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<document id="BDDEF75926C0ECA3939C9C51169187B8" ID-DOI="10.5281/zenodo.177977" ID-GBIF-Dataset="68d70702-9696-4c34-8197-83f16ce26020" ID-ISSN="1175-5326" ID-Zenodo-Dep="177977" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459878007176" checkinUser="plazi" docAuthor="Jagt, John W. M., Zonova, Tatiana D. &amp; Jagt-Yazykova, Elena A." docDate="2007" docId="8E2E1A5C62238D18FF63C8CFFF40FBBD" docLanguage="en" docName="zt01545p047.pdf" docOrigin="Zootaxa 1545" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Pollicipes bruennichi Withers 1914" docType="treatment" docVersion="10" lastPageNumber="40" masterDocId="7217622462248D1FFFF4C858FFA3FFF9" masterDocTitle="A review of the brachylepadomorph cirripede genus Pycnolepas, including the first record of an Early Cretaceous species from the Russian Far East *" masterLastPageNumber="47" masterPageNumber="33" pageNumber="40" updateTime="1698227215383" updateUser="plazi">
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<mods:title id="5AE050095891E9C75EC28D16B1BFADAC">A review of the brachylepadomorph cirripede genus Pycnolepas, including the first record of an Early Cretaceous species from the Russian Far East *</mods:title>
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<mods:namePart id="CFE6F7DDC3C1629C67C1DBC49518026E">Zonova, Tatiana D.</mods:namePart>
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<mods:date id="F01E84A9C017694C6A94FDE841AF4E5C">2007</mods:date>
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<treatment id="8E2E1A5C62238D18FF63C8CFFF40FBBD" ID-DOI="http://doi.org/10.5281/zenodo.6247756" ID-GBIF-Taxon="119370373" ID-Zenodo-Dep="6247756" LSID="urn:lsid:plazi:treatment:8E2E1A5C62238D18FF63C8CFFF40FBBD" httpUri="http://treatment.plazi.org/id/8E2E1A5C62238D18FF63C8CFFF40FBBD" lastPageNumber="40" pageId="7" pageNumber="40">
<subSubSection id="4E9DF8C162238D18FF63C8CFFAA5FF05" pageId="7" pageNumber="40" type="nomenclature">
<paragraph id="0638AB4A62238D18FF63C8CFFDC1FF48" blockId="7.[151,610,151,177]" box="[151,610,151,177]" pageId="7" pageNumber="40">
<heading id="5D701C2662238D18FF63C8CFFDC1FF48" bold="true" box="[151,610,151,177]" fontSize="11" level="1" pageId="7" pageNumber="40" reason="1">
<emphasis id="34F3775862238D18FF63C8CFFDC1FF48" bold="true" box="[151,610,151,177]" pageId="7" pageNumber="40">
<emphasis id="34F3775862238D18FF63C8CFFE83FF48" bold="true" box="[151,288,151,177]" italics="true" pageId="7" pageNumber="40">Pycnolepas</emphasis>
<taxonomicName id="C187D0C962238D18FED3C8CFFDC1FF48" authority="Withers, 1914" authorityName="Withers" authorityYear="1914" box="[295,610,151,177]" class="Maxillopoda" family="Pollicipedidae" genus="Pollicipes" kingdom="Animalia" order="Pedunculata" pageId="7" pageNumber="40" phylum="Arthropoda" rank="species" species="bruennichi">
<emphasis id="34F3775862238D18FED3C8CFFE0EFF48" bold="true" box="[295,429,151,177]" italics="true" pageId="7" pageNumber="40">bruennichi</emphasis>
<bibRefCitation id="6216D6BB62238D18FE40C8CFFDC1FF48" author="Withers" box="[436,610,151,177]" pageId="7" pageNumber="46" refString="Withers, T. H. (1914) Some Cretaceous and Tertiary Cirripedia referred to Pollicipes. The Annals and Magazine of Natural History, (8) 14, 167 - 206." type="journal article" year="1914">Withers, 1914</bibRefCitation>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
<paragraph id="0638AB4A62238D18FF63C8BAFAA5FF05" blockId="7.[151,1437,226,1092]" box="[151,1286,226,252]" pageId="7" pageNumber="40">
<emphasis id="34F3775862238D18FF63C8BAFE38FF05" bold="true" box="[151,411,226,252]" pageId="7" pageNumber="40">Original description.</emphasis>
<bibRefCitation id="6216D6BB62238D18FE57C8BAFAA3FF05" author="Withers" box="[419,1280,226,252]" pageId="7" pageNumber="46" refString="Withers, T. H. (1914) Some Cretaceous and Tertiary Cirripedia referred to Pollicipes. The Annals and Magazine of Natural History, (8) 14, 167 - 206." type="journal article" year="1914">
Withers (1914: 181, pl. 7, figs 59; pl. 8, fig. 6, as
<emphasis id="34F3775862238D18FC00C8BAFB56FF02" box="[1012,1269,226,251]" italics="true" pageId="7" pageNumber="40">Pycnolepas brünnichi</emphasis>
)
</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="4E9DF8C162238D18FF32C952FEF8FE8D" pageId="7" pageNumber="40" type="materials_examined">
<paragraph id="0638AB4A62238D18FF32C952FCA6FEDD" blockId="7.[151,1437,226,1092]" box="[198,773,266,292]" pageId="7" pageNumber="40">
<emphasis id="34F3775862238D18FF32C952FEAAFEDD" bold="true" box="[198,265,266,292]" pageId="7" pageNumber="40">
<typeStatus id="D93C15E862238D18FF32C952FEA2FEDD" box="[198,257,266,292]" pageId="7" pageNumber="40">Type</typeStatus>
.
</emphasis>
<typeStatus id="D93C15E862238D18FEE6C952FEDCFEDD" box="[274,383,266,292]" pageId="7" pageNumber="40" type="holotype">Holotype</typeStatus>
is NHM I.16625, a right scutum.
</paragraph>
<paragraph id="0638AB4A62238D18FF32C96AFEF8FE8D" blockId="7.[151,1437,226,1092]" pageId="7" pageNumber="40">
<emphasis id="34F3775862238D18FF32C96AFDB2FEB5" bold="true" box="[198,529,306,332]" pageId="7" pageNumber="40">Locality and stratigraphy.</emphasis>
Fakse, Sjaelland (eastern
<collectingCountry id="7E90EBDA62238D18FCB1C96AFC1BFEB5" box="[837,952,306,332]" name="Denmark" pageId="7" pageNumber="40">Denmark</collectingCountry>
); middle Danian (
<taxonomicName id="C187D0C962238D18FB64C96AFF74FE8D" authority="Zone" authorityName="Zone" class="Echinoidea" family="Psychocidaridae" genus="Tylocidaris" kingdom="Animalia" order="Cidaroida" pageId="7" pageNumber="40" phylum="Echinodermata" rank="species" species="bruennichi">
<emphasis id="34F3775862238D18FB64C96AFA3FFEB2" box="[1168,1436,306,331]" italics="true" pageId="7" pageNumber="40">Tylocidaris bruennichi</emphasis>
Zone
</taxonomicName>
; c. 63 Ma).
</paragraph>
</subSubSection>
<subSubSection id="4E9DF8C162238D18FF32C9DAFF40FBBD" pageId="7" pageNumber="40" type="discussion">
<paragraph id="0638AB4A62238D18FF32C9DAFF48FD4D" blockId="7.[151,1437,226,1092]" pageId="7" pageNumber="40">
<emphasis id="34F3775862238D18FF32C9DAFE9CFE65" bold="true" box="[198,319,386,412]" pageId="7" pageNumber="40">Remarks.</emphasis>
Valves of this species are strongly ridged both transversely and longitudinally; scutum subtriangular, with broad wall-sided apicobasal ridge, occasionally broader than tergo-lateral portion, tergum with apical portion only slightly curved towards scutum and a similar, yet much narrower, near-straight apicobasal ridge. Height of imbricating plates exceeding width. On tergum, longitudinal ridges subdued by prominent growth lines. Thousands of isolated, yet associated, valves are known from the middle Danian of Fakse, and allow the species to be reconstructed; however, upper latera invariably are rare. Postulated to have been attached either to certain scleractinian corals and/or logs (
<bibRefCitation id="6216D6BB62238D18FCB9CA2AFB03FD75" author="Donovan" box="[845,1184,626,652]" pageId="7" pageNumber="44" refString="Donovan, S. K. &amp; Jakobsen, S. L. (2004) An unusual crinoid-barnacle association in the type area of the Danian (Paleocene), Denmark. Lethaia, 37, 407 - 415." type="journal article" year="2004">Donovan &amp; Jakobsen, 2004</bibRefCitation>
), substrates now dissolved.
</paragraph>
<paragraph id="0638AB4A62238D18FF32CA9AFC06FC5D" blockId="7.[151,1437,226,1092]" pageId="7" pageNumber="40">
<bibRefCitation id="6216D6BB62238D18FF32CA9AFEDAFD25" author="Withers" box="[198,377,706,732]" pageId="7" pageNumber="46" refString="Withers, T. H. (1914) Some Cretaceous and Tertiary Cirripedia referred to Pollicipes. The Annals and Magazine of Natural History, (8) 14, 167 - 206." type="journal article" year="1914">Withers (1914)</bibRefCitation>
proposed
<emphasis id="34F3775862238D18FE0DCA9AFCAAFD22" box="[505,777,706,731]" italics="true" pageId="7" pageNumber="40">
Pycnolepas
<taxonomicName id="C187D0C962238D18FD7ECA9AFCAAFD22" box="[650,777,706,731]" class="Echinoidea" family="Psychocidaridae" genus="Tylocidaris" kingdom="Animalia" order="Cidaroida" pageId="7" pageNumber="40" phylum="Echinodermata" rank="species" species="bruennichi">bruennichi</taxonomicName>
</emphasis>
as a replacement name for
<taxonomicName id="C187D0C962238D18FBAECA9AFF72FCFD" authority="Darwin, 1851" authorityName="Darwin" authorityYear="1851" class="Maxillopoda" family="Pollicipedidae" genus="Pollicipes" kingdom="Animalia" order="Pedunculata" pageId="7" pageNumber="40" phylum="Arthropoda" rank="species" species="elegans">
<emphasis id="34F3775862238D18FBAECA9AFA90FD22" box="[1114,1331,706,731]" italics="true" pageId="7" pageNumber="40">Pollicipes elegans</emphasis>
<bibRefCitation id="6216D6BB62238D18FAC8CA9AFF72FCFD" author="Darwin" pageId="7" pageNumber="44" refString="Darwin, C. R. (1851) A monograph on the fossil Lepadidae, or, pedunculated cirripedes of Great Britain, vi + 88 pp. Palaeontographical Society, London." type="book" year="1851">Darwin, 1851</bibRefCitation>
</taxonomicName>
(76, pl. 4, fig. 9ad),
<emphasis id="34F3775862238D18FE25CAB2FE5EFCFA" box="[465,509,746,771]" italics="true" pageId="7" pageNumber="40">non</emphasis>
<bibRefCitation id="6216D6BB62238D18FDF0CAB2FD48FCFD" author="Lesson" box="[516,747,746,772]" pageId="7" pageNumber="45" refString="Lesson, R. - P. (1830) Zoologie, 2 (1) (II). Mollusques, annelides et vers, 1. Mollusques. In: Duperrey, L. I. (Ed.). Vo yag e autour du monde, execute par ordre du roy, sur la corvette de sa majeste, La Coquille, pendant les annees 1822, 1823, 1824 et 1825, sous le ministere et conformement aux instructions du S. E. M. le Marquis de Clermont-Tonnerre, ministere de la marine et publie sous les auspices de son excellence Mgr. Le Cte de Chabrol, ministere de la marine et des colonies, pp. 239 - 448. A. Bertrand, Paris." type="journal article" year="1830">Lesson (1830: 441)</bibRefCitation>
, and later (
<bibRefCitation id="6216D6BB62238D18FC9FCAB2FBB0FCFD" author="Withers" box="[875,1043,746,772]" pageId="7" pageNumber="46" refString="Withers, T. H. (1935) Catalogue of fossil Cirripedia in the Department of Geology. Vol. II. Cretaceous. Trustees of the British Museum [Natural History], London, xiv + 535 pp." type="book" year="1935">Withers, 1935</bibRefCitation>
) showed
<taxonomicName id="C187D0C962238D18FB70CAB2FA83FCFA" box="[1156,1312,746,772]" class="Echinoidea" family="Psychocidaridae" genus="Tylocidaris" kingdom="Animalia" order="Cidaroida" pageId="7" pageNumber="40" phylum="Echinodermata" rank="species" species="bruennichi">
<emphasis id="34F3775862238D18FB70CAB2FB30FCFA" box="[1156,1171,746,771]" italics="true" pageId="7" pageNumber="40">P</emphasis>
.
<emphasis id="34F3775862238D18FB55CAB2FA83FCFA" box="[1185,1312,746,771]" italics="true" pageId="7" pageNumber="40">bruennichi</emphasis>
</taxonomicName>
to be confined to strata of early Paleocene (Danian) age.
<bibRefCitation id="6216D6BB62238D18FD39CB4AFCDDFCD5" author="Drygant" box="[717,894,786,812]" pageId="7" pageNumber="44" refString="Drygant, D. M. (1966) Usonogie rakoobraznye verkhnemelovykh otlozhenii Volyno-Podol'skoi Plity. Paleontologicheskii Sbornik, 1966 / 3, 113 - 117." type="journal article" year="1966">Drygant (1966</bibRefCitation>
: 116, fig. 8a, b) recorded a single carina from upper Maastrichtian levels at Briukhovichi, Volhynia-Podolia (now western
<collectingCountry id="7E90EBDA62238D18FBD8CB62FB2FFCAD" box="[1068,1164,826,852]" name="Ukraine" pageId="7" pageNumber="40">Ukraine</collectingCountry>
) which he assigned to
<taxonomicName id="C187D0C962238D18FF63CB3AFE57FC85" authority="Darwin" authorityName="Darwin" box="[151,500,866,892]" class="Maxillopoda" family="Brachylepadidae" genus="Pycnolepas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="7" pageNumber="40" phylum="Arthropoda" rank="species" species="elegans">
<emphasis id="34F3775862238D18FF63CB3AFEDCFC82" box="[151,383,866,891]" italics="true" pageId="7" pageNumber="40">Pycnolepas elegans</emphasis>
(Darwin)
</taxonomicName>
. This, however, clearly represents a typical carina of the calanticine
<taxonomicName id="C187D0C962238D18FAFACB3AFE46FC5D" authority="Bosquet, 1854" authorityName="Bosquet" authorityYear="1854" class="Maxillopoda" family="Calanticidae" genus="Scillaelepas" kingdom="Animalia" order="Pedunculata" pageId="7" pageNumber="40" phylum="Arthropoda" rank="species" species="darwiniana">
<emphasis id="34F3775862238D18FAFACB3AFEBDFC5A" italics="true" pageId="7" pageNumber="40">Scillaelepas darwiniana</emphasis>
(
<bibRefCitation id="6216D6BB62238D18FED9CBD2FE7FFC5D" author="Bosquet" box="[301,476,906,932]" pageId="7" pageNumber="44" refString="Bosquet, J. (1854) Les Crustaces fossiles du Terrain Cretace du Limbourg. Verhandelingen uitgegeven door de Commissie belast met het vervaardigen eener geologische beschrijving en kaart van Nederland, 2, 1 - 127 [10 - 137]. A. C. Kruseman, Haarlem." type="journal article" year="1854">Bosquet, 1854</bibRefCitation>
)
</taxonomicName>
, as already noted by
<bibRefCitation id="6216D6BB62238D18FD2FCBD2FC02FC5D" author="Alekseev" box="[731,929,906,932]" pageId="7" pageNumber="44" refString="Alekseev, A. S. (1979) Verkhnemelovye usonogie raki (Cirripedia, Thoracica) evropeiskoi chasti SSSR i sopredel'nykh raionov. Moskva, Moskovskii Gosudarstvennyi Universitet im. M. V. Lomonosova, 488 pp. (unpublished PhD thesis)." type="book" year="1979">Alekseev (1979)</bibRefCitation>
.
</paragraph>
<paragraph id="0638AB4A62238D18FF32CBEAFF40FBBD" blockId="7.[151,1437,226,1092]" pageId="7" pageNumber="40">
<emphasis id="34F3775862238D18FF32CBEAFEFCFC35" bold="true" box="[198,351,946,972]" pageId="7" pageNumber="40">Occurrence.</emphasis>
Lower and middle Danian of
<collectingCountry id="7E90EBDA62238D18FD3DCBEAFC95FC35" box="[713,822,946,972]" name="Denmark" pageId="7" pageNumber="40">Denmark</collectingCountry>
(
<bibRefCitation id="6216D6BB62238D18FCB2CBEAFC52FC35" author="Withers" box="[838,1009,946,972]" pageId="7" pageNumber="46" refString="Withers, T. H. (1914) Some Cretaceous and Tertiary Cirripedia referred to Pollicipes. The Annals and Magazine of Natural History, (8) 14, 167 - 206." type="journal article" year="1914">Withers, 1914</bibRefCitation>
,
<bibRefCitation id="6216D6BB62238D18FC0ACBEAFB9FFC35" author="Withers" box="[1022,1084,946,972]" pageId="7" pageNumber="46" refString="Withers, T. H. (1935) Catalogue of fossil Cirripedia in the Department of Geology. Vol. II. Cretaceous. Trustees of the British Museum [Natural History], London, xiv + 535 pp." type="book" year="1935">1935</bibRefCitation>
;
<bibRefCitation id="6216D6BB62238D18FBBECBEAFAA0FC35" author="Jakobsen" box="[1098,1283,946,972]" pageId="7" pageNumber="45" refString="Jakobsen, S. L. (2003) A new preparatory approach of decapod and thoracican crustaceans from the Middle Danian at Fakse, Denmark. Contributions to Zoology, 72, 141 - 145." type="journal article" year="2003">Jakobsen, 2003</bibRefCitation>
;
<bibRefCitation id="6216D6BB62238D18FAE5CBEAFEF3FC0D" author="Donovan" pageId="7" pageNumber="44" refString="Donovan, S. K. &amp; Jakobsen, S. L. (2004) An unusual crinoid-barnacle association in the type area of the Danian (Paleocene), Denmark. Lethaia, 37, 407 - 415." type="journal article" year="2004">Donovan &amp; Jakobsen, 2004</bibRefCitation>
;
<bibRefCitation id="6216D6BB62238D18FEA8CB82FD11FC0D" author="Jakobsen" box="[348,690,986,1012]" pageId="7" pageNumber="45" refString="Jakobsen, S. L. &amp; Feldmann, R. M. (2004) Epibionts on Dromiopsis rugosa (Decapoda: Brachyura) from the late Middle Danian limestones at Fakse quarry, Denmark: novel preparation techniques yield amazing results. Journal of Paleontology, 78, 953 - 960." type="journal article" year="2004">Jakobsen &amp; Feldmann, 2004</bibRefCitation>
), southern
<collectingCountry id="7E90EBDA62238D18FCC7CB82FC33FC0D" box="[819,912,986,1012]" name="Sweden" pageId="7" pageNumber="40">Sweden</collectingCountry>
(Limhamn area), northeast
<collectingCountry id="7E90EBDA62238D18FB2DCB82FA9DFC0D" box="[1241,1342,986,1012]" name="Belgium" pageId="7" pageNumber="40">Belgium</collectingCountry>
and the southeast
<collectingCountry id="7E90EBDA62238D18FEE4CC5AFE3EFBE5" box="[272,413,1026,1052]" name="Netherlands" pageId="7" pageNumber="40">Netherlands</collectingCountry>
(
<bibRefCitation id="6216D6BB62238D18FE45CC5AFD14FBE5" author="Jagt" box="[433,695,1026,1052]" pageId="7" pageNumber="45" refString="Jagt, J. W. M. &amp; Collins, J. S. H. (1988) The biostratigraphy of the Geulhem Member (Early Palaeocene), with reference to the occurrence of Pycnolepas bruennichi Withers, 1914 (Crustacea, Cirripedia). In: Jagt, J. W. M. &amp; Janssen, A. W. (Eds.). Faunal and stratigraphical aspects of the Early Palaeocene (Danian) in the SE Netherlands and NE Belgium. Mededelingen van de Werkgroep voor Tertiaire en Kwartaire Geologie, 25, 175 - 196." type="journal article" year="1988">Jagt &amp; Collins, 1988</bibRefCitation>
;
<bibRefCitation id="6216D6BB62238D18FD33CC5AFCE5FBE5" author="Jagt" box="[711,838,1026,1052]" pageId="7" pageNumber="45" refString="Jagt, J. W. M. (1996) Late Maastrichtian and Early Palaeocene index macrofossils in the Maastrichtian type area (SE Netherlands, NE Belgium). In: Brinkhuis, H. &amp; Smit, J. (Eds.). The Geulhemmerberg Cretaceous / Tertiary boundary section (Maastrichtian type area, SE Netherlands). Geologie en Mijnbouw, 75, 153 - 162." type="journal article" year="1996">Jagt, 1996</bibRefCitation>
) and the Crimean Peninsula,
<collectingCountry id="7E90EBDA62238D18FB41CC5AFAB0FBE5" box="[1205,1299,1026,1052]" name="Ukraine" pageId="7" pageNumber="40">Ukraine</collectingCountry>
(
<bibRefCitation id="6216D6BB62238D18FADCCC5AFF75FBBD" author="Alekseev" pageId="7" pageNumber="44" refString="Alekseev, A. S. (1979) Verkhnemelovye usonogie raki (Cirripedia, Thoracica) evropeiskoi chasti SSSR i sopredel'nykh raionov. Moskva, Moskovskii Gosudarstvennyi Universitet im. M. V. Lomonosova, 488 pp. (unpublished PhD thesis)." type="book" year="1979">Alekseev, 1979</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>