicName LSID="http://zoobank.org/83D47279-D75A-4BEB-80E9-9F3BA89F0A30" authority="Viertler & Klopfstein & Jouault & Spasojevic, 2022" authorityName="Viertler & Klopfstein & Jouault & Spasojevic" authorityYear="2022" class="Insecta" family="Ichneumonidae" genus="Madma" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Madma oisella" order="Hymenoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="oisella" status="sp. nov.">Madma oisella</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="19">sp. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="19" type="description">
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<paragraph pageId="0" pageNumber="19">
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<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Holotype of Madma oisella sp. nov. A lateral view of whole specimen B fore tibia, black triangle points to tooth on apical margin C latero-ventral view showing propodeum and bilobed posterior transverse carina on mesosternum (I) and extended small lobe on metapleuron (II) D lateral view of head E first tergite and sternite F frontal view of face G ovipositor H Detailed photo of fore wing. Scale bar: 2 mm (A); 0.5 mm (B, D-G); 1 mm (H)." figureDoi="10.3897/jhr.89.80163.figure5" httpUri="https://binary.pensoft.net/fig/651025" pageId="0" pageNumber="19">Figs 5</figureCitation>
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<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Interpretative line drawing of Madma oisella sp. nov. Dotted lines indicate uncertain interpretations. Different detailed photographs were used as templates to create this drawing. Scale bar: 1 mm." figureDoi="10.3897/jhr.89.80163.figure6" httpUri="https://binary.pensoft.net/fig/651026" pageId="0" pageNumber="19">, 6</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="19" type="etymology">
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<paragraph pageId="0" pageNumber="19">Etymology.</paragraph>
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<paragraph pageId="0" pageNumber="19">
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Derived from the locality name: Oise. The word is transformed to resemble the ending of the French word for Miss
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<normalizedToken originalValue="“Madmoiselle”">"Madmoiselle"</normalizedToken>
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and latinised.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="19" type="materials_examined">
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<paragraph pageId="0" pageNumber="19">Material examined.</paragraph>
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<paragraph pageId="0" pageNumber="19">
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<materialsCitation accessionNumber="A71347" collectingDate="2022-01-01" collectingDateMax="2022-12-31" collectingDateMin="2022-01-01" collectorName="Oise, Creil, Chevriere, Le Quesnoy, G. De Ploeg, A. Nel" country="France" latitude="49.335" location="France" longLatPrecision="1" longitude="2.6746945" specimenCount="1" typeStatus="Holotype">
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<collectingCountry name="France">France</collectingCountry>
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•
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<emphasis bold="true" italics="true" pageId="0" pageNumber="19">
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<typeStatus>Holotype</typeStatus>
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</emphasis>
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female;
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<collectorName>Oise</collectorName>
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department, region of
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<collectorName>Creil</collectorName>
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,
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<normalizedToken originalValue="Chevrière">
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<collectorName>Chevriere</collectorName>
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</normalizedToken>
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,
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<collectorName>Le Quesnoy</collectorName>
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;
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<geoCoordinate degrees="49" direction="north" minutes="20" orientation="latitude" precision="1" seconds="06.0" value="49.335">49°20'06.0"N</geoCoordinate>
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,
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<geoCoordinate degrees="2" direction="east" minutes="40" orientation="longitude" precision="1" seconds="28.9" value="2.6746945">2°40'28.9"E</geoCoordinate>
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; 1998-2000;
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<collectorName>
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G. De
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<normalizedToken originalValue="Ploëg">Ploeg</normalizedToken>
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</collectorName>
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and
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<collectorName>A. Nel</collectorName>
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leg.; in amber; MNHN.F.
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<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/A71347">A71347</accessionNumber>
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(PA2189).
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</materialsCitation>
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</paragraph>
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<caption ID-Zenodo-Dep="6336562" doi="10.3897/jhr.89.80163.figure5" httpUri="https://binary.pensoft.net/fig/651025" pageId="0" pageNumber="19" start="Figure 5" startId="F5">
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<paragraph pageId="0" pageNumber="19">
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<emphasis bold="true" pageId="0" pageNumber="19">Figure 5.</emphasis>
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<typeStatus>Holotype</typeStatus>
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of
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<taxonomicName authorityName="Viertler & Klopfstein & Jouault & Spasojevic" authorityYear="2022" class="Insecta" family="Ichneumonidae" genus="Madma" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Madma oisella" order="Hymenoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="oisella">
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<emphasis italics="true" pageId="0" pageNumber="19">Madma oisella</emphasis>
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</taxonomicName>
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sp. nov.
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<emphasis bold="true" pageId="0" pageNumber="19">A</emphasis>
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lateral view of whole specimen
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<emphasis bold="true" pageId="0" pageNumber="19">B</emphasis>
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fore tibia, black triangle points to tooth on apical margin
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<emphasis bold="true" pageId="0" pageNumber="19">C</emphasis>
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latero-ventral view showing propodeum and bilobed posterior transverse carina on mesosternum (I) and extended small lobe on metapleuron (II)
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<emphasis bold="true" pageId="0" pageNumber="19">D</emphasis>
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lateral view of head
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<emphasis bold="true" pageId="0" pageNumber="19">E</emphasis>
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first tergite and sternite
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<emphasis bold="true" pageId="0" pageNumber="19">F</emphasis>
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frontal view of face
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<emphasis bold="true" pageId="0" pageNumber="19">G</emphasis>
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ovipositor
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<emphasis bold="true" pageId="0" pageNumber="19">H</emphasis>
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Detailed photo of fore wing. Scale bar:
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<quantity metricMagnitude="-3" metricUnit="m" metricValue="2.0" unit="mm" value="2.0">2 mm</quantity>
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(
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<emphasis bold="true" pageId="0" pageNumber="19">A</emphasis>
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);
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<quantity metricMagnitude="-4" metricUnit="m" metricValue="5.0" unit="mm" value="0.5">0.5 mm</quantity>
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(
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<emphasis bold="true" pageId="0" pageNumber="19">B, D-G</emphasis>
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);
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<quantity metricMagnitude="-3" metricUnit="m" metricValue="1.0" unit="mm" value="1.0">1 mm</quantity>
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(
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<emphasis bold="true" pageId="0" pageNumber="19">H</emphasis>
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).
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="19" type="type condition">
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<paragraph pageId="0" pageNumber="19">Type condition.</paragraph>
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<paragraph pageId="0" pageNumber="19">Lateral aspects clearly visible through amber; dorsal and ventral aspects partly visible through irregular surface of amber. Complete, except few apical flagellomeres at surface of amber with their ventral side missing. Body translucent, thus parts of body from opposite side often visible below cuticle, as well as some inner structures. Body surface sculpture and carinae weakly visible; outer surface partly covered with dark material, which probably represents remains of organic matter.</paragraph>
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<caption ID-Zenodo-Dep="6336564" doi="10.3897/jhr.89.80163.figure6" httpUri="https://binary.pensoft.net/fig/651026" pageId="0" pageNumber="19" start="Figure 6" startId="F6">
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<paragraph pageId="0" pageNumber="19">
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<emphasis bold="true" pageId="0" pageNumber="19">Figure 6.</emphasis>
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Interpretative line drawing of
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<taxonomicName authorityName="Viertler & Klopfstein & Jouault & Spasojevic" authorityYear="2022" class="Insecta" family="Ichneumonidae" genus="Madma" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Madma oisella" order="Hymenoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="oisella">
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<emphasis italics="true" pageId="0" pageNumber="19">Madma oisella</emphasis>
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</taxonomicName>
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sp. nov. Dotted lines indicate uncertain interpretations. Different detailed photographs were used as templates to create this drawing. Scale bar: 1 mm.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="19" type="diagnosis">
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<paragraph pageId="0" pageNumber="19">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="19">See genus diagnosis.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="19" type="description">
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<paragraph pageId="0" pageNumber="19">Description.</paragraph>
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<paragraph pageId="0" pageNumber="19">Body around 4.9 mm. Color difficult to interpret, except tip of mandibles darker, hind tibiae clearly darker at base and apex (dark-light-dark), antennae uniformly colored and T1 dark.</paragraph>
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<paragraph pageId="0" pageNumber="19">
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<emphasis bold="true" pageId="0" pageNumber="19">
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<emphasis italics="true" pageId="0" pageNumber="19">Head</emphasis>
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.
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</emphasis>
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Mandibles moderately large, weakly tapered from base to apex, not twisted, with two oblique and subequal teeth; outer surface without strong sculpture, smooth or sparsely pubescent. Malar space moderately long, 0.9
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<normalizedToken originalValue="×">x</normalizedToken>
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mandibular width at base, smooth, seemingly without subocular sulcus. Labrum not clearly exposed below clypeus. Clypeus in frontal view subquadrate, transversely undivided and separated from face with weak clypeal groove, in lateral view clearly convex; apical margin simple, truncate, without tooth or tubercles. Face weakly convex with weak median swelling, without upper process. Eyes large, in lateral view 0.85
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<normalizedToken originalValue="×">x</normalizedToken>
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head height; inner margin of eyes straight opposite antennal sockets, in frontal view parallel to each other. Modification between antennal sockets absent. Frons without strong impressions for scape. Ocelli of normal size, separated from eyes by more than their diameter. Vertex steeply declivous behind ocelli with straight surface to occipital carina. Occipital carina complete and dorsally evenly convex, high on head, in lateral view almost reaching height of dorsal eye margin. Maxillary palps with five segments, labial palps not visible. Scape about as long as wide, without extended membranous area; pedicel clearly smaller than scape. Flagellum 3.9 mm with 27 segments; first flagellomere 6.25
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<normalizedToken originalValue="×">x</normalizedToken>
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as long as wide; median and apical flagellomeres longer than wide; tip of apical flagellomere unmodified.
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</paragraph>
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<paragraph pageId="0" pageNumber="19">
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<emphasis bold="true" pageId="0" pageNumber="19">
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<emphasis italics="true" pageId="0" pageNumber="19">Mesosoma</emphasis>
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.
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</emphasis>
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Pronotum in profile on left side higher than long, on right side about as high as long (specimen clearly viewed only in slight diagonal position), without latero-ventral posteriorly projecting lobe; pronotal collar not visible; anterior vertical part clearly extended dorsally. Epomia weakly visible, present ventrally, extending laterally but unclear how far. Modifications near base of notauli absent. Mesosternum with epicnemial carina extending laterally on mesopleuron at least to mid-height of pronotum, possibly longer, but dorsal part not visible clearly through amber; not modified behind fore coxae. Posterior transverse carina absent laterally, present in front of mid coxae as a raised flange and in between as a low carina. Mesopleuron evenly convex with sternaulus weakly visible, extending from ventrolateral anterior margin of mesopleuron to posterior margin of mesopleuron clearly above mid-height of mid coxa, bowed upwards around mid-height. Mesopleural furrow with horizontal impression extending from around mid-height to episternal scrobe. Mesoscutum evenly shagreened and matt. Notauli deeply impressed anteriorly, extending posteriorly past centre of mesoscutum but unclear how far. Carina along lateral margin of mesoscutum evenly raised and extending to anterior margin of scutellum. Scutellum in profile flat, smooth, without carinae. Postscutellum evenly convex in profile. Propodeum about as long as high; structure difficult to assess but either smooth or punctured. Metapleuron as long as high, with anteroventral corner extended to a small lobe, without juxtacoxal carina and at most with few rugae lateroventrally. Submetapleural carina difficult to interpret, present only in anterior half or completely absent. Propodeal carination complete, although presence of medial portion of anterior transverse carina uncertain; area basalis shorter than area superomedia; the latter similar in size to area petiolaris. Spiracle subcircular, separated from pleural carina by its own diameter, touching dorsal half of pleural portion of lateral longitudinal carina. Dorsal margin of metacoxal cavities above ventral margin of metasomal cavity. Legs simple, not unusually stout or slender. Fore tibia with small apical tooth. Mid and hind tibia with two long slender spurs. Hind tibia with fringe of parallel setae on inner apex; 1st tarsomere 1.1
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<normalizedToken originalValue="×">x</normalizedToken>
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as long as wide; 4th tarsomere apically more or less evenly truncated. Tarsal claws without modification.
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</paragraph>
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<paragraph pageId="0" pageNumber="19">
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<emphasis bold="true" pageId="0" pageNumber="19">
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<emphasis italics="true" pageId="0" pageNumber="19">Wings</emphasis>
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.
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</emphasis>
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Fore wing 3.8 mm. Areolet oblique pentagonal, 1.5-2.2
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<normalizedToken originalValue="×">x</normalizedToken>
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as wide as long, with 3Rs 0.8
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<normalizedToken originalValue="×">x</normalizedToken>
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2Rs, 2 + 3M 2.1
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<normalizedToken originalValue="×">x</normalizedToken>
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2Rs, 4M 0.3
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<normalizedToken originalValue="×">x</normalizedToken>
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2 + 3M; 3rs-m with two bullae. 2m-cu clearly curved outwards, with two bullae. 1Rs + M absent. 1m-cu at level of 1M + 1Rs. Pterostigma 4.3
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<normalizedToken originalValue="×">x</normalizedToken>
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as long as wide. Cell 2R1 2.1
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<normalizedToken originalValue="×">x</normalizedToken>
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as long as pterostigma, 3.4
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<normalizedToken originalValue="×">x</normalizedToken>
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as long as wide. 2Cu 0.8
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<normalizedToken originalValue="×">x</normalizedToken>
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1M + 1Rs, 0.8
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<normalizedToken originalValue="×">x</normalizedToken>
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r-rs. 3Cu about same length as 2cu-a. Hind wing with M + Cu curved distally. 1Cu around same length as cu-a. 1Rs 1.2
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<normalizedToken originalValue="×">x</normalizedToken>
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as long as rs-m.
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</paragraph>
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<paragraph pageId="0" pageNumber="19">
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<emphasis bold="true" pageId="0" pageNumber="19">
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<emphasis italics="true" pageId="0" pageNumber="19">Metasoma</emphasis>
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.
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</emphasis>
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Depressed to cylindrical. S1 reaches around middle of T1, anteriorly with a median keel, centrally with a weak median swelling. Presence of laterotergite 1 unclear; if present then membranous on entire length. T1 in dorsal view about 2-2.5
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<normalizedToken originalValue="×">x</normalizedToken>
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as long as wide, petiolate, but sides evenly tapering from apex to base as tergite relatively short, in lateral view continuously expanding in profile; spiracle at around 0.6
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<normalizedToken originalValue="×">x</normalizedToken>
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T1 length in profile; dorso-lateral carina complete, above spiracle; latero-median carinae long, almost reaching posterior margin, parallel to each other; sculpture shagreened or punctured, but could be an artefact. T2 in dorsal view transverse; sculpture if present not strong and even on entire tergite length; latero-median carinae absent; impressions in anterior half either absent or shallow; thyridium present and shape unclear. Dorsal surface of T3 and remaining tergites evenly convex. T3 and T4 with posterolateral corners rounded. S6 unclear, seems transverse with simple apical margin. T7 conspicuously shorter than T6. T8 short, not elongated in horn or boss. Ovipositor sheaths 0.2
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
metasoma length, parallel-sided, with dense short setae. Ovipositor straight and compressed, parallel sided until tip where dorsal valve expanding into nodus; tip of lower valve with at least four distinct oblique teeth.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</subSection>
|
||
</document> |