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<document id="6FE9848A6482ED98593EB7F6BBE74553" ID-CLB-Dataset="8487" ID-DOI="10.11646/zootaxa.4853.2.4" ID-GBIF-Dataset="d47312c2-2225-42f8-8445-be0999dbfc04" ID-ISSN="1175-5326" ID-Zenodo-Dep="4410589" ID-ZooBank="4F99C503-C467-4514-84F3-486B6D65C87A" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1609602039661" checkinUser="plazi" docAuthor="Trujano-Ortega, Marysol, Callaghan, Curtis J., García-Vázquez, Uri Omar, Luis-Martínez, Armando, Ávalos-Hernández, Omar &amp; Llorente-Bousquets, Jorge" docDate="2020" docId="B516E37D59344C0BFF194AC24333FC48" docLanguage="en" docName="zootaxa.4853.2.4.pdf" docOrigin="Zootaxa 4853 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Emesis planeca Trujano-Ortega &amp; Callaghan &amp; García-Vázquez &amp; Luis-Martínez &amp; Ávalos-Hernández &amp; Llorente-Bousquets 2020, n. comb." docType="treatment" docVersion="8" lastPageNumber="225" masterDocId="492F9B0559304C0CFF8E4A1D477AFF81" masterDocTitle="Emesis planeca n. comb. (Lepidoptera: Riodinidae): a new combination revealed by molecular evidence with a description of its morphological variation" masterLastPageNumber="234" masterPageNumber="218" pageNumber="222" updateTime="1698851236129" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="A13EFCCA9E9472FF299A0AA79311D08C">Emesis planeca n. comb. (Lepidoptera: Riodinidae): a new combination revealed by molecular evidence with a description of its morphological variation</mods:title>
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<mods:namePart id="D97413FB858993E02F771D6563843DCA">Trujano-Ortega, Marysol</mods:namePart>
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<mods:affiliation id="9BEC44B85AAAE1F2838829BD83C2E640">Facultad de Estudios Superiores Zaragoza, Universidad Nacional Autónoma de México, Batalla 5 de Mayo s / n, Ejército de Oriente, Ciudad de México, 09230, México. urigarcia @ gmail. com; https: // orcid. org / 0000 - 0003 - 2963 - 0981</mods:affiliation>
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<mods:affiliation id="5F9BFBE39D28BE6BBC6C20D4DACBE2AC">alm @ ciencias. unam. mx; https: // orcid. org / 0000 - 0002 - 1044 - 3986</mods:affiliation>
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<mods:namePart id="D66434C430A43CAF1DF5726D687C289B">Ávalos-Hernández, Omar</mods:namePart>
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<mods:affiliation id="306D73A00D4A810859BEE05A6EF2E549">omaravalosh @ ciencias. unam. mx; https: // orcid. org / 0000 - 0002 - 5476 - 9400</mods:affiliation>
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<mods:affiliation id="73FF3DAB8EA2BA444CD6A51F5D669D8C">enantia @ prodigy. net. mx; https: // orcid. org / 0000 - 0002 - 9542 - 2358</mods:affiliation>
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<treatment id="B516E37D59344C0BFF194AC24333FC48" ID-DOI="http://doi.org/10.5281/zenodo.4498875" ID-GBIF-Taxon="177209367" ID-Zenodo-Dep="4498875" LSID="urn:lsid:plazi:treatment:B516E37D59344C0BFF194AC24333FC48" httpUri="http://treatment.plazi.org/id/B516E37D59344C0BFF194AC24333FC48" lastPageId="7" lastPageNumber="225" pageId="4" pageNumber="222">
<subSubSection id="75A501E059344C08FF194AC24341FF78" box="[151,1083,223,250]" pageId="4" pageNumber="222" type="nomenclature">
<paragraph id="3D00526B59344C08FF194AC24341FF78" blockId="4.[151,1083,223,285]" box="[151,1083,223,250]" pageId="4" pageNumber="222">
<heading id="6648E50759344C08FF194AC24341FF78" bold="true" box="[151,1083,223,250]" fontSize="11" level="1" pageId="4" pageNumber="222" reason="1">
<emphasis id="0FCB8E7959344C08FF194AC24341FF78" bold="true" box="[151,1083,223,250]" pageId="4" pageNumber="222">
<taxonomicName id="FABF29E859344C08FF194AC24456FF78" authority="(R. de la Maza E. &amp; J. de la Maza E.)" authorityName="Trujano-Ortega &amp; Callaghan &amp; García-Vázquez &amp; Luis-Martínez &amp; Ávalos-Hernández &amp; Llorente-Bousquets" authorityYear="2020" baseAuthorityName="R. de la Maza E. &amp; J. de la Maza E." box="[151,812,223,250]" class="Insecta" family="Lycaenidae" genus="Emesis" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="222" phylum="Arthropoda" rank="species" species="planeca" status="comb. nov.">
<emphasis id="0FCB8E7959344C08FF194AC2462EFF7B" bold="true" box="[151,340,223,250]" italics="true" pageId="4" pageNumber="222">Emesis planeca</emphasis>
(R. de la Maza E. &amp; J. de la Maza E.)
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<taxonomicNameLabel id="14F8330259344C08FCBA4AC244E5FF78" box="[820,927,223,249]" pageId="4" pageNumber="222" rank="species">n. comb.</taxonomicNameLabel>
(
<emphasis id="0FCB8E7959344C08FC3E4AC24350FF7B" bold="true" box="[944,1066,223,250]" italics="true" pageId="4" pageNumber="222">Apodemia</emphasis>
).
</emphasis>
</heading>
</paragraph>
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<subSubSection id="75A501E059344C08FF194B1E4666FE9C" box="[151,284,259,285]" pageId="4" pageNumber="222" type="description">
<paragraph id="3D00526B59344C08FF194B1E4666FE9C" blockId="4.[151,1083,223,285]" box="[151,284,259,285]" pageId="4" pageNumber="222">
<figureCitation id="A5844EEE59344C08FF194B1E4799FE9C" box="[151,227,259,285]" captionStart="FIGURE 1" captionStartId="1.[151,250,1942,1966]" captionTargetBox="[159,1428,1252,1912]" captionTargetId="figure-460@1.[151,1436,1242,1916]" captionTargetPageId="1" captionText="FIGURE 1. Emesis planeca n. comb. Dorsal (left) and ventral (right) views. AF males, GL females. México: Michoacán: Múgica, El Marqués, 500 m elev. leg. L. L. González Cota. Collection catalog numbers (MZFC LEP):A, 429994; B, 429990; C, 429991; D, 429992; E, 429993; F, 429988; G, 429998; H, 429995; I, 429999; J, 429997; K, 429996; L 354625." figureDoi="http://doi.org/10.5281/zenodo.4410591" httpUri="https://zenodo.org/record/4410591/files/figure.png" pageId="4" pageNumber="222">Figs. 1</figureCitation>
,
<figureCitation id="A5844EEE59344C08FF7F4B1E4666FE9C" box="[241,284,259,285]" captionStart-0="FIGURE 3" captionStart-1="FIGURE 4" captionStart-2="FIGURE 5" captionStart-3="FIGURE 6" captionStart-4="FIGURE 7" captionStartId-0="5.[151,250,1943,1967]" captionStartId-1="6.[151,250,1284,1308]" captionStartId-2="7.[151,250,1880,1904]" captionStartId-3="8.[151,250,1944,1968]" captionStartId-4="9.[151,250,1111,1135]" captionTargetBox-0="[271,1327,201,1904]" captionTargetBox-1="[152,1436,184,1260]" captionTargetBox-2="[157,1426,1001,1847]" captionTargetBox-3="[194,1395,181,1916]" captionTargetBox-4="[154,1435,184,1084]" captionTargetId-0="figure-29@5.[260,1327,181,1919]" captionTargetId-1="figure-22@6.[152,1436,181,1260]" captionTargetId-2="figure-447@7.[151,1436,993,1856]" captionTargetId-3="figure-22@8.[193,1395,181,1920]" captionTargetId-4="figure-29@9.[151,1436,181,1087]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionTargetPageId-2="7" captionTargetPageId-3="8" captionTargetPageId-4="9" captionText-0="FIGURE 3. Wing of Emesis planeca n. comb. FW, forewing; HW, hindwing. Vein abbreviations (black lettering): Sc subcostal, R radial, M median, Cu cubital, A anal.Arrows in dorsal view point at: band of six white spots in the postmedial region and; line of small black dots (six dots in forewing, five dots in hindwing) in the submarginal area." captionText-1="FIGURE 4. Male labial palpus and latero - ventral view of the head of Emesis planeca n. comb." captionText-2="FIGURE 5. Male genitalia of Emesis planeca n. comb. (MZFC LEP 429988). Arrows in lateral view point at: two types of cornuti and; dorsal process of the valvae with many setae. In ventral view this process looks squared, wide and projected outwards." captionText-3="FIGURE 6. Female genitalia of Emesis planeca n. comb. (MZFC LEP 429998). Arrows in ventral and lateral views point at: antrum (wide, strongly sclerotized, and funnel-shaped) and; the single long invaginated signum with numerous tiny spines." captionText-4="FIGURE 7. Geographic distribution of Emesis planeca n. comb. and other Emesis species in Michoacán, Mexico. Black lines represent state limits." figureDoi-0="http://doi.org/10.5281/zenodo.4410595" figureDoi-1="http://doi.org/10.5281/zenodo.4410597" figureDoi-2="http://doi.org/10.5281/zenodo.4410599" figureDoi-3="http://doi.org/10.5281/zenodo.4410601" figureDoi-4="http://doi.org/10.5281/zenodo.4410603" httpUri-0="https://zenodo.org/record/4410595/files/figure.png" httpUri-1="https://zenodo.org/record/4410597/files/figure.png" httpUri-2="https://zenodo.org/record/4410599/files/figure.png" httpUri-3="https://zenodo.org/record/4410601/files/figure.png" httpUri-4="https://zenodo.org/record/4410603/files/figure.png" pageId="4" pageNumber="222">37</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="75A501E059344C08FF194B564612FE74" pageId="4" pageNumber="222" type="diagnosis">
<paragraph id="3D00526B59344C08FF194B564612FE74" blockId="4.[151,1437,331,1942]" pageId="4" pageNumber="222">
<emphasis id="0FCB8E7959344C08FF194B564670FEE4" bold="true" box="[151,266,331,357]" pageId="4" pageNumber="222">Diagnosis</emphasis>
. Postmedial region in dorsal and ventral views of both wings with a band of white spots, more evident in ventral view; submarginal area in dorsal and ventral views of both wings with a line of small black dots, six dots in dorsal view of forewing, five dots in dorsal view of hindwing, seven dots in ventral view of forewing and six dots in ventral view of hindwing (
<figureCitation id="A5844EEE59344C08FD9E4BAA4526FE50" box="[528,604,439,465]" captionStart="FIGURE 1" captionStartId="1.[151,250,1942,1966]" captionTargetBox="[159,1428,1252,1912]" captionTargetId="figure-460@1.[151,1436,1242,1916]" captionTargetPageId="1" captionText="FIGURE 1. Emesis planeca n. comb. Dorsal (left) and ventral (right) views. AF males, GL females. México: Michoacán: Múgica, El Marqués, 500 m elev. leg. L. L. González Cota. Collection catalog numbers (MZFC LEP):A, 429994; B, 429990; C, 429991; D, 429992; E, 429993; F, 429988; G, 429998; H, 429995; I, 429999; J, 429997; K, 429996; L 354625." figureDoi="http://doi.org/10.5281/zenodo.4410591" httpUri="https://zenodo.org/record/4410591/files/figure.png" pageId="4" pageNumber="222">Figs. 1</figureCitation>
,
<figureCitation id="A5844EEE59344C08FDE44BAA4500FE50" box="[618,634,439,465]" captionStart="FIGURE 3" captionStartId="5.[151,250,1943,1967]" captionTargetBox="[271,1327,201,1904]" captionTargetId="figure-29@5.[260,1327,181,1919]" captionTargetPageId="5" captionText="FIGURE 3. Wing of Emesis planeca n. comb. FW, forewing; HW, hindwing. Vein abbreviations (black lettering): Sc subcostal, R radial, M median, Cu cubital, A anal.Arrows in dorsal view point at: band of six white spots in the postmedial region and; line of small black dots (six dots in forewing, five dots in hindwing) in the submarginal area." figureDoi="http://doi.org/10.5281/zenodo.4410595" httpUri="https://zenodo.org/record/4410595/files/figure.png" pageId="4" pageNumber="222">3</figureCitation>
). These characters distinguish
<taxonomicName id="FABF29E859344C08FC5E4BA54305FE50" authorityName="Trujano-Ortega &amp; Callaghan &amp; García-Vázquez &amp; Luis-Martínez &amp; Ávalos-Hernández &amp; Llorente-Bousquets" authorityYear="2020" baseAuthorityName="Trujano-Ortega &amp; Callaghan &amp; García-Vázquez &amp; Luis-Martínez &amp; Ávalos-Hernández &amp; Llorente-Bousquets" baseAuthorityYear="2020" box="[976,1151,439,465]" class="Insecta" family="Lycaenidae" genus="Emesis" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="222" phylum="Arthropoda" rank="species" species="planeca" status="comb. nov.">
<emphasis id="0FCB8E7959344C08FC5E4BA54305FE50" box="[976,1151,439,465]" italics="true" pageId="4" pageNumber="222">Emesis planeca</emphasis>
</taxonomicName>
<emphasis id="0FCB8E7959344C08FB0B4BAA4390FE50" bold="true" box="[1157,1258,439,465]" pageId="4" pageNumber="222">
<taxonomicNameLabel id="14F8330259344C08FB0B4BAA4390FE50" box="[1157,1258,439,465]" pageId="4" pageNumber="222" rank="species">n. comb.</taxonomicNameLabel>
</emphasis>
from all known
<taxonomicName id="FABF29E859344C08FF194BC64673FE74" box="[151,265,475,501]" class="Insecta" family="Riodinidae" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="222" phylum="Arthropoda" rank="tribe" tribe="Emesidini">Emesidini</taxonomicName>
species.
</paragraph>
</subSubSection>
<subSubSection id="75A501E059344C0BFF494BE24333FC48" lastPageId="7" lastPageNumber="225" pageId="4" pageNumber="222" type="description">
<paragraph id="3D00526B59344C08FF494BE2438BFCB8" blockId="4.[151,1437,331,1942]" pageId="4" pageNumber="222">
<emphasis id="0FCB8E7959344C08FF494BE246DEFD98" bold="true" box="[199,420,511,538]" pageId="4" pageNumber="222">
Description.
<emphasis id="0FCB8E7959344C08FEEC4BE246E7FD9B" bold="true" box="[354,413,511,538]" italics="true" pageId="4" pageNumber="222">Male</emphasis>
.
</emphasis>
(
<figureCitation id="A5844EEE59344C08FE384BE2468CFD98" box="[438,502,511,537]" captionStart="FIGURE 1" captionStartId="1.[151,250,1942,1966]" captionTargetBox="[159,1428,1252,1912]" captionTargetId="figure-460@1.[151,1436,1242,1916]" captionTargetPageId="1" captionText="FIGURE 1. Emesis planeca n. comb. Dorsal (left) and ventral (right) views. AF males, GL females. México: Michoacán: Múgica, El Marqués, 500 m elev. leg. L. L. González Cota. Collection catalog numbers (MZFC LEP):A, 429994; B, 429990; C, 429991; D, 429992; E, 429993; F, 429988; G, 429998; H, 429995; I, 429999; J, 429997; K, 429996; L 354625." figureDoi="http://doi.org/10.5281/zenodo.4410591" httpUri="https://zenodo.org/record/4410591/files/figure.png" pageId="4" pageNumber="222">Fig. 1</figureCitation>
AF). Most of the examined specimens are smaller than the original description, with wingspan of
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(n=7). Additional forewing length measurements are: a) wing base to anterior apex
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(n=7, ẍ = 13.5 ±0.5), b) anterior apex to anal angle
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(n=7, ẍ = 9.35 ±0.5), and c) anal angle to wing base
<quantity id="FA47FF8E59344C08FEFE4876457FFD07" box="[368,517,619,646]" metricMagnitude="-3" metricUnit="m" metricValue="8.61" metricValueMax="9.1" metricValueMin="8.12" pageId="4" pageNumber="222" unit="mm" value="8.61" valueMax="9.1" valueMin="8.12">8.129.1 mm</quantity>
(n=7, ẍ = 8.12 ±0.33).
<emphasis id="0FCB8E7959344C08FC9E487144E1FD04" box="[784,923,619,645]" italics="true" pageId="4" pageNumber="222">Wing shape:</emphasis>
Forewing is triangular with the costal margin slightly curved, apex with a small curve extending into the distal margin, thus appearing slightly convex. The internal margin is straight and shorter than the costal margin. Hindwing is more rounded than forewing, apex and distal margin convex and anal margin straight (
<figureCitation id="A5844EEE59344C08FDE348CA45C9FD70" box="[621,691,727,753]" captionStart="FIGURE 3" captionStartId="5.[151,250,1943,1967]" captionTargetBox="[271,1327,201,1904]" captionTargetId="figure-29@5.[260,1327,181,1919]" captionTargetPageId="5" captionText="FIGURE 3. Wing of Emesis planeca n. comb. FW, forewing; HW, hindwing. Vein abbreviations (black lettering): Sc subcostal, R radial, M median, Cu cubital, A anal.Arrows in dorsal view point at: band of six white spots in the postmedial region and; line of small black dots (six dots in forewing, five dots in hindwing) in the submarginal area." figureDoi="http://doi.org/10.5281/zenodo.4410595" httpUri="https://zenodo.org/record/4410595/files/figure.png" pageId="4" pageNumber="222">Fig. 3</figureCitation>
).
<emphasis id="0FCB8E7959344C08FD4448C54456FD70" box="[714,812,728,753]" italics="true" pageId="4" pageNumber="222">Venation</emphasis>
: forewing with four radial veins originating in the last third of the discal cell; subcostal vein clearly separated and parallel to R
<subScript id="A13B502E59344C08FC31491344BDFC9D" attach="left" box="[959,967,782,796]" fontSize="6" pageId="4" pageNumber="222">1</subScript>
. Vein R
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reaches the margin of the wing at the apex; 3A of hindwing slightly truncated, reaching the inner margin closer to the base (
<figureCitation id="A5844EEE59344C08FB1149024399FCB8" box="[1183,1251,799,825]" captionStart="FIGURE 3" captionStartId="5.[151,250,1943,1967]" captionTargetBox="[271,1327,201,1904]" captionTargetId="figure-29@5.[260,1327,181,1919]" captionTargetPageId="5" captionText="FIGURE 3. Wing of Emesis planeca n. comb. FW, forewing; HW, hindwing. Vein abbreviations (black lettering): Sc subcostal, R radial, M median, Cu cubital, A anal.Arrows in dorsal view point at: band of six white spots in the postmedial region and; line of small black dots (six dots in forewing, five dots in hindwing) in the submarginal area." figureDoi="http://doi.org/10.5281/zenodo.4410595" httpUri="https://zenodo.org/record/4410595/files/figure.png" pageId="4" pageNumber="222">Fig. 3</figureCitation>
).
</paragraph>
<paragraph id="3D00526B59344C08FF49495E43A2FB90" blockId="4.[151,1437,331,1942]" pageId="4" pageNumber="222">
<emphasis id="0FCB8E7959344C08FF49495E467EFCDC" box="[199,260,835,861]" italics="true" pageId="4" pageNumber="222">Head</emphasis>
(
<figureCitation id="A5844EEE59344C08FE9D495E4622FCDC" box="[275,344,835,861]" captionStart="FIGURE 4" captionStartId="6.[151,250,1284,1308]" captionTargetBox="[152,1436,184,1260]" captionTargetId="figure-22@6.[152,1436,181,1260]" captionTargetPageId="6" captionText="FIGURE 4. Male labial palpus and latero - ventral view of the head of Emesis planeca n. comb." figureDoi="http://doi.org/10.5281/zenodo.4410597" httpUri="https://zenodo.org/record/4410597/files/figure.png" pageId="4" pageNumber="222">Fig. 4</figureCitation>
). Eyes bare, with orange scales at the edge, the frons orange. Labial palpi small, close to the head and directed upward, hidden in dorsal view. The length of the second segment of labial palpi is at least two and a half the length of the third segment. Labial palpi white with a few orange scales on the external side of the third segment. Antennae almost two thirds the length of the wing, ringed, dark brown flagellomeres with white scales at their bases. Antennal club formed by the last 15 flagellomeres, which are white in ventral view and in some cases dark orange. Flagellomeres are mostly dark brown in dorsal view, but apical segments are orange to brownish.
</paragraph>
<paragraph id="3D00526B59344C08FF494E0643F6F988" blockId="4.[151,1437,331,1942]" pageId="4" pageNumber="222">
<emphasis id="0FCB8E7959344C08FF494E06464BFBB4" box="[199,305,1051,1077]" italics="true" pageId="4" pageNumber="222">Genitalia</emphasis>
(
<figureCitation id="A5844EEE59344C08FECC4E0646FDFBB7" box="[322,391,1051,1078]" captionStart="FIGURE 5" captionStartId="7.[151,250,1880,1904]" captionTargetBox="[157,1426,1001,1847]" captionTargetId="figure-447@7.[151,1436,993,1856]" captionTargetPageId="7" captionText="FIGURE 5. Male genitalia of Emesis planeca n. comb. (MZFC LEP 429988). Arrows in lateral view point at: two types of cornuti and; dorsal process of the valvae with many setae. In ventral view this process looks squared, wide and projected outwards." figureDoi="http://doi.org/10.5281/zenodo.4410599" httpUri="https://zenodo.org/record/4410599/files/figure.png" pageId="4" pageNumber="222">Fig. 5</figureCitation>
). Uncus wide with squared margin and a clear notch in the posterior margin in lateral view, with large setae. In ventral view the edge appears slightly concave in the middle with two knobs at the center, differing from the small cleavage with sharp edges described by
<bibRefCitation id="592E2F9A59344C08FCC04E7943B0FBFC" author="De la Maza, E. R. &amp; De la Maza, R. J." box="[846,1226,1123,1150]" pageId="4" pageNumber="222" pagination="31 - 36" refId="ref7729" refString="De la Maza, E. R. &amp; De la Maza, R. J. (2017) Una nueva especie de Apodemia C. y R. Felder, de la cuenca inferior del Rio Balsas, Michoacan, Mexico (Lepidoptera-Riodinidae). Revista de la Sociedad Mexicana de Lepidopterologia, Nueva Serie, 4, 31 - 36." type="journal article" year="2017">De la Maza &amp; De la Maza (2017)</bibRefCitation>
. Gnathi are sickleshaped, narrower towards the apex, distal ends in a sharp tip pointing laterally, rounder in ventral view as in the original description. In ventral view, gnathi may or may not converge toward the middle. Tegumen wide, curved in the anterior margin and with weakly sclerotized areas appearing as membranes which
<bibRefCitation id="592E2F9A59344C08FB0E4ED24279FB68" author="Hall, J. P. W." box="[1152,1283,1231,1257]" pageId="4" pageNumber="222" pagination="144 - 158" refId="ref8090" refString="Hall, J. P. W. (2008) Theope revisited: a synopsis of new discoveries, with the description of three new species (Lepidoptera: Riodinidae: Nymphidiini). Proceeding of the Entomology Society of Washington, 110 (1), 144 - 158. https: // doi. org / 10.4289 / 0013 - 8797 - 110.1.144" type="journal article" year="2008">Hall (2008)</bibRefCitation>
named “windows”. Vinculum narrow and straight, sometimes slightly inclining, with a curve right before turning into the saccus, which is strongly projected anteriorly, swollen in the middle portion and the posterior margin. Valvae divided in a ventral and a dorsal process, the dorsal process larger and straight and the ventral with rounded tips; both with conspicuous setae. Dorsal process is shorter than gnathi in lateral view. In ventral view, the dorsal process looks squared, wide and projected outwards; the ventral process is rounded, short, and turned inwards. Aedeagus is large, sigmoid shaped, curved before the genital capsule, with two
<typeStatus id="E204ECC959344C08FCBB4FB5440AFA43" box="[821,880,1448,1474]" pageId="4" pageNumber="222">types</typeStatus>
of cornuti, one
<typeStatus id="E204ECC959344C08FB914FB54335FA43" box="[1055,1103,1448,1474]" pageId="4" pageNumber="222">type</typeStatus>
is laminar, larger, sclerotized, and in an anterior position in relation to the second
<typeStatus id="E204ECC959344C08FD524FD14476FA67" box="[732,780,1484,1510]" pageId="4" pageNumber="222">type</typeStatus>
which is a series of small indented evenly distributed cornuti. Aedeagus joined to the capsule by a broad and strongly sclerotized strap-like pedicel.
</paragraph>
<paragraph id="3D00526B59344C08FF494C0E44B3F918" blockId="4.[151,1437,331,1942]" pageId="4" pageNumber="222">
<emphasis id="0FCB8E7959344C08FF494C0E465FF9AC" bold="true" box="[199,293,1555,1582]" pageId="4" pageNumber="222">
<emphasis id="0FCB8E7959344C08FF494C0E4664F9AF" bold="true" box="[199,286,1555,1582]" italics="true" pageId="4" pageNumber="222">Female</emphasis>
.
</emphasis>
(
<figureCitation id="A5844EEE59344C08FEBA4C0E460CF9AC" box="[308,374,1555,1581]" captionStart="FIGURE 1" captionStartId="1.[151,250,1942,1966]" captionTargetBox="[159,1428,1252,1912]" captionTargetId="figure-460@1.[151,1436,1242,1916]" captionTargetPageId="1" captionText="FIGURE 1. Emesis planeca n. comb. Dorsal (left) and ventral (right) views. AF males, GL females. México: Michoacán: Múgica, El Marqués, 500 m elev. leg. L. L. González Cota. Collection catalog numbers (MZFC LEP):A, 429994; B, 429990; C, 429991; D, 429992; E, 429993; F, 429988; G, 429998; H, 429995; I, 429999; J, 429997; K, 429996; L 354625." figureDoi="http://doi.org/10.5281/zenodo.4410591" httpUri="https://zenodo.org/record/4410591/files/figure.png" pageId="4" pageNumber="222">Fig. 1</figureCitation>
GL). Slightly larger than males, but smaller than in the original description, wingspan
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(n=7). Forewing length: a) wing base to anterior apex
<quantity id="FA47FF8E59344C08FCAE4C2A44ABF9D3" box="[800,977,1591,1618]" metricMagnitude="-2" metricUnit="m" metricValue="1.433" metricValueMax="1.55" metricValueMin="1.316" pageId="4" pageNumber="222" unit="mm" value="14.33" valueMax="15.5" valueMin="13.16">13.1615.5 mm</quantity>
(n=7, ẍ = 14.17 ±10.2); b) anterior apex to anal angle
<quantity id="FA47FF8E59344C08FEBF4C4646AFF9F7" box="[305,469,1627,1654]" metricMagnitude="-3" metricUnit="m" metricValue="9.58" metricValueMax="10.56" metricValueMin="8.6" pageId="4" pageNumber="222" unit="mm" value="9.58" valueMax="10.56" valueMin="8.6">8.610.56 mm</quantity>
(n=7, ẍ =9.46 ±0.9) and, c) anal angle to wing base
<quantity id="FA47FF8E59344C08FBA14C4643C4F9F7" box="[1071,1214,1627,1654]" metricMagnitude="-2" metricUnit="m" metricValue="1.0125" metricValueMax="1.125" metricValueMin="0.9" pageId="4" pageNumber="222" unit="mm" value="10.125" valueMax="11.25" valueMin="9.0">911.25 mm</quantity>
(n=7, ẍ =10 ±0.7).
<emphasis id="0FCB8E7959344C08FF194C9D4658F918" box="[151,290,1663,1689]" italics="true" pageId="4" pageNumber="222">Wing shape:</emphasis>
forewing more convex in the distal margin than in the male.
</paragraph>
<paragraph id="3D00526B59344C08FF494CBE461DF814" blockId="4.[151,1437,331,1942]" pageId="4" pageNumber="222">
<emphasis id="0FCB8E7959344C08FF494CBE464BF93C" box="[199,305,1699,1725]" italics="true" pageId="4" pageNumber="222">Genitalia</emphasis>
(
<figureCitation id="A5844EEE59344C08FECA4CBE46F1F93F" box="[324,395,1699,1726]" captionStart="FIGURE 6" captionStartId="8.[151,250,1944,1968]" captionTargetBox="[194,1395,181,1916]" captionTargetId="figure-22@8.[193,1395,181,1920]" captionTargetPageId="8" captionText="FIGURE 6. Female genitalia of Emesis planeca n. comb. (MZFC LEP 429998). Arrows in ventral and lateral views point at: antrum (wide, strongly sclerotized, and funnel-shaped) and; the single long invaginated signum with numerous tiny spines." figureDoi="http://doi.org/10.5281/zenodo.4410601" httpUri="https://zenodo.org/record/4410601/files/figure.png" pageId="4" pageNumber="222">Fig. 6</figureCitation>
). Anal papillae are rounded. With a membranous sterigma. Ostium bursae heptagonal not sclerotized in the dorsal and ventral regions, slightly sclerotized in the lateral edges. Ostium bursae joined the antrum which is wide and strongly sclerotized and funnel-shaped, smooth anteriorly and wrinkled posteriorly. Ductus bursae are membranous, slender, and straight, slightly longer than corpus bursae. Corpus bursae elongated and ovalshaped, with a single long invaginated signum forming a slightly inward curved sharp blade, placed at the middle and laterally in the corpus bursae. The signum has numerous tiny spines posteriorly directed on the ovoid opening to the corpus wall.
</paragraph>
<caption id="69C002E359354C09FF194D8A436DF876" ID-DOI="http://doi.org/10.5281/zenodo.4410595" ID-Zenodo-Dep="4410595" httpUri="https://zenodo.org/record/4410595/files/figure.png" pageId="5" pageNumber="223" startId="5.[151,250,1943,1967]" targetBox="[271,1327,201,1904]" targetPageId="5">
<paragraph id="3D00526B59354C09FF194D8A436DF876" blockId="5.[151,1436,1943,2040]" pageId="5" pageNumber="223">
<emphasis id="0FCB8E7959354C09FF194D8A4668F82E" bold="true" box="[151,274,1943,1967]" pageId="5" pageNumber="223">FIGURE 3.</emphasis>
Wing of
<taxonomicName id="FABF29E859354C09FEE14D854576F831" authorityName="Trujano-Ortega &amp; Callaghan &amp; García-Vázquez &amp; Luis-Martínez &amp; Ávalos-Hernández &amp; Llorente-Bousquets" authorityYear="2020" baseAuthorityName="Trujano-Ortega &amp; Callaghan &amp; García-Vázquez &amp; Luis-Martínez &amp; Ávalos-Hernández &amp; Llorente-Bousquets" baseAuthorityYear="2020" box="[367,524,1944,1968]" class="Insecta" family="Lycaenidae" genus="Emesis" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="223" phylum="Arthropoda" rank="species" species="planeca" status="comb. nov.">
<emphasis id="0FCB8E7959354C09FEE14D854576F831" box="[367,524,1944,1968]" italics="true" pageId="5" pageNumber="223">Emesis planeca</emphasis>
</taxonomicName>
<emphasis id="0FCB8E7959354C09FD9C4D854514F831" bold="true" box="[530,622,1944,1968]" pageId="5" pageNumber="223">
<taxonomicNameLabel id="14F8330259354C09FD9C4D854514F831" box="[530,622,1944,1968]" pageId="5" pageNumber="223" rank="species">n. comb.</taxonomicNameLabel>
</emphasis>
FW, forewing; HW, hindwing. Vein abbreviations (black lettering): Sc subcostal, R radial, M median, Cu cubital, A anal. Arrows in dorsal view point at: band of six white spots in the postmedial region and; line of small black dots (six dots in forewing, five dots in hindwing) in the submarginal area.
</paragraph>
</caption>
<caption id="69C002E359364C0AFF194F194316FA9D" ID-DOI="http://doi.org/10.5281/zenodo.4410597" ID-Zenodo-Dep="4410597" box="[151,1132,1284,1308]" httpUri="https://zenodo.org/record/4410597/files/figure.png" pageId="6" pageNumber="224" startId="6.[151,250,1284,1308]" targetBox="[152,1436,184,1260]" targetPageId="6">
<paragraph id="3D00526B59364C0AFF194F194316FA9D" blockId="6.[151,1132,1284,1308]" box="[151,1132,1284,1308]" pageId="6" pageNumber="224">
<emphasis id="0FCB8E7959364C0AFF194F194669FA9D" bold="true" box="[151,275,1284,1308]" pageId="6" pageNumber="224">FIGURE 4.</emphasis>
Male labial palpus and latero - ventral view of the head of
<taxonomicName id="FABF29E859364C0AFCE54F184373FA9D" authorityName="Trujano-Ortega &amp; Callaghan &amp; García-Vázquez &amp; Luis-Martínez &amp; Ávalos-Hernández &amp; Llorente-Bousquets" authorityYear="2020" baseAuthorityName="Trujano-Ortega &amp; Callaghan &amp; García-Vázquez &amp; Luis-Martínez &amp; Ávalos-Hernández &amp; Llorente-Bousquets" baseAuthorityYear="2020" box="[875,1033,1284,1308]" class="Insecta" family="Lycaenidae" genus="Emesis" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="224" phylum="Arthropoda" rank="species" species="planeca" status="comb. nov.">
<emphasis id="0FCB8E7959364C0AFCE54F184373FA9D" box="[875,1033,1284,1308]" italics="true" pageId="6" pageNumber="224">Emesis planeca</emphasis>
</taxonomicName>
<taxonomicNameLabel id="14F8330259364C0AFB9E4F194316FA9D" box="[1040,1132,1284,1308]" pageId="6" pageNumber="224" rank="species">n. comb.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="3D00526B59364C0AFF494F574330F824" blockId="6.[151,1437,1354,2035]" pageId="6" pageNumber="224">
<emphasis id="0FCB8E7959364C0AFF494F5746A3FAE5" bold="true" box="[199,473,1354,1381]" pageId="6" pageNumber="224">Wing pattern variation</emphasis>
(
<figureCitation id="A5844EEE59364C0AFE694F574549FAE4" box="[487,563,1354,1381]" captionStart="FIGURE 1" captionStartId="1.[151,250,1942,1966]" captionTargetBox="[159,1428,1252,1912]" captionTargetId="figure-460@1.[151,1436,1242,1916]" captionTargetPageId="1" captionText="FIGURE 1. Emesis planeca n. comb. Dorsal (left) and ventral (right) views. AF males, GL females. México: Michoacán: Múgica, El Marqués, 500 m elev. leg. L. L. González Cota. Collection catalog numbers (MZFC LEP):A, 429994; B, 429990; C, 429991; D, 429992; E, 429993; F, 429988; G, 429998; H, 429995; I, 429999; J, 429997; K, 429996; L 354625." figureDoi="http://doi.org/10.5281/zenodo.4410591" httpUri="https://zenodo.org/record/4410591/files/figure.png" pageId="6" pageNumber="224">Figs. 1</figureCitation>
,
<figureCitation id="A5844EEE59364C0AFDCE4F56452AFAE4" box="[576,592,1355,1381]" captionStart="FIGURE 3" captionStartId="5.[151,250,1943,1967]" captionTargetBox="[271,1327,201,1904]" captionTargetId="figure-29@5.[260,1327,181,1919]" captionTargetPageId="5" captionText="FIGURE 3. Wing of Emesis planeca n. comb. FW, forewing; HW, hindwing. Vein abbreviations (black lettering): Sc subcostal, R radial, M median, Cu cubital, A anal.Arrows in dorsal view point at: band of six white spots in the postmedial region and; line of small black dots (six dots in forewing, five dots in hindwing) in the submarginal area." figureDoi="http://doi.org/10.5281/zenodo.4410595" httpUri="https://zenodo.org/record/4410595/files/figure.png" pageId="6" pageNumber="224">3</figureCitation>
).
<taxonomicName id="FABF29E859364C0AFDEA4F564468FAE5" authorityName="Trujano-Ortega &amp; Callaghan &amp; García-Vázquez &amp; Luis-Martínez &amp; Ávalos-Hernández &amp; Llorente-Bousquets" authorityYear="2020" baseAuthorityName="Trujano-Ortega &amp; Callaghan &amp; García-Vázquez &amp; Luis-Martínez &amp; Ávalos-Hernández &amp; Llorente-Bousquets" baseAuthorityYear="2020" box="[612,786,1354,1380]" class="Insecta" family="Lycaenidae" genus="Emesis" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="224" phylum="Arthropoda" rank="species" species="planeca" status="comb. nov.">
<emphasis id="0FCB8E7959364C0AFDEA4F564468FAE5" box="[612,786,1354,1380]" italics="true" pageId="6" pageNumber="224">Emesis planeca</emphasis>
</taxonomicName>
<emphasis id="0FCB8E7959364C0AFC964F564406FAE4" bold="true" box="[792,892,1355,1381]" pageId="6" pageNumber="224">
<taxonomicNameLabel id="14F8330259364C0AFC964F564406FAE4" box="[792,892,1355,1381]" pageId="6" pageNumber="224" rank="species">n. comb.</taxonomicNameLabel>
</emphasis>
does not show evident sexual dimorphism. Wing pattern in both wings and views are consistent, with little variation listed below:
<emphasis id="0FCB8E7959364C0AFB804F7343E0FA09" box="[1038,1178,1390,1416]" italics="true" pageId="6" pageNumber="224">Dorsal view.</emphasis>
Forewing; background color is originally described as dull cinnamon brown (p. 31), as in MZFC 429990 (
<figureCitation id="A5844EEE59364C0AFBCF4F8F43E4FA2C" box="[1089,1182,1426,1453]" captionStart="FIGURE 1" captionStartId="1.[151,250,1942,1966]" captionTargetBox="[159,1428,1252,1912]" captionTargetId="figure-460@1.[151,1436,1242,1916]" captionTargetPageId="1" captionText="FIGURE 1. Emesis planeca n. comb. Dorsal (left) and ventral (right) views. AF males, GL females. México: Michoacán: Múgica, El Marqués, 500 m elev. leg. L. L. González Cota. Collection catalog numbers (MZFC LEP):A, 429994; B, 429990; C, 429991; D, 429992; E, 429993; F, 429988; G, 429998; H, 429995; I, 429999; J, 429997; K, 429996; L 354625." figureDoi="http://doi.org/10.5281/zenodo.4410591" httpUri="https://zenodo.org/record/4410591/files/figure.png" pageId="6" pageNumber="224">Fig. 1 B</figureCitation>
), however, most of the examined specimens present a light ochraceous tawny (15.Y.O.i; in
<bibRefCitation id="592E2F9A59364C0AFC364FAB4325FA50" author="Ridgway, R." box="[952,1119,1462,1489]" pageId="6" pageNumber="224" refId="ref9733" refString="Ridgway, R. (1912) Color standards and color nomenclature. Published by the author, Washington, D. C., 43 pp. https: // doi. org / 10.5962 / bhl. title. 62375" type="book" year="1912">Ridgway 1912</bibRefCitation>
) background. The forewing fringe is bronze, with white scales only in cell R
<subScript id="A13B502E59364C0AFD354FF045AFFA7A" attach="left" box="[699,725,1517,1531]" fontSize="6" pageId="6" pageNumber="224">4+5</subScript>
at the apex. Discal cell with four lines of faint dark transversal scales, only the fourth line at the distal end of the discal cell is clearly visible and straight, with the same four lines in Cu
<subScript id="A13B502E59364C0AFF5A4C2847A6F9C2" attach="left" box="[212,220,1589,1603]" fontSize="6" pageId="6" pageNumber="224">2</subScript>
, where the fourth line aligns with the fourth line of the discal cell. The post-medial area presents an irregular median row of white spots with dark brown scales in the basal edge, as described by
<bibRefCitation id="592E2F9A59364C0AFBE54C5A479FF904" author="De la Maza, E. R. &amp; De la Maza, R. J." pageId="6" pageNumber="224" pagination="31 - 36" refId="ref7729" refString="De la Maza, E. R. &amp; De la Maza, R. J. (2017) Una nueva especie de Apodemia C. y R. Felder, de la cuenca inferior del Rio Balsas, Michoacan, Mexico (Lepidoptera-Riodinidae). Revista de la Sociedad Mexicana de Lepidopterologia, Nueva Serie, 4, 31 - 36." type="journal article" year="2017">De la Maza &amp; De la Maza (2017)</bibRefCitation>
. The row was originally described as reaching the posterior margin of M
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(p. 32), but all examined specimens show six spots as far as Cu
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(
<figureCitation id="A5844EEE59364C0AFE574C934564F928" box="[473,542,1678,1705]" captionStart="FIGURE 3" captionStartId="5.[151,250,1943,1967]" captionTargetBox="[271,1327,201,1904]" captionTargetId="figure-29@5.[260,1327,181,1919]" captionTargetPageId="5" captionText="FIGURE 3. Wing of Emesis planeca n. comb. FW, forewing; HW, hindwing. Vein abbreviations (black lettering): Sc subcostal, R radial, M median, Cu cubital, A anal.Arrows in dorsal view point at: band of six white spots in the postmedial region and; line of small black dots (six dots in forewing, five dots in hindwing) in the submarginal area." figureDoi="http://doi.org/10.5281/zenodo.4410595" httpUri="https://zenodo.org/record/4410595/files/figure.png" pageId="6" pageNumber="224">Fig. 3</figureCitation>
). Since this is a distinctive character of the species, it will be described in detail (
<figureCitation id="A5844EEE59364C0AFF2E4CAF4799F94C" box="[160,227,1714,1741]" captionStart="FIGURE 3" captionStartId="5.[151,250,1943,1967]" captionTargetBox="[271,1327,201,1904]" captionTargetId="figure-29@5.[260,1327,181,1919]" captionTargetPageId="5" captionText="FIGURE 3. Wing of Emesis planeca n. comb. FW, forewing; HW, hindwing. Vein abbreviations (black lettering): Sc subcostal, R radial, M median, Cu cubital, A anal.Arrows in dorsal view point at: band of six white spots in the postmedial region and; line of small black dots (six dots in forewing, five dots in hindwing) in the submarginal area." figureDoi="http://doi.org/10.5281/zenodo.4410595" httpUri="https://zenodo.org/record/4410595/files/figure.png" pageId="6" pageNumber="224">Fig. 3</figureCitation>
): first two spots are triangular, small, sharp and evident (cells R
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y R
<subScript id="A13B502E59364C0AFC504CD84482F952" attach="left" box="[990,1016,1733,1747]" fontSize="6" pageId="6" pageNumber="224">4+5</subScript>
); third and fourth spots (M
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y M
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) are large and visible, although barely visible in some specimens (
<figureCitation id="A5844EEE59364C0AFCCB4CCB44AAF970" box="[837,976,1750,1777]" captionStart="FIGURE 1" captionStartId="1.[151,250,1942,1966]" captionTargetBox="[159,1428,1252,1912]" captionTargetId="figure-460@1.[151,1436,1242,1916]" captionTargetPageId="1" captionText="FIGURE 1. Emesis planeca n. comb. Dorsal (left) and ventral (right) views. AF males, GL females. México: Michoacán: Múgica, El Marqués, 500 m elev. leg. L. L. González Cota. Collection catalog numbers (MZFC LEP):A, 429994; B, 429990; C, 429991; D, 429992; E, 429993; F, 429988; G, 429998; H, 429995; I, 429999; J, 429997; K, 429996; L 354625." figureDoi="http://doi.org/10.5281/zenodo.4410591" httpUri="https://zenodo.org/record/4410591/files/figure.png" pageId="6" pageNumber="224">Fig. 1 B y L</figureCitation>
); the fourth spot is larger and close to the distal margin; fifth and sixth spots (M
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y Cu
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) are generally small and faint, and in some specimens only as scattered white scales (
<figureCitation id="A5844EEE59364C0AFEA34D0346B6F8B8" box="[301,460,1822,1849]" captionStart="FIGURE 1" captionStartId="1.[151,250,1942,1966]" captionTargetBox="[159,1428,1252,1912]" captionTargetId="figure-460@1.[151,1436,1242,1916]" captionTargetPageId="1" captionText="FIGURE 1. Emesis planeca n. comb. Dorsal (left) and ventral (right) views. AF males, GL females. México: Michoacán: Múgica, El Marqués, 500 m elev. leg. L. L. González Cota. Collection catalog numbers (MZFC LEP):A, 429994; B, 429990; C, 429991; D, 429992; E, 429993; F, 429988; G, 429998; H, 429995; I, 429999; J, 429997; K, 429996; L 354625." figureDoi="http://doi.org/10.5281/zenodo.4410591" httpUri="https://zenodo.org/record/4410591/files/figure.png" pageId="6" pageNumber="224">Fig. 1 B, H, L</figureCitation>
); white spot in Cu
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is close to the wing base. Marginal and distal post-medial areas with an orange band, limited in the basal edge by an irregular line of brown-reddish scales in the post-medial area, from R
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to Cu
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. Submarginal area with a line of six small black dots sometimes faint at the apex or in Cu
<subScript id="A13B502E59364C0AFB6C4D644390F806" attach="left" box="[1250,1258,1913,1927]" fontSize="6" pageId="6" pageNumber="224">2</subScript>
, in this case are barely visible as reddish scales (
<figureCitation id="A5844EEE59364C0AFD8E4D9745EEF824" box="[512,660,1930,1957]" captionStart="FIGURE 1" captionStartId="1.[151,250,1942,1966]" captionTargetBox="[159,1428,1252,1912]" captionTargetId="figure-460@1.[151,1436,1242,1916]" captionTargetPageId="1" captionText="FIGURE 1. Emesis planeca n. comb. Dorsal (left) and ventral (right) views. AF males, GL females. México: Michoacán: Múgica, El Marqués, 500 m elev. leg. L. L. González Cota. Collection catalog numbers (MZFC LEP):A, 429994; B, 429990; C, 429991; D, 429992; E, 429993; F, 429988; G, 429998; H, 429995; I, 429999; J, 429997; K, 429996; L 354625." figureDoi="http://doi.org/10.5281/zenodo.4410591" httpUri="https://zenodo.org/record/4410591/files/figure.png" pageId="6" pageNumber="224">Fig. 1 F, H, J</figureCitation>
). Veins marked along the wing in grey.
</paragraph>
<paragraph id="3D00526B59364C0BFF494DB3442DFE9C" blockId="6.[151,1437,1354,2035]" lastBlockId="7.[151,1437,151,969]" lastPageId="7" lastPageNumber="225" pageId="6" pageNumber="224">
Hindwing; with the same pattern as forewing.
<bibRefCitation id="592E2F9A59364C0AFD504DB2431EF848" author="De la Maza, E. R. &amp; De la Maza, R. J." box="[734,1124,1966,1993]" pageId="6" pageNumber="224" pagination="31 - 36" refId="ref7729" refString="De la Maza, E. R. &amp; De la Maza, R. J. (2017) Una nueva especie de Apodemia C. y R. Felder, de la cuenca inferior del Rio Balsas, Michoacan, Mexico (Lepidoptera-Riodinidae). Revista de la Sociedad Mexicana de Lepidopterologia, Nueva Serie, 4, 31 - 36." type="journal article" year="2017">De la Maza &amp; De la Maza (2017)</bibRefCitation>
mention seven white spots reaching Cu
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Cu
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; however, specimens examined show only six spots as described for forewing, although less defined in hindwing (
<figureCitation id="A5844EEE59374C0BFEFE4A8A46C5FF30" box="[368,447,151,177]" captionStart="FIGURE 1" captionStartId="1.[151,250,1942,1966]" captionTargetBox="[159,1428,1252,1912]" captionTargetId="figure-460@1.[151,1436,1242,1916]" captionTargetPageId="1" captionText="FIGURE 1. Emesis planeca n. comb. Dorsal (left) and ventral (right) views. AF males, GL females. México: Michoacán: Múgica, El Marqués, 500 m elev. leg. L. L. González Cota. Collection catalog numbers (MZFC LEP):A, 429994; B, 429990; C, 429991; D, 429992; E, 429993; F, 429988; G, 429998; H, 429995; I, 429999; J, 429997; K, 429996; L 354625." figureDoi="http://doi.org/10.5281/zenodo.4410591" httpUri="https://zenodo.org/record/4410591/files/figure.png" pageId="7" pageNumber="225">Figs. 1</figureCitation>
,
<figureCitation id="A5844EEE59374C0BFE404A8A46A4FF30" box="[462,478,151,177]" captionStart="FIGURE 3" captionStartId="5.[151,250,1943,1967]" captionTargetBox="[271,1327,201,1904]" captionTargetId="figure-29@5.[260,1327,181,1919]" captionTargetPageId="5" captionText="FIGURE 3. Wing of Emesis planeca n. comb. FW, forewing; HW, hindwing. Vein abbreviations (black lettering): Sc subcostal, R radial, M median, Cu cubital, A anal.Arrows in dorsal view point at: band of six white spots in the postmedial region and; line of small black dots (six dots in forewing, five dots in hindwing) in the submarginal area." figureDoi="http://doi.org/10.5281/zenodo.4410595" httpUri="https://zenodo.org/record/4410595/files/figure.png" pageId="7" pageNumber="225">3</figureCitation>
). Postmedial area with a band of reddish scales more evident than in forewing, from the apex to the tornus; sometimes not clearly defined (
<figureCitation id="A5844EEE59374C0BFD774AA64422FF54" box="[761,856,187,213]" captionStart="FIGURE 1" captionStartId="1.[151,250,1942,1966]" captionTargetBox="[159,1428,1252,1912]" captionTargetId="figure-460@1.[151,1436,1242,1916]" captionTargetPageId="1" captionText="FIGURE 1. Emesis planeca n. comb. Dorsal (left) and ventral (right) views. AF males, GL females. México: Michoacán: Múgica, El Marqués, 500 m elev. leg. L. L. González Cota. Collection catalog numbers (MZFC LEP):A, 429994; B, 429990; C, 429991; D, 429992; E, 429993; F, 429988; G, 429998; H, 429995; I, 429999; J, 429997; K, 429996; L 354625." figureDoi="http://doi.org/10.5281/zenodo.4410591" httpUri="https://zenodo.org/record/4410591/files/figure.png" pageId="7" pageNumber="225">Fig. 1 D</figureCitation>
) or with lunulae (
<figureCitation id="A5844EEE59374C0BFBAC4AA643BEFF54" box="[1058,1220,187,213]" captionStart="FIGURE 1" captionStartId="1.[151,250,1942,1966]" captionTargetBox="[159,1428,1252,1912]" captionTargetId="figure-460@1.[151,1436,1242,1916]" captionTargetPageId="1" captionText="FIGURE 1. Emesis planeca n. comb. Dorsal (left) and ventral (right) views. AF males, GL females. México: Michoacán: Múgica, El Marqués, 500 m elev. leg. L. L. González Cota. Collection catalog numbers (MZFC LEP):A, 429994; B, 429990; C, 429991; D, 429992; E, 429993; F, 429988; G, 429998; H, 429995; I, 429999; J, 429997; K, 429996; L 354625." figureDoi="http://doi.org/10.5281/zenodo.4410591" httpUri="https://zenodo.org/record/4410591/files/figure.png" pageId="7" pageNumber="225">Fig. 1 B, G, L</figureCitation>
). Submarginal area with a line of five small black dots clearly defined, more evident than in forewing (
<figureCitation id="A5844EEE59374C0BFBCF4AC243FCFF78" box="[1089,1158,223,249]" captionStart="FIGURE 3" captionStartId="5.[151,250,1943,1967]" captionTargetBox="[271,1327,201,1904]" captionTargetId="figure-29@5.[260,1327,181,1919]" captionTargetPageId="5" captionText="FIGURE 3. Wing of Emesis planeca n. comb. FW, forewing; HW, hindwing. Vein abbreviations (black lettering): Sc subcostal, R radial, M median, Cu cubital, A anal.Arrows in dorsal view point at: band of six white spots in the postmedial region and; line of small black dots (six dots in forewing, five dots in hindwing) in the submarginal area." figureDoi="http://doi.org/10.5281/zenodo.4410595" httpUri="https://zenodo.org/record/4410595/files/figure.png" pageId="7" pageNumber="225">Fig. 3</figureCitation>
). The hindwing fringe is entirely checkered bronze and white from distal margin to Cu
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.
</paragraph>
<paragraph id="3D00526B59374C0BFF494B3A4541FE50" blockId="7.[151,1437,151,969]" pageId="7" pageNumber="225">
<emphasis id="0FCB8E7959374C0BFF494B3A4629FEC1" box="[199,339,295,321]" italics="true" pageId="7" pageNumber="225">Ventral view</emphasis>
. With more intense colors in the orange range ochraceous orange (13 O.Y.O) and cadmium orange (13.OY.o; in
<bibRefCitation id="592E2F9A59374C0BFEE54B564569FEE4" author="Ridgway, R." box="[363,531,331,357]" pageId="7" pageNumber="225" refId="ref9733" refString="Ridgway, R. (1912) Color standards and color nomenclature. Published by the author, Washington, D. C., 43 pp. https: // doi. org / 10.5962 / bhl. title. 62375" type="book" year="1912">Ridgway 1912</bibRefCitation>
), differing from the not-iridescent pearl salmon color described by
<bibRefCitation id="592E2F9A59374C0BFB714B564614FE08" author="De la Maza, E. R. &amp; De la Maza, R. J." pageId="7" pageNumber="225" pagination="31 - 36" refId="ref7729" refString="De la Maza, E. R. &amp; De la Maza, R. J. (2017) Una nueva especie de Apodemia C. y R. Felder, de la cuenca inferior del Rio Balsas, Michoacan, Mexico (Lepidoptera-Riodinidae). Revista de la Sociedad Mexicana de Lepidopterologia, Nueva Serie, 4, 31 - 36." type="journal article" year="2017">De la Maza &amp; De la Maza (2017)</bibRefCitation>
. White spots dimmed in dorsal view (as in M
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and Cu
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) are clearly visible in ventral view in all specimens (
<figureCitation id="A5844EEE59374C0BFE924B8E46E7FE2C" box="[284,413,403,429]" captionStart="FIGURE 1" captionStartId="1.[151,250,1942,1966]" captionTargetBox="[159,1428,1252,1912]" captionTargetId="figure-460@1.[151,1436,1242,1916]" captionTargetPageId="1" captionText="FIGURE 1. Emesis planeca n. comb. Dorsal (left) and ventral (right) views. AF males, GL females. México: Michoacán: Múgica, El Marqués, 500 m elev. leg. L. L. González Cota. Collection catalog numbers (MZFC LEP):A, 429994; B, 429990; C, 429991; D, 429992; E, 429993; F, 429988; G, 429998; H, 429995; I, 429999; J, 429997; K, 429996; L 354625." figureDoi="http://doi.org/10.5281/zenodo.4410591" httpUri="https://zenodo.org/record/4410591/files/figure.png" pageId="7" pageNumber="225">Fig. 1 B, L</figureCitation>
). Two evident black dots in Cu
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in
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both wings, therefore submarginal area has seven black dots in forewing and six in hindwing.
</paragraph>
<paragraph id="3D00526B59374C0BFF494BC74333FC48" blockId="7.[151,1437,151,969]" pageId="7" pageNumber="225">
<emphasis id="0FCB8E7959374C0BFF494BC746A3FE74" bold="true" box="[199,473,474,501]" pageId="7" pageNumber="225">Habitat and phenology</emphasis>
. All examined specimens are from the
<typeStatus id="E204ECC959374C0BFC004BC644C4FE74" box="[910,958,475,501]" pageId="7" pageNumber="225">type</typeStatus>
locality (
<bibRefCitation id="592E2F9A59374C0BFBA94BC642EAFE74" author="De la Maza, E. R. &amp; De la Maza, R. J." box="[1063,1424,475,501]" pageId="7" pageNumber="225" pagination="31 - 36" refId="ref7729" refString="De la Maza, E. R. &amp; De la Maza, R. J. (2017) Una nueva especie de Apodemia C. y R. Felder, de la cuenca inferior del Rio Balsas, Michoacan, Mexico (Lepidoptera-Riodinidae). Revista de la Sociedad Mexicana de Lepidopterologia, Nueva Serie, 4, 31 - 36." type="journal article" year="2017">De la Maza &amp; De la Maza 2017</bibRefCitation>
), El
<collectingRegion id="FF7B9C8959374C0BFF374BE24664FD98" box="[185,286,511,537]" country="Italy" name="Marche" pageId="7" pageNumber="225">Marqués</collectingRegion>
, municipality of Múgica, in the central state of
<collectingRegion id="FF7B9C8959374C0BFCC94BE244BDFD98" box="[839,967,511,537]" country="Mexico" name="Michoacan de Ocampo" pageId="7" pageNumber="225">Michoacán</collectingRegion>
. The site is
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a.s.l., forming part of the morphotectonic province of Balsas Basin (
<bibRefCitation id="592E2F9A59374C0BFD29483E4491FDBC" author="Ferrusquia-Villafranca, I." box="[679,1003,547,573]" pageId="7" pageNumber="225" pagination="3 - 107" refId="ref7987" refString="Ferrusquia-Villafranca, I. (1993) Geology of Mexico: a synopsis. In: Ramamoorthy, T. P., Bye, R., Lot, A. &amp; Fa, J. (Eds.), Biological diversity of Mexico: origins and distribution, Oxford University Press, New York, New York, pp. 3 - 107." type="book chapter" year="1993">Ferrusquía-Villafranca 1993</bibRefCitation>
). The dominant vegetation is Tropical Deciduous Forest, however, Tropical Subdeciduous Forest can be found along the margin of nearby rivers areas (
<bibRefCitation id="592E2F9A59374C0BFF1148764625FD04" author="Rzedowski, J." box="[159,351,619,645]" pageId="7" pageNumber="225" refId="ref9866" refString="Rzedowski, J. (1978) Vegetacion de Mexico. Editorial Limusa, Mexico, D. F., 432 pp." type="book" year="1978">Rzedowski 1978</bibRefCitation>
;
<bibRefCitation id="592E2F9A59374C0BFEE4487645EBFD04" author="Martinez-Cruz, J. &amp; Mendez-Toribio, M. &amp; Cortes-Flores, J. &amp; Coba-Perez, P. &amp; Cornejo-Tenorio, G. &amp; Ibarra-Manriquez, G." box="[362,657,619,645]" pageId="7" pageNumber="225" pagination="1216 - 1234" refId="ref9120" refString="Martinez-Cruz, J., Mendez-Toribio, M., Cortes-Flores, J., Coba-Perez, P., Cornejo-Tenorio, G. &amp; Ibarra-Manriquez, G. (2013) Estructura y diversidad de los bosques estacionales desaparecidos por la construccion de la presa Gral. Francisco J. Mugica, en la Depresion del Balsas, Michoacan, Mexico. Revista Mexicana de Biodiversidad, 84, 1216 - 1234. https: // doi. org / 10.7550 / rmb. 33799" type="journal article" year="2013">
Martínez-Cruz
<emphasis id="0FCB8E7959374C0BFD9B4876453CFD04" box="[533,582,619,645]" italics="true" pageId="7" pageNumber="225">et al</emphasis>
. 2013
</bibRefCitation>
). Dominant tree species are
<taxonomicName id="FABF29E859374C0BFC44487643C9FD04" box="[970,1203,619,645]" class="Insecta" family="Aphrophoridae" genus="Cordia" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="225" phylum="Arthropoda" rank="species" species="elaeagnoides">
<emphasis id="0FCB8E7959374C0BFC44487643C9FD04" box="[970,1203,619,645]" italics="true" pageId="7" pageNumber="225">Cordia elaeagnoides</emphasis>
</taxonomicName>
,
<taxonomicName id="FABF29E859374C0BFB4F48764797FD28" class="Magnoliopsida" family="Euphorbiaceae" genus="Euphorbia" kingdom="Plantae" order="Malpighiales" pageId="7" pageNumber="225" phylum="Tracheophyta" rank="species" species="schlechtendalii">
<emphasis id="0FCB8E7959374C0BFB4F48764797FD28" italics="true" pageId="7" pageNumber="225">Euphorbia schlechtendalii</emphasis>
</taxonomicName>
,
<taxonomicName id="FABF29E859374C0BFF75489246D5FD28" authorityName="S.Watson" authorityYear="1889" box="[251,431,655,681]" class="Aves" family="Bernieridae" genus="Randia" kingdom="Animalia" order="Passeriformes" pageId="7" pageNumber="225" phylum="Chordata" rank="species" species="thurberi">
<emphasis id="0FCB8E7959374C0BFF75489246D5FD28" box="[251,431,655,681]" italics="true" pageId="7" pageNumber="225">Randia thurberi</emphasis>
</taxonomicName>
,
<emphasis id="0FCB8E7959374C0BFE33489245C4FD28" box="[445,702,655,681]" italics="true" pageId="7" pageNumber="225">
<taxonomicName id="FABF29E859374C0BFE33489245C0FD28" authorityName="Standley ex Leavenworth" authorityYear="1946" box="[445,698,655,681]" class="Magnoliopsida" family="Euphorbiaceae" genus="Manihot" kingdom="Plantae" order="Malpighiales" pageId="7" pageNumber="225" phylum="Tracheophyta" rank="species" species="tomatophylla">Manihot tomatophylla</taxonomicName>
,
</emphasis>
and
<taxonomicName id="FABF29E859374C0BFD7B48924277FD28" authority="(Martinez-Cruz et al. 2013)" baseAuthorityName="Martinez-Cruz" baseAuthorityYear="2013" box="[757,1293,655,681]" class="Magnoliopsida" family="Fabaceae" genus="Acacia" kingdom="Plantae" order="Fabales" pageId="7" pageNumber="225" phylum="Tracheophyta" rank="species" species="picachensis">
<emphasis id="0FCB8E7959374C0BFD7B489244B7FD28" box="[757,973,655,681]" italics="true" pageId="7" pageNumber="225">Acacia picachensis</emphasis>
(
<bibRefCitation id="592E2F9A59374C0BFC554892427FFD28" author="Martinez-Cruz, J. &amp; Mendez-Toribio, M. &amp; Cortes-Flores, J. &amp; Coba-Perez, P. &amp; Cornejo-Tenorio, G. &amp; Ibarra-Manriquez, G." box="[987,1285,655,681]" pageId="7" pageNumber="225" pagination="1216 - 1234" refId="ref9120" refString="Martinez-Cruz, J., Mendez-Toribio, M., Cortes-Flores, J., Coba-Perez, P., Cornejo-Tenorio, G. &amp; Ibarra-Manriquez, G. (2013) Estructura y diversidad de los bosques estacionales desaparecidos por la construccion de la presa Gral. Francisco J. Mugica, en la Depresion del Balsas, Michoacan, Mexico. Revista Mexicana de Biodiversidad, 84, 1216 - 1234. https: // doi. org / 10.7550 / rmb. 33799" type="journal article" year="2013">
Martínez-Cruz
<emphasis id="0FCB8E7959374C0BFB07489243C0FD28" box="[1161,1210,655,681]" italics="true" pageId="7" pageNumber="225">et al</emphasis>
. 2013
</bibRefCitation>
)
</taxonomicName>
. The climate in the region is semiarid, warm, with summer rains (BS1(h) xigw”); annual median temperature is 26.5° C; total annual rainfall is
<quantity id="FA47FF8E59374C0BFEEE48CB46AEFD70" box="[352,468,726,753]" metricMagnitude="-1" metricUnit="m" metricValue="6.255" pageId="7" pageNumber="225" unit="mm" value="625.5">625.5 mm</quantity>
and the rainy season extends from June to September (
<bibRefCitation id="592E2F9A59374C0BFBC148CA4392FD70" author="Garcia, E." box="[1103,1256,727,753]" pageId="7" pageNumber="225" refId="ref8054" refString="Garcia, E. (2004) Modificaciones al sistema de clasificacion climatica de Koppen. Instituto de Geografia, Universidad Nacional Autonoma de Mexico. Mexico, D. F., 90 pp." type="book" year="2004">García, 2004</bibRefCitation>
). All specimens were collected by L. L. González Cota between March and April of 1997, with just one specimen from
<date id="490174AB59374C0BFAA748E647A8FCB8" pageId="7" pageNumber="225" value="1996-12">December 1996</date>
. A Mariposa database query (
<bibRefCitation id="592E2F9A59374C0BFDAC49024432FCB8" author="Luis-Martinez, A. &amp; Llorente-Bousquets, J. &amp; Vargas-Fernandez, I." box="[546,840,798,825]" pageId="7" pageNumber="225" pagination="269 - 294" refId="ref8770" refString="Luis-Martinez, A., Llorente-Bousquets, J. &amp; Vargas-Fernandez, I. (2005) Una megabase de datos de mariposas y la regionalizacion biogeografica de Mexico. In: Llorente-Bousquets, J. &amp; Morrone, J. J. (Eds.), Regionalizacion geografica en Iberoamerica y topicos afines: Primeras Jornadas Biogeograficas de la Red Iberoamericana de Biogeografia y Entomologia Sistematica (RIBES XII. X-CYTED). Universidad Nacional Autonoma de Mexico, Mexico, D. F., pp. 269 - 294." type="book chapter" year="2005">
Luis-Martínez
<emphasis id="0FCB8E7959374C0BFD4349024585FCB8" box="[717,767,799,825]" italics="true" pageId="7" pageNumber="225">et al</emphasis>
. 2005
</bibRefCitation>
) showed that the same collector captured specimens from other families in that site from
<date id="490174AB59374C0BFDA9495E45A7FCDC" box="[551,733,835,861]" pageId="7" pageNumber="225" value="1996-09">September 1996</date>
to
<date id="490174AB59374C0BFD71495E4409FCDC" box="[767,883,835,861]" pageId="7" pageNumber="225" value="1997-06">June 1997</date>
. Therefore, we infer that adults of
<taxonomicName id="FABF29E859374C0BFB60495E42E7FCDC" authorityName="Trujano-Ortega &amp; Callaghan &amp; García-Vázquez &amp; Luis-Martínez &amp; Ávalos-Hernández &amp; Llorente-Bousquets" authorityYear="2020" baseAuthorityName="Trujano-Ortega &amp; Callaghan &amp; García-Vázquez &amp; Luis-Martínez &amp; Ávalos-Hernández &amp; Llorente-Bousquets" baseAuthorityYear="2020" box="[1262,1437,835,861]" class="Insecta" family="Lycaenidae" genus="Emesis" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="225" phylum="Arthropoda" rank="species" species="planeca" status="comb. nov.">
<emphasis id="0FCB8E7959374C0BFB60495E42E7FCDC" box="[1262,1437,835,861]" italics="true" pageId="7" pageNumber="225">Emesis planeca</emphasis>
</taxonomicName>
<emphasis id="0FCB8E7959374C0BFF19497A4787FC00" bold="true" box="[151,253,871,897]" pageId="7" pageNumber="225">
<taxonomicNameLabel id="14F8330259374C0BFF19497A4787FC00" box="[151,253,871,897]" pageId="7" pageNumber="225" rank="species">n. comb.</taxonomicNameLabel>
</emphasis>
fly during the dry season of the year. It is sympatric with
<taxonomicName id="FABF29E859374C0BFC05497A4329FC00" authority="Godman" authorityName="Godman" box="[907,1107,871,897]" class="Insecta" family="Lycaenidae" genus="Emesis" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="225" phylum="Arthropoda" rank="species" species="poeas">
<emphasis id="0FCB8E7959374C0BFC05497A4491FC01" box="[907,1003,871,896]" italics="true" pageId="7" pageNumber="225">E. poeas</emphasis>
Godman
</taxonomicName>
and probably also with other species of
<taxonomicName id="FABF29E859374C0BFE8349964626FC25" authorityName="Fabricius" authorityYear="1807" box="[269,348,907,932]" class="Insecta" family="Lycaenidae" genus="Emesis" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="225" phylum="Arthropoda" rank="genus">
<emphasis id="0FCB8E7959374C0BFE8349964626FC25" box="[269,348,907,932]" italics="true" pageId="7" pageNumber="225">Emesis</emphasis>
</taxonomicName>
distributed in the state like
<taxonomicName id="FABF29E859374C0BFD1A499644F6FC24" authority="(Cramer)" baseAuthorityName="Cramer" box="[660,908,907,933]" class="Insecta" family="Lycaenidae" genus="Emesis" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="225" phylum="Arthropoda" rank="species" species="mandana">
<emphasis id="0FCB8E7959374C0BFD1A49964461FC24" box="[660,795,907,933]" italics="true" pageId="7" pageNumber="225">E. mandana</emphasis>
(Cramer)
</taxonomicName>
,
<taxonomicName id="FABF29E859374C0BFC1949964393FC24" authority="Godman &amp; Salvin" authorityName="Godman &amp; Salvin" authorityYear="1886" box="[919,1257,907,933]" class="Insecta" family="Lycaenidae" genus="Emesis" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="225" phylum="Arthropoda" rank="species" species="saturata">
<emphasis id="0FCB8E7959374C0BFC1949964369FC25" box="[919,1043,907,932]" italics="true" pageId="7" pageNumber="225">E. saturata</emphasis>
Godman &amp; Salvin
</taxonomicName>
,
<taxonomicName id="FABF29E859374C0BFB7A4996421FFC24" box="[1268,1381,907,933]" class="Insecta" family="Lycaenidae" genus="Emesis" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="225" phylum="Arthropoda" rank="species" species="tenedia">
<emphasis id="0FCB8E7959374C0BFB7A4996421FFC24" box="[1268,1381,907,933]" italics="true" pageId="7" pageNumber="225">E. tenedia</emphasis>
</taxonomicName>
, and
<taxonomicName id="FABF29E859374C0BFF1949B246A7FC48" authority="Godman &amp; Salvin" authorityName="Godman &amp; Salvin" authorityYear="1886" box="[151,477,943,969]" class="Insecta" family="Lycaenidae" genus="Emesis" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="225" phylum="Arthropoda" rank="species" species="vulpina">
<emphasis id="0FCB8E7959374C0BFF1949B24670FC48" box="[151,266,943,969]" italics="true" pageId="7" pageNumber="225">E. vulpina</emphasis>
Godman &amp; Salvin
</taxonomicName>
(
<figureCitation id="A5844EEE59374C0BFE6249B2454BFC48" box="[492,561,943,969]" captionStart="FIGURE 7" captionStartId="9.[151,250,1111,1135]" captionTargetBox="[154,1435,184,1084]" captionTargetId="figure-29@9.[151,1436,181,1087]" captionTargetPageId="9" captionText="FIGURE 7. Geographic distribution of Emesis planeca n. comb. and other Emesis species in Michoacán, Mexico. Black lines represent state limits." figureDoi="http://doi.org/10.5281/zenodo.4410603" httpUri="https://zenodo.org/record/4410603/files/figure.png" pageId="7" pageNumber="225">Fig. 7</figureCitation>
), but more sampling is required to confirm this.
</paragraph>
</subSubSection>
</treatment>
</document>